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<document id="36612461BCB5D59290F998057E66F1F2" ID-CLB-Dataset="22413" ID-DOI="10.11646/zootaxa.4768.3.2" ID-GBIF-Dataset="43c18a9c-5545-4f3d-b285-956094e43d43" ID-ISSN="1175-5326" ID-Zenodo-Dep="3784036" ID-ZooBank="A005B93A-2070-46F5-A06E-6F6268A207EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588576355177" checkinUser="plazi" docAuthor="Hu, Fang-Shuo" docDate="2020" docId="997687A2FFCBFF9AFACC3597546BA1B3" docLanguage="en" docName="zootaxa.4768.3.2.pdf" docOrigin="Zootaxa 4768 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Miobdelus taiwanensis subsp. taiwanensis taiwanensis Smetana 2001" docType="treatment" docVersion="6" lastPageNumber="341" masterDocId="654FFFDAFFCCFF9DFA5B34445127A409" masterDocTitle="New distributional records of Staphylinina in Taiwan, including a new species of Miobdelus Sharp (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)" masterLastPageNumber="360" masterPageNumber="334" pageNumber="341" updateTime="1698838343837" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F83247A65B41B51578AF090B27D03DC3">New distributional records of Staphylinina in Taiwan, including a new species of Miobdelus Sharp (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)</mods:title>
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<mods:namePart id="0095E627FC64CC56372F2D189168AA03">Hu, Fang-Shuo</mods:namePart>
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<mods:date id="33B7F6A58D1C739106C14186A87B8D7F">2020</mods:date>
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<treatment id="997687A2FFCBFF9AFACC3597546BA1B3" ID-DOI="http://doi.org/10.5281/zenodo.3794840" ID-GBIF-Taxon="163885976" ID-Zenodo-Dep="3794840" LSID="urn:lsid:plazi:treatment:997687A2FFCBFF9AFACC3597546BA1B3" httpUri="http://treatment.plazi.org/id/997687A2FFCBFF9AFACC3597546BA1B3" lastPageNumber="341" pageId="7" pageNumber="341">
<subSubSection id="59C5653FFFCBFF9AFACC359753D8A5E4" box="[151,767,467,494]" pageId="7" pageNumber="341" type="nomenclature">
<paragraph id="116036B4FFCBFF9AFACC359753D8A5E4" blockId="7.[151,767,467,529]" box="[151,767,467,494]" pageId="7" pageNumber="341">
<heading id="4A2881D8FFCBFF9AFACC359753D8A5E4" bold="true" box="[151,767,467,494]" fontSize="11" level="1" pageId="7" pageNumber="341" reason="1">
<taxonomicName id="D6DF4D37FFCBFF9AFACC359753D8A5E4" ID-CoL="8YDXG" authority="taiwanensis Smetana, 2001" authorityName="taiwanensis Smetana" authorityYear="2001" box="[151,767,467,494]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="subSpecies" species="taiwanensis" subSpecies="taiwanensis">
<emphasis id="23ABEAA6FFCBFF9AFACC359753D8A5E4" bold="true" box="[151,767,467,494]" pageId="7" pageNumber="341">
<emphasis id="23ABEAA6FFCBFF9AFACC35975366A5E4" bold="true" box="[151,577,467,494]" italics="true" pageId="7" pageNumber="341">Miobdelus taiwanensis taiwanensis</emphasis>
<bibRefCitation id="754E4B45FFCBFF9AF813359753D8A5E4" author="Smetana, A." box="[584,767,467,493]" pageId="7" pageNumber="341" pagination="161 - 201" refId="ref17474" refString="Smetana, A. (2001) Contribution to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part 1. The review of the genus Miobdelus. Folia Heyrovskyana, 9 (3 - 4), 161 - 201." type="journal article" year="2001">Smetana, 2001</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="59C5653FFFCBFF9AFACC35B351CCA618" box="[151,235,503,529]" pageId="7" pageNumber="341" type="description">
