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<document id="89BA2E94332F165FD99C441CC3118585" ID-CLB-Dataset="264218" ID-DOI="10.11646/zootaxa.5331.1.1" ID-GBIF-Dataset="ebee8044-9130-4ce6-9c05-c0f7c65e7f1e" ID-ISSN="1175-5326" ID-Zenodo-Dep="8260962" ID-ZooBank="85D2BD10-1F94-44DF-98C9-9312A28EE584" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1692348832794" checkinUser="plazi" docAuthor="Massa, Bruno" docDate="2023" docId="981CAA6F2C4715267EC84AC48D4BF834" docLanguage="en" docName="zootaxa.5331.1.1.pdf" docOrigin="Zootaxa 5331 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5331.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Plangiopsidini Cadena-Castaneda 2015" docType="treatment" docVersion="10" lastPageNumber="13" masterDocId="6425D2172C4C152A7E5F4B3B8C20FFCC" masterDocTitle="Biodiversity hotspot in the Sangha Trinational Protected Area revealed through Orthoptera Tettigoniidae" masterLastPageNumber="67" masterPageNumber="1" pageNumber="12" updateTime="1699592501176" updateUser="ExternalLinkService">
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<mods:title id="E998A7393FA04AFB3B56D3FCBA126B9B">Biodiversity hotspot in the Sangha Trinational Protected Area revealed through Orthoptera Tettigoniidae</mods:title>
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<treatment id="981CAA6F2C4715267EC84AC48D4BF834" ID-DOI="http://doi.org/10.5281/zenodo.8270432" ID-GBIF-Taxon="212448484" ID-Zenodo-Dep="8270432" LSID="urn:lsid:plazi:treatment:981CAA6F2C4715267EC84AC48D4BF834" httpUri="http://treatment.plazi.org/id/981CAA6F2C4715267EC84AC48D4BF834" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="58AF48F22C4715217EC84AC48EF5FDD5" box="[151,725,511,537]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="100A1B792C4715217EC84AC48EF5FDD5" blockId="11.[151,725,511,537]" box="[151,725,511,537]" pageId="11" pageNumber="12">
<heading id="4B42AC152C4715217EC84AC48EF5FDD5" bold="true" box="[151,725,511,537]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="3">
<emphasis id="22C1C76B2C4715217EC84AC48EF5FDD5" bold="true" box="[151,725,511,537]" pageId="11" pageNumber="12">
Tribe
<taxonomicName id="D7B560FA2C4715217EBD4AC48EF5FDD5" ID-CoL="8PTGB" authority="Cadena-Castaneda, 2015" authorityName="Cadena-Castaneda" authorityYear="2015" box="[226,725,511,537]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Plangiopsidini">
Plangiopsidini
<bibRefCitation id="742466882C4715217FC24AC48EF5FDD5" author="Cadena-Castaneda, O." box="[413,725,511,537]" pageId="11" pageNumber="12" pagination="409 - 414" refId="ref39139" refString="Cadena-Castaneda, O. (2015) Suprageneric taxa derived from Amblycoryphini sensu lato (Orthoptera: Tettigonioidea: Phaneropteridae: Phaneropterinae). Zootaxa, 3904 (3), 409 - 414. https: // doi. org / 10.11646 / zootaxa. 3904.3.6" type="journal article" year="2015">Cadeña-Castaneda, 2015</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="58AF48F22C4715267EC8497C8D4BF834" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="100A1B792C4715217EC8497C8FD0FC06" blockId="11.[151,1437,583,2014]" pageId="11" pageNumber="12">
According to
<bibRefCitation id="742466882C4715217F68497C8E7BFDAE" author="Cadena-Castaneda, O." box="[311,603,583,610]" pageId="11" pageNumber="12" pagination="409 - 414" refId="ref39139" refString="Cadena-Castaneda, O. (2015) Suprageneric taxa derived from Amblycoryphini sensu lato (Orthoptera: Tettigonioidea: Phaneropteridae: Phaneropterinae). Zootaxa, 3904 (3), 409 - 414. https: // doi. org / 10.11646 / zootaxa. 3904.3.6" type="journal article" year="2015">Cadeña-Castaneda (2015)</bibRefCitation>
the tribe
<taxonomicName id="D7B560FA2C4715217C91497C8F50FDAD" authorityName="Cadena-Castaneda" authorityYear="2015" box="[718,880,583,609]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Plangiopsidini">Plangiopsidini</taxonomicName>
is diagnosed based on the following characters: face oval, eyes subcircular, pronotal disc flat and with small denticulations on lateral carina, lateral lobes compact, curving progressively to the humeral sinus, tegmen lanceolate, fore coxae armed with a spine, tympanic organs exposed in the exterior side and conchate in the interior side, meso- and metasternum with lateral lobes rounded or acuminate, tenth tergite without modification, male cerci simple and armed with apical spine. Females with ovipositor as long as a third of the length of the hind femora, curving sharply upwards, valves with crenulate margins, upper valve with cuticular invaginations from the base to the apex (variable character). Among these characters, only small denticulations on the lateral carina are absent in some species (e.g.,
<taxonomicName id="D7B560FA2C4715217AD548788976FC91" authorityName="Karsch" authorityYear="1896" box="[1162,1366,835,861]" class="Insecta" family="Tettigoniidae" genus="Plangiopsis" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="adeps">
<emphasis id="22C1C76B2C4715217AD548788976FC91" box="[1162,1366,835,861]" italics="true" pageId="11" pageNumber="12">Plangiopsis adeps</emphasis>
</taxonomicName>
), thus we could remove this character from those belonging to the tribe. Presently only the genera
<taxonomicName id="D7B560FA2C4715217AE6485C8CEFFC69" authority="Karsch, 1889" authorityName="Karsch" authorityYear="1889" class="Insecta" family="Tettigoniidae" genus="Plangiopsis" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217AE6485C891DFC4D" box="[1209,1341,871,897]" italics="true" pageId="11" pageNumber="12">Plangiopsis</emphasis>
Karsch, 1889
</taxonomicName>
(
<figureCitation id="888E07FC2C4715217EBF48B08D46FC6A" box="[224,358,907,934]" captionStart="FIGURE 5" captionStartId="12.[152,255,1705,1729]" captionTargetBox="[277,1310,244,1681]" captionTargetId="figure-54@12.[277,1310,237,1681]" captionTargetPageId="12" captionText="FIGURE 5. Frontal view of the head of Plangiopsis nouabalensis n. sp. (a), P. foraminata Karsch, 1892 (b), P. adeps Karsch, 1896 (c) and P. semiconchata Karsch, 1889 (d), belonging to the tribe Plangiopsidini Cadeña-Castaneda, 2015. Arrows indicate the lateral dimples on the fastigium of vertex." figureDoi="http://doi.org/10.5281/zenodo.8260983" httpUri="https://zenodo.org/record/8260983/files/figure.png" pageId="11" pageNumber="12">Figs. 5a–5d</figureCitation>
