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<mods:title id="B7680F08A225EB9BEBE5E4FE6DC8DACC">Records and redescription of a mygalomorph spider genus ignored for over 100 years with a new species: the genus Atmetochilus Simon, 1887 (Araneae Nemesiidae) in Thailand</mods:title>
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Kunsete &amp; Warrit
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(
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A–B, 3, 4, 5, 9A–F, 10A–B, 11C; 
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<emphasis id="23E9E371FFF7FFE1FF0FFC50899901E1" bold="true" box="[151,391,906,932]" pageId="3" pageNumber="524">Material Examined.</emphasis>
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<specimenCode id="413B9718FFF7FFE1FBC5FC2D8D110654" box="[1117,1295,1015,1041]" collectionCode="ARA-" pageId="3" pageNumber="524">ARA-2018-010</specimenCode>
<date id="652319A3FFF7FFE1FA82FC2C8D8F0654" box="[1306,1425,1014,1041]" pageId="3" pageNumber="524" value="2018-01-20">
<collectingDate id="7567E04BFFF7FFE1FA82FC2C8D8F0654" box="[1306,1425,1014,1041]" pageId="3" pageNumber="524" value="2018-01-20">20/1/2018</collectingDate>
</date>
], 
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.
</paragraph>
<paragraph id="11223F63FFF7FFE1FF5FFBE48B440638" blockId="3.[151,1437,906,2013]" pageId="3" pageNumber="524">
<emphasis id="23E9E371FFF7FFE1FF5FFBE4899A061D" bold="true" box="[199,388,1086,1112]" pageId="3" pageNumber="524">Other material.</emphasis>
<materialsCitation id="A1F5353EFFF7FFE1FE15FBE5898C0638" ID-GBIF-Occurrence="3014900306" country="Thailand" elevation="23" latitude="14.028138" location="Phra Thaen Dong Rang Forest" longLatPrecision="1" longitude="99.783195" pageId="3" pageNumber="524" specimenCount="1" specimenCount-male="1" stateProvince="Kanchanaburi">
<specimenCount id="079BF4EAFFF7FFE1FE15FBE589A3061C" box="[397,445,1087,1113]" pageId="3" pageNumber="524" type="male">1 ♂</specimenCount>
: 
<collectingCountry id="698A7FF3FFF7FFE1FE53FBE58A2D061C" box="[459,563,1087,1113]" name="Thailand" pageId="3" pageNumber="524">Thailand</collectingCountry>
: 
<collectingRegion id="D359F181FFF7FFE1FDD8FBE58B52061C" box="[576,844,1087,1113]" country="Thailand" name="Kanchanaburi" pageId="3" pageNumber="524">Kanchanaburi province</collectingRegion>
: 
<location id="144269B8FFF7FFE1FCC2FBE58CA6061C" LSID="urn:lsid:plazi:treatment:99348E75FFF7FFE8FF0FFCC48AD20747:144269B8FFF7FFE1FCC2FBE58CA6061C" box="[858,1208,1087,1113]" country="Thailand" latitude="14.028138" longLatPrecision="1" longitude="99.783195" name="Phra Thaen Dong Rang Forest" pageId="3" pageNumber="524" stateProvince="Kanchanaburi">Phra Thaen Dong Rang Forest</location>
park, 
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; 
<geoCoordinate id="74A959A4FFF7FFE1FF0FFBB8892B0638" box="[151,309,1122,1149]" degrees="99" direction="east" minutes="46" orientation="longitude" pageId="3" pageNumber="524" precision="1" seconds="59.5" value="99.783195">99°46’59.5”E</geoCoordinate>
, 
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<elevation id="9AB0D850FFF7FFE1FED8FBB9898C0638" box="[320,402,1123,1149]" metricMagnitude="1" metricUnit="m" metricValue="2.29" pageId="3" pageNumber="524" unit="m" value="22.9">22.9 m</elevation>
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</materialsCitation>
: 
<materialsCitation id="A1F5353EFFF7FFE1FE05FBB98B490638" ID-GBIF-Occurrence="3014900308" box="[413,855,1122,1149]" collectingDate="2015-07-31" collectionCode="CUMZ" country="Thailand" pageId="3" pageNumber="524" specimenCode="ARA-2017-120" specimenCount="1" specimenCount-male="1" stateProvince="Kanchanaburi">
<specimenCount id="079BF4EAFFF7FFE1FE05FBB989AC0638" box="[413,434,1123,1149]" pageId="3" pageNumber="524" type="male">♂</specimenCount>
[
<specimenCode id="413B9718FFF7FFE1FE27FBB98A6C0638" box="[447,626,1123,1149]" collectionCode="ARA-" pageId="3" pageNumber="524">ARA-2017-120</specimenCode>
<date id="652319A3FFF7FFE1FDE1FBB88AEE0638" box="[633,752,1122,1149]" pageId="3" pageNumber="524" value="2015-07-31">
<collectingDate id="7567E04BFFF7FFE1FDE1FBB88AEE0638" box="[633,752,1122,1149]" pageId="3" pageNumber="524" value="2015-07-31">31/7/2015</collectingDate>
</date>
], 
<collectionCode id="778CA7A6FFF7FFE1FC9AFBB98B490638" box="[770,855,1123,1149]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="3" pageNumber="524">CUMZ</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="59876CE8FFF7FFE1FF5FFB5D8A120680" pageId="3" pageNumber="524" type="etymology">
<paragraph id="11223F63FFF7FFE1FF5FFB5D8A120680" blockId="3.[151,1437,906,2013]" pageId="3" pageNumber="524">
<emphasis id="23E9E371FFF7FFE1FF5FFB5D895006E4" bold="true" box="[199,334,1159,1185]" pageId="3" pageNumber="524">Etymology.</emphasis>
Named after Mr. Chaowalit Songsangchote, a great spider lover, the best travelling companion, the best of our laboratory supporters.
</paragraph>
</subSubSection>
<subSubSection id="59876CE8FFF7FFE1FF5FFB148CDC07A0" pageId="3" pageNumber="524" type="diagnosis">
<paragraph id="11223F63FFF7FFE1FF5FFB148CDC07A0" blockId="3.[151,1437,906,2013]" pageId="3" pageNumber="524">
<emphasis id="23E9E371FFF7FFE1FF5FFB14895C06AD" bold="true" box="[199,322,1230,1256]" pageId="3" pageNumber="524">Diagnosis.</emphasis>
Females are most similar to those of 
<taxonomicName id="D69D44E0FFF7FFE1FD70FB158B5506AC" authorityName="Simon" authorityYear="1887" box="[744,843,1231,1257]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="23E9E371FFF7FFE1FD70FB158B5506AC" box="[744,843,1231,1257]" italics="true" pageId="3" pageNumber="524">A. fossor</emphasis>
</taxonomicName>
and can be distinguished by the carapace being dark brown (not red), no spines on the prolateral palpal patella of palp, or legs I, II (present in 
<taxonomicName id="D69D44E0FFF7FFE1FB09FB298CE80748" authorityName="Simon" authorityYear="1887" box="[1169,1270,1267,1293]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="23E9E371FFF7FFE1FB09FB298CE80748" box="[1169,1270,1267,1293]" italics="true" pageId="3" pageNumber="524">A. fossor</emphasis>
</taxonomicName>
). Females differ from those of 
<taxonomicName id="D69D44E0FFF7FFE1FEC6FACD89C60775" authorityName="Pocock, Tenasserim" authorityYear="1900" box="[350,472,1303,1329]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="atriceps">
<emphasis id="23E9E371FFF7FFE1FEC6FACD89C60775" box="[350,472,1303,1329]" italics="true" pageId="3" pageNumber="524">A. atriceps</emphasis>
</taxonomicName>
by metatarsus I being without scopula for its length and that on metatarsus II only in its distal half (for scopula and spine characters comparisons, see 
<figureCitation id="89A623E6FFF7FFE1FCE0FAE18BA20710" box="[888,956,1339,1365]" captionStart="FIGURE 9" captionStartId="16.[151,250,929,953]" captionTargetBox="[151,1436,181,905]" captionTargetId="figure-24@16.[151,1436,181,905]" captionTargetPageId="16" captionText="FIGURE 9 A–L. Atmetochilus, prolateral views of patella and femur of palp, legs I and II; A–C. A. songsangchotei sp. nov. male holotype (ARA-2017-116); D–F. A. songsangchotei sp. nov. female paratype (ARA-2017-117); G–I. A. fossor female neotype (ARA-2018-132); J–L. A. atriceps female (ARA-2018-173)." figureDoi="http://doi.org/10.5281/zenodo.4397102" httpUri="https://zenodo.org/record/4397102/files/figure.png" pageId="3" pageNumber="524">Figs 9</figureCitation>
D–L and 10A–F). Males differ those of 
<taxonomicName id="D69D44E0FFF7FFE1FA1CFAE188E0073C" authorityName="Zonstein and Marusik" authorityYear="2016" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="koponeni">
<emphasis id="23E9E371FFF7FFE1FA1CFAE188E0073C" italics="true" pageId="3" pageNumber="524">A. koponeni</emphasis>
</taxonomicName>
, 
<taxonomicName id="D69D44E0FFF7FFE1FE93FA858992073C" authorityName="Zonstein and Marusik" authorityYear="2016" box="[267,396,1375,1401]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="lehtineni">
<emphasis id="23E9E371FFF7FFE1FE93FA858992073C" box="[267,396,1375,1401]" italics="true" pageId="3" pageNumber="524">A. lehtineni</emphasis>
</taxonomicName>
and 
<taxonomicName id="D69D44E0FFF7FFE1FE5AFA858A7D073D" authorityName="Zonstein and Marusik" authorityYear="2016" box="[450,611,1375,1401]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="23E9E371FFF7FFE1FE5AFA858A7D073D" box="[450,611,1375,1401]" italics="true" pageId="3" pageNumber="524">A. sumatranus</emphasis>
</taxonomicName>
by the tip of tibial spur on leg I having three straight adjacent megaspines grouped in one row on spur and differ from those of 
<taxonomicName id="D69D44E0FFF7FFE1FD46FA598B7D07D8" authorityName="Zonstein and Marusik" authorityYear="2016" box="[734,867,1411,1437]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="koponeni">
<emphasis id="23E9E371FFF7FFE1FD46FA598B7D07D8" box="[734,867,1411,1437]" italics="true" pageId="3" pageNumber="524">A. koponeni</emphasis>
</taxonomicName>
, 
<taxonomicName id="D69D44E0FFF7FFE1FCE8FA598BEE07D8" authorityName="Zonstein and Marusik" authorityYear="2016" box="[880,1008,1411,1437]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="lehtineni">
<emphasis id="23E9E371FFF7FFE1FCE8FA598BEE07D8" box="[880,1008,1411,1437]" italics="true" pageId="3" pageNumber="524">A. lehtineni</emphasis>
</taxonomicName>
and 
<taxonomicName id="D69D44E0FFF7FFE1FBBEFA598CD907D9" authorityName="Zonstein and Marusik" authorityYear="2016" box="[1062,1223,1411,1437]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="23E9E371FFF7FFE1FBBEFA598CD907D9" box="[1062,1223,1411,1437]" italics="true" pageId="3" pageNumber="524">A. sumatranus</emphasis>
</taxonomicName>
by the long curved embolus. Moreover, female spermathecae differ from those of 
<taxonomicName id="D69D44E0FFF7FFE1FCCBFA7D8BC70784" authorityName="Zonstein and Marusik" authorityYear="2016" box="[851,985,1447,1473]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="koponeni">
<emphasis id="23E9E371FFF7FFE1FCCBFA7D8BC70784" box="[851,985,1447,1473]" italics="true" pageId="3" pageNumber="524">A. koponeni</emphasis>
</taxonomicName>
and 
<taxonomicName id="D69D44E0FFF7FFE1FB96FA7D8C910784" authorityName="Zonstein and Marusik" authorityYear="2016" box="[1038,1167,1447,1473]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="lehtineni">
<emphasis id="23E9E371FFF7FFE1FB96FA7D8C910784" box="[1038,1167,1447,1473]" italics="true" pageId="3" pageNumber="524">A. lehtineni</emphasis>
</taxonomicName>
by the head of the spermathecae being relatively longer, semicircular, dome-shaped with strong subdistal constriction.
