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<document id="2E63E86159D139A40C4E98C4462F1FD4" ID-DOI="10.3897/zookeys.1220.130885" ID-ZooBank="7A7FE0E0-7BAA-4DB2-8061-A51BE183924F" ID-publisher-id="130885" URI-arpha="21435FA6-2D5B-5E72-AE9D-E32EF9111E8C" URI-zoobank="http://zoobank.org/7A7FE0E0-7BAA-4DB2-8061-A51BE183924F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1733757385579" checkinUser="pensoft" docAuthor="Hibino, Yusuke &amp; Ho, Hsuan-Ching" docDate="2024" docId="97FDEBB46C8C531E9B1D47FDB09E671F" docLanguage="en" docName="ZooKeys 1220: 45-62" docOrigin="ZooKeys 1220" docSource="https://zookeys.pensoft.net/article/130885/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Yirrkala kaupii" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="62" masterDocId="21435FA62D5B5E72AE9DE32EF9111E8C" masterDocTitle="Species of the snake eel genus Yirrkala Whitley, 1940 from Taiwan, with descriptions of a new species and two new records (Anguilliformes, Ophichthidae)" masterLastPageNumber="62" masterPageNumber="45" pageNumber="45" updateTime="1733772390322" updateUser="admin">
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<mods:title id="60AE4EE7E84552F5694860C71D001E38">Species of the snake eel genus Yirrkala Whitley, 1940 from Taiwan, with descriptions of a new species and two new records (Anguilliformes, Ophichthidae)</mods:title>
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<mods:namePart id="D35E7F4E43E72E557F06305B7FC4E2C7">Hibino, Yusuke</mods:namePart>
<mods:nameIdentifier id="CED24C6065DE2A39C96DDF408B575C3D" type="ORCID">0000-0002-5670-3851</mods:nameIdentifier>
<mods:affiliation id="D3C441270C3CA6D547127FDA34C20624">Kitakyushu Museum of Natural History and Human History, Fukuoka 805 - 0071, Japan</mods:affiliation>
</mods:name>
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<mods:namePart id="894667731CABA4D82B3FDF31D4AFF374">Ho, Hsuan-Ching</mods:namePart>
<mods:nameIdentifier id="25B53B546322C88328C9CB8E9D922C75" type="ORCID">0000-0003-1154-601X</mods:nameIdentifier>
<mods:affiliation id="DE8BB9D44F473B9F3ECB69E97C2BDAEB">Department and Graduate Institute of Aquaculture, National Kaohsiung University of Science and Technology, Kaohsiung 81157, Taiwan &amp; Australian Museum, Sydney 2010, Australia</mods:affiliation>
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<mods:date id="DFE4BAA08B820E64666805CF4F14375B">2024</mods:date>
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<mods:number id="8CF134B93651000482EAAB2BC7EE77CA">2024-12-09</mods:number>
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<treatment id="97FDEBB46C8C531E9B1D47FDB09E671F" ID-arpha="97FDEBB4-6C8C-531E-9B1D-47FDB09E671F" LSID="urn:lsid:plazi:treatment:97FDEBB46C8C531E9B1D47FDB09E671F" httpUri="http://treatment.plazi.org/id/97FDEBB46C8C531E9B1D47FDB09E671F" scope_family="Ophichthidae" scope_order="Anguilliformes">
<subSubSection id="FF95EB31EA0571C5ECDA6BE2B6DD7B70" type="nomenclature">
<paragraph id="2AE42615D5BAF5358157D62587B76123">
<taxonomicName id="2337EFF3C699DBF7F172E516E698DF05" ID-CoL="5CRTW" authority="(Bleeker, 1858)" baseAuthorityName="Bleeker" baseAuthorityYear="1858" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">
<emphasis id="D7510880478899D1B51F14D56ECE0A71" italics="true">Yirrkala kaupii</emphasis>
(