) and
<taxonomicName id="D7B560FA2C4715217FF948B08E96FC69" authority="Bolivar, 1906" authorityName="Bolivar" authorityYear="1906" box="[422,694,907,933]" class="Insecta" family="Tettigoniidae" genus="Plangiola" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217FF948B08E34FC69" box="[422,532,907,933]" italics="true" pageId="11" pageNumber="12">Plangiola</emphasis>
<bibRefCitation id="742466882C4715217C4248B08E96FC69" author="Bolivar, I." box="[541,694,907,933]" pageId="11" pageNumber="12" pagination="327 - 377" refId="ref39056" refString="Bolivar, I. (1906) Fasgonurideos de la Guinea Espanona. Memorias de la Real Sociedad Espanola de Historia Natural, 1, 327 - 377. [http: // bibdigital. rjb. csic. es / ing / Libro. php? Libro = 1342]" type="journal article" year="1906">Bolívar, 1906</bibRefCitation>
</taxonomicName>
(
<figureCitation id="888E07FC2C4715217C9848B08F39FC69" box="[711,793,907,933]" captionStart="FIGURE 6" captionStartId="13.[152,255,1633,1657]" captionTargetBox="[277,1310,184,1609]" captionTargetId="figure-16@13.[277,1310,181,1609]" captionTargetPageId="13" captionText="FIGURE 6. Frontal view of the head of species, whose genera are here included in the tribe Plangiopsidini: Bongeia puncticollis Sĵstedt, 1902 (a), Plangiola herbacea Bolívar, 1906 (b), Afrotapiena n. gen. minor Bolívar, 1906 (before Tapiena minor) (c), Itokiia silvarum Sĵstedt, 1902 (d). Arrows indicate the lateral dimples on the fastigium of vertex." figureDoi="http://doi.org/10.5281/zenodo.8260985" httpUri="https://zenodo.org/record/8260985/files/figure.png" pageId="11" pageNumber="12">Fig. 6d</figureCitation>
) are listed within the tribe
<taxonomicName id="D7B560FA2C4715217A1148B088D4FC69" authorityName="Cadena-Castaneda" authorityYear="2015" box="[1102,1268,907,933]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Plangiopsidini">Plangiopsidini</taxonomicName>
.
<taxonomicName id="D7B560FA2C4715217AA048B0894DFC69" authorityName="Bolivar" authorityYear="1906" box="[1279,1389,907,933]" class="Insecta" family="Tettigoniidae" genus="Plangiola" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217AA048B0894DFC69" box="[1279,1389,907,933]" italics="true" pageId="11" pageNumber="12">Plangiola</emphasis>
</taxonomicName>
is a little different, having eyes more elongate than in the species of
<taxonomicName id="D7B560FA2C4715217D3A48948FC9FC05" authorityName="Karsch" authorityYear="1889" box="[869,1001,943,969]" class="Insecta" family="Tettigoniidae" genus="Plangiopsis" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217D3A48948FC9FC05" box="[869,1001,943,969]" italics="true" pageId="11" pageNumber="12">Plangiopsis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="100A1B792C4715217E9848E88E4FFB6D" blockId="11.[151,1437,583,2014]" pageId="11" pageNumber="12">
Another character that has not been previously considered is the shape of the fastigium of the vertex, which is a little raised, with a rather wide space between the antennae, different from the tribe
<taxonomicName id="D7B560FA2C4715217A2748CC8D7AFBF9" authority="Brunner von Wattenwyl, 1878" authorityName="Brunner von Wattenwyl" authorityYear="1878" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Holochlorini">
Holochlorini
<bibRefCitation id="742466882C4715217B5048C38D7AFBF9" author="Brunner von Wattenwyl, C." pageId="11" pageNumber="12" refId="ref39107" refString="Brunner von Wattenwyl, C. (1878) Monographie der Phaneropteriden. Vienna, 401 pp. [http: // www. biodiversitylibrary. org / item / 37136]" type="book" year="1878">Brunner von Wattenwyl, 1878</bibRefCitation>
</taxonomicName>
, where the fastigium of the vertex is raised, narrow, and the space between the antennae is very small (e.g., genera
<taxonomicName id="D7B560FA2C4715217F2F4F048DD1FB95" authorityName="Stal" authorityYear="1873" box="[368,497,1087,1113]" class="Insecta" family="Tettigoniidae" genus="Holochlora" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217F2F4F048DD1FB95" box="[368,497,1087,1113]" italics="true" pageId="11" pageNumber="12">Holochlora</emphasis>
</taxonomicName>
,
<taxonomicName id="D7B560FA2C4715217C5F4F048E76FB95" authorityName="Stal" authorityYear="1874" box="[512,598,1087,1113]" class="Insecta" family="Tettigoniidae" genus="Arantia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217C5F4F048E76FB95" box="[512,598,1087,1113]" italics="true" pageId="11" pageNumber="12">Arantia</emphasis>
</taxonomicName>
,
<taxonomicName id="D7B560FA2C4715217C3A4F7B8EF7FB95" authorityName="Karsch" authorityYear="1889" box="[613,727,1088,1113]" class="Insecta" family="Tettigoniidae" genus="Dapanera" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217C3A4F7B8EF7FB95" box="[613,727,1088,1113]" italics="true" pageId="11" pageNumber="12">Dapanera</emphasis>
</taxonomicName>
, etc.) (
<figureCitation id="888E07FC2C4715217D774F048F91FB95" box="[808,945,1087,1113]" captionStart="FIGURE 7" captionStartId="14.[152,255,1741,1765]" captionTargetBox="[171,1416,194,1716]" captionTargetId="figure-33@14.[171,1416,181,1717]" captionTargetPageId="14" captionText="FIGURE 7. Frontal and dorsal view of the head of the genera Arantia Stål, 1874 (a, b) and Holochlora Stål, 1873 (c, d), both belonging to the tribe Holochlorini Brunner von Wattenwyl, 1878. Arrows indicate the narrowness of fastigium." figureDoi="http://doi.org/10.5281/zenodo.8260991" httpUri="https://zenodo.org/record/8260991/files/figure.png" pageId="11" pageNumber="12">Figs. 7a–7d</figureCitation>
). The fastigium of the vertex in the species of the tribe
<taxonomicName id="D7B560FA2C4715217F404F588DE1FBB1" authorityName="Cadena-Castaneda" authorityYear="2015" box="[287,449,1123,1149]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Plangiopsidini">Plangiopsidini</taxonomicName>
shows clearly two lateral dimples, not so evident in the species belonging to the tribe
<taxonomicName id="D7B560FA2C4715217EC84FBC8D06FB6D" authorityName="Brunner von Wattenwyl" authorityYear="1878" box="[151,294,1159,1185]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Holochlorini">Holochlorini</taxonomicName>
(