</paragraph>
</subSubSection>
<subSubSection id="59876CE8FFF7FFE8FF5FFA348B47066F" lastPageId="10" lastPageNumber="531" pageId="3" pageNumber="524" type="description">
<paragraph id="11223F63FFF7FFE1FF5FFA348B74044C" blockId="3.[151,1437,906,2013]" box="[199,874,1518,1546]" pageId="3" pageNumber="524">
<emphasis id="23E9E371FFF7FFE1FF5FFA348947044D" bold="true" box="[199,345,1518,1544]" pageId="3" pageNumber="524">Description.</emphasis>
Male (
<typeStatus id="CE2681C1FFF7FFE1FE30FA358A0F044F" box="[424,529,1519,1546]" pageId="3" pageNumber="524" type="holotype">holotype</typeStatus>
): Body (
<figureCitation id="89A623E6FFF7FFE1FDE8FA358AD6044C" box="[624,712,1519,1545]" captionStart="FIGURE 3" captionStartId="7.[151,250,1955,1979]" captionTargetBox="[187,1402,186,1931]" captionTargetId="figure-18@7.[186,1402,181,1931]" captionTargetPageId="7" captionText="FIGURE 3 A–D. Atmetochilus songsangchotei sp. nov. male holotype (ARA-2017-116) habitus; A (left). dorsal habitus, A (right). ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium." figureDoi="http://doi.org/10.5281/zenodo.4397088" httpUri="https://zenodo.org/record/4397088/files/figure.png" pageId="3" pageNumber="524">Fig. 3A</figureCitation>
) length 22.26.
</paragraph>
<paragraph id="11223F63FFF7FFE1FF5FF9C98AB70414" blockId="3.[151,1437,906,2013]" pageId="3" pageNumber="524">Color in nature: Carapace, chelicerae, legs and abdomen black, abdomen covered by pale white spots and ventral abdomen and spinnerets pale white overall.</paragraph>
<paragraph id="11223F63FFF7FFE1FF55F9818B0404DC" blockId="3.[151,1437,906,2013]" pageId="3" pageNumber="524">Color in alcohol: Carapace, chelicerae and legs brown, abdomen dorsally pale gray and covered with pale white spots; ventral abdomen and spinnerets pale white overall.</paragraph>
<paragraph id="11223F63FFF7FFE6FF5FF97988F00290" blockId="3.[151,1437,906,2013]" lastBlockId="4.[151,1437,150,573]" lastPageId="4" lastPageNumber="525" pageId="3" pageNumber="524">
Carapace (
<figureCitation id="89A623E6FFF7FFE1FED9F979898304F8" box="[321,413,1699,1725]" captionStart="FIGURE 3" captionStartId="7.[151,250,1955,1979]" captionTargetBox="[187,1402,186,1931]" captionTargetId="figure-18@7.[186,1402,181,1931]" captionTargetPageId="7" captionText="FIGURE 3 A–D. Atmetochilus songsangchotei sp. nov. male holotype (ARA-2017-116) habitus; A (left). dorsal habitus, A (right). ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium." figureDoi="http://doi.org/10.5281/zenodo.4397088" httpUri="https://zenodo.org/record/4397088/files/figure.png" pageId="3" pageNumber="524">Fig. 3B</figureCitation>
): 10.50 long, 9.00 wide. Chelicerae 4.30 long. Eyes (
<figureCitation id="89A623E6FFF7FFE1FB90F9798C7A04F8" box="[1032,1124,1699,1725]" captionStart="FIGURE 3" captionStartId="7.[151,250,1955,1979]" captionTargetBox="[187,1402,186,1931]" captionTargetId="figure-18@7.[186,1402,181,1931]" captionTargetPageId="7" captionText="FIGURE 3 A–D. Atmetochilus songsangchotei sp. nov. male holotype (ARA-2017-116) habitus; A (left). dorsal habitus, A (right). ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium." figureDoi="http://doi.org/10.5281/zenodo.4397088" httpUri="https://zenodo.org/record/4397088/files/figure.png" pageId="3" pageNumber="524">Fig. 3C</figureCitation>
): tubercle 2 rows with low mound, 4 eyes in each row; anterior row procurved; posterior row slightly recurved, almost straight. Eye size and interdistances: AME 0.36, AME-AME 0.30, ALE 0.45, ALE-ALE 1.20, PME 0.48, PME-PME 0.75, PLE 0.45, PLE-PLE 1.23, AME-ALE 0.21, AME-PME 0.18, AME-PLE 0.36, ALE-PME 0.36, ALE-PLE 0.30, PME-PLE 0 (contiguous). Eye to fovea 6.30. Fovea (
<figureCitation id="89A623E6FFF7FFE1FDCEF8E98AB10508" box="[598,687,1843,1869]" captionStart="FIGURE 3" captionStartId="7.[151,250,1955,1979]" captionTargetBox="[187,1402,186,1931]" captionTargetId="figure-18@7.[186,1402,181,1931]" captionTargetPageId="7" captionText="FIGURE 3 A–D. Atmetochilus songsangchotei sp. nov. male holotype (ARA-2017-116) habitus; A (left). dorsal habitus, A (right). ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium." figureDoi="http://doi.org/10.5281/zenodo.4397088" httpUri="https://zenodo.org/record/4397088/files/figure.png" pageId="3" pageNumber="524">Fig. 3B</figureCitation>
): strongly procurved to U-shaped. 3 rows of setae on fovea. Rastellum of about 30 robust down-curved setae on anterior margin of chelicerae. Cheliceral furrow with 9 promarginal teeth in one row, mesobasal denticles present. Fang ventrally smooth without teeth. Stridulating strikers absent. Intercheliceral tumescence absent. Labium (
<figureCitation id="89A623E6FFF7FFE1FD11F8458AFB05FC" box="[649,741,1951,1977]" captionStart="FIGURE 3" captionStartId="7.[151,250,1955,1979]" captionTargetBox="[187,1402,186,1931]" captionTargetId="figure-18@7.[186,1402,181,1931]" captionTargetPageId="7" captionText="FIGURE 3 A–D. Atmetochilus songsangchotei sp. nov. male holotype (ARA-2017-116) habitus; A (left). dorsal habitus, A (right). ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium." figureDoi="http://doi.org/10.5281/zenodo.4397088" httpUri="https://zenodo.org/record/4397088/files/figure.png" pageId="3" pageNumber="524">Fig. 3D</figureCitation>
): 1.50 long, 1.05 wide, without cuspules. Maxillae (
<figureCitation id="89A623E6FFF7FFE1FAACF8458D9105FC" box="[1332,1423,1951,1977]" captionStart="FIGURE 3" captionStartId="7.[151,250,1955,1979]" captionTargetBox="[187,1402,186,1931]" captionTargetId="figure-18@7.[186,1402,181,1931]" captionTargetPageId="7" captionText="FIGURE 3 A–D. Atmetochilus songsangchotei sp. nov. male holotype (ARA-2017-116) habitus; A (left). dorsal habitus, A (right). ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium." figureDoi="http://doi.org/10.5281/zenodo.4397088" httpUri="https://zenodo.org/record/4397088/files/figure.png" pageId="3" pageNumber="524">Fig. 3D</figureCitation>
): 4.00 long, 2.00 wide; serrula absent; 20 tiny cuspules each on posterior lobe, absent on heel. Sternum (
<figureCitation id="89A623E6FFF7FFE1FAABF8198D900598" box="[1331,1422,1987,2013]" captionStart="FIGURE 1" captionStartId="5.[151,250,1942,1966]" captionTargetBox="[281,1307,183,1919]" captionTargetId="figure-18@5.[281,1307,181,1919]" captionTargetPageId="5" captionText="FIGURE 1A–F. Atmetochilus and Damarchus bifidus. Sternum.A. Atmetochilus songsangchotei sp. nov. male holotype (ARA- 2017-116); B. A. songsangchotei sp. nov. female paratype (ARA-2017-117); C. A. fossor female neotype (ARA-2018-132); D. A. atriceps female (ARA-2018-173); E. A. atriceps female lectotype; F. Damarchus bifidus type specimen." figureDoi="http://doi.org/10.5281/zenodo.4397084" httpUri="https://zenodo.org/record/4397084/files/figure.png" pageId="3" pageNumber="524">Fig. 1A</figureCitation>
): 5.00 long, 5.00 wide, posterior and medial sternal sigilla centrally confluent, anterior sigilla small, oval, on upper margin.