<paragraph id="116036B4FFCBFF9AFACC35B351CCA618" blockId="7.[151,767,467,529]" box="[151,235,503,529]" pageId="7" pageNumber="341">
(
<figureCitation id="89E42A31FFCBFF9AFAFB35B351C4A618" box="[160,227,503,529]" captionStart="FIGURES 8–13" captionStartId="11.[151,264,1862,1887]" captionTargetBox="[151,1412,185,1837]" captionTargetId="figure@11.[151,1435,181,1837]" captionTargetPageId="11" captionText="FIGURES 8–13. 8. Miobdelus humilis sp. nov.: male, paratype. 9. Miobdelus taiwanensis taiwanensis: male. 10. Nelmanwasus ornatus: male. 11–12. Sphaerobulbus miwai comb. nov.: male, lectotype of Staphylinus (Xanthocypus) miwai and attached labels; male. 13. Naddia taiwanensis: female. Scale bar: 3 mm" figureDoi="http://doi.org/10.5281/zenodo.3784040" httpUri="https://zenodo.org/record/3784040/files/figure.png" pageId="7" pageNumber="341">Fig. 9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="59C5653FFFCBFF9AFACC3678538AA670" pageId="7" pageNumber="341" type="materials_examined">
<paragraph id="116036B4FFCBFF9AFACC3678538AA670" blockId="7.[151,1437,571,1466]" pageId="7" pageNumber="341">
<emphasis id="23ABEAA6FFCBFF9AFACC367850A7A65E" bold="true" box="[151,384,572,599]" pageId="7" pageNumber="341">Material examined.</emphasis>
<materialsCitation id="A1B73CE9FFCBFF9AFBDD367855E5A65E" ID-GBIF-Occurrence="2609798921" box="[390,1218,571,599]" collectionCode="V" collectorName="Anmashan" country="Taiwan" location="Daxue Mt." pageId="7" pageNumber="341" specimenCount="1" specimenCount-male="1" stateProvince="Taichung">
<emphasis id="23ABEAA6FFCBFF9AFBDD36785327A65E" bold="true" box="[390,512,572,599]" pageId="7" pageNumber="341">
<collectingCountry id="69C87624FFCBFF9AFBDD367850DBA65E" box="[390,508,572,599]" name="Taiwan" pageId="7" pageNumber="341">TAIWAN</collectingCountry>
:
</emphasis>
<collectingRegion id="D31BF856FFCBFF9AF85D3678538EA65F" box="[518,681,572,598]" country="Taiwan" name="Taichung" pageId="7" pageNumber="341">Taichung City</collectingRegion>
:
<specimenCount id="07D9FD3DFFCBFF9AF8E9367953D8A65F" box="[690,767,572,599]" pageId="7" pageNumber="341" type="male">1 male</specimenCount>
,
<collectorName id="BC2A5362FFCBFF9AF952367852A6A65F" box="[777,897,572,598]" pageId="7" pageNumber="341">Anmashan</collectorName>
[=
<location id="1400606FFFCBFF9AF9C736795530A65E" LSID="urn:lsid:plazi:treatment:997687A2FFCBFF9AFACC3597546BA1B3:1400606FFFCBFF9AF9C736795530A65E" box="[924,1047,573,599]" country="Taiwan" name="Daxue Mt." pageId="7" pageNumber="341" stateProvince="Taichung">Daxue Mt.</location>
(
<emphasis id="23ABEAA6FFCBFF9AFE7D367F555DA65B" bold="true" box="[1062,1146,571,594]" pageId="7" pageNumber="341">±ũƜ</emphasis>
)],
<date id="65611074FFCBFF9AFEC1367955E5A65E" box="[1178,1218,573,599]" pageId="7" pageNumber="341" value="1990-05-01">
1/
<collectionCode id="77CEAE71FFCBFF9AFEF7367955E5A65E" box="[1196,1218,573,599]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="341" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="A1B73CE9FFCBFF9AFE993679538EA670" ID-GBIF-Occurrence="2609798949" collectionCode="NMNS" collectorName="C. C. Chiang" country="Taiwan" pageId="7" pageNumber="341" specimenCode="ENT 583-288" specimenCount="1" stateProvince="Taichung City">
/1990,
<collectorName id="BC2A5362FFCBFF9AFF4A367854BFA65F" box="[1297,1432,572,598]" pageId="7" pageNumber="341">C.C.Chiang</collectorName>
,
<collectingMethod id="C89E4EA3FFCBFF9AFACC361B5008A673" box="[151,303,607,634]" pageId="7" pageNumber="341">Sweeping net</collectingMethod>
[