<figureCitation id="888E07FC2C4715217F694FBC8D9CFB6E" box="[310,444,1159,1186]" captionStart="FIGURE 5" captionStartId="12.[152,255,1705,1729]" captionTargetBox="[277,1310,244,1681]" captionTargetId="figure-54@12.[277,1310,237,1681]" captionTargetPageId="12" captionText="FIGURE 5. Frontal view of the head of Plangiopsis nouabalensis n. sp. (a), P. foraminata Karsch, 1892 (b), P. adeps Karsch, 1896 (c) and P. semiconchata Karsch, 1889 (d), belonging to the tribe Plangiopsidini Cadeña-Castaneda, 2015. Arrows indicate the lateral dimples on the fastigium of vertex." figureDoi="http://doi.org/10.5281/zenodo.8260983" httpUri="https://zenodo.org/record/8260983/files/figure.png" pageId="11" pageNumber="12">Figs. 5a–5d</figureCitation>
,
<figureCitation id="888E07FC2C4715217F984FBC8E2FFB6D" box="[455,527,1159,1185]" captionStart="FIGURE 6" captionStartId="13.[152,255,1633,1657]" captionTargetBox="[277,1310,184,1609]" captionTargetId="figure-16@13.[277,1310,181,1609]" captionTargetPageId="13" captionText="FIGURE 6. Frontal view of the head of species, whose genera are here included in the tribe Plangiopsidini: Bongeia puncticollis Sĵstedt, 1902 (a), Plangiola herbacea Bolívar, 1906 (b), Afrotapiena n. gen. minor Bolívar, 1906 (before Tapiena minor) (c), Itokiia silvarum Sĵstedt, 1902 (d). Arrows indicate the lateral dimples on the fastigium of vertex." figureDoi="http://doi.org/10.5281/zenodo.8260985" httpUri="https://zenodo.org/record/8260985/files/figure.png" pageId="11" pageNumber="12">6a–6d</figureCitation>
,
<figureCitation id="888E07FC2C4715217C454FBC8E42FB6D" box="[538,610,1159,1185]" captionStart="FIGURE 7" captionStartId="14.[152,255,1741,1765]" captionTargetBox="[171,1416,194,1716]" captionTargetId="figure-33@14.[171,1416,181,1717]" captionTargetPageId="14" captionText="FIGURE 7. Frontal and dorsal view of the head of the genera Arantia Stål, 1874 (a, b) and Holochlora Stål, 1873 (c, d), both belonging to the tribe Holochlorini Brunner von Wattenwyl, 1878. Arrows indicate the narrowness of fastigium." figureDoi="http://doi.org/10.5281/zenodo.8260991" httpUri="https://zenodo.org/record/8260991/files/figure.png" pageId="11" pageNumber="12">7a–7d</figureCitation>
).
</paragraph>
<paragraph id="100A1B792C4715217E984F908F2CF9B9" blockId="11.[151,1437,583,2014]" pageId="11" pageNumber="12">
When Sĵstedt (1902) described the genus
<taxonomicName id="D7B560FA2C4715217CFF4F978EDDFB09" box="[672,765,1196,1221]" class="Insecta" family="Tettigoniidae" genus="Bongeia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217CFF4F978EDDFB09" box="[672,765,1196,1221]" italics="true" pageId="11" pageNumber="12">Bongeia</emphasis>
</taxonomicName>
(
<typeStatus id="CF0EA5DB2C4715217D554F908F1CFB09" box="[778,828,1195,1221]" pageId="11" pageNumber="12">type</typeStatus>
species:
<taxonomicName id="D7B560FA2C4715217DFF4F978861FB09" authorityName="Sjostedt" authorityYear="1902" box="[928,1089,1195,1221]" class="Insecta" family="Tettigoniidae" genus="Bongeia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="22C1C76B2C4715217DFF4F978861FB09" box="[928,1089,1195,1221]" italics="true" pageId="11" pageNumber="12">B. puncticollis</emphasis>
</taxonomicName>
from
<collectingCountry id="68A25BE92C4715217ADD4F9088DBFB09" box="[1154,1275,1195,1221]" name="Cameroon" pageId="11" pageNumber="12">Cameroon</collectingCountry>
), he wrote that it is most closely related to the genus
<taxonomicName id="D7B560FA2C4715217C664FEB8FCCFB25" authority="Brunner von Wattenwyl, 1878" authorityName="Brunner von Wattenwyl" authorityYear="1878" box="[569,1004,1231,1258]" class="Insecta" family="Cerambycidae" genus="Tapeina" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217C664FEB8EB2FB25" box="[569,658,1232,1257]" italics="true" pageId="11" pageNumber="12">Tapeina</emphasis>
<bibRefCitation id="742466882C4715217CC64FEB8FCCFB25" author="Brunner von Wattenwyl, C." box="[665,1004,1231,1258]" pageId="11" pageNumber="12" refId="ref39107" refString="Brunner von Wattenwyl, C. (1878) Monographie der Phaneropteriden. Vienna, 401 pp. [http: // www. biodiversitylibrary. org / item / 37136]" type="book" year="1878">Brunner von Wattenwyl, 1878</bibRefCitation>
</taxonomicName>
. Because the genus
<taxonomicName id="D7B560FA2C4715217A944FEB8D4FFAC1" authority="Lepeletier &amp; Audinet-Serville" authorityName="Lepeletier &amp; Audinet-Serville" class="Insecta" family="Cerambycidae" genus="Tapeina" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217A944FEB8904FB25" box="[1227,1316,1232,1257]" italics="true" pageId="11" pageNumber="12">Tapeina</emphasis>
Lepeletier &amp; Audinet-Serville
</taxonomicName>
in Latreille, 1828 already existed for a neotropical
<taxonomicName id="D7B560FA2C4715217D9E4FC8893AFAC1" authority=", Bolivar (1906)" authorityName="Bolivar" authorityYear="1906" box="[961,1306,1267,1293]" class="Insecta" family="Cerambycidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">
Cerambycidae,
<bibRefCitation id="742466882C4715217A2C4FC8893AFAC1" author="Bolivar, I." box="[1139,1306,1267,1293]" pageId="11" pageNumber="12" pagination="327 - 377" refId="ref39056" refString="Bolivar, I. (1906) Fasgonurideos de la Guinea Espanona. Memorias de la Real Sociedad Espanola de Historia Natural, 1, 327 - 377. [http: // bibdigital. rjb. csic. es / ing / Libro. php? Libro = 1342]" type="journal article" year="1906">Bolívar (1906)</bibRefCitation>
</taxonomicName>
renamed it as
<taxonomicName id="D7B560FA2C4715217EE74E238D31FAFD" authorityName="Bolivar" authorityYear="1906" box="[184,273,1304,1329]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217EE74E238D31FAFD" box="[184,273,1304,1329]" italics="true" pageId="11" pageNumber="12">Tapiena</emphasis>
</taxonomicName>
and described
<taxonomicName id="D7B560FA2C4715217FE04E238E45FAFD" authorityName="Bolivar" authorityYear="1906" box="[447,613,1304,1329]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="22C1C76B2C4715217FE04E238E45FAFD" box="[447,613,1304,1329]" italics="true" pageId="11" pageNumber="12">Tapiena minor</emphasis>
</taxonomicName>
from
<collectingCountry id="68A25BE92C4715217CF24E2C8ED6FAFD" box="[685,758,1303,1329]" name="Ghana" pageId="11" pageNumber="12">Ghana</collectingCountry>
(West Africa), the only African species for this genus. The genus