<bibRefCitation id="A62E59DCB34F20DA0928648CA1C73E47" DOI="10.5962/bhl.title.144153" author="Bleeker P" issue="8" journalOrPublisher="Acta Societatis Regiae Scientiarum Indo-Neêrlandicae" pagination="1-4" refId="B1" refString="Bleeker P (1858) Twaalfde bijdrage tot de kennis der vischfauna van Celebes. Visschen van Manado. Acta Societatis Regiae Scientiarum Indo-Neerlandicae 5 (8): 1 - 4. https: // doi. org / 10.5962 / bhl. title. 144153" title="Twaalfde bijdrage tot de kennis der vischfauna van Celebes. Visschen van Manado." volume="5" year="1858">Bleeker, 1858</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="87F5D2A123DD01717899CB7AC54F2FED" type="description">
<paragraph id="D935A7942745096E92985365E0061753">
<figureCitation id="9FACAD5E33FC63AD6CD83D074BE2CCFE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawing of lateral head A Yirrkala nkust sp. nov., holotype, NMMB-P 38652, 496 mm TL B Yirrkala kaupii, NMMB-P 38424, 328 mm TL C Yirrkala omanensis, KMNH VR 100616, 216 mm TL. Arrows indicate ethmoid = 1 st supraorbital (lefts), interorbital (middles) and mid-temporal (rights) pores." figureDoi="10.3897/zookeys.1220.130885.figure3" httpUri="https://binary.pensoft.net/fig/1194275">Figs 3 B</figureCitation>
,
<figureCitation id="47E050360498D88D393FD2639B622CEB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Preserved specimen of Yirrkala kaupii, NMMB-P 36108, 239 mm TL A whole body (dorsal view) B whole body (ventral view) C enlarged view of head (lateral). AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure4" httpUri="https://binary.pensoft.net/fig/1194276">4</figureCitation>
,
<tableCitation id="D3BDEEE11537F18CF7A1EC8983A935EE" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Counts and measurements of three Yirrkala species newly collected from Taiwan, with type information." httpUri="http://table.plazi.org/id/BB17082F8F822513CD451454E60E62F5" tableUuid="BB17082F8F822513CD451454E60E62F5">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="9903E3F8E30EE1DEF1EAAB20E6450620" type="reference_group">
<paragraph id="B3C8E0FE2CADE5EA3A57FF2A4493C3C9">
<treatmentCitationGroup id="B817E649A2D8260D664765324896148C">
<treatmentCitation id="2420B68A4BB70349A8E6379C96675BF6" author="Bleeker P" page="3" year="1858">
<taxonomicName id="3E69F36276559BCC9B453EB8929A138F" ID-CoL="4YT3P" authority="Bleeker, 1858: 3" authorityName="Bleeker" authorityPageNumber="3" authorityYear="1858" family="Ophichthidae" genus="Sphagebranchus" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">
<emphasis id="1A37D0539A44B3C1E2674D9CCDD91F85" italics="true">Sphagebranchus kaupii</emphasis>
<comment id="AD72CB76E3965D3974FFE506CD1E548C">
<bibRefCitation id="34BDC0A42417967F198AC4DE48EA7E66" DOI="10.5962/bhl.title.144153" author="Bleeker P" issue="8" journalOrPublisher="Acta Societatis Regiae Scientiarum Indo-Neêrlandicae" pagination="1-4" refId="B1" refString="Bleeker P (1858) Twaalfde bijdrage tot de kennis der vischfauna van Celebes. Visschen van Manado. Acta Societatis Regiae Scientiarum Indo-Neerlandicae 5 (8): 1 - 4. https: // doi. org / 10.5962 / bhl. title. 144153" title="Twaalfde bijdrage tot de kennis der vischfauna van Celebes. Visschen van Manado." volume="5" year="1858">Bleeker, 1858: 3</bibRefCitation>
(type locality: Manado,
<collectingRegion id="F2626BB432CF151FF4B4F4C83282FE26" country="Indonesia" name="Sulawesi Utara">Sulawesi</collectingRegion>
,
<collectingCountry id="9B997BAB014F13B420CC5A5F22679A2B" name="Indonesia">Indonesia</collectingCountry>
);
