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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.198.3030" ID-GBIF-Dataset="02b922d7-51af-41a6-b9d3-00419437b323" ID-PMC="PMC3368254" ID-Pensoft-Pub="1313-2970-198-1" ID-PubMed="22707904" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-198-1" ModsDocOrigin="ZooKeys 198" ModsDocTitle="Serpula and Spiraserpula (Polychaeta, Serpulidae) from the Tropical Western Atlantic and Gulf of Guinea" checkinTime="1451248986690" checkinUser="pensoft" docAuthor="Bastida-Zavala, Rolando" docDate="2012" docId="995EB0A83FE669E2A6B91D8F99755F13" docLanguage="en" docName="ZooKeys 198: 1-23" docOrigin="ZooKeys 198" docSource="http://dx.doi.org/10.3897/zookeys.198.3030" docTitle="Serpula vermicularis" docType="treatment" docVersion="3" lastPageNumber="9" masterDocId="BD52B970F00CFF92FF91B24B5634EF13" masterDocTitle="Serpula and Spiraserpula (Polychaeta, Serpulidae) from the Tropical Western Atlantic and Gulf of Guinea" masterLastPageNumber="23" masterPageNumber="1" pageNumber="8" updateTime="1668153950847" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Serpula and Spiraserpula (Polychaeta, Serpulidae) from the Tropical Western Atlantic and Gulf of Guinea</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bastida-Zavala, Rolando</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>198</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>23</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.198.3030</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.198.3030</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-198-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152035877" LSID="urn:lsid:plazi:treatment:995EB0A83FE669E2A6B91D8F99755F13" httpUri="http://treatment.plazi.org/id/995EB0A83FE669E2A6B91D8F99755F13" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection pageId="7" pageNumber="8" type="nomenclature">
<paragraph pageId="7" pageNumber="8">
<taxonomicName class="Polychaeta" family="Serpulidae" genus="Serpula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serpula vermicularis" order="Sabellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="vermicularis">Serpula cf. vermicularis</taxonomicName>
Figs 1
<normalizedToken originalValue="EF2HJ">E-F2H-J</normalizedToken>
56
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="material examined">
<paragraph pageId="7" pageNumber="8">Material examined.</paragraph>
<paragraph pageId="7" pageNumber="8">
Nigeria. Five specimens (UMML 22.545), RV Pillsbury, sta. 248, Southeast of Lagos,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="4.0833335">4°05'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="5.6666665">5°40'E</geoCoordinate>
, 10-foot try-net, 33 m, May 13, 1965.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph pageId="7" pageNumber="8">Description.</paragraph>
<paragraph pageId="7" pageNumber="8">
Tubes missing. Body light brown (preserved material only). TL= 18.5 mm (n=4, r:9.4-18.5,
<normalizedToken originalValue="µ">µ</normalizedToken>
=36.6
<normalizedToken originalValue="±">+/-</normalizedToken>
10.3); THW= 1.7 mm (n=5, r:1.2-1.7,
<normalizedToken originalValue="µ">µ</normalizedToken>
=1.6
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2). Thoracic membranes and opercular peduncles of all the specimens damaged. Branchial crown with 27 radioles (n=4, r:19-27,
<normalizedToken originalValue="µ">µ</normalizedToken>
=23.8
<normalizedToken originalValue="±">+/-</normalizedToken>
3.4) left, and 25 right (n=4, r:17-25,
<normalizedToken originalValue="µ">µ</normalizedToken>
=22.5
<normalizedToken originalValue="±">+/-</normalizedToken>
3.7); lacking inter-radiolar membrane.