</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FFF058A480358" blockId="4.[151,1437,150,573]" pageId="4" pageNumber="525">Spination: No spines on leg tarsi. Leg I: femur, p2; patella, 0; tibia, p4 r1; metatarsus, p1 v4 with small group of short velvet-like setae on top margin.</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FFEFD8C9703E8" blockId="4.[151,1437,150,573]" pageId="4" pageNumber="525">
Tibial spur and metatarsus (
<figureCitation id="89A623E6FFF0FFE6FE64FEFD8A200304" box="[508,574,295,321]" captionStart="FIGURE 4" captionStartId="8.[151,250,1725,1749]" captionTargetBox="[157,1430,192,1701]" captionTargetId="figure-24@8.[151,1436,181,1701]" captionTargetPageId="8" captionText="FIGURE 4 A–H. Atmetochilus songsangchotei sp. nov., male. A–F. Holotype (ARA-2017-116). A–C. Palpal tibia and tarsus, retrolateral, ventral and prolateral view; D–F. Tibia, metatarsus and tarsus I showing tibial spur, prolateral, retroventral and retrolateral views; G–H. Male paratype (ARA-2017-120), tibial spur, retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.4397090" httpUri="https://zenodo.org/record/4397090/files/figure.png" pageId="4" pageNumber="525">Fig. 4</figureCitation>
D–H): leg I with dense group of short dark setae at ventral base of metatarsus with subbasal bulge, tibial spur, ventral, opposing tip distally unsclerotized; tibial spur stalk- shaped with 3 straight adjacent short spines (2–3 times longer than wide) on wide digitiform or long triangular spines tip of the stalk (
<figureCitation id="89A623E6FFF0FFE6FAF7FEB588BB03E8" captionStart="FIGURE 4" captionStartId="8.[151,250,1725,1749]" captionTargetBox="[157,1430,192,1701]" captionTargetId="figure-24@8.[151,1436,181,1701]" captionTargetPageId="8" captionText="FIGURE 4 A–H. Atmetochilus songsangchotei sp. nov., male. A–F. Holotype (ARA-2017-116). A–C. Palpal tibia and tarsus, retrolateral, ventral and prolateral view; D–F. Tibia, metatarsus and tarsus I showing tibial spur, prolateral, retroventral and retrolateral views; G–H. Male paratype (ARA-2017-120), tibial spur, retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.4397090" httpUri="https://zenodo.org/record/4397090/files/figure.png" pageId="4" pageNumber="525">Fig. 4</figureCitation>
A–C). Tibia with 2 spines on ventral, one spine on prolateral and 3 spines on retrolateral.
</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FFE6D8BD50078" blockId="4.[151,1437,150,573]" pageId="4" pageNumber="525">
Leg II: tibia and patella without spines. metatarsus scopulate, v6 p1, tarsus scopula covers entire ventral, without spines. Leg III: tibia with 12 spines including, v1 r5 p2 d1; patella, r3, metatarsus with 10 spines, v5 r2 p3, tarsus scopulate ventrally. Leg IV: tibia with 12 spines, v5 r2 p5, metatarsus with 11 spines, v3 r5 p3, tarsus scopulate ventrally. Preening combs entirely absent. Legs (
<tableCitation id="5C1F0AD8FFF0FFE6FD20FDF98B120078" box="[696,780,547,573]" captionStart="TABLE 1" captionStartId="4.[151,239,620,644]" captionText="TABLE 1. Atmetochilus songsangchotei sp. nov., holotype male" pageId="4" pageNumber="525">Table 1</tableCitation>
), formula: 1243.
</paragraph>
<caption id="45E26FEBFFF0FFE6FF0FFDB68B5400C1" box="[151,842,620,645]" pageId="4" pageNumber="525">
<paragraph id="11223F63FFF0FFE6FF0FFDB68B5400C1" blockId="4.[151,842,620,645]" box="[151,842,620,645]" pageId="4" pageNumber="525">
<emphasis id="23E9E371FFF0FFE6FF0FFDB6891400C1" bold="true" box="[151,266,620,644]" pageId="4" pageNumber="525">TABLE 1.</emphasis>
<taxonomicName id="D69D44E0FFF0FFE6FE89FDB68A5700C1" authority="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit, 2020" authorityName="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit" authorityYear="2020" box="[273,585,620,644]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="525" phylum="Arthropoda" rank="species" species="songsangchotei" status="sp. nov.">
<emphasis id="23E9E371FFF0FFE6FE89FDB68A5700C1" box="[273,585,620,644]" italics="true" pageId="4" pageNumber="525">Atmetochilus songsangchotei</emphasis>
</taxonomicName>
<emphasis id="23E9E371FFF0FFE6FDD7FDB78ABB00C0" bold="true" box="[591,677,621,645]" pageId="4" pageNumber="525">
<taxonomicNameLabel id="38DA5E0AFFF0FFE6FDD7FDB78ABB00C0" box="[591,677,621,645]" pageId="4" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male
</paragraph>
</caption>
<paragraph id="11223F63FFF0FFE6FEC5FD4F8D320133" pageId="4" pageNumber="525">
<table id="639DCDC3FFF0001DFF3EFD4F8D320133" box="[166,1324,661,886]" colsContinueIn="16.[166,1325,1330,1555]" gridcols="7" gridrows="6" pageId="4" pageNumber="525">
<tr id="AFAD3D21FFF0001DFF3EFD4F8D3200E8" box="[166,1324,661,685]" gridrow="0" pageId="4" pageNumber="525" rowspan-0="1">
<th id="EC7C545DFFF0001DFEC5FD4F898100E8" box="[349,415,661,685]" gridcol="1" gridrow="0" pageId="4" pageNumber="525">Femur</th>
<th id="EC7C545DFFF0001DFD8DFD4F8A4400E8" box="[533,602,661,685]" gridcol="2" gridrow="0" pageId="4" pageNumber="525">Patella</th>
<th id="EC7C545DFFF0001DFD54FD4F8B1F00E8" box="[716,769,661,685]" gridcol="3" gridrow="0" pageId="4" pageNumber="525">Tibia</th>
<th id="EC7C545DFFF0001DFC1CFD4F8BED00E8" box="[900,1011,661,685]" gridcol="4" gridrow="0" pageId="4" pageNumber="525">Metatarsus</th>
<th id="EC7C545DFFF0001DFBA4FD4F8C6000E8" box="[1084,1150,661,685]" gridcol="5" gridrow="0" pageId="4" pageNumber="525">Tarsus</th>
<th id="EC7C545DFFF0001DFB6BFD4F8D3200E8" box="[1267,1324,661,685]" gridcol="6" gridrow="0" pageId="4" pageNumber="525">Total</th>
</tr>
<tr id="AFAD3D21FFF0001DFF3EFD678D320090" box="[166,1324,701,725]" gridrow="1" pageId="4" pageNumber="525">
<th id="EC7C545DFFF0001DFF3EFD6788F00090" box="[166,238,701,725]" gridcol="0" gridrow="1" pageId="4" pageNumber="525">Palp</th>
<td id="EC7C545DFFF0001DFEC5FD6789810090" box="[349,415,701,725]" gridcol="1" gridrow="1" pageId="4" pageNumber="525">4.50</td>
<td id="EC7C545DFFF0001DFD8DFD678A440090" box="[533,602,701,725]" gridcol="2" gridrow="1" pageId="4" pageNumber="525">2.22</td>
<td id="EC7C545DFFF0001DFD54FD678B1F0090" box="[716,769,701,725]" gridcol="3" gridrow="1" pageId="4" pageNumber="525">3.66</td>
<td id="EC7C545DFFF0001DFC1CFD678BED0090" box="[900,1011,701,725]" gridcol="4" gridrow="1" pageId="4" pageNumber="525">-</td>
<td id="EC7C545DFFF0001DFBA4FD678C600090" box="[1084,1150,701,725]" gridcol="5" gridrow="1" pageId="4" pageNumber="525">1.60</td>
<td id="EC7C545DFFF0001DFB6BFD678D320090" box="[1267,1324,701,725]" gridcol="6" gridrow="1" pageId="4" pageNumber="525">11.98</td>
</tr>
<tr id="AFAD3D21FFF0001DFF3EFD3F8D3200BB" box="[166,1324,741,766]" gridrow="2" pageId="4" pageNumber="525">
<th id="EC7C545DFFF0001DFF3EFD3F88F000BB" box="[166,238,741,766]" gridcol="0" gridrow="2" pageId="4" pageNumber="525">Leg I</th>
<td id="EC7C545DFFF0001DFEC5FD3F898100BB" box="[349,415,741,766]" gridcol="1" gridrow="2" pageId="4" pageNumber="525">8.40</td>
<td id="EC7C545DFFF0001DFD8DFD3F8A4400BB" box="[533,602,741,766]" gridcol="2" gridrow="2" pageId="4" pageNumber="525">4.20</td>
<td id="EC7C545DFFF0001DFD54FD3F8B1F00BB" box="[716,769,741,766]" gridcol="3" gridrow="2" pageId="4" pageNumber="525">6.00</td>
<td id="EC7C545DFFF0001DFC1CFD3F8BED00BB" box="[900,1011,741,766]" gridcol="4" gridrow="2" pageId="4" pageNumber="525">6.30</td>
<td id="EC7C545DFFF0001DFBA4FD3F8C6000BB" box="[1084,1150,741,766]" gridcol="5" gridrow="2" pageId="4" pageNumber="525">3.66</td>
<td id="EC7C545DFFF0001DFB6BFD3F8D3200BB" box="[1267,1324,741,766]" gridcol="6" gridrow="2" pageId="4" pageNumber="525">28.56</td>