<treatmentCitation id="FE9D441E119EC9BB93F00123ECD86D78" author="Bleeker P" page="70" year="1864">
<bibRefCitation id="655C81AAEBC7C57CEFE2DD5B30B29C8B" author="Bleeker P" firstAuthor="Bleeker" journalOrPublisher="Murènes, Synbranches, Leptocéphales (anterior part)" page="70" refId="B3" refString="Bleeker P (1864) Atlas ichthyologique des Indes Orientales Néêrlandaises, publié sous les auspices du Gouvernement colonial néêrlandaises. Tome IV. Murènes, Synbranches, Leptocéphales (anterior part), 72 pp." title="Atlas ichthyologique des Indes Orientales Néêrlandaises, publié sous les auspices du Gouvernement colonial néêrlandaises. Tome IV." year="1864">Bleeker 1864: 70</bibRefCitation>
</treatmentCitation>
(Manado).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="F5249FCB5E060D41529A27F8DE11FB20">
<treatmentCitationGroup id="B567F3FA1A68E1E222A35E775DED9413">
<taxonomicName id="3E206A9EF1B7148E2E7A8A40323C58D6" baseAuthorityName="Bleeker" baseAuthorityYear="1858" family="Ophichthidae" genus="Ophichthys" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupi">
<emphasis id="9358DDD53EFB436845AEB43195EF0D40" italics="true">Ophichthys kaupi</emphasis>
</taxonomicName>
<comment id="5379D223119996BF4C48F0BC87228058">
:
<treatmentCitation id="588F289C0AE1F7BE008A2B578F2F50C3" author="Günther A" page="86" year="1870">
<bibRefCitation id="FB9620DE42287DC02DA1B905D294D90D" author="Günther A" firstAuthor="Gunther" journalOrPublisher="Taylor and Francis, London" page="86" refId="B7" refString="Günther A (1870) Catalogue of the Fishes in the British Museum, Vol. 8. Taylor and Francis, London, 549 pp." title="Catalogue of the Fishes in the British Museum, Vol. 8." year="1870">Günther 1870: 86</bibRefCitation>
</treatmentCitation>
(
<collectingRegion id="A0FCFBB07DEEC6508AD2044B3116A9B2" country="Indonesia" name="Sulawesi Utara">Celebes</collectingRegion>
).
</comment>
</treatmentCitationGroup>
</paragraph>
<paragraph id="199FCE89CAAECF9E197720E8B135B757">
<treatmentCitationGroup id="7691847D2BCB5D4F33D077E17B1B03E6">
<taxonomicName id="4F5C8B86C3767297E87D4C4CAE4DE662" authorityName="Bleeker" authorityYear="1858" family="Ophichthidae" genus="Sphagebranchus" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupi">
<emphasis id="C934B2AE0106D7776CD1066DFADE548E" italics="true">Sphagebranchus kaupi</emphasis>
</taxonomicName>
<comment id="41743C13EA18413A0C2BFFEA432FD092">
:
<treatmentCitation id="AEC5C5EA096DCF530703C5E4F0C2C2E2" author="Weber M &amp; de Beaufort LF" page="325" year="1916">
<bibRefCitation id="3C108416E51B8AACD50B3FF18E12A935" DOI="10.1163/9789004590434" author="Weber M &amp; de Beaufort LF" firstAuthor="Weber" journalOrPublisher="Brill, Leiden" page="325" refId="B30" refString="Weber M, de Beaufort LF (1916) The Fishes of the Indo-Australian Archipelago. III. Ostariophysi: II Cyprinoidea, Apodes, Synbranchi. E. J. Brill, Leiden, 455 pp. https://doi.org/10.1163/9789004590434" title="The Fishes of the Indo-Australian Archipelago. III. Ostariophysi: II Cyprinoidea, Apodes, Synbranchi. E. J." year="1916">Weber and de Beaufort 1916: 325</bibRefCitation>
</treatmentCitation>
(
<collectingRegion id="CB58C3FFF72B314DC6464294BC848C14" country="Indonesia" name="Sulawesi Utara">Celebes</collectingRegion>
).
</comment>
</treatmentCitationGroup>
</paragraph>
<paragraph id="D92F703FE82D77A3D3DBB68419CBDC67">
<treatmentCitationGroup id="01D7F4938168B7FC2D4CB055C6F2361A">
<comment id="21269448E60F11A1C3B9C8B8D8478C3F">
?