<taxonomicName id="D7B560FA2C4715217E804E078D18FA99" authorityName="Bolivar" authorityYear="1906" box="[223,312,1340,1365]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217E804E078D18FA99" box="[223,312,1340,1365]" italics="true" pageId="11" pageNumber="12">Tapiena</emphasis>
</taxonomicName>
is an east Asian-Indonesian genus belonging to the tribe
<taxonomicName id="D7B560FA2C4715217DF04E008861FA99" authorityName="Brunner von Wattenwyl" authorityYear="1878" box="[943,1089,1339,1365]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Holochlorini">Holochlorini</taxonomicName>
, with only one African species; actually the genus is heterogeneous, but the fastigium of the
<typeStatus id="CF0EA5DB2C4715217D1A4E5B8F55FAB6" box="[837,885,1376,1402]" pageId="11" pageNumber="12">type</typeStatus>
species,
<taxonomicName id="D7B560FA2C4715217D824E5B8CFBFA51" authority="(Brunner von Wattenwyl, 1878)" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1878" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acutangula">
<emphasis id="22C1C76B2C4715217D824E5B8857FAB5" box="[989,1143,1375,1401]" italics="true" pageId="11" pageNumber="12">T. acutangula</emphasis>
(
<bibRefCitation id="742466882C4715217AD94E648CF3FA51" author="Brunner von Wattenwyl, C." pageId="11" pageNumber="12" refId="ref39107" refString="Brunner von Wattenwyl, C. (1878) Monographie der Phaneropteriden. Vienna, 401 pp. [http: // www. biodiversitylibrary. org / item / 37136]" type="book" year="1878">Brunner von Wattenwyl, 1878</bibRefCitation>
)
</taxonomicName>
, is narrow, like in the other
<taxonomicName id="D7B560FA2C4715217C4C4EB88E86FA51" authorityName="Brunner von Wattenwyl" authorityYear="1878" box="[531,678,1411,1437]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="tribe" tribe="Holochlorini">Holochlorini</taxonomicName>
. The African species,
<taxonomicName id="D7B560FA2C4715217DC34EBF8FDAFA51" authorityName="Bolivar" authorityYear="1906" box="[924,1018,1412,1437]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="22C1C76B2C4715217DC34EBF8FDAFA51" box="[924,1018,1412,1437]" italics="true" pageId="11" pageNumber="12">T. minor</emphasis>
</taxonomicName>
, recently again recorded from
<collectingCountry id="68A25BE92C4715217B0C4EB889BCFA51" box="[1363,1436,1411,1437]" name="Ghana" pageId="11" pageNumber="12">Ghana</collectingCountry>
by
<bibRefCitation id="742466882C4715217EE24E9C8DA5FA0D" author="Naskrecki, P." box="[189,389,1447,1473]" pageId="11" pageNumber="12" pagination="63 - 68" refId="ref40431" refString="Naskrecki, P. (2008) The katydids of the Atewa Range Forest Reserve, Ghana. RAP Bulletin of Biological Assessment, 47, 63 - 68. [http: // www. bioone. org / doi / abs / 10.1896 / 054.047.0112] https: // doi. org / 10.1896 / 054.047.0112" type="journal article" year="2008">Naskrecki (2008)</bibRefCitation>
, differs in this character from
<taxonomicName id="D7B560FA2C4715217CB84E938FA3FA0D" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1878" box="[743,899,1447,1473]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acutangula">
<emphasis id="22C1C76B2C4715217CB84E938FA3FA0D" box="[743,899,1447,1473]" italics="true" pageId="11" pageNumber="12">T. acutangula</emphasis>
</taxonomicName>
, with its fastigium wide between the antennae (
<figureCitation id="888E07FC2C4715217EC04EF08CD1FA29" box="[159,241,1483,1509]" captionStart="FIGURE 6" captionStartId="13.[152,255,1633,1657]" captionTargetBox="[277,1310,184,1609]" captionTargetId="figure-16@13.[277,1310,181,1609]" captionTargetPageId="13" captionText="FIGURE 6. Frontal view of the head of species, whose genera are here included in the tribe Plangiopsidini: Bongeia puncticollis Sĵstedt, 1902 (a), Plangiola herbacea Bolívar, 1906 (b), Afrotapiena n. gen. minor Bolívar, 1906 (before Tapiena minor) (c), Itokiia silvarum Sĵstedt, 1902 (d). Arrows indicate the lateral dimples on the fastigium of vertex." figureDoi="http://doi.org/10.5281/zenodo.8260985" httpUri="https://zenodo.org/record/8260985/files/figure.png" pageId="11" pageNumber="12">Fig. 6c</figureCitation>
), much like that of other genera included in the tribe
<taxonomicName id="D7B560FA2C4715217D1B4EF08FCAFA29" authorityName="Cadena-Castaneda" authorityYear="2015" box="[836,1002,1483,1509]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Plangiopsidini">Plangiopsidini</taxonomicName>
. This difference leaves little doubt that
<taxonomicName id="D7B560FA2C4715217EC84ECB8D1AF9C5" authorityName="Bolivar" authorityYear="1906" box="[151,314,1520,1545]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="22C1C76B2C4715217EC84ECB8D1AF9C5" box="[151,314,1520,1545]" italics="true" pageId="11" pageNumber="12">Tapiena minor</emphasis>
</taxonomicName>
does not actually belong to the tribe
<taxonomicName id="D7B560FA2C4715217C8A4ED48F47F9C5" authorityName="Brunner von Wattenwyl" authorityYear="1878" box="[725,871,1519,1545]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Holochlorini">Holochlorini</taxonomicName>
, but instead to the tribe
<taxonomicName id="D7B560FA2C4715217A2F4ED48935F9C5" authorityName="Cadena-Castaneda" authorityYear="2015" box="[1136,1301,1519,1545]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Plangiopsidini">Plangiopsidini</taxonomicName>
, with which it shares the other characters of the tribe. Consequently,
<taxonomicName id="D7B560FA2C4715217D474D2F8F58F9E1" authorityName="Bolivar" authorityYear="1906" box="[792,888,1556,1581]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="22C1C76B2C4715217D474D2F8F58F9E1" box="[792,888,1556,1581]" italics="true" pageId="11" pageNumber="12">T. minor</emphasis>
</taxonomicName>
does not belong to the genus
<taxonomicName id="D7B560FA2C4715217A904D2F8908F9E1" authorityName="Bolivar" authorityYear="1906" box="[1231,1320,1556,1581]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217A904D2F8908F9E1" box="[1231,1320,1556,1581]" italics="true" pageId="11" pageNumber="12">Tapiena</emphasis>
</taxonomicName>
, but to an undescribed genus, described below as
<taxonomicName id="D7B560FA2C4715217C064D0C8EFEF99D" authority="Massa, 2023" authorityName="Massa" authorityYear="2023" box="[601,734,1591,1617]" class="Insecta" family="Tettigoniidae" genus="Afrotapiena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="22C1C76B2C4715217C064D0C8EFEF99D" box="[601,734,1591,1617]" italics="true" pageId="11" pageNumber="12">Afrotapiena</emphasis>