</tr>
<tr id="AFAD3D21FFF0001DFF3EFCD78D320163" box="[166,1324,781,806]" gridrow="3" pageId="4" pageNumber="525">
<th id="EC7C545DFFF0001DFF3EFCD788F00163" box="[166,238,781,806]" gridcol="0" gridrow="3" pageId="4" pageNumber="525">Leg II</th>
<td id="EC7C545DFFF0001DFEC5FCD789810163" box="[349,415,781,806]" gridcol="1" gridrow="3" pageId="4" pageNumber="525">6.18</td>
<td id="EC7C545DFFF0001DFD8DFCD78A440163" box="[533,602,781,806]" gridcol="2" gridrow="3" pageId="4" pageNumber="525">3.90</td>
<td id="EC7C545DFFF0001DFD54FCD78B1F0163" box="[716,769,781,806]" gridcol="3" gridrow="3" pageId="4" pageNumber="525">4.74</td>
<td id="EC7C545DFFF0001DFC1CFCD78BED0163" box="[900,1011,781,806]" gridcol="4" gridrow="3" pageId="4" pageNumber="525">4.50</td>
<td id="EC7C545DFFF0001DFBA4FCD78C600163" box="[1084,1150,781,806]" gridcol="5" gridrow="3" pageId="4" pageNumber="525">2.70</td>
<td id="EC7C545DFFF0001DFB6BFCD78D320163" box="[1267,1324,781,806]" gridcol="6" gridrow="3" pageId="4" pageNumber="525">22.02</td>
</tr>
<tr id="AFAD3D21FFF0001DFF3EFCEF8D32010B" box="[166,1324,821,846]" gridrow="4" pageId="4" pageNumber="525">
<th id="EC7C545DFFF0001DFF3EFCEF88F0010B" box="[166,238,821,846]" gridcol="0" gridrow="4" pageId="4" pageNumber="525">Leg III</th>
<td id="EC7C545DFFF0001DFEC5FCEF8981010B" box="[349,415,821,846]" gridcol="1" gridrow="4" pageId="4" pageNumber="525">4.50</td>
<td id="EC7C545DFFF0001DFD8DFCEF8A44010B" box="[533,602,821,846]" gridcol="2" gridrow="4" pageId="4" pageNumber="525">2.10</td>
<td id="EC7C545DFFF0001DFD54FCEF8B1F010B" box="[716,769,821,846]" gridcol="3" gridrow="4" pageId="4" pageNumber="525">2.76</td>
<td id="EC7C545DFFF0001DFC1CFCEF8BED010B" box="[900,1011,821,846]" gridcol="4" gridrow="4" pageId="4" pageNumber="525">3.60</td>
<td id="EC7C545DFFF0001DFBA4FCEF8C60010B" box="[1084,1150,821,846]" gridcol="5" gridrow="4" pageId="4" pageNumber="525">1.92</td>
<td id="EC7C545DFFF0001DFB6BFCEF8D32010B" box="[1267,1324,821,846]" gridcol="6" gridrow="4" pageId="4" pageNumber="525">14.88</td>
</tr>
<tr id="AFAD3D21FFF0001DFF3EFC848D320133" box="[166,1324,862,886]" gridrow="5" pageId="4" pageNumber="525">
<th id="EC7C545DFFF0001DFF3EFC8488F00133" box="[166,238,862,886]" gridcol="0" gridrow="5" pageId="4" pageNumber="525">Leg IV</th>
<td id="EC7C545DFFF0001DFEC5FC8489810133" box="[349,415,862,886]" gridcol="1" gridrow="5" pageId="4" pageNumber="525">5.82</td>
<td id="EC7C545DFFF0001DFD8DFC848A440133" box="[533,602,862,886]" gridcol="2" gridrow="5" pageId="4" pageNumber="525">2.52</td>
<td id="EC7C545DFFF0001DFD54FC848B1F0133" box="[716,769,862,886]" gridcol="3" gridrow="5" pageId="4" pageNumber="525">4.20</td>
<td id="EC7C545DFFF0001DFC1CFC848BED0133" box="[900,1011,862,886]" gridcol="4" gridrow="5" pageId="4" pageNumber="525">5.10</td>
<td id="EC7C545DFFF0001DFBA4FC848C600133" box="[1084,1150,862,886]" gridcol="5" gridrow="5" pageId="4" pageNumber="525">2.10</td>
<td id="EC7C545DFFF0001DFB6BFC848D320133" box="[1267,1324,862,886]" gridcol="6" gridrow="5" pageId="4" pageNumber="525">19.74</td>
</tr>
</table>
</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FFC7F89F101A1" blockId="4.[151,1437,933,2004]" pageId="4" pageNumber="525">Scopula: entire on tarsi I–IV and metatarsi I–II, not divided, absent on cymbium, metatarsi III–IV, tibiae I–IV, patellae I–IV and femora I–IV.</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FFC378B0E066E" blockId="4.[151,1437,933,2004]" pageId="4" pageNumber="525">Trichobothria: one zigzag row with 12–18 per row on tarsi I–IV, one row with 6–10 on metatarsi I–IV and two rows of 6–10 on tibiae I, III; none on patellae or femora.</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FFBEF8BF7060A" blockId="4.[151,1437,933,2004]" box="[199,1001,1077,1104]" pageId="4" pageNumber="525">Claws: S-shaped row of teeth with 10 teeth per row on all paired claws.</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FFB838B0F06D2" blockId="4.[151,1437,933,2004]" pageId="4" pageNumber="525">
Palp (
<figureCitation id="89A623E6FFF0FFE6FE90FB8389540631" box="[264,330,1113,1140]" captionStart="FIGURE 4" captionStartId="8.[151,250,1725,1749]" captionTargetBox="[157,1430,192,1701]" captionTargetId="figure-24@8.[151,1436,181,1701]" captionTargetPageId="8" captionText="FIGURE 4 A–H. Atmetochilus songsangchotei sp. nov., male. A–F. Holotype (ARA-2017-116). A–C. Palpal tibia and tarsus, retrolateral, ventral and prolateral view; D–F. Tibia, metatarsus and tarsus I showing tibial spur, prolateral, retroventral and retrolateral views; G–H. Male paratype (ARA-2017-120), tibial spur, retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.4397090" httpUri="https://zenodo.org/record/4397090/files/figure.png" pageId="4" pageNumber="525">Fig. 4</figureCitation>
A–C); cymbium 1.60 with long setae on tip and without spines. Bulb 1.25 spherical, narrow on tip, embolus 1.65 elongated, slightly curved and sharp at tip.
</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FFB7B8AF2069A" blockId="4.[151,1437,933,2004]" pageId="4" pageNumber="525">Spinnerets: PMS: 1.25 long, 0.20 diameter, PLS: 4.50 long, 0.40 diameter, length of basal, medial and apical segments 1.50, 1.00, 2.00; apical segment digitiform.</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FFB3389120709" blockId="4.[151,1437,933,2004]" pageId="4" pageNumber="525">
Variation. The male tibial spur varies in the number of large megaspines on the tip of the spur. Another specimen (
<specimenCount id="079BF4EAFFF0FFE6FF4FFAD488F2076D" box="[215,236,1294,1320]" pageId="4" pageNumber="525" type="male">♂</specimenCount>
[ARA-2017-120 31/7/58] has two large megaspines on the tibial spur on left leg I (
<figureCitation id="89A623E6FFF0FFE6FB17FAD78CCF076D" box="[1167,1233,1293,1320]" captionStart="FIGURE 4" captionStartId="8.[151,250,1725,1749]" captionTargetBox="[157,1430,192,1701]" captionTargetId="figure-24@8.[151,1436,181,1701]" captionTargetPageId="8" captionText="FIGURE 4 A–H. Atmetochilus songsangchotei sp. nov., male. A–F. Holotype (ARA-2017-116). A–C. Palpal tibia and tarsus, retrolateral, ventral and prolateral view; D–F. Tibia, metatarsus and tarsus I showing tibial spur, prolateral, retroventral and retrolateral views; G–H. Male paratype (ARA-2017-120), tibial spur, retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.4397090" httpUri="https://zenodo.org/record/4397090/files/figure.png" pageId="4" pageNumber="525">Fig. 4</figureCitation>
G–H) but three on right leg I.
</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FFA8F8BCB0735" blockId="4.[151,1437,933,2004]" box="[199,981,1365,1392]" pageId="4" pageNumber="525">
Female (
<typeStatus id="CE2681C1FFF0FFE6FEB0FA8C89910735" box="[296,399,1366,1392]" pageId="4" pageNumber="525" type="paratype">paratype</typeStatus>
). Body (
<figureCitation id="89A623E6FFF0FFE6FE76FA8F8A560735" box="[494,584,1365,1392]" captionStart="FIGURE 5" captionStartId="9.[151,250,1976,2000]" captionTargetBox="[220,1392,189,1953]" captionTargetId="figure-18@9.[196,1392,181,1953]" captionTargetPageId="9" captionText="FIGURE 5 A–D. Atmetochilus songsangchotei sp. nov. female paratype (ARA-2017-117) habitus; A (left) dorsal habitus, A (right) ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium." figureDoi="http://doi.org/10.5281/zenodo.4397092" httpUri="https://zenodo.org/record/4397092/files/figure.png" pageId="4" pageNumber="525">Fig. 5A</figureCitation>
). length 28.09±4.20 (23.46–34.79).