<taxonomicName id="B5B9A5C7556E2E77C514CD57029FAA0A" ID-CoL="P8NC" baseAuthorityName="Bleeker" baseAuthorityYear="1858" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupi">
<emphasis id="4AD9F921E231964CF9E7D1F052405206" italics="true">Yirrkala kaupi</emphasis>
</taxonomicName>
:
<treatmentCitation id="B25365A0362D8CC45683FAAF74928C27" author="McCosker JE" page="16" year="1977">
<bibRefCitation id="32A5163BBD620EBBDA03BFDB921FFBDA" author="McCosker JE" firstAuthor="McCosker" page="16" refId="B16" refString="McCosker JE (1977) The osteology, classification, and relationships of the eel family Ophichthidae. Proceedings of the California Academy of Sciences (Series 4) 41: 1123." year="1977">McCosker 1977: 16</bibRefCitation>
</treatmentCitation>
(listed; specimen collected from
<collectingCountry id="5E1F7427D7AB6DDAC538744EC28ED490" name="Philippines">Philippines</collectingCountry>
);
<treatmentCitation id="2190FDB19DAC77599E6C6F497B6C6572" author="McCosker JE" page="339" year="2014">
<bibRefCitation id="4D9DF42413D9AEF0B51CA72E737B6F68" author="McCosker JE" editor="Williams GC &amp; Gosliner TM" firstAuthor="McCosker" journalOrPublisher="The 2011 Hearst Philippine Biodiversity Expedition. California Academy of Sciences, San Francisco" page="339" pagination="333-340" refId="B20" refString="McCosker JE (2014) A gigantic deepwater worm eel (Anguilliformes: Ophichthidae) from the Verde Island Passage, Philippine Archipelago. In: Williams GC, Gosliner TM (Eds) The Coral Triangle. The 2011 Hearst Philippine Biodiversity Expedition. California Academy of Sciences, San Francisco, 333340." title="A gigantic deepwater worm eel (Anguilliformes: Ophichthidae) from the Verde Island Passage, Philippine Archipelago." volumeTitle="The Coral Triangle." year="2014">McCosker 2014: 339</bibRefCitation>
</treatmentCitation>
(listed).
</comment>
</treatmentCitationGroup>
</paragraph>
<paragraph id="848CC40896880CA0ECF2B756BBB39D7E">
<treatmentCitationGroup id="EE62D568D9E07ADBE5D50FAA03F7E53C">
<taxonomicName id="93D2A8D0CE0DE4DF7DAF60432758ED3E" ID-CoL="5CRTW" baseAuthorityName="Bleeker" baseAuthorityYear="1858" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">
<emphasis id="13DDBCB5ADD81FD0676152E2129F43DE" italics="true">Yirrkala kaupii</emphasis>
</taxonomicName>
<comment id="BA6F80E971C99C4B5C4BBF225626859F">
:
<treatmentCitation id="F63E339949721F76BCCBE8D5AE120CFC" author="Kottelat M &amp; Whitten AJ &amp; Kartikasari SN &amp; Wirjoatmodjo S" page="13" year="1993">
<bibRefCitation id="14B2154DF019A1DD3CA626838C81D6AC" author="Kottelat M &amp; Whitten AJ &amp; Kartikasari SN &amp; Wirjoatmodjo S" etAl="et al." firstAuthor="Kottelat" page="13" refId="B14" refString="Kottelat M, Whitten AJ, Kartikasari SN, Wirjoatmodjo S (1993) Freshwater Fishes of Western Indonesia and Sulawesi. Periplus Editions, Hong Kong, [xxxviii +] 259 pp [84 pls]." year="1993">Kottelat et al. 1993: 13</bibRefCitation>
</treatmentCitation>
(listed,
<collectingCountry id="7E8F69B49CC28C979102D1777DEC989B" name="Indonesia">Indonesia</collectingCountry>
);
<treatmentCitation id="B103B307F39CAB8837F33129FEB64826" author="Smith DG &amp; McCosker JE" page="1669" year="1999">
<bibRefCitation id="114DE0790C79368A03D85692A47BF246" author="Smith DG &amp; McCosker JE" editor="Carpenter KE &amp; Niem VH" firstAuthor="Smith" journalOrPublisher="The Living Marine Resources of the Western Central Pacific. Batoid Fishes, Chimeras and Bony Fishes Part 1 (Elopidae to Linophrynidae). Vol. 3. FAO, Rome" page="1669" pagination="1662-1669" refId="B29" refString="Smith DG, McCosker JE (1999) Family Ophichthidae. In: Carpenter KE, Niem VH (Eds) FAO species identification guide for fisheries purposes. The Living Marine Resources of the Western Central Pacific. Batoid Fishes, Chimeras and Bony Fishes Part 1 (Elopidae to Linophrynidae). Vol. 3. FAO, Rome, 16621669." title="Family Ophichthidae." volumeTitle="FAO species identification guide for fisheries purposes." year="1999">Smith and McCosker 1999: 1669</bibRefCitation>