</paragraph>
<paragraph pageId="7" pageNumber="8">Peduncle smooth, with insertion on left (n=2) or right (n=2); lacking constriction between it and operculum, its position represented only by a slight change in color (Figs 1E, 2I). Club-shaped pseudoperculum present in all specimens.</paragraph>
<paragraph pageId="7" pageNumber="8">
Operculum with short, shallow symmetrical funnel; lacking bulbous basal part (Figs 1E, 2I). OL= 2 mm (n=4, r:1.3-2.1,
<normalizedToken originalValue="µ">µ</normalizedToken>
=1.9
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4), OD= 1.8 mm (n=4, r:1.1-1.8,
<normalizedToken originalValue="µ">µ</normalizedToken>
=1.6
<normalizedToken originalValue="±">+/-</normalizedToken>
0.3). Interradial grooves 1/4 of funnel length (Fig. 2H). Funnel with up to 39 radii (n=4, r:33-39,
<normalizedToken originalValue="µ">µ</normalizedToken>
=36.3
<normalizedToken originalValue="±">+/-</normalizedToken>
3.2) with blunt tips (Fig. 1
<normalizedToken originalValue="EF">E-F</normalizedToken>
). Opercular inner surface lacking tubercles (Figs 1
<normalizedToken originalValue="EF">E-F</normalizedToken>
, 2I).
</paragraph>
<paragraph pageId="7" pageNumber="8">
Collar thick, with ventral and dorsal lobes short. Thorax consists of seven chaetigers. Collar chaetal fascicles symmetrical with regard to size and composition unlike in
<taxonomicName class="Polychaeta" family="Serpulidae" genus="Serpula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serpula vossae" order="Sabellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="vossae">Serpula vossae</taxonomicName>
sp. n. Bayonet chaetae with two blunt-elongate teeth, distal blade smooth, lacking proximal rasp (Fig. 2J); hooded (capillary) chaetae present.
</paragraph>
<paragraph pageId="7" pageNumber="8">Thoracic membranes apparently well developed (membranes damaged), narrowing toward posterior thorax, fused ventrally, forming a short apron. Remaining six thoracic chaetigers with hooded (limbate) chaetae of two sizes; saw-shaped uncini.</paragraph>
<paragraph pageId="8" pageNumber="9">
<pageBreakToken pageId="8" pageNumber="9" start="start">Abdomen</pageBreakToken>
with anterior achaetous region. Anterior and middle abdominal chaetigers with flat-trumpet chaetae. Posterior chaetigers with
<normalizedToken originalValue="capillary">'capillary'</normalizedToken>
chaetae. Anterior and posterior uncini saw-shaped.
</paragraph>
<caption pageId="8" pageNumber="9">
<paragraph pageId="8" pageNumber="9">
Figure 5. Exploratory analysis of the number of opercular radii and body length ratio:
<taxonomicName class="Polychaeta" family="Serpulidae" genus="Serpula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serpula madrigalae" order="Sabellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="madrigalae">Serpula madrigalae</taxonomicName>
sp. n. (n= 1, only for reference),
<taxonomicName class="Polychaeta" family="Serpulidae" genus="Serpula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serpula vossae" order="Sabellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="vossae">Serpula vossae</taxonomicName>
sp. n. (n= 7), and
<taxonomicName class="Polychaeta" family="Serpulidae" genus="Serpula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serpula" order="Sabellida" pageId="8" pageNumber="9" phylum="Annelida" rank="genus">Serpula</taxonomicName>
cf. vermicularis (n= 4).