<collectionCode id="77CEAE71FFCBFF9AFB67361B50B7A670" box="[316,400,607,633]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="7" pageNumber="341">NMNS</collectionCode>
<specimenCode id="41799ECFFFCBFF9AFBCC36245313A670" box="[407,564,606,634]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="7" pageNumber="341">ENT 583-288</specimenCode>
] (
<collectionCode id="77CEAE71FFCBFF9AF811361B5386A670" box="[586,673,607,633]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="7" pageNumber="341">NMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="59C5653FFFCBFF9AFA9C36C55256A787" pageId="7" pageNumber="341" type="diagnosis">
<paragraph id="116036B4FFCBFF9AFA9C36C552F5A72E" blockId="7.[151,1437,571,1466]" pageId="7" pageNumber="341">
<emphasis id="23ABEAA6FFCBFF9AFA9C36C55065A695" bold="true" box="[199,322,641,668]" pageId="7" pageNumber="341">Diagnosis.</emphasis>
In
<collectingCountry id="69C87624FFCBFF9AFB3E36C6509DA695" box="[357,442,642,668]" name="Taiwan" pageId="7" pageNumber="341">Taiwan</collectingCountry>
,
<taxonomicName id="D6DF4D37FFCBFF9AFB9F36C7534AA695" authorityName="Smetana" authorityYear="2001" box="[452,621,642,668]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="23ABEAA6FFCBFF9AFB9F36C750FCA695" box="[452,475,643,668]" italics="true" pageId="7" pageNumber="341">M</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AFBB236C7534AA695" box="[489,621,643,668]" italics="true" pageId="7" pageNumber="341">taiwanensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="D6DF4D37FFCBFF9AF8A736C75508A695" authority="Smetana, 2001" authorityName="Smetana" authorityYear="2001" box="[764,1071,642,668]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="23ABEAA6FFCBFF9AF8A736C75234A695" box="[764,787,643,668]" italics="true" pageId="7" pageNumber="341">M</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AF97A36C652A4A695" box="[801,899,642,668]" italics="true" pageId="7" pageNumber="341">insularis</emphasis>
<bibRefCitation id="754E4B45FFCBFF9AF9D236C65508A695" author="Smetana, A." box="[905,1071,642,668]" pageId="7" pageNumber="341" pagination="161 - 201" refId="ref17474" refString="Smetana, A. (2001) Contribution to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part 1. The review of the genus Miobdelus. Folia Heyrovskyana, 9 (3 - 4), 161 - 201." type="journal article" year="2001">Smetana, 2001</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="D6DF4D37FFCBFF9AFE3836C75455A695" authority="Smetana, 2001" authorityName="Smetana" authorityYear="2001" box="[1123,1394,642,668]" class="Insecta" family="Staphylinidae" genus="Dinothenarus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="choui">
<emphasis id="23ABEAA6FFCBFF9AFE3836C7555DA695" box="[1123,1146,643,668]" italics="true" pageId="7" pageNumber="341">M</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AFED336C655E1A695" box="[1160,1222,642,668]" italics="true" pageId="7" pageNumber="341">choui</emphasis>
<bibRefCitation id="754E4B45FFCBFF9AFE9636C65455A695" author="Smetana, A." box="[1229,1394,642,668]" pageId="7" pageNumber="341" pagination="161 - 201" refId="ref17474" refString="Smetana, A. (2001) Contribution to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part 1. The review of the genus Miobdelus. Folia Heyrovskyana, 9 (3 - 4), 161 - 201." type="journal article" year="2001">Smetana, 2001</bibRefCitation>
</taxonomicName>
but can be distinguished by presence of the patch of yellow tomentose pubescence in the middle of tergite 6.