</treatmentCitation>
(listed);
<treatmentCitation id="0FA8DCF89203EC1A6CAD05F5E85BCB13" author="Miesen FW &amp; Droppelmann F &amp; Hüllen S &amp; Hadiaty RK &amp; Herder F" page="81" year="2016">
<bibRefCitation id="67D5DCEE2B8CC9A15A929C2E5A491290" author="Miesen FW &amp; Droppelmann F &amp; Hüllen S &amp; Hadiaty RK &amp; Herder F" etAl="et al." firstAuthor="Miesen" issue="2" journalOrPublisher="Bonn Zoological Bulletin" page="81" pagination="77-106" refId="B25" refString="Miesen FW, Droppelmann F, Hüllen S, Hadiaty RK, Herder F (2016) An annotated checklist of the inland fishes of Sulawesi. Bonn Zoological Bulletin 64 (2): 77106." title="An annotated checklist of the inland fishes of Sulawesi." volume="64" year="2016">Miesen et al. 2016: 81</bibRefCitation>
</treatmentCitation>
(listed,
<collectingCountry id="D4F58CDDFC61A0182DEB0821F494B032" name="Indonesia">Indonesia</collectingCountry>
);
<treatmentCitation id="BEAACFE959BDFF1F81E97C3888DDD6CA" author="Jamandre BW" page="160" year="2023">
<bibRefCitation id="3D18D8222734998C2E6F5299438C3C53" DOI="10.11646/zootaxa.5301.2.1" author="Jamandre BW" firstAuthor="Jamandre" issue="2" journalOrPublisher="Zootaxa" page="160" pagination="151-181" refId="B12" refString="Jamandre BW (2023) Freshwater fishes of the Philippines: a provisional checklist. Zootaxa 5301 (2): 151181. https://doi.org/10.11646/zootaxa.5301.2.1" title="Freshwater fishes of the Philippines: a provisional checklist." volume="5301" year="2023">Jamandre 2023: 160</bibRefCitation>
</treatmentCitation>
(listed,
<collectingCountry id="7F42FC36518CD7C6F15C0C3A3E394BB3" name="Indonesia">Indonesia</collectingCountry>
).
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAJAG" type="material">
<paragraph id="0C54EBE0D3D321761B31174C292A55D1">
<heading id="CE42C630E5D962955B03336E016B9EC2" reason="title">Materials examined.</heading>
</paragraph>
<paragraph id="8671858BB7549E888E8F833FC423BA96">
<materialsCitation id="12FEA2DA0E01C549C3403D1469B68FD6" collectingDate="2021-05-10" collectingMethod="trap net / fyke net" collectionCode="NMMB-P" country="Taiwan" elevation="1" latitude="23.936666" location="jointed mouth of Mu-Gua River and Hualien River" longLatPrecision="125" longitude="121.60833" municipality="River" specimenCode="NMMB-P 36108" specimenCount="1" stateProvince="Hualien">
<specimenCode id="995207E51113B0CE055078257E0BE4C2">NMMB-P 36108</specimenCode>
, 239 mm
<abbrev id="ABBRID0EGJAG" xlink_title="Total length">TL</abbrev>
, ca
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENJAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[121.608333,23.936667]}">
<geoCoordinate id="803C44F66BBA5E67DCEFA2B0A0BE3297" degrees="23" direction="north" minutes="56.2" orientation="latitude" precision="92" value="23.936666">23°56.2'N</geoCoordinate>
,
<geoCoordinate id="4F7C34E9C93C5621EFD1A0B78CC16115" degrees="121" direction="east" minutes="36.5" orientation="longitude" precision="92" value="121.60833">121°36.5'E</geoCoordinate>
</named-content>
,
<location id="95460B5535D96CC458EE95C11AA4BA3D" LSID="urn:lsid:plazi:treatment:97FDEBB46C8C531E9B1D47FDB09E671F:95460B5535D96CC458EE95C11AA4BA3D" country="Taiwan" latitude="23.936666" longLatPrecision="125" longitude="121.60833" municipality="River" name="jointed mouth of Mu-Gua River and Hualien River" stateProvince="Hualien">jointed mouth of Mu-Gua River and Hualien River</location>
,
<collectingRegion id="1C760E0549DDDAD181B839AEF0B087BF" country="Taiwan" name="Hualien">Hualien</collectingRegion>
, eastern
<collectingCountry id="27CADD635D3BFAA88263A5F576787E12" name="Taiwan">Taiwan</collectingCountry>
, ca
<quantity id="1EA266D51BE515BE77BB36C46AE78381" metricMagnitude="0" metricUnit="m" metricValue="1.0" unit="m" value="1.0">