</paragraph>
</caption>
<caption pageId="8" pageNumber="9">
<paragraph pageId="8" pageNumber="9">
Figure 6. Distribution of
<taxonomicName class="Polychaeta" family="Serpulidae" genus="Serpula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serpula madrigalae" order="Sabellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="madrigalae">Serpula madrigalae</taxonomicName>
sp. n.,
<taxonomicName class="Polychaeta" family="Serpulidae" genus="Serpula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serpula vossae" order="Sabellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="vossae">Serpula vossae</taxonomicName>
sp. n., and
<taxonomicName class="Polychaeta" family="Serpulidae" genus="Serpula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serpula" order="Sabellida" pageId="8" pageNumber="9" phylum="Annelida" rank="genus">Serpula</taxonomicName>
cf. vermicularis.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="variation">
<paragraph pageId="8" pageNumber="9">Variation.</paragraph>
<paragraph pageId="8" pageNumber="9">Two specimens with a hyaline circle in radii tip (Fig. 1F). One specimen with few inconspicuous tubercles in interior funnel surface.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="distribution">
<paragraph pageId="8" pageNumber="9">Distribution.</paragraph>
<paragraph pageId="8" pageNumber="9">Nigeria, Gulf of Guinea (Fig. 6).</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="ecology">
<paragraph pageId="8" pageNumber="9">Ecology.</paragraph>
<paragraph pageId="8" pageNumber="9">Sublittoral, 33 m.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="remarks">
<paragraph pageId="8" pageNumber="9">Remarks.</paragraph>
<paragraph pageId="8" pageNumber="9">
<taxonomicName class="Polychaeta" family="Serpulidae" genus="Serpula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serpula" order="Sabellida" pageId="8" pageNumber="9" phylum="Annelida" rank="genus">Serpula</taxonomicName>
cf. vermicularis resembles the nominal species; unfortunately, the tubes of all the specimens are missing. There are some differences with the nominal species, particularly with regard to the number of radii:
<taxonomicName class="Polychaeta" family="Serpulidae" genus="Serpula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serpula" order="Sabellida" pageId="8" pageNumber="9" phylum="Annelida" rank="genus">Serpula</taxonomicName>
cf. vermicularis has 33-39 opercular radii (Fig. 1F, 2I, 5), while
<bibRefCitation author="Zibrowius, H" journalOrPublisher="Recueil des travaux de la Station Marine d'Endoume, Bulletin" pageId="10" pageNumber="11" pagination="81 - 253" title="Etude morphologique, systematique et ecologique, des Serpulidae (Annelida Polychaeta) de la region de Marseille." volume="43" year="1968">Zibrowius (1968)</bibRefCitation>
recorded specimens from Marseille with more than 40 opercular radii, and
<bibRefCitation author="Kupriyanova, EK" journalOrPublisher="Sarsia" pageId="10" pageNumber="11" pagination="203 - 236" title="Serpulidae (Annelida, Polychaeta) of the Arctic Ocean." volume="82" year="1997">Kupriyanova and Jirkov (1997)</bibRefCitation>
recorded specimens from Norway and Iceland with a mean of 50.8 opercular radii; and the proximal rasp of the bayonet chaetae:
<taxonomicName class="Polychaeta" family="Serpulidae" genus="Serpula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serpula" order="Sabellida" pageId="8" pageNumber="9" phylum="Annelida" rank="genus">Serpula</taxonomicName>
cf. vermicularis lacks a proximal rasp (Fig. 2J), while
<bibRefCitation author="Rioja, E" journalOrPublisher=", Naturales, Madrid," pageId="10" pageNumber="11" pagination="1 - 471" title="Estudio de los poliquetos de la Peninsula Iberica. Memorias de la Academia de Ciencias Exactas y Fisicas." volume="2" year="1931">Rioja (1931)</bibRefCitation>
and Kupriyanova (1999, Table 1) mentioned that their specimens have a proximal rasp.
</paragraph>
<paragraph pageId="8" pageNumber="9">
<bibRefCitation pageId="8" pageNumber="9">Zibrowius (1973)</bibRefCitation>
recorded several specimens as
<taxonomicName class="Polychaeta" family="Serpulidae" genus="Serpula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serpula vermicularis" order="Sabellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="vermicularis">Serpula vermicularis</taxonomicName>
, from Western Africa (from Angola to Morocco); unfortunately the description was too brief and did not included figures; however,
<bibRefCitation pageId="8" pageNumber="9">Zibrowius (1973)</bibRefCitation>
mentioned that the specimens that he reviewed showed considerable variation.
</paragraph>
</subSubSection>
</treatment>
</document>