<taxonomicName id="D6DF4D37FFCBFF9AFF7D36E0508FA6E8" authority="taiwanensis" authorityName="taiwanensis Smetana" authorityYear="2001" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="subSpecies" species="taiwanensis" subSpecies="taiwanensis">
<emphasis id="23ABEAA6FFCBFF9AFF7D36E0508FA6E8" italics="true" pageId="7" pageNumber="341">Miobdelus taiwanensis taiwanensis</emphasis>
</taxonomicName>
can be distinguished from the subspecies
<taxonomicName id="D6DF4D37FFCBFF9AF9D5368C546BA6E8" authority="Smetana, 2001" authorityName="Smetana" authorityYear="2001" box="[910,1356,711,738]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="subSpecies" species="taiwanensis" subSpecies="apicalis">
<emphasis id="23ABEAA6FFCBFF9AF9D5368C5282A6E8" box="[910,933,712,737]" italics="true" pageId="7" pageNumber="341">M</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AF9EE368C55BCA6E8" box="[949,1179,711,737]" italics="true" pageId="7" pageNumber="341">taiwanensis apicalis</emphasis>
<bibRefCitation id="754E4B45FFCBFF9AFEFF3683546BA6E8" author="Smetana, A." box="[1188,1356,711,737]" pageId="7" pageNumber="341" pagination="161 - 201" refId="ref17474" refString="Smetana, A. (2001) Contribution to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part 1. The review of the genus Miobdelus. Folia Heyrovskyana, 9 (3 - 4), 161 - 201." type="journal article" year="2001">Smetana, 2001</bibRefCitation>
</taxonomicName>
by the relatively narrower median lobe, especially on the apical portion and presence of a rounded apical emargination as well as the fact that
<taxonomicName id="D6DF4D37FFCBFF9AFB233749538BA72F" authority="taiwanensis" authorityName="taiwanensis Smetana" authorityYear="2001" box="[376,684,781,807]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="subSpecies" species="taiwanensis" subSpecies="taiwanensis">
<emphasis id="23ABEAA6FFCBFF9AFB23374950A8A72F" box="[376,399,781,806]" italics="true" pageId="7" pageNumber="341">M</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AFBC63749538BA72F" box="[413,684,781,806]" italics="true" pageId="7" pageNumber="341">taiwanensis taiwanensis</emphasis>
</taxonomicName>
is restricted to Daxue Mt.
</paragraph>
<paragraph id="116036B4FFCBFF9AFA9C376A5256A787" blockId="7.[151,1437,571,1466]" pageId="7" pageNumber="341">
<emphasis id="23ABEAA6FFCBFF9AFA9C376A506CA740" bold="true" box="[199,331,814,841]" pageId="7" pageNumber="341">Bionomics.</emphasis>
Most specimens of
<taxonomicName id="D6DF4D37FFCBFF9AF86A377453FBA740" authorityName="Smetana" authorityYear="2001" box="[561,732,816,842]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="23ABEAA6FFCBFF9AF86A3774536FA740" box="[561,584,816,841]" italics="true" pageId="7" pageNumber="341">M</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AF803377453FBA740" box="[600,732,816,841]" italics="true" pageId="7" pageNumber="341">taiwanensis</emphasis>
</taxonomicName>
were collected by yellow pan trap set on higher mountains in mature broadleaved forests or by sifting forest floor litter or moss on fallen trees (
<bibRefCitation id="754E4B45FFCBFF9AFE7D371655ECA765" author="Smetana, A." box="[1062,1227,850,876]" pageId="7" pageNumber="341" pagination="161 - 201" refId="ref17474" refString="Smetana, A. (2001) Contribution to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part 1. The review of the genus Miobdelus. Folia Heyrovskyana, 9 (3 - 4), 161 - 201." type="journal article" year="2001">Smetana 2001</bibRefCitation>
). The collection of the specimen in this study by sweep net was probably accidental.