<elevation id="C68D51E876496DAE205D4B7843BC23AA" metricMagnitude="0" metricUnit="m" metricValue="1.0" unit="m" value="1.0">1 m</elevation>
</quantity>
,
<collectingMethod id="00030D28E6DE972829BF675795F45344">trap net</collectingMethod>
(
<collectingMethod id="AF9A6A1C7C894B0602F617774D456E85">fyke net</collectingMethod>
),
<date id="B425AA248D295CDDF8F9F51389FB195C" value="2021-05-10">
<collectingDate id="BD4023E9DB817E0198494F60990C4FA6" value="2021-05-10">10 May 2021</collectingDate>
</date>
</materialsCitation>
<materialsCitation id="EB9C652C52E64C6D439847B54FF1C753" collectingDate="2023-05-26" collectionCode="NMMB-P" country="Taiwan" county="May" latitude="23.465" location="Jin-pu village" longLatPrecision="125" longitude="121.495" municipality="Fengbin township" specimenCode="NMMB-P 38424, NMMB-P 38425" specimenCount="1" stateProvince="Taitung">
<specimenCode id="A314111CC3CAF805484163AFC6E6EDF0">NMMB-P 38424</specimenCode>
, 328 mm
<abbrev id="ABBRID0ESJAG" xlink_title="Total length">TL</abbrev>
,
<specimenCode id="567199E13880457D77DA601814D47CD2">NMMB-P 38425</specimenCode>
, 331 mm
<abbrev id="ABBRID0EWJAG" xlink_title="Total length">TL</abbrev>
, ca
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4JAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[121.495000,23.465000]}">
<geoCoordinate id="41E6B4225BE3CC78F5DEC43F442677B4" degrees="23" direction="north" minutes="27.9" orientation="latitude" precision="92" value="23.465">23°27.9'N</geoCoordinate>
,
<geoCoordinate id="1E1B8C5E694F3C73DD5F57248D139A82" degrees="121" direction="east" minutes="29.7" orientation="longitude" precision="92" value="121.495">121°29.7'E</geoCoordinate>
</named-content>
,
<location id="7E7E27B84ACB6CFEF903B51C8B5D7189" LSID="urn:lsid:plazi:treatment:97FDEBB46C8C531E9B1D47FDB09E671F:7E7E27B84ACB6CFEF903B51C8B5D7189" country="Taiwan" county="May" latitude="23.465" longLatPrecision="125" longitude="121.495" municipality="Fengbin township" name="Jin-pu village" stateProvince="Taitung">Jin-pu village</location>
,
<collectingMunicipality id="B87CB1893D629362F47649CD077AD765">Fengbin township</collectingMunicipality>
,
<collectingRegion id="B07D0E3A8D26DC743591C150BEE39BEE" country="Taiwan" name="Taitung">Taitung</collectingRegion>
,
<collectingDate id="1803C63B6269FE6D0464D19415BC07A8">
<date id="D00DFC5B89344BD14255E516EA8AD8E2" value="1926-05">26 May</date>
, 2023
</collectingDate>
, collected after preserving in wine for several years
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECKAG" type="diagnosis">
<paragraph id="844765CA0E015534383319D7DF041B1F">
<heading id="4C54CCC7C6368B8C761952B3EF85BAB1" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="6B0E176121D0F344EA61ED97C6B5D987">
An elongate species of
<taxonomicName id="B729F6648D477E8D53C4BA2531A7EA3D" authorityName="Whitley" authorityYear="1940" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="genus">
<emphasis id="C2B9BA42D85D6DBD45E9BD0DD3499BB7" italics="true">Yirrkala</emphasis>
</taxonomicName>
with the following combination of characters: body bicolored, dark brown dorsally and pale ventrally, lateral line pores margined as pale blank;
<abbrev id="ABBRID0EPKAG" xlink_title="supraorbital pores">SO</abbrev>
1 + 3; dorsal-fin origin behind gill opening; lateral-line pores before anus 6365; predorsal vertebrae 1718, preanal 6466, and total 151156;
<abbrev id="ABBRID0ETKAG" xlink_title="Mean vertebral formula">MVF</abbrev>
17-65 - 154.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXKAG" type="description">
<paragraph id="9DFA0BA0790B9B7C302233919D9E747A">
<heading id="F3D9FA4FBD01398B377F35F97B4623DA" reason="title">Description of Taiwanese specimens.</heading>
</paragraph>
<paragraph id="BBEA7B3FB9192F316C05067DBBD5540B">
New record. Body elongate, subcylindrical, tip of tail laterally compressed and extremely pointed (Fig.