</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FFAA38B2107F2" blockId="4.[151,1437,933,2004]" pageId="4" pageNumber="525">Color in nature: Carapace, chelicerae, legs and abdomen black; abdomen covered by pale white spots and ventral abdomen and spinnerets entirely with pale white overall.</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FFA1B8A8A07BA" blockId="4.[151,1437,933,2004]" pageId="4" pageNumber="525">Color in alcohol: Carapace, chelicerae and legs dark brown, abdomen dorsally black with pale white spots and pale gray on ventral abdomen and spinnerets.</paragraph>
<paragraph id="11223F63FFF0FFE6FF5FF9D08B010596" blockId="4.[151,1437,933,2004]" pageId="4" pageNumber="525">
Carapace: (
<figureCitation id="89A623E6FFF0FFE6FED0F9D389BC0461" box="[328,418,1545,1572]" captionStart="FIGURE 5" captionStartId="9.[151,250,1976,2000]" captionTargetBox="[220,1392,189,1953]" captionTargetId="figure-18@9.[196,1392,181,1953]" captionTargetPageId="9" captionText="FIGURE 5 A–D. Atmetochilus songsangchotei sp. nov. female paratype (ARA-2017-117) habitus; A (left) dorsal habitus, A (right) ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium." figureDoi="http://doi.org/10.5281/zenodo.4397092" httpUri="https://zenodo.org/record/4397092/files/figure.png" pageId="4" pageNumber="525">Fig. 5B</figureCitation>
): 10.33±1.19 (9.00–11.88) long, 7.94±0.73 (7.2–8.75) wide. Chelicerae 4.77±0.68 (4.136.00) long. Eyes (
<figureCitation id="89A623E6FFF0FFE6FEF1F9F789DB040D" box="[361,453,1581,1608]" captionStart="FIGURE 5" captionStartId="9.[151,250,1976,2000]" captionTargetBox="[220,1392,189,1953]" captionTargetId="figure-18@9.[196,1392,181,1953]" captionTargetPageId="9" captionText="FIGURE 5 A–D. Atmetochilus songsangchotei sp. nov. female paratype (ARA-2017-117) habitus; A (left) dorsal habitus, A (right) ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium." figureDoi="http://doi.org/10.5281/zenodo.4397092" httpUri="https://zenodo.org/record/4397092/files/figure.png" pageId="4" pageNumber="525">Fig. 5C</figureCitation>
): tubercle with 2 rows on low mound, 4 eyes on each row. Anterior row procurved, posterior row slightly recurved. Eye sizes and interdistances: AME 0.42±0.03 (0.39–0.48), AME-AME 0.15, ALE 0.65±0.05 (0.60–0.75), ALE-ALE 1.22±0.23 (0.90–1.44), PME 0.50±0.04 (0.45–0.57), PME-PME 0.75±0.24 (0.34–0.90), PLE 0.67±0.38 (0.45–1.44), PLE-PLE 1.42±0.13 (1.20–1.50), AME-ALE 0.25±0.16 (0.09–0.51), AME-PME 0.15±0.04 (0.12–0.21), AME-PLE 0.35±0.09 (0.24–0.45), ALE-PME 0.34±0.07 (0.30–0.45), ALE-PLE 0.38±0.12 (0.30–0.57), PME-PLE 0 (contiguous). Eye to fovea 6.58±0.81 (5.5–7.5). Fovea (
<figureCitation id="89A623E6FFF0FFE6FB3DF93B8CE504B9" box="[1189,1275,1761,1788]" captionStart="FIGURE 5" captionStartId="9.[151,250,1976,2000]" captionTargetBox="[220,1392,189,1953]" captionTargetId="figure-18@9.[196,1392,181,1953]" captionTargetPageId="9" captionText="FIGURE 5 A–D. Atmetochilus songsangchotei sp. nov. female paratype (ARA-2017-117) habitus; A (left) dorsal habitus, A (right) ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium." figureDoi="http://doi.org/10.5281/zenodo.4397092" httpUri="https://zenodo.org/record/4397092/files/figure.png" pageId="4" pageNumber="525">Fig. 5B</figureCitation>
) strongly procurved to U-shaped. Setae absent in fovea. “Rastellum” of 25 robust down-curved setae on anterior margin of chelicerae. Cheliceral furrow with 8 promarginal teeth in one row, mesobasal denticles present. Fang ventrally smooth without teeth. Stridulatory strikers absent. Labium (
<figureCitation id="89A623E6FFF0FFE6FD7DF8978B5C052D" box="[741,834,1869,1896]" captionStart="FIGURE 5" captionStartId="9.[151,250,1976,2000]" captionTargetBox="[220,1392,189,1953]" captionTargetId="figure-18@9.[196,1392,181,1953]" captionTargetPageId="9" captionText="FIGURE 5 A–D. Atmetochilus songsangchotei sp. nov. female paratype (ARA-2017-117) habitus; A (left) dorsal habitus, A (right) ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium." figureDoi="http://doi.org/10.5281/zenodo.4397092" httpUri="https://zenodo.org/record/4397092/files/figure.png" pageId="4" pageNumber="525">Fig. 5D</figureCitation>
): 1.62±0.84 (1.20–3.12) long, 1.63±0.43 (0.90–1.84) wide without cuspules. Maxillae (
<figureCitation id="89A623E6FFF0FFE6FD8DF8AB8A6F05C9" box="[533,625,1905,1932]" captionStart="FIGURE 5" captionStartId="9.[151,250,1976,2000]" captionTargetBox="[220,1392,189,1953]" captionTargetId="figure-18@9.[196,1392,181,1953]" captionTargetPageId="9" captionText="FIGURE 5 A–D. Atmetochilus songsangchotei sp. nov. female paratype (ARA-2017-117) habitus; A (left) dorsal habitus, A (right) ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium." figureDoi="http://doi.org/10.5281/zenodo.4397092" httpUri="https://zenodo.org/record/4397092/files/figure.png" pageId="4" pageNumber="525">Fig. 5D</figureCitation>
): 4.32 long, 3.00 wide with 33 cuspules each on proximal anterior lobe. Sternum (
<figureCitation id="89A623E6FFF0FFE6FE90F84F897F05F5" box="[264,353,1941,1968]" captionStart="FIGURE 1" captionStartId="5.[151,250,1942,1966]" captionTargetBox="[281,1307,183,1919]" captionTargetId="figure-18@5.[281,1307,181,1919]" captionTargetPageId="5" captionText="FIGURE 1A–F. Atmetochilus and Damarchus bifidus. Sternum.A. Atmetochilus songsangchotei sp. nov. male holotype (ARA- 2017-116); B. A. songsangchotei sp. nov. female paratype (ARA-2017-117); C. A. fossor female neotype (ARA-2018-132); D. A. atriceps female (ARA-2018-173); E. A. atriceps female lectotype; F. Damarchus bifidus type specimen." figureDoi="http://doi.org/10.5281/zenodo.4397084" httpUri="https://zenodo.org/record/4397084/files/figure.png" pageId="4" pageNumber="525">Fig. 1B</figureCitation>
): 5.55±0.62 (4.80–6.32) long, 5.19±0.49 (4.50–5.76) wide. posterior and medial sternal sigilla centrally confluent, anterior sigilla small in upper margin.
</paragraph>
<caption id="45E26FEBFFF1FFE7FF0FF84C8CCC05B2" ID-DOI="http://doi.org/10.5281/zenodo.4397084" ID-Zenodo-Dep="4397084" httpUri="https://zenodo.org/record/4397084/files/figure.png" pageId="5" pageNumber="526" startId="5.[151,250,1942,1966]" targetBox="[281,1307,183,1919]" targetPageId="5">
<paragraph id="11223F63FFF1FFE7FF0FF84C8CCC05B2" blockId="5.[151,1437,1942,2039]" pageId="5" pageNumber="526">
<emphasis id="23E9E371FFF1FFE7FF0FF84C895F05EA" bold="true" box="[151,321,1942,1967]" pageId="5" pageNumber="526">
FIGURE 1 A 
<emphasis id="23E9E371FFF1FFE7FEB9F84D893005EB" bold="true" box="[289,302,1943,1966]" italics="true" pageId="5" pageNumber="526">–</emphasis>
F.
</emphasis>
<taxonomicName id="D69D44E0FFF1FFE7FEDEF84D89D405EA" authorityName="Simon" authorityYear="1887" box="[326,458,1943,1967]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="526" phylum="Arthropoda" rank="genus">
<emphasis id="23E9E371FFF1FFE7FEDEF84D89D405EA" box="[326,458,1943,1967]" italics="true" pageId="5" pageNumber="526">Atmetochilus</emphasis>
</taxonomicName>
and 
<taxonomicName id="D69D44E0FFF1FFE7FE61F84D8AA705EA" authorityName="Gravely" authorityYear="1935" baseAuthorityName=": Zonstein &amp; Marusik" baseAuthorityYear="2014" box="[505,697,1943,1967]" class="Arachnida" family="Nemesiidae" genus="Damarchus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="23E9E371FFF1FFE7FE61F84D8AA705EA" box="[505,697,1943,1967]" italics="true" pageId="5" pageNumber="526">Damarchus bifidus</emphasis>
</taxonomicName>
. Sternum. A. 
<taxonomicName id="D69D44E0FFF1FFE7FCD9F84D8C7705EA" authority="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit, 2020" authorityName="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit" authorityYear="2020" box="[833,1129,1943,1967]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="songsangchotei" status="sp. nov.">
<emphasis id="23E9E371FFF1FFE7FCD9F84D8C7705EA" box="[833,1129,1943,1967]" italics="true" pageId="5" pageNumber="526">Atmetochilus songsangchotei</emphasis>
</taxonomicName>
<emphasis id="23E9E371FFF1FFE7FBF5F84D8CA205EA" bold="true" box="[1133,1212,1943,1967]" pageId="5" pageNumber="526">
<taxonomicNameLabel id="38DA5E0AFFF1FFE7FBF5F84D8CA205EA" box="[1133,1212,1943,1967]" pageId="5" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype (ARA-2017-116); B. 
<taxonomicName id="D69D44E0FFF1FFE7FEB3F86189FB0596" authority="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit, 2020" authorityName="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit" authorityYear="2020" box="[299,485,1979,2003]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="songsangchotei" status="sp. nov.">
<emphasis id="23E9E371FFF1FFE7FEB3F86189FB0596" box="[299,485,1979,2003]" italics="true" pageId="5" pageNumber="526">A. songsangchotei</emphasis>
</taxonomicName>
<emphasis id="23E9E371FFF1FFE7FE73F8618A250596" bold="true" box="[491,571,1979,2003]" pageId="5" pageNumber="526">
<taxonomicNameLabel id="38DA5E0AFFF1FFE7FE73F8618A250596" box="[491,571,1979,2003]" pageId="5" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female paratype (ARA-2017-117); C. 
<taxonomicName id="D69D44E0FFF1FFE7FC5CF8618C000596" authorityName="Simon" authorityYear="1887" box="[964,1054,1979,2003]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="23E9E371FFF1FFE7FC5CF8618C000596" box="[964,1054,1979,2003]" italics="true" pageId="5" pageNumber="526">A. fossor</emphasis>
</taxonomicName>
female neotype (ARA-2018-132); D. 
<taxonomicName id="D69D44E0FFF1FFE7FF0FF805891B05B3" authorityName="Pocock, Tenasserim" authorityYear="1900" box="[151,261,2015,2038]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="atriceps">
<emphasis id="23E9E371FFF1FFE7FF0FF805891B05B3" box="[151,261,2015,2038]" italics="true" pageId="5" pageNumber="526">A. atriceps</emphasis>
</taxonomicName>
female (ARA-2018-173); E. 
<taxonomicName id="D69D44E0FFF1FFE7FDA8F8058A8005B3" authorityName="Pocock, Tenasserim" authorityYear="1900" box="[560,670,2015,2038]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="atriceps">
<emphasis id="23E9E371FFF1FFE7FDA8F8058A8005B3" box="[560,670,2015,2038]" italics="true" pageId="5" pageNumber="526">A. atriceps</emphasis>
</taxonomicName>
female lectotype; F. 