</paragraph>
</subSubSection>
<subSubSection id="59C5653FFFCBFF9AFA9C37D25355A7DC" pageId="7" pageNumber="341" type="distribution">
<paragraph id="116036B4FFCBFF9AFA9C37D25355A7DC" blockId="7.[151,1437,571,1466]" pageId="7" pageNumber="341">
<emphasis id="23ABEAA6FFCBFF9AFA9C37D25046A7B8" bold="true" box="[199,353,918,945]" pageId="7" pageNumber="341">Distribution.</emphasis>
All known specimens of the subspecies were collected from central
<collectingCountry id="69C87624FFCBFF9AFE0E37D35580A7B8" box="[1109,1191,919,945]" name="Taiwan" pageId="7" pageNumber="341">Taiwan</collectingCountry>
at the
<typeStatus id="CE648816FFCBFF9AFEAB37DC5407A7BB" box="[1264,1312,920,946]" pageId="7" pageNumber="341">type</typeStatus>
locality on Daxue Mt. (
<bibRefCitation id="754E4B45FFCBFF9AFB4637FE5098A7DD" author="Smetana, A." box="[285,447,954,980]" pageId="7" pageNumber="341" pagination="161 - 201" refId="ref17474" refString="Smetana, A. (2001) Contribution to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part 1. The review of the genus Miobdelus. Folia Heyrovskyana, 9 (3 - 4), 161 - 201." type="journal article" year="2001">Smetana 2001</bibRefCitation>
and this study).
</paragraph>
</subSubSection>
<subSubSection id="59C5653FFFCBFF9AFA9C3799546BA1B3" pageId="7" pageNumber="341" type="discussion">
<paragraph id="116036B4FFCBFF9AFA9C37995561A013" blockId="7.[151,1437,571,1466]" pageId="7" pageNumber="341">
<emphasis id="23ABEAA6FFCBFF9AFA9C3799501CA7FE" bold="true" box="[199,315,989,1015]" pageId="7" pageNumber="341">Remarks.</emphasis>
<bibRefCitation id="754E4B45FFCBFF9AFB1C379950D9A7FE" author="Smetana, A." box="[327,510,989,1015]" pageId="7" pageNumber="341" pagination="161 - 201" refId="ref17474" refString="Smetana, A. (2001) Contribution to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part 1. The review of the genus Miobdelus. Folia Heyrovskyana, 9 (3 - 4), 161 - 201." type="journal article" year="2001">Smetana (2001)</bibRefCitation>
indicated there were two
<typeStatus id="CE648816FFCBFF9AF96337995284A7FE" box="[824,931,989,1015]" pageId="7" pageNumber="341" type="paratype">paratypes</typeStatus>
deposited in NMNS and two
<typeStatus id="CE648816FFCBFF9AFF543799545DA7FE" box="[1295,1402,989,1015]" pageId="7" pageNumber="341" type="paratype">paratypes</typeStatus>
in NHMW, but these
<typeStatus id="CE648816FFCBFF9AFB31304450F2A013" box="[362,469,1024,1050]" pageId="7" pageNumber="341" type="paratype">paratypes</typeStatus>
were not found in these two institutions at the moment.