<figureCitation id="79A8B97BDF934E4F915F87B3F12CD571" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Preserved specimen of Yirrkala kaupii, NMMB-P 36108, 239 mm TL A whole body (dorsal view) B whole body (ventral view) C enlarged view of head (lateral). AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure4" httpUri="https://binary.pensoft.net/fig/1194276">4 A, B</figureCitation>
). Head moderate in size,
<quantity id="26DD56A8990EFEAC7898A868BF086E86" metricMagnitude="-1" metricUnit="m" metricValue="3.2384999999999997" metricValueMax="3.4798" metricValueMin="2.9972" unit="in" value="12.75" valueMax="13.7" valueMin="11.8">11.813.7 in</quantity>
<abbrev id="ABBRID0EBLAG" xlink_title="Total length">TL</abbrev>
; preanal length shorter than tail,
<quantity id="00FFA4C3A1D4BDCE1E8F1D4F8D6F9C89" metricMagnitude="-2" metricUnit="m" metricValue="2.794" unit="in" value="1.1">1.1 in</quantity>
tail length and
<quantity id="1C13FAAF871C1D87A809885D4DCB9E5A" metricMagnitude="-2" metricUnit="m" metricValue="5.334" unit="in" value="2.1">2.1 in</quantity>
<abbrev id="ABBRID0EFLAG" xlink_title="Total length">TL</abbrev>
.
</paragraph>
<caption id="B4992DC2AECB2C3BCB8862D42588D5A4" ID-DOI="10.3897/zookeys.1220.130885.figure4" ID-arpha="7AF57FFF-2A91-5E14-A508-1449C60FE71C" httpUri="https://binary.pensoft.net/fig/1194276" startId="F4">
<paragraph id="3BC614DA21809725179A7335AF6F734A">
<label id="AC284DE906DF2BE7709E0DF902935E3B">Figure 4.</label>
</paragraph>
<paragraph id="EFDFEECE880B1B08B63D8D7963897A60">
Preserved specimen of
<taxonomicName id="E78508A0DE0E9D84244511119EA95FDD" baseAuthorityName="Bleeker" baseAuthorityYear="1858" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">
<emphasis id="D61A7BA6A64C0D4D01D7C52E5BDA2E16" italics="true">Yirrkala kaupii</emphasis>
</taxonomicName>
,
<specimenCode id="D85339F19D236F257CDAF202456FF2AB">NMMB-P 36108</specimenCode>
, 239 mm
<abbrev id="ABBRID0E3LAG" xlink_title="Total length">TL</abbrev>
<emphasis id="208A5569286BAB49D0C2894A36BDBB53" bold="true">A</emphasis>
whole body (dorsal view)
<emphasis id="A1D15EDDE01BBC986FC88447ACCE7AEC" bold="true">B</emphasis>
whole body (ventral view)
<emphasis id="94DDD47ECFFBFC2310F10B8C72A1077A" bold="true">C</emphasis>
enlarged view of head (lateral).
<emphasis id="D9E06A0BB5D30BCE12BFE22BEEF30922" bold="true">AN</emphasis>
anus
<emphasis id="63BF995107570838E45DC172CE58537C" bold="true">DO</emphasis>
dorsal-fin origin.