<taxonomicName id="D69D44E0FFF1FFE7FCEAF8058C2D05B2" authorityName="Gravely" authorityYear="1935" baseAuthorityName=": Zonstein &amp; Marusik" baseAuthorityYear="2014" box="[882,1075,2015,2039]" class="Arachnida" family="Nemesiidae" genus="Damarchus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="bifidus">
<emphasis id="23E9E371FFF1FFE7FCEAF8058C2D05B2" box="[882,1075,2015,2039]" italics="true" pageId="5" pageNumber="526">Damarchus bifidus</emphasis>
</taxonomicName>
type specimen.
</paragraph>
</caption>
<caption id="45E26FEBFFF2FFE4FF0FF87F8CC105A4" ID-DOI="http://doi.org/10.5281/zenodo.4397086" ID-Zenodo-Dep="4397086" httpUri="https://zenodo.org/record/4397086/files/figure.png" pageId="6" pageNumber="527" startId="6.[151,250,1957,1981]" targetBox="[386,1207,181,1933]" targetPageId="6">
<paragraph id="11223F63FFF2FFE4FF0FF87F8CC105A4" blockId="6.[151,1437,1957,2017]" pageId="6" pageNumber="527">
<emphasis id="23E9E371FFF2FFE4FF0FF87F895705F8" bold="true" box="[151,329,1957,1981]" pageId="6" pageNumber="527">
FIGURE 2 A 
<emphasis id="23E9E371FFF2FFE4FEBCF87F892F05F9" bold="true" box="[292,305,1957,1980]" italics="true" pageId="6" pageNumber="527">–</emphasis>
C.
</emphasis>
Collecting sites and 
<taxonomicName id="D69D44E0FFF2FFE4FD86F87F8ABC05F8" authorityName="Simon" authorityYear="1887" box="[542,674,1957,1981]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
<emphasis id="23E9E371FFF2FFE4FD86F87F8ABC05F8" box="[542,674,1957,1981]" italics="true" pageId="6" pageNumber="527">Atmetochilus</emphasis>
</taxonomicName>
habitats; A. 
<emphasis id="23E9E371FFF2FFE4FCBAF87F8BED05F8" box="[802,1011,1957,1981]" italics="true" pageId="6" pageNumber="527">A. songsangchotei—</emphasis>
Phra Thaen Dong Rang Forest Park; B. 
<emphasis id="23E9E371FFF2FFE4FA1EF87F88F505A4" italics="true" pageId="6" pageNumber="527">A. fossor—</emphasis>
Roadside hill, Dawei. 4/5/2018; C. 
<taxonomicName id="D69D44E0FFF2FFE4FDCBF8138B3B05A4" authority="Roadside" authorityName="Roadside" box="[595,805,1993,2017]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="atriceps">
<emphasis id="23E9E371FFF2FFE4FDCBF8138ADF05A4" box="[595,705,1993,2017]" italics="true" pageId="6" pageNumber="527">A. atriceps</emphasis>
Roadside
</taxonomicName>
hill, Tenasserim mountain range, 2/5/2018.
</paragraph>
</caption>
<caption id="45E26FEBFFF3FFE5FF0FF8798BBC059A" ID-DOI="http://doi.org/10.5281/zenodo.4397088" ID-Zenodo-Dep="4397088" httpUri="https://zenodo.org/record/4397088/files/figure.png" pageId="7" pageNumber="528" startId="7.[151,250,1955,1979]" targetBox="[187,1402,186,1931]" targetPageId="7">
<paragraph id="11223F63FFF3FFE5FF0FF8798BBC059A" blockId="7.[151,1436,1955,2016]" pageId="7" pageNumber="528">
<emphasis id="23E9E371FFF3FFE5FF0FF879895005FE" bold="true" box="[151,334,1955,1979]" pageId="7" pageNumber="528">
FIGURE 3 A 
<emphasis id="23E9E371FFF3FFE5FEB1F879892805FF" bold="true" box="[297,310,1955,1978]" italics="true" pageId="7" pageNumber="528">–</emphasis>
D.
</emphasis>
<taxonomicName id="D69D44E0FFF3FFE5FECFF8798A9C05FE" authority="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit, 2020" authorityName="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit" authorityYear="2020" box="[343,642,1955,1979]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="songsangchotei" status="sp. nov.">
<emphasis id="23E9E371FFF3FFE5FECFF8798A9C05FE" box="[343,642,1955,1979]" italics="true" pageId="7" pageNumber="528">Atmetochilus songsangchotei</emphasis>
</taxonomicName>
<emphasis id="23E9E371FFF3FFE5FD12F8798AC305FE" bold="true" box="[650,733,1955,1979]" pageId="7" pageNumber="528">
<taxonomicNameLabel id="38DA5E0AFFF3FFE5FD12F8798AC305FE" box="[650,733,1955,1979]" pageId="7" pageNumber="528" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype (ARA-2017-116) habitus; A (left). dorsal habitus, A (right). ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium.
</paragraph>
</caption>
<caption id="45E26FEBFFFCFFEAFF0FF9678C160558" ID-DOI="http://doi.org/10.5281/zenodo.4397090" ID-Zenodo-Dep="4397090" httpUri="https://zenodo.org/record/4397090/files/figure.png" pageId="8" pageNumber="529" startId="8.[151,250,1725,1749]" targetBox="[157,1430,192,1701]" targetPageId="8">
<paragraph id="11223F63FFFCFFEAFF0FF9678C160558" blockId="8.[151,1437,1725,1821]" pageId="8" pageNumber="529">
<emphasis id="23E9E371FFFCFFEAFF0FF96789520490" bold="true" box="[151,332,1725,1749]" pageId="8" pageNumber="529">FIGURE 4 A–H.</emphasis>
<taxonomicName id="D69D44E0FFFCFFEAFECAF9678A650490" authority="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit, 2020" authorityName="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit" authorityYear="2020" box="[338,635,1725,1749]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="songsangchotei" status="sp. nov.">
<emphasis id="23E9E371FFFCFFEAFECAF9678A650490" box="[338,635,1725,1749]" italics="true" pageId="8" pageNumber="529">Atmetochilus songsangchotei</emphasis>
</taxonomicName>
<emphasis id="23E9E371FFFCFFEAFD19F9678ACC0490" bold="true" box="[641,722,1725,1749]" pageId="8" pageNumber="529">
<taxonomicNameLabel id="38DA5E0AFFFCFFEAFD19F9678ACC0490" box="[641,722,1725,1749]" pageId="8" pageNumber="529" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 
<emphasis id="23E9E371FFFCFFEAFC86F9678B4E0490" bold="true" box="[798,848,1725,1749]" pageId="8" pageNumber="529">A–F.</emphasis>
Holotype (ARA-2017-116). 
<emphasis id="23E9E371FFFCFFEAFBE0F9678CAE0490" bold="true" box="[1144,1200,1725,1749]" pageId="8" pageNumber="529">A–C.</emphasis>
Palpal tibia and tarsus, retrolateral, ventral and prolateral view; 
<emphasis id="23E9E371FFFCFFEAFDA2F93B8A7604BC" bold="true" box="[570,616,1761,1785]" pageId="8" pageNumber="529">D–F</emphasis>
. Tibia, metatarsus and tarsus I showing tibial spur, prolateral, retroventral and retrolateral views; 
<emphasis id="23E9E371FFFCFFEAFECDF8DF89970558" bold="true" box="[341,393,1797,1821]" pageId="8" pageNumber="529">G–H</emphasis>
. Male paratype (ARA-2017-120), tibial spur, retrolateral view.
</paragraph>
</caption>
<paragraph id="11223F63FFFCFFEAFF5FF8918C0A05B0" blockId="8.[151,1437,1867,2038]" pageId="8" pageNumber="529">
Spination and scopula: palp: tibia with 5 spines, v3 p2, tarsus scopulate, v2, leg I: Patella and femur without spines, tibia, v1, metatarsus scopula covers all, v4, tarsus scopula on ventral without spines. leg II: patella and femur without spines, tibia, v4, metatarsus scopulate on its distal half, v6, tarsus scopula covers all, without spines. leg III: tibia, r5 p2 d1, leg III: metatarsus has 17 spines, v6 r5 p4 d2, leg III: tibia, r5 p2 d1, leg III: metatarsus, p4 r6 v7. Preening combs on distal ventral metatarsi I–III. Legs (
<tableCitation id="5C1F0AD8FFFCFFEAFC99F8018B4605B0" box="[769,856,2011,2037]" captionStart="TABLE 2" captionStartId="10.[151,239,151,175]" captionText="TABLE 2. Atmetochilus songsangchotei sp. nov., paratype female, length of pedipalp and leg segments (n=5)." pageId="8" pageNumber="529">Table 2</tableCitation>
), formula: 4123.
</paragraph>
<caption id="45E26FEBFFFDFFEBFF0FF8628B8205B1" ID-DOI="http://doi.org/10.5281/zenodo.4397092" ID-Zenodo-Dep="4397092" httpUri="https://zenodo.org/record/4397092/files/figure.png" pageId="9" pageNumber="530" startId="9.[151,250,1976,2000]" targetBox="[220,1392,189,1953]" targetPageId="9">
<paragraph id="11223F63FFFDFFEBFF0FF8628B8205B1" blockId="9.[151,1436,1976,2037]" pageId="9" pageNumber="530">
<emphasis id="23E9E371FFFDFFEBFF0FF86289530595" bold="true" box="[151,333,1976,2001]" pageId="9" pageNumber="530">FIGURE 5 A–D.</emphasis>
<taxonomicName id="D69D44E0FFFDFFEBFECDF8628A610595" authority="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit, 2020" authorityName="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit" authorityYear="2020" box="[341,639,1976,2000]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="songsangchotei" status="sp. nov.">
<emphasis id="23E9E371FFFDFFEBFECDF8628A610595" box="[341,639,1976,2000]" italics="true" pageId="9" pageNumber="530">Atmetochilus songsangchotei</emphasis>
</taxonomicName>
<emphasis id="23E9E371FFFDFFEBFD1FF8638AC70594" bold="true" box="[647,729,1977,2001]" pageId="9" pageNumber="530">
<taxonomicNameLabel id="38DA5E0AFFFDFFEBFD1FF8638AC70594" box="[647,729,1977,2001]" pageId="9" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female paratype (ARA-2017-117) habitus; A (left) dorsal habitus, A (right) ventral habitus; B. Carapace; C. Eye tubercle; D. Maxillae &amp; labium.