</taxonomicName>
<emphasis id="22C1C76B2C4715217CB94D0C8F16F99D" bold="true" box="[742,822,1591,1617]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="39F27A102C4715217CB94D0C8F16F99D" box="[742,822,1591,1617]" pageId="11" pageNumber="12" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
Therefore, the genus
<taxonomicName id="D7B560FA2C4715217A6E4D0388AAF99D" authorityName="Bolivar" authorityYear="1906" box="[1073,1162,1592,1617]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217A6E4D0388AAF99D" box="[1073,1162,1592,1617]" italics="true" pageId="11" pageNumber="12">Tapiena</emphasis>
</taxonomicName>
is definitively excluded from Africa, distributed only in
<collectingCountry id="68A25BE92C4715217FA24D678E41F9B9" box="[509,609,1627,1654]" name="China" pageId="11" pageNumber="12">east Asia</collectingCountry>
and
<collectingCountry id="68A25BE92C4715217CC74D608F29F9B9" box="[664,777,1627,1653]" name="Indonesia" pageId="11" pageNumber="12">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="100A1B792C4715217E984D448856F8E5" blockId="11.[151,1437,583,2014]" pageId="11" pageNumber="12">
Concerning the genus
<taxonomicName id="D7B560FA2C4715217F9F4DBB8E3DF955" box="[448,541,1664,1689]" class="Insecta" family="Tettigoniidae" genus="Bongeia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217F9F4DBB8E3DF955" box="[448,541,1664,1689]" italics="true" pageId="11" pageNumber="12">Bongeia</emphasis>
</taxonomicName>
(
<typeStatus id="CF0EA5DB2C4715217C744D448E7DF955" box="[555,605,1663,1689]" pageId="11" pageNumber="12">type</typeStatus>
species
<taxonomicName id="D7B560FA2C4715217CE54DBB8F7CF955" authorityName="Sjostedt" authorityYear="1902" box="[698,860,1663,1689]" class="Insecta" family="Tettigoniidae" genus="Bongeia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="22C1C76B2C4715217CE54DBB8F7CF955" box="[698,860,1663,1689]" italics="true" pageId="11" pageNumber="12">B. puncticollis</emphasis>
</taxonomicName>
; see
<figureCitation id="888E07FC2C4715217DCC4D448FC3F955" box="[915,995,1663,1689]" captionStart="FIGURE 6" captionStartId="13.[152,255,1633,1657]" captionTargetBox="[277,1310,184,1609]" captionTargetId="figure-16@13.[277,1310,181,1609]" captionTargetPageId="13" captionText="FIGURE 6. Frontal view of the head of species, whose genera are here included in the tribe Plangiopsidini: Bongeia puncticollis Sĵstedt, 1902 (a), Plangiola herbacea Bolívar, 1906 (b), Afrotapiena n. gen. minor Bolívar, 1906 (before Tapiena minor) (c), Itokiia silvarum Sĵstedt, 1902 (d). Arrows indicate the lateral dimples on the fastigium of vertex." figureDoi="http://doi.org/10.5281/zenodo.8260985" httpUri="https://zenodo.org/record/8260985/files/figure.png" pageId="11" pageNumber="12">Fig. 6a</figureCitation>
), it was placed within Psyrae (presently tribe
<taxonomicName id="D7B560FA2C4715217E8D4D988D46F971" authorityName="Brunner von Wattenwyl" authorityYear="1878" box="[210,358,1699,1725]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Holochlorini">Holochlorini</taxonomicName>
), like
<taxonomicName id="D7B560FA2C4715217FF64D9F8E22F971" authorityName="Bolivar" authorityYear="1906" box="[425,514,1700,1725]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217FF64D9F8E22F971" box="[425,514,1700,1725]" italics="true" pageId="11" pageNumber="12">Tapiena</emphasis>
</taxonomicName>
, but also in this case the fastigium, as well as other characters, are more similar to those of the other genera included in the tribe
<taxonomicName id="D7B560FA2C4715217CF84DFC8F6DF92D" authorityName="Cadena-Castaneda" authorityYear="2015" box="[679,845,1735,1761]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Plangiopsidini">Plangiopsidini</taxonomicName>
. Furthermore, the genus
<taxonomicName id="D7B560FA2C4715217A384DFC88F1F92E" authority="Sj" authorityName="Sj" box="[1127,1233,1735,1762]" class="Insecta" family="Tettigoniidae" genus="Itokiia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4715217A384DFC8890F92D" box="[1127,1200,1735,1761]" italics="true" pageId="11" pageNumber="12">Itokiia</emphasis>
Sj
</taxonomicName>
̂stedt, 1902 (
<typeStatus id="CF0EA5DB2C4715217B354DFC89BCF92D" box="[1386,1436,1735,1761]" pageId="11" pageNumber="12">type</typeStatus>
species:
<taxonomicName id="D7B560FA2C4715217EA84DD78DB0F8CA" authority="Sj" authorityName="Sjostedt" authorityYear="1902" box="[247,400,1771,1798]" class="Insecta" family="Tettigoniidae" genus="Itokiia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="silvarum">
<emphasis id="22C1C76B2C4715217EA84DD78D50F8C9" box="[247,368,1771,1797]" italics="true" pageId="11" pageNumber="12">I. silvarum</emphasis>
Sj
</taxonomicName>
̂stedt, 1902; see
<figureCitation id="888E07FC2C4715217C0D4DD08E84F8C9" box="[594,676,1771,1797]" captionStart="FIGURE 6" captionStartId="13.[152,255,1633,1657]" captionTargetBox="[277,1310,184,1609]" captionTargetId="figure-16@13.[277,1310,181,1609]" captionTargetPageId="13" captionText="FIGURE 6. Frontal view of the head of species, whose genera are here included in the tribe Plangiopsidini: Bongeia puncticollis Sĵstedt, 1902 (a), Plangiola herbacea Bolívar, 1906 (b), Afrotapiena n. gen. minor Bolívar, 1906 (before Tapiena minor) (c), Itokiia silvarum Sĵstedt, 1902 (d). Arrows indicate the lateral dimples on the fastigium of vertex." figureDoi="http://doi.org/10.5281/zenodo.8260985" httpUri="https://zenodo.org/record/8260985/files/figure.png" pageId="11" pageNumber="12">Fig. 6d</figureCitation>
), was also included within Psyrae (
<taxonomicName id="D7B560FA2C4715217A704DD088E4F8C9" authorityName="Brunner von Wattenwyl" authorityYear="1878" box="[1071,1220,1771,1797]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="tribe" tribe="Holochlorini">Holochlorini</taxonomicName>
), but its characters, including the fastigium of the vertex, suggests it also belongs to the tribe
<taxonomicName id="D7B560FA2C4715217D934C348852F8E5" authorityName="Cadena-Castaneda" authorityYear="2015" box="[972,1138,1807,1833]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Plangiopsidini">Plangiopsidini</taxonomicName>
.