</paragraph>
</caption>
<paragraph id="877C491B3F951506B9EB4CE67DFA7355">
Snout relatively longer, more than twice eye diameter, its tip pointed; distinct median groove ventrally on snout but the groove mostly not teared, its anterior tip beyond ethmoid pore; slope of dorsal surface of snout smooth, without notch or hump. Anterior nostril tubular, moderate in length, tube length about equal to pupil diameter; posterior nostril oval in shape with an inner valve, located on anteroventral margin of eye, opening ventrally, covered by a flap extending slightly below edge of mouth gape. Eye moderate in size, covered by a transparent skin; center of eye anterior to mid-jaw (Fig.
<figureCitation id="E1D5F7E07108BD33937A162FAAFFBFE2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Preserved specimen of Yirrkala kaupii, NMMB-P 36108, 239 mm TL A whole body (dorsal view) B whole body (ventral view) C enlarged view of head (lateral). AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure4" httpUri="https://binary.pensoft.net/fig/1194276">4 C</figureCitation>
). Mouth inferior, distance from tip of snout to anterior tip of lower jaw slightly longer than eye diameter; lower jaw short, its tip relatively pointed, slightly beyond or reaching posterior base of anterior-nostril tube (Fig.
<figureCitation id="B95A0B2DE02FC8CDF6E309094E6C24DF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Preserved specimen of Yirrkala kaupii, NMMB-P 36108, 239 mm TL A whole body (dorsal view) B whole body (ventral view) C enlarged view of head (lateral). AN anus DO dorsal-fin origin." figureDoi="10.3897/zookeys.1220.130885.figure4" httpUri="https://binary.pensoft.net/fig/1194276">4 C</figureCitation>
); rictus short, posterior end of gape slightly behind a vertical through posterior margin of eye; lips smooth with folds, the fold along upper lip extending from second infraorbital pore to posterior rictus, but absent in
<specimenCount id="47277E28CA122794949AC1E458BC9118" count="1" type="generic">one specimen</specimenCount>
(
<specimenCode id="D5AC1755CEE20B5F4B177175A73D4014">
<collectionCode id="4A6E2F08B3E671A8FAEBD25DF2F498DA">NMMB-P</collectionCode>
38425
</specimenCode>
); the fold along lower lip extending from position between first and second mandibular pores to anterior of first preopercular pore. Gill openings positioned ventrolaterally of breast, not close each side; shape of opening curved, diameter more than twice eye diameter.
</paragraph>
<paragraph id="CD73029F48B15DD41ADA1B18C528A2F8">
Sensory pores on head developed, arrangement of those pores as follows (Fig.
<figureCitation id="55973930BA72DA80610E8F2C2C6B02DE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawing of lateral head A Yirrkala nkust sp. nov., holotype, NMMB-P 38652, 496 mm TL B Yirrkala kaupii, NMMB-P 38424, 328 mm TL C Yirrkala omanensis, KMNH VR 100616, 216 mm TL. Arrows indicate ethmoid = 1 st supraorbital (lefts), interorbital (middles) and mid-temporal (rights) pores." figureDoi="10.3897/zookeys.1220.130885.figure3" httpUri="https://binary.pensoft.net/fig/1194275">3 B</figureCitation>
): 1 (ethmoid) + 3 on supraorbital, 3 + 3 on infraorbital, 4 or 5 on lower jaw, 2 on preopercle, and 3 on supratemporal, one of those on mid-temporal; a single median interorbital pore.
</paragraph>
<paragraph id="CBC04EF8B14AC2039B525AA9AA51CF76">Lateral-line pores small but obvious. Lateral line almost complete except for near tip of tail, eight or nine anterior to a vertical through gill opening, 1618 anterior to dorsal-fin origin, 6365 anterior to mid anus, and total 116138.</paragraph>
<paragraph id="39383F51895C2AE7B933142E7B42429F">
Teeth pointed, slightly recurved posteriorly; teeth on maxilla, vomer and dentary uniserial; maxilla comprising 19 (right) / 25 (left) teeth and mandible comprising 21 / 23 teeth in
<specimenCode id="6AA25BED1C85C14F4C03C9E59A8FDB6C">
<collectionCode id="5D6E08EA4417CF2DB514C1951292CD7A">NMMB-P</collectionCode>
36108
</specimenCode>
; intermaxillary with four or five teeth arranged in two rows or a chevron shape.
</paragraph>
<paragraph id="0140AAE6DECBBB7B98F6459ECFA61460">Median fins low