</paragraph>
<paragraph id="116036B4FFCBFF9AFA9C3066546BA1B3" blockId="7.[151,1437,571,1466]" pageId="7" pageNumber="341">
<bibRefCitation id="754E4B45FFCBFF9AFA9C3066505DA035" author="Smetana, A." box="[199,378,1058,1084]" pageId="7" pageNumber="341" pagination="161 - 201" refId="ref17474" refString="Smetana, A. (2001) Contribution to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part 1. The review of the genus Miobdelus. Folia Heyrovskyana, 9 (3 - 4), 161 - 201." type="journal article" year="2001">Smetana (2001)</bibRefCitation>
described two subspecies of
<taxonomicName id="D6DF4D37FFCBFF9AF89E30675248A035" authorityName="Smetana" authorityYear="2001" box="[709,879,1058,1084]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="23ABEAA6FFCBFF9AF89E306753FBA035" box="[709,732,1059,1084]" italics="true" pageId="7" pageNumber="341">M</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AF8B030675248A035" box="[747,879,1059,1084]" italics="true" pageId="7" pageNumber="341">taiwanensis</emphasis>
</taxonomicName>
as well two similar species,
<taxonomicName id="D6DF4D37FFCBFF9AFEEF3067541BA035" authorityName="Smetana" authorityYear="2001" box="[1204,1340,1058,1084]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="23ABEAA6FFCBFF9AFEEF306755ECA035" box="[1204,1227,1059,1084]" italics="true" pageId="7" pageNumber="341">M</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AFE813066541BA035" box="[1242,1340,1058,1084]" italics="true" pageId="7" pageNumber="341">insularis</emphasis>
</taxonomicName>
(including three subspecies) and
<taxonomicName id="D6DF4D37FFCBFF9AFBE630015306A057" authorityName="Smetana" authorityYear="2001" box="[445,545,1092,1119]" class="Insecta" family="Staphylinidae" genus="Dinothenarus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="choui">
<emphasis id="23ABEAA6FFCBFF9AFBE6300150F3A057" box="[445,468,1093,1118]" italics="true" pageId="7" pageNumber="341">M</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AFBB830005306A057" box="[483,545,1092,1118]" italics="true" pageId="7" pageNumber="341">choui</emphasis>
</taxonomicName>
; all of them were collected from high mountain areas on
<collectingCountry id="69C87624FFCBFF9AFEF430005426A057" box="[1199,1281,1092,1118]" name="Taiwan" pageId="7" pageNumber="341">Taiwan</collectingCountry>
island but on different mountains. However, the species or subspecies status need to be reassessed. Pleistocene glacial oscillations and the resulting geographic isolation on mountains can be major drivers of speciation in animals on
<collectingCountry id="69C87624FFCBFF9AFF1130CE54BBA0AD" box="[1354,1436,1162,1188]" name="Taiwan" pageId="7" pageNumber="341">Taiwan</collectingCountry>
island, especially animals living in high mountain habitats (see following references). Such research has been applied many times to the family
<taxonomicName id="D6DF4D37FFCBFF9AFBAA308B5234A0E0" authority="Latreille, 1804" authorityName="Latreille" authorityYear="1804" box="[497,787,1231,1257]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="family">Lucanidae Latreille, 1804</taxonomicName>
, where highly morphologically different beetles can either be treated as different subspecies (etc.,
<taxonomicName id="D6DF4D37FFCBFF9AF80830B753D0A105" authorityName="Kurosawa" authorityYear="1966" box="[595,759,1266,1292]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="kanoi">
<emphasis id="23ABEAA6FFCBFF9AF80830B753D0A105" box="[595,759,1266,1292]" italics="true" pageId="7" pageNumber="341">Lucanus kanoi</emphasis>
</taxonomicName>
<taxonomicName id="D6DF4D37FFCBFF9AF8A430B652D9A105" authority="Kurosawa, 1966" authorityName="Kurosawa" authorityYear="1966" box="[767,1022,1266,1292]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="kanoi">kanoi Kurosawa, 1966</taxonomicName>
and