</paragraph>
<paragraph id="100A1B792C4715267E984C088E61FF19" blockId="11.[151,1437,583,2014]" lastBlockId="12.[151,1436,151,213]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Thus, the characters of the tribe
<taxonomicName id="D7B560FA2C4715217C704C088FCDF882" authority="Cadena-Castaneda, 2015" authorityName="Cadena-Castaneda" authorityYear="2015" box="[559,1005,1843,1870]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Plangiopsidini">
Plangiopsidini
<bibRefCitation id="742466882C4715217C884C088FCDF882" author="Cadena-Castaneda, O." box="[727,1005,1843,1870]" pageId="11" pageNumber="12" pagination="409 - 414" refId="ref39139" refString="Cadena-Castaneda, O. (2015) Suprageneric taxa derived from Amblycoryphini sensu lato (Orthoptera: Tettigonioidea: Phaneropteridae: Phaneropterinae). Zootaxa, 3904 (3), 409 - 414. https: // doi. org / 10.11646 / zootaxa. 3904.3.6" type="journal article" year="2015">Cadeña-Castaneda, 2015</bibRefCitation>
</taxonomicName>
should be: face oval, eyes subcircular, rounded or elongate, anterior margin of pronotum incurved, posterior margin rounded, pronotal disc flat with or without small denticulations on lateral carina, lateral lobes compact, curving progressively to the humeral sinus, higher than long or as high as long, tegmen lanceolate, second pair of wings longer than tegmina, speculum on right tegmen present or absent, fore coxae armed with a spine, conchate tympana in inner side, open in the outer side, tenth tergite without modification, male cerci simple with or without apical spine, styli present. Ovipositor gently upcurved or curving sharply upwards.
</paragraph>
<caption id="44CA4BF12C4015267EC74D928E46F8C4" ID-DOI="http://doi.org/10.5281/zenodo.8260983" ID-Zenodo-Dep="8260983" httpUri="https://zenodo.org/record/8260983/files/figure.png" pageId="12" pageNumber="13" startId="12.[152,255,1705,1729]" targetBox="[277,1310,244,1681]" targetPageId="12" targetType="figure">
<paragraph id="100A1B792C4015267EC74D928E46F8C4" blockId="12.[151,1436,1705,1801]" pageId="12" pageNumber="13">
<emphasis id="22C1C76B2C4015267EC74D928D38F90D" bold="true" box="[152,280,1705,1729]" pageId="12" pageNumber="13">FIGURE 5.</emphasis>
Frontal view of the head of
<taxonomicName id="D7B560FA2C4015267C664D928F1DF90D" authority="Massa, 2023" authorityName="Massa" authorityYear="2023" box="[569,829,1705,1729]" class="Insecta" family="Tettigoniidae" genus="Plangiopsis" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nouabalensis" status="sp. nov.">
<emphasis id="22C1C76B2C4015267C664D928F1DF90D" box="[569,829,1705,1729]" italics="true" pageId="12" pageNumber="13">Plangiopsis nouabalensis</emphasis>
</taxonomicName>
<emphasis id="22C1C76B2C4015267D1C4D928F5CF90D" bold="true" box="[835,892,1705,1729]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="39F27A102C4015267D1C4D928F5CF90D" box="[835,892,1705,1729]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(a),
<taxonomicName id="D7B560FA2C4015267DF34D9188E3F90D" authority="Karsch, 1892" authorityName="Karsch" authorityYear="1892" box="[940,1219,1705,1729]" class="Insecta" family="Tettigoniidae" genus="Pangiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="foraminata">
<emphasis id="22C1C76B2C4015267DF34D918816F90D" box="[940,1078,1705,1729]" italics="true" pageId="12" pageNumber="13">P. foraminata</emphasis>
Karsch, 1892
</taxonomicName>
(b),
<taxonomicName id="D7B560FA2C4015267AAB4D918CEAF929" authority="Karsch, 1896" authorityName="Karsch" authorityYear="1896" class="Insecta" family="Tettigoniidae" genus="Plangiopsis" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="adeps">
<emphasis id="22C1C76B2C4015267AAB4D918968F90D" box="[1268,1352,1705,1729]" italics="true" pageId="12" pageNumber="13">P. adeps</emphasis>
Karsch, 1896
</taxonomicName>
(c) and
<taxonomicName id="D7B560FA2C4015267F424DF58E6DF929" authority="Karsch, 1889" authorityName="Karsch" authorityYear="1889" box="[285,589,1741,1765]" class="Insecta" family="Tettigoniidae" genus="Pangiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="semiconchata">
<emphasis id="22C1C76B2C4015267F424DF58DE0F929" box="[285,448,1741,1765]" italics="true" pageId="12" pageNumber="13">P. semiconchata</emphasis>
Karsch, 1889
</taxonomicName>
(d), belonging to the tribe
<taxonomicName id="D7B560FA2C4015267D044DF688D2F929" authority="Cadena-Castaneda, 2015" authorityName="Cadena-Castaneda" authorityYear="2015" box="[859,1266,1741,1765]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Plangiopsidini">
Plangiopsidini
<bibRefCitation id="742466882C4015267DAB4DF688D2F929" author="Cadena-Castaneda, O." box="[1012,1266,1741,1765]" pageId="12" pageNumber="13" pagination="409 - 414" refId="ref39139" refString="Cadena-Castaneda, O. (2015) Suprageneric taxa derived from Amblycoryphini sensu lato (Orthoptera: Tettigonioidea: Phaneropteridae: Phaneropterinae). Zootaxa, 3904 (3), 409 - 414. https: // doi. org / 10.11646 / zootaxa. 3904.3.6" type="journal article" year="2015">Cadeña-Castaneda, 2015</bibRefCitation>
</taxonomicName>
. Arrows indicate the lateral dimples on the fastigium of vertex.