</paragraph>
</caption>
<caption id="45E26FEBFFFEFFE8FF0FFF4D8D2A02EA" box="[151,1332,151,176]" pageId="10" pageNumber="531">
<paragraph id="11223F63FFFEFFE8FF0FFF4D8D2A02EA" blockId="10.[151,1332,151,176]" box="[151,1332,151,176]" pageId="10" pageNumber="531">
<emphasis id="23E9E371FFFEFFE8FF0FFF4D891402EA" bold="true" box="[151,266,151,175]" pageId="10" pageNumber="531">TABLE 2.</emphasis>
<taxonomicName id="D69D44E0FFFEFFE8FE89FF4D8A5702EA" authority="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit, 2020" authorityName="Kunsete &amp; Sivayyapram &amp; Traiyasut &amp; Thanoosing &amp; Khamwan &amp; Warrit" authorityYear="2020" box="[273,585,151,175]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="531" phylum="Arthropoda" rank="species" species="songsangchotei" status="sp. nov.">
<emphasis id="23E9E371FFFEFFE8FE89FF4D8A5702EA" box="[273,585,151,175]" italics="true" pageId="10" pageNumber="531">Atmetochilus songsangchotei</emphasis>
</taxonomicName>
<emphasis id="23E9E371FFFEFFE8FDD7FF428ABB02F5" bold="true" box="[591,677,152,176]" pageId="10" pageNumber="531">
<taxonomicNameLabel id="38DA5E0AFFFEFFE8FDD7FF428ABB02F5" box="[591,677,152,176]" pageId="10" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype female, length of pedipalp and leg segments (n=5).
</paragraph>
</caption>
<paragraph id="11223F63FFFEFFE8FEA1FF1A8D680010" pageId="10" pageNumber="531">
<table id="639DCDC3FFFE001DFF3EFF1A8D680010" box="[166,1398,192,597]" gridcols="7" gridrows="6" pageId="10" pageNumber="531" rowsContinueIn="11.[166,1324,984,1210]">
<tr id="AFAD3D21FFFE001DFF3EFF1A8D68029D" box="[166,1398,192,216]" gridrow="0" pageId="10" pageNumber="531" rowspan-0="1">
<th id="EC7C545DFFFE001DFEA1FF1A89B1029D" box="[313,431,192,216]" gridcol="1" gridrow="0" pageId="10" pageNumber="531">Femur</th>
<th id="EC7C545DFFFE001DFE75FF1A8A7D029D" box="[493,611,192,216]" gridcol="2" gridrow="0" pageId="10" pageNumber="531">Patella</th>
<th id="EC7C545DFFFE001DFD3FFF1A8B00029D" box="[679,798,192,216]" gridcol="3" gridrow="0" pageId="10" pageNumber="531">Tibia</th>
<th id="EC7C545DFFFE001DFCCEFF1A8BD3029D" box="[854,973,192,216]" gridcol="4" gridrow="0" pageId="10" pageNumber="531">Metatarsus</th>
<th id="EC7C545DFFFE001DFBB8FF1A8C88029D" box="[1056,1174,192,216]" gridcol="5" gridrow="0" pageId="10" pageNumber="531">Tarsus</th>
<th id="EC7C545DFFFE001DFB7EFF1A8D68029D" box="[1254,1398,192,216]" gridcol="6" gridrow="0" pageId="10" pageNumber="531">Total</th>
</tr>
<tr id="AFAD3D21FFFE001DFF3EFF328D680361" box="[166,1398,232,292]" gridrow="1" pageId="10" pageNumber="531">
<th id="EC7C545DFFFE001DFF3EFF3288F00361" box="[166,238,232,292]" gridcol="0" gridrow="1" pageId="10" pageNumber="531">Palp</th>
<td id="EC7C545DFFFE001DFEA1FF3289B10361" box="[313,431,232,292]" gridcol="1" gridrow="1" pageId="10" pageNumber="531">5.36±2.20 (3.50–9.60)</td>
<td id="EC7C545DFFFE001DFE75FF328A7D0361" box="[493,611,232,292]" gridcol="2" gridrow="1" pageId="10" pageNumber="531">2.63±0.43 (2.63–0.43)</td>
<td id="EC7C545DFFFE001DFD3FFF328B000361" box="[679,798,232,292]" gridcol="3" gridrow="1" pageId="10" pageNumber="531">3.29±0.30 (3.29–0.30)</td>
<td id="EC7C545DFFFE001DFCCEFF328BD30361" box="[854,973,232,292]" gridcol="4" gridrow="1" pageId="10" pageNumber="531">-</td>
<td id="EC7C545DFFFE001DFBB8FF328C880361" box="[1056,1174,232,292]" gridcol="5" gridrow="1" pageId="10" pageNumber="531">3.15±0.35 (3.15–0.35)</td>
<td id="EC7C545DFFFE001DFB7EFF328D680361" box="[1254,1398,232,292]" gridcol="6" gridrow="1" pageId="10" pageNumber="531">14.44±2.70 (11.70–19.44)</td>
</tr>
<tr id="AFAD3D21FFFE001DFF3EFEEE8D680335" box="[166,1398,308,368]" gridrow="2" pageId="10" pageNumber="531">
<th id="EC7C545DFFFE001DFF3EFEEE88F00335" box="[166,238,308,368]" gridcol="0" gridrow="2" pageId="10" pageNumber="531">Leg I</th>
<td id="EC7C545DFFFE001DFEA1FEEE89B10335" box="[313,431,308,368]" gridcol="1" gridrow="2" pageId="10" pageNumber="531">7.78±1.22 (6.16–9.60)</td>
<td id="EC7C545DFFFE001DFE75FEEE8A7D0335" box="[493,611,308,368]" gridcol="2" gridrow="2" pageId="10" pageNumber="531">4.08±0.67 (3.12–4.8)</td>
<td id="EC7C545DFFFE001DFD3FFEEE8B000335" box="[679,798,308,368]" gridcol="3" gridrow="2" pageId="10" pageNumber="531">4.59±1.39 (1.92–5.60)</td>
<td id="EC7C545DFFFE001DFCCEFEEE8BD30335" box="[854,973,308,368]" gridcol="4" gridrow="2" pageId="10" pageNumber="531">4.50±0.60 (3.60–5.04)</td>
<td id="EC7C545DFFFE001DFBB8FEEE8C880335" box="[1056,1174,308,368]" gridcol="5" gridrow="2" pageId="10" pageNumber="531">3.15±0.35 (3.15–0.35)</td>
<td id="EC7C545DFFFE001DFB7EFEEE8D680335" box="[1254,1398,308,368]" gridcol="6" gridrow="2" pageId="10" pageNumber="531">23.81±2.55 (20.72–26.70)</td>
</tr>
<tr id="AFAD3D21FFFE001DFF3EFE5A8D6803F8" box="[166,1398,384,445]" gridrow="3" pageId="10" pageNumber="531">
<th id="EC7C545DFFFE001DFF3EFE5A88F003F8" box="[166,238,384,445]" gridcol="0" gridrow="3" pageId="10" pageNumber="531">Leg II</th>
<td id="EC7C545DFFFE001DFEA1FE5A89B103F8" box="[313,431,384,445]" gridcol="1" gridrow="3" pageId="10" pageNumber="531">6.78±0.75 (5.92–7.60)</td>
<td id="EC7C545DFFFE001DFE75FE5A8A7D03F8" box="[493,611,384,445]" gridcol="2" gridrow="3" pageId="10" pageNumber="531">3.99±0.49 (3.28–4.32)</td>
<td id="EC7C545DFFFE001DFD3FFE5A8B0003F8" box="[679,798,384,445]" gridcol="3" gridrow="3" pageId="10" pageNumber="531">4.02±0.91 (2.40–4.96)</td>
<td id="EC7C545DFFFE001DFCCEFE5A8BD303F8" box="[854,973,384,445]" gridcol="4" gridrow="3" pageId="10" pageNumber="531">4.10±0.52 (3.44–4.80)</td>
<td id="EC7C545DFFFE001DFBB8FE5A8C8803F8" box="[1056,1174,384,445]" gridcol="5" gridrow="3" pageId="10" pageNumber="531">2.71±0.28 (2.40–3.04)</td>
<td id="EC7C545DFFFE001DFB7EFE5A8D6803F8" box="[1254,1398,384,445]" gridcol="6" gridrow="3" pageId="10" pageNumber="531">21.60±2.04 (18.72–24.30)</td>
</tr>
<tr id="AFAD3D21FFFE001DFF3EFE178D68004C" box="[166,1398,461,521]" gridrow="4" pageId="10" pageNumber="531">
<th id="EC7C545DFFFE001DFF3EFE1788F0004C" box="[166,238,461,521]" gridcol="0" gridrow="4" pageId="10" pageNumber="531">Leg III</th>
<td id="EC7C545DFFFE001DFEA1FE1789B1004C" box="[313,431,461,521]" gridcol="1" gridrow="4" pageId="10" pageNumber="531">5.44±0.53 (4.50–5.84)</td>
<td id="EC7C545DFFFE001DFE75FE178A7D004C" box="[493,611,461,521]" gridcol="2" gridrow="4" pageId="10" pageNumber="531">3.60±0.46 (3.00–4.16)</td>
<td id="EC7C545DFFFE001DFD3FFE178B00004C" box="[679,798,461,521]" gridcol="3" gridrow="4" pageId="10" pageNumber="531">3.08±0.42 (2.40–3.44)</td>
<td id="EC7C545DFFFE001DFCCEFE178BD3004C" box="[854,973,461,521]" gridcol="4" gridrow="4" pageId="10" pageNumber="531">4.08±0.53 (3.42–4.64)</td>
<td id="EC7C545DFFFE001DFBB8FE178C88004C" box="[1056,1174,461,521]" gridcol="5" gridrow="4" pageId="10" pageNumber="531">2.39±0.21 (2.10–2.64)</td>
<td id="EC7C545DFFFE001DFB7EFE178D68004C" box="[1254,1398,461,521]" gridcol="6" gridrow="4" pageId="10" pageNumber="531">18.59±1.82 (15.42–20.48)</td>
</tr>
<tr id="AFAD3D21FFFE001DFF3EFDC38D680010" box="[166,1398,537,597]" gridrow="5" pageId="10" pageNumber="531">
<th id="EC7C545DFFFE001DFF3EFDC388F00010" box="[166,238,537,597]" gridcol="0" gridrow="5" pageId="10" pageNumber="531">Leg IV</th>
<td id="EC7C545DFFFE001DFEA1FDC389B10010" box="[313,431,537,597]" gridcol="1" gridrow="5" pageId="10" pageNumber="531">8.06±0.91 (6.88–9.00)</td>
<td id="EC7C545DFFFE001DFE75FDC38A7D0010" box="[493,611,537,597]" gridcol="2" gridrow="5" pageId="10" pageNumber="531">4.27±0.49 (3.60–4.80)</td>