<emphasis id="23ABEAA6FFCBFF9AFE6E30B75562A105" box="[1077,1093,1267,1292]" italics="true" pageId="7" pageNumber="341">L</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AFE0F30B65547A105" box="[1108,1120,1266,1292]" italics="true" pageId="7" pageNumber="341">k</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AFE3430B75590A105" box="[1135,1207,1267,1292]" italics="true" pageId="7" pageNumber="341">piceus</emphasis>
Kurosawa, 1966;
<taxonomicName id="D6DF4D37FFCBFF9AFFDD30B750FAA126" authority="ogakii Imanishi, 1990" authorityName="Imanishi" authorityYear="1990" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="subSpecies" species="ogakii" subSpecies="ogakii">
<emphasis id="23ABEAA6FFCBFF9AFFDD30B754B1A105" box="[1414,1430,1267,1292]" italics="true" pageId="7" pageNumber="341">L</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AFACC3150500BA127" box="[151,300,1300,1326]" italics="true" pageId="7" pageNumber="341">ogakii ogakii</emphasis>
Imanishi, 1990
</taxonomicName>
and
<emphasis id="23ABEAA6FFCBFF9AF84D31515301A127" box="[534,550,1301,1326]" italics="true" pageId="7" pageNumber="341">L</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AF86E31515364A127" box="[565,579,1301,1326]" italics="true" pageId="7" pageNumber="341">o</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AF808315053D5A127" box="[595,754,1300,1326]" italics="true" pageId="7" pageNumber="341">chuyunshanus</emphasis>
Sakaino &amp; Yu, 1993;
<taxonomicName id="D6DF4D37FFCBFF9AF9AB3150501EA158" authority="Bomans &amp; Ratti., 1973" authorityName="Bomans &amp; Ratti." authorityYear="1973" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="subSpecies" species="davidis" subSpecies="cheni">
<emphasis id="23ABEAA6FFCBFF9AF9AB3150541FA127" box="[1008,1336,1300,1326]" italics="true" pageId="7" pageNumber="341">Prismognathus davidis cheni</emphasis>
Bomans &amp; Ratti., 1973
</taxonomicName>
and
<emphasis id="23ABEAA6FFCBFF9AFB2A317C50A5A158" box="[369,386,1336,1361]" italics="true" pageId="7" pageNumber="341">P</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AFBCA317350B8A158" box="[401,415,1335,1361]" italics="true" pageId="7" pageNumber="341">d</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AFBF6317C533AA158" box="[429,541,1336,1361]" italics="true" pageId="7" pageNumber="341">nigerrimu</emphasis>
Sakaino &amp; Yu,
<quantity id="D6279B51FFCBFF9AF89031735238A158" box="[715,799,1335,1361]" metricMagnitude="1" metricUnit="m" metricValue="5.06222" pageId="7" pageNumber="341" unit="in" value="1993.0">1993 in</quantity>
<bibRefCitation id="754E4B45FFCBFF9AF97D317352C1A15B" author="Tsai, C. - L. &amp; Yeh, W. - B." box="[806,998,1335,1362]" pageId="7" pageNumber="341" pagination="0156600" refId="ref18231" refString="Tsai, C. - L. &amp; Yeh, W. - B. (2016) Subspecific differentiation events of montane stag beetles (Coleoptera, Lucanidae) endemic to Formosa Island. PLoS One, 11 (6), e 0156600. https: // doi. org / 10.1371 / journal. pone. 0156600" type="journal article" year="2016">Tsai &amp; Yeh 2016</bibRefCitation>
) or one species (
<taxonomicName id="D6DF4D37FFCBFF9AFEFC317C50EFA17D" authority="Planet, 1899 in Huang &amp; Lin 2010" authorityName="Planet, 1899 in Huang &amp; Lin" authorityYear="2010" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="23ABEAA6FFCBFF9AFEFC317C5590A158" box="[1191,1207,1336,1361]" italics="true" pageId="7" pageNumber="341">L</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AFE9E3173546FA158" box="[1221,1352,1335,1361]" italics="true" pageId="7" pageNumber="341">formosanus</emphasis>
Planet,
<quantity id="D6279B51FFCBFF9AFACC311E51CBA17D" box="[151,236,1370,1396]" metricMagnitude="1" metricUnit="m" metricValue="4.82346" pageId="7" pageNumber="341" unit="in" value="1899.0">1899 in</quantity>