</paragraph>
</caption>
<paragraph id="100A1B792C4015267E984C758D4BF834" blockId="12.[151,1437,1870,2040]" pageId="12" pageNumber="13">
The following genera therefore are considered to belong to the tribe
<taxonomicName id="D7B560FA2C4015267DEB4C75887AF8A4" authorityName="Cadena-Castaneda" authorityYear="2015" box="[948,1114,1870,1896]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" tribe="Plangiopsidini">Plangiopsidini</taxonomicName>
:
<taxonomicName id="D7B560FA2C4015267A3C4C7588C7F8A4" authorityName="Karsch" authorityYear="1889" box="[1123,1255,1870,1896]" class="Insecta" family="Tettigoniidae" genus="Plangiopsis" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4015267A3C4C7588C7F8A4" box="[1123,1255,1870,1896]" italics="true" pageId="12" pageNumber="13">Plangiopsis</emphasis>
</taxonomicName>
(four species:
<taxonomicName id="D7B560FA2C4015267BD74C748DF0F840" authority="Karsch, 1889" authorityName="Karsch" authorityYear="1889" class="Insecta" family="Tettigoniidae" genus="Pangiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="semiconchata">
<emphasis id="22C1C76B2C4015267BD74C748D11F840" italics="true" pageId="12" pageNumber="13">P. semiconchata</emphasis>
Karsch, 1889
</taxonomicName>
,
<taxonomicName id="D7B560FA2C4015267F844C488F31F840" authority="Karsch, 1892" authorityName="Karsch" authorityYear="1892" box="[475,785,1906,1932]" class="Insecta" family="Tettigoniidae" genus="Pangiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="foraminata">
<emphasis id="22C1C76B2C4015267F844C488E52F840" box="[475,626,1906,1932]" italics="true" pageId="12" pageNumber="13">P. foraminata</emphasis>
Karsch, 1892
</taxonomicName>
,
<taxonomicName id="D7B560FA2C4015267D434C488837F840" authority="Karsch, 1896" authorityName="Karsch" authorityYear="1896" box="[796,1047,1906,1932]" class="Insecta" family="Tettigoniidae" genus="Plangiopsis" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="adeps">
<emphasis id="22C1C76B2C4015267D434C488F58F840" box="[796,888,1906,1932]" italics="true" pageId="12" pageNumber="13">P. adeps</emphasis>
Karsch, 1896
</taxonomicName>
,
<taxonomicName id="D7B560FA2C4015267A7D4C4888F1F840" authority="Massa, 2023" authorityName="Massa" authorityYear="2023" box="[1058,1233,1906,1932]" class="Insecta" family="Tettigoniidae" genus="Plangiopsis" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nouabalensis" status="sp. nov.">
<emphasis id="22C1C76B2C4015267A7D4C4888F1F840" box="[1058,1233,1906,1932]" italics="true" pageId="12" pageNumber="13">P. nouabalensis</emphasis>
</taxonomicName>
<emphasis id="22C1C76B2C4015267A874C498936F840" bold="true" box="[1240,1302,1906,1932]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="39F27A102C4015267A874C498936F840" box="[1240,1302,1906,1932]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
),
<taxonomicName id="D7B560FA2C4015267B714C4989BCF840" authorityName="Bolivar" authorityYear="1906" box="[1326,1436,1906,1932]" class="Insecta" family="Tettigoniidae" genus="Plangiola" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4015267B714C4989BCF840" box="[1326,1436,1906,1932]" italics="true" pageId="12" pageNumber="13">Plangiola</emphasis>
</taxonomicName>
(only one species:
<taxonomicName id="D7B560FA2C4015267F3A4CAC8DC7F87C" authorityName="Bolivar" authorityYear="1906" box="[357,487,1942,1968]" class="Insecta" family="Tettigoniidae" genus="Plangiola" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="herbacea">
<emphasis id="22C1C76B2C4015267F3A4CAC8DC7F87C" box="[357,487,1942,1968]" italics="true" pageId="12" pageNumber="13">P. herbacea</emphasis>
</taxonomicName>
),
<taxonomicName id="D7B560FA2C4015267FA14CAC8E7BF87C" box="[510,603,1943,1968]" class="Insecta" family="Tettigoniidae" genus="Bongeia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4015267FA14CAC8E7BF87C" box="[510,603,1943,1968]" italics="true" pageId="12" pageNumber="13">Bongeia</emphasis>
</taxonomicName>
(two species:
<taxonomicName id="D7B560FA2C4015267CA64CAC8FBAF87C" authorityName="Sjostedt" authorityYear="1902" box="[761,922,1942,1968]" class="Insecta" family="Tettigoniidae" genus="Bongeia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="22C1C76B2C4015267CA64CAC8FBAF87C" box="[761,922,1942,1968]" italics="true" pageId="12" pageNumber="13">B. puncticollis</emphasis>
</taxonomicName>
,
<taxonomicName id="D7B560FA2C4015267DF94CAC88F1F87C" authority="Ebner, 1943" authorityName="Ebner" authorityYear="1943" box="[934,1233,1942,1968]" class="Insecta" family="Tettigoniidae" genus="Bongeia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brevicauda">
<emphasis id="22C1C76B2C4015267DF94CAC8861F87C" box="[934,1089,1942,1968]" italics="true" pageId="12" pageNumber="13">B. brevicauda</emphasis>
Ebner, 1943
</taxonomicName>
),
<taxonomicName id="D7B560FA2C4015267ABB4CAD890DF87C" box="[1252,1325,1942,1968]" class="Insecta" family="Tettigoniidae" genus="Itokiia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="22C1C76B2C4015267ABB4CAD890DF87C" box="[1252,1325,1942,1968]" italics="true" pageId="12" pageNumber="13">Itokiia</emphasis>
</taxonomicName>
(only one species:
<taxonomicName id="D7B560FA2C4015267EA74C808D52F818" authorityName="Sjostedt" authorityYear="1902" box="[248,370,1978,2004]" class="Insecta" family="Tettigoniidae" genus="Itokiia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="silvarum">
<emphasis id="22C1C76B2C4015267EA74C808D52F818" box="[248,370,1978,2004]" italics="true" pageId="12" pageNumber="13">I. silvarum</emphasis>