<td id="EC7C545DFFFE001DFD3FFDC38B000010" box="[679,798,537,597]" gridcol="3" gridrow="5" pageId="10" pageNumber="531">5.36±0.51 (4.64–5.70)</td>
<td id="EC7C545DFFFE001DFCCEFDC38BD30010" box="[854,973,537,597]" gridcol="4" gridrow="5" pageId="10" pageNumber="531">6.59±0.72 (5.36–7.5)</td>
<td id="EC7C545DFFFE001DFBB8FDC38C880010" box="[1056,1174,537,597]" gridcol="5" gridrow="5" pageId="10" pageNumber="531">2.93±0.34 (2.50–3.50)</td>
<td id="EC7C545DFFFE001DFB7EFDC38D680010" box="[1254,1398,537,597]" gridcol="6" gridrow="5" pageId="10" pageNumber="531">27.20±2.44 (23.20–29.70)</td>
</tr>
</table>
</paragraph>
<paragraph id="11223F63FFFEFFE8FF5FFD5E8C2F0086" blockId="10.[151,1437,644,1283]" pageId="10" pageNumber="531">Scopula: entire on tarsus of palp, legs I–II, relatively short, sparse cover on distal half on metatarsi II, not divided, absent on metatarsi I, III and IV, tibiae I–IV, patellae I–IV and femora I–IV.</paragraph>
<paragraph id="11223F63FFFEFFE8FF5FFD168988014E" blockId="10.[151,1437,644,1283]" pageId="10" pageNumber="531">Trichobothria: one zigzag row of 12–16 on tarsi I–IV, one row of 5–16 on metatarsi I–IV and two rows, each of 5–11 on tibiae I, III.</paragraph>
<paragraph id="11223F63FFFEFFE8FF5FFCCE8CCA0132" blockId="10.[151,1437,644,1283]" pageId="10" pageNumber="531">Claws: 1 row of teeth on prolateral face of palpal claw. 2 rows of teeth on both faces of paired claws of leg I–IV but different number per face, including 3 teeth on inner side, 4–6 teeth on outer side of Leg I–II, 2 teeth on inner side, 3–4 teeth on outer side of leg III and one tooth on inner side, 3 teeth on outer side of leg IV.</paragraph>
<paragraph id="11223F63FFFEFFE8FF5FFC5A8C0901FB" blockId="10.[151,1437,644,1283]" pageId="10" pageNumber="531">
Spermathecae (ARA-2017-117) (
<figureCitation id="89A623E6FFFEFFE8FDA2FC5A8A8101DE" box="[570,671,896,923]" captionStart="FIGURE 11" captionStartId="18.[151,250,1959,1983]" captionTargetBox="[225,1337,190,1935]" captionTargetId="figure-24@18.[225,1362,181,1935]" captionTargetPageId="18" captionText="FIGURE 11 A–D. Atmetochilus, spermathecae: A. A. fossor neotype; B. A. atriceps; C. A. songsangchotei sp. nov. paratype (ARA-2017-117)." figureDoi="http://doi.org/10.5281/zenodo.4397106" httpUri="https://zenodo.org/record/4397106/files/figure.png" pageId="10" pageNumber="531">Fig. 11C</figureCitation>
): head of spermathecae dome-shaped or semicircular with cap darkly sclerotized, neck of stalk 0.75, base separated by 1.50, stalk base wider than tip.
</paragraph>
<paragraph id="11223F63FFFEFFE8FF5FFC128B47066F" blockId="10.[151,1437,644,1283]" pageId="10" pageNumber="531">Spinnerets: PMS: 1.28±0.22 (1.60–2.50) long, 0.64±0.11 (0.50–0.75) diameter, PLS: 4.95±0.91 (4.16–6.45) long, 1.00±0.17 (0.80–1.25) diameter, length of basal, medial and apical segments 1.99±0.22 (1.60–2.50), 1.30±0.16 (1.15–1.50), 1.66±0.54 (0.96–2.45); apical segment digitiform.</paragraph>
</subSubSection>
<subSubSection id="59876CE8FFFEFFE8FF5FFBEE88C90636" pageId="10" pageNumber="531" type="distribution">
<paragraph id="11223F63FFFEFFE8FF5FFBEE88C90636" blockId="10.[151,1437,644,1283]" pageId="10" pageNumber="531">
<emphasis id="23E9E371FFFEFFE8FF5FFBEE897F060B" bold="true" box="[199,353,1076,1102]" pageId="10" pageNumber="531">Distribution.</emphasis>
Known only from the 
<typeStatus id="CE2681C1FFFEFFE8FDE8FBEF8ABE060A" box="[624,672,1077,1103]" pageId="10" pageNumber="531">type</typeStatus>
locality (Phra Thaen Dong rang Forest Park, 
<collectingRegion id="D359F181FFFEFFE8FB20FBEE88D40636" country="Thailand" name="Kanchanaburi" pageId="10" pageNumber="531">Kanchanaburi province</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="59876CE8FFFEFFE8FF5FFBA78AD20747" pageId="10" pageNumber="531" type="discussion">
<paragraph id="11223F63FFFEFFE8FF5FFBA78AD20747" blockId="10.[151,1437,644,1283]" pageId="10" pageNumber="531">
<emphasis id="23E9E371FFFEFFE8FF5FFBA7892A06D2" bold="true" box="[199,308,1149,1175]" pageId="10" pageNumber="531">Remarks</emphasis>
. This material was taken during field collecting from 
<quantity id="D6659286FFFEFFE8FC10FBA68C3C06D3" box="[904,1058,1148,1175]" metricMagnitude="1" metricUnit="m" metricValue="5.12191" metricValueMax="5.125719999999999" metricValueMin="5.1181" pageId="10" pageNumber="531" unit="in" value="2016.5" valueMax="2018.0" valueMin="2015.0">2015–2018 in</quantity>
<collectingCountry id="698A7FF3FFFEFFE8FBB0FBA68C9106D3" box="[1064,1167,1148,1174]" name="Thailand" pageId="10" pageNumber="531">Thailand</collectingCountry>
. We found 
<taxonomicName id="D69D44E0FFFEFFE8FA92FBA68D8206D3" authorityName="Simon" authorityYear="1887" box="[1290,1436,1148,1174]" class="Arachnida" family="Nemesiidae" genus="Atmetochilus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="531" phylum="Arthropoda" rank="genus">
<emphasis id="23E9E371FFFEFFE8FA92FBA68D8206D3" box="[1290,1436,1148,1174]" italics="true" pageId="10" pageNumber="531">Atmetochilus</emphasis>
</taxonomicName>
from Phra Thaen Dong Rang Forest Park, 
<collectingRegion id="D359F181FFFEFFE8FDEDFB7A8B6506FF" box="[629,891,1184,1210]" country="Thailand" name="Kanchanaburi" pageId="10" pageNumber="531">Kanchanaburi province</collectingRegion>
(
<figureCitation id="89A623E6FFFEFFE8FC13FB7A8BF806FE" box="[907,998,1184,1211]" captionStart="FIGURE 2" captionStartId="6.[151,250,1957,1981]" captionTargetBox="[386,1207,181,1933]" captionTargetId="figure-24@6.[381,1207,181,1933]" captionTargetPageId="6" captionText="FIGURE 2 A–C. Collecting sites and Atmetochilus habitats; A. A. songsangchotei—Phra Thaen Dong Rang Forest Park; B. A. fossor—Roadside hill, Dawei. 4/5/2018; C. A. atriceps Roadside hill, Tenasserim mountain range, 2/5/2018." figureDoi="http://doi.org/10.5281/zenodo.4397086" httpUri="https://zenodo.org/record/4397086/files/figure.png" pageId="10" pageNumber="531">Fig. 2A</figureCitation>
). In the three visits—1 
<superScript id="E6E8922BFFFEFFE8FB7FFB7B8CEC06EA" attach="left" box="[1255,1266,1185,1199]" fontSize="6" pageId="10" pageNumber="531">st</superScript>
round: 
<date id="652319A3FFFEFFE8FAD2FB7B88CC069A" pageId="10" pageNumber="531" value="2015-07-31">31 July 2015</date>
, 2 
<superScript id="E6E8922BFFFEFFE8FF73FB1E88E50697" attach="left" box="[235,251,1220,1234]" fontSize="6" pageId="10" pageNumber="531">nd</superScript>
round: 
<date id="652319A3FFFEFFE8FEC9FB1F8A5E069A" box="[337,576,1220,1247]" pageId="10" pageNumber="531" value="2017-02-11" valueMax="2017-02-13" valueMin="2017-02-11">11–13 February 2017</date>
and 3 
<superScript id="E6E8922BFFFEFFE8FD1BFB1E8A8F0697" attach="left" box="[643,657,1220,1234]" fontSize="6" pageId="10" pageNumber="531">rd</superScript>
round: 
<date id="652319A3FFFEFFE8FD7FFB1F8BD7069A" box="[743,969,1221,1247]" pageId="10" pageNumber="531" value="2018-01-19" valueMax="2018-01-20" valueMin="2018-01-19">19–20 January 2018</date>
— 
<specimenCount id="079BF4EAFFFEFFE8FC7EFB1F8C2E069B" box="[998,1072,1220,1247]" pageId="10" pageNumber="531" type="male">2 male</specimenCount>
individuals, 
<specimenCount id="079BF4EAFFFEFFE8FB58FB1F8D31069B" box="[1216,1327,1220,1247]" pageId="10" pageNumber="531" type="female">29 female</specimenCount>
individuals and 
<specimenCount id="079BF4EAFFFEFFE8FF75FB3289740747" box="[237,362,1256,1282]" pageId="10" pageNumber="531" type="juvenile">26 juvenile</specimenCount>
individuals were taken in total.
</paragraph>
</subSubSection>
</treatment>
</document>