<bibRefCitation id="754E4B45FFCBFF9AFAA9311E50EFA17D" author="Huang, J. P. &amp; Lin, C. - P." box="[242,456,1370,1396]" pageId="7" pageNumber="341" pagination="1149 - 1161" refId="ref16929" refString="Huang, J. P. &amp; Lin, C. - P. (2010) Diversification in subtropical mountains: Phylogeography, Pleistocene demographic expansion, and evolution of polyphenic mandibles in Taiwanese stag beetle, Lucanus formosanus. Molecular Phylogenetics and Evolution, 57, 1149 - 1161. https: // doi. org / 10.1016 / j. ympev. 2010.10.012" type="journal article" year="2010">Huang &amp; Lin 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="D6DF4D37FFCBFF9AFB89311E553DA17D" authority="Bates, 1866 in Tsai et al. 2014" box="[466,1050,1370,1396]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="swinhoei">
<emphasis id="23ABEAA6FFCBFF9AFB89311E53E6A17D" box="[466,705,1370,1396]" italics="true" pageId="7" pageNumber="341">Neolucanus swinhoei</emphasis>
Bates,
<quantity id="D6279B51FFCBFF9AF949311E5241A17D" box="[786,870,1370,1396]" metricMagnitude="1" metricUnit="m" metricValue="4.73964" pageId="7" pageNumber="341" unit="in" value="1866.0">1866 in</quantity>
<bibRefCitation id="754E4B45FFCBFF9AF937311E553DA17D" author="Tsai, C. - L. &amp; Wan, X. &amp; Yeh, W. - B." box="[876,1050,1370,1396]" pageId="7" pageNumber="341" pagination="245 - 259" refId="ref18154" refString="Tsai, C. - L., Wan, X. &amp; Yeh, W. - B. (2014) Differentiation in stag beetles, Neolucanus swinhoei complex (Coleoptera: Lucanidae): Four major lineages caused by periodical Pleistocene glaciations and separation by a mountain range. Molecular Phylogenetics and Evolution, 78, 245 - 259. https: // doi. org / 10.1016 / j. ympev. 2014.05.004" type="journal article" year="2014">
Tsai
<emphasis id="23ABEAA6FFCBFF9AF9FB311F52F6A17D" box="[928,977,1370,1396]" italics="true" pageId="7" pageNumber="341">et al</emphasis>
. 2014
</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="D6DF4D37FFCBFF9AFE7F311F5006A19E" authority="Sakaino, 1992 in Tsai &amp; Yeh 2016" authorityName="Sakaino, 1992 in Tsai &amp; Yeh" authorityYear="2016" class="Insecta" family="Lucanidae" genus="Prismognathus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="species" species="piluensis">
<emphasis id="23ABEAA6FFCBFF9AFE7F311F5512A17D" box="[1060,1077,1371,1396]" italics="true" pageId="7" pageNumber="341">P</emphasis>
.
<emphasis id="23ABEAA6FFCBFF9AFE18311E5580A17D" box="[1091,1191,1370,1396]" italics="true" pageId="7" pageNumber="341">piluensis</emphasis>
Sakaino,
<quantity id="D6279B51FFCBFF9AFF4F311E544FA17D" box="[1300,1384,1370,1396]" metricMagnitude="1" metricUnit="m" metricValue="5.05968" pageId="7" pageNumber="341" unit="in" value="1992.0">1992 in</quantity>
<bibRefCitation id="754E4B45FFCBFF9AFF35311E5006A19E" author="Tsai, C. - L. &amp; Yeh, W. - B." pageId="7" pageNumber="341" pagination="0156600" refId="ref18231" refString="Tsai, C. - L. &amp; Yeh, W. - B. (2016) Subspecific differentiation events of montane stag beetles (Coleoptera, Lucanidae) endemic to Formosa Island. PLoS One, 11 (6), e 0156600. https: // doi. org / 10.1371 / journal. pone. 0156600" type="journal article" year="2016">Tsai &amp; Yeh 2016</bibRefCitation>
</taxonomicName>
). In such cases, the definition of species or subspecies should combine genetic study and biogeography.
<taxonomicName id="D6DF4D37FFCBFF9AFACC31DB502AA1B0" authorityName="Sharp" authorityYear="1889" box="[151,269,1439,1465]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="23ABEAA6FFCBFF9AFACC31DB502AA1B0" box="[151,269,1439,1465]" italics="true" pageId="7" pageNumber="341">Miobdelus</emphasis>
</taxonomicName>
in
<collectingCountry id="69C87624FFCBFF9AFB6A31DB50A4A1B0" box="[305,387,1439,1465]" name="Taiwan" pageId="7" pageNumber="341">Taiwan</collectingCountry>
is a good topic for the study of speciation but this is outside of the scope of this paper.
</paragraph>
</subSubSection>
</treatment>
</document>
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