</taxonomicName>
) and
<taxonomicName id="D7B560FA2C4015267FEA4C818E1AF818" authority="Massa, 2023" authorityName="Massa" authorityYear="2023" box="[437,570,1978,2004]" class="Insecta" family="Tettigoniidae" genus="Afrotapiena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="22C1C76B2C4015267FEA4C818E1AF818" box="[437,570,1978,2004]" italics="true" pageId="12" pageNumber="13">Afrotapiena</emphasis>
</taxonomicName>
<emphasis id="22C1C76B2C4015267C1D4C818EB1F818" bold="true" box="[578,657,1978,2004]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="39F27A102C4015267C1D4C818EB1F818" box="[578,657,1978,2004]" pageId="12" pageNumber="13" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
(only one species:
<taxonomicName id="D7B560FA2C4015267D314C818FF1F818" authorityName="Bolivar" authorityYear="1906" box="[878,977,1978,2004]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="22C1C76B2C4015267D314C818FF1F818" box="[878,977,1978,2004]" italics="true" pageId="12" pageNumber="13">A. minor</emphasis>
</taxonomicName>
) (see
<figureCitation id="888E07FC2C4015267A474C8188BEF819" box="[1048,1182,1978,2005]" captionStart="FIGURE 5" captionStartId="12.[152,255,1705,1729]" captionTargetBox="[277,1310,244,1681]" captionTargetId="figure-54@12.[277,1310,237,1681]" captionTargetPageId="12" captionText="FIGURE 5. Frontal view of the head of Plangiopsis nouabalensis n. sp. (a), P. foraminata Karsch, 1892 (b), P. adeps Karsch, 1896 (c) and P. semiconchata Karsch, 1889 (d), belonging to the tribe Plangiopsidini Cadeña-Castaneda, 2015. Arrows indicate the lateral dimples on the fastigium of vertex." figureDoi="http://doi.org/10.5281/zenodo.8260983" httpUri="https://zenodo.org/record/8260983/files/figure.png" pageId="12" pageNumber="13">Figs. 5a–5d</figureCitation>
,
<figureCitation id="888E07FC2C4015267AF44C8188D2F818" box="[1195,1266,1978,2004]" captionStart="FIGURE 6" captionStartId="13.[152,255,1633,1657]" captionTargetBox="[277,1310,184,1609]" captionTargetId="figure-16@13.[277,1310,181,1609]" captionTargetPageId="13" captionText="FIGURE 6. Frontal view of the head of species, whose genera are here included in the tribe Plangiopsidini: Bongeia puncticollis Sĵstedt, 1902 (a), Plangiola herbacea Bolívar, 1906 (b), Afrotapiena n. gen. minor Bolívar, 1906 (before Tapiena minor) (c), Itokiia silvarum Sĵstedt, 1902 (d). Arrows indicate the lateral dimples on the fastigium of vertex." figureDoi="http://doi.org/10.5281/zenodo.8260985" httpUri="https://zenodo.org/record/8260985/files/figure.png" pageId="12" pageNumber="13">6a–6d</figureCitation>
,
<figureCitation id="888E07FC2C4015267AA04C818965F818" box="[1279,1349,1978,2004]" captionStart="FIGURE 8" captionStartId="16.[152,255,1943,1967]" captionTargetBox="[188,1401,190,1895]" captionTargetId="figure-33@16.[186,1401,181,1919]" captionTargetPageId="16" captionText="FIGURE 8. Habitus in lateral view of Plangiopsis nouabalensis n. sp. (a), P. semiconchata Karsch, 1889 (b), P. foraminata Karsch, 1892 (c), Plangiopsis adeps Karsch, 1896 (d)." figureDoi="http://doi.org/10.5281/zenodo.8260993" httpUri="https://zenodo.org/record/8260993/files/figure.png" pageId="12" pageNumber="13">8a–8d</figureCitation>
,
<figureCitation id="888E07FC2C4015267B0D4C8189B7F818" box="[1362,1431,1978,2004]" captionStart="FIGURE 9" captionStartId="17.[152,255,1744,1768]" captionTargetBox="[151,1435,204,1720]" captionTargetId="figure-16@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 9. Habitus in lateral view of Bongeia puncticollis Sĵstedt, 1902 (a), Itokiia silvarum Sĵstedt, 1902 (b), Tapiena (presently Afrotapiena n. gen.) minor Bolívar, 1906, syntype (c)." figureDoi="http://doi.org/10.5281/zenodo.8260999" httpUri="https://zenodo.org/record/8260999/files/figure.png" pageId="12" pageNumber="13">9a–9c</figureCitation>
,
<figureCitation id="888E07FC2C4015267EC84CE58CD5F834" box="[151,245,2014,2040]" captionStart="FIGURE 10" captionStartId="18.[152,255,1943,1967]" captionTargetBox="[263,1323,181,1919]" captionTargetId="figure-33@18.[263,1323,181,1919]" captionTargetPageId="18" captionText="FIGURE 10. Lateral view of the headand pronotum of species here considered belonging to the tribe Plangiopsidini: Plangiopsis nouabalensis n. sp. (a), Plangiopsis semiconchata Karsch, 1889 (b), P. adeps Karsch, 1896 (c), Bongeia puncticollis Sĵstedt, 1902 (d), Plangiola herbacea Bolívar, 1906 (e), Afrotapiena n. gen. (previously Tapiena) minor Bolívar, 1906 (f)." figureDoi="http://doi.org/10.5281/zenodo.8261003" httpUri="https://zenodo.org/record/8261003/files/figure.png" pageId="12" pageNumber="13">10a–10f</figureCitation>
,
<figureCitation id="888E07FC2C4015267F5E4CE58D7FF834" box="[257,351,2014,2040]" captionStart="FIGURE 11" captionStartId="19.[152,255,1943,1967]" captionTargetBox="[239,1349,181,1919]" captionTargetId="figure-16@19.[239,1349,181,1919]" captionTargetPageId="19" captionText="FIGURE 11. Dorsal view of head, pronotum and stridulatory area of Plangiopsis nouabalensis n. sp. (a), P. semiconchata Karsch, 1889 (b), P. foraminata Karsch, 1892 (c), P. adeps Karsch, 1896 (d), Bongeia puncticollis Sjöstedt, 1902 (♀ holotypus) (e), Itokiia silvarum Sjöstedt, 1902 (♁ holotypus) (f)." figureDoi="http://doi.org/10.5281/zenodo.8261007" httpUri="https://zenodo.org/record/8261007/files/figure.png" pageId="12" pageNumber="13">11a–11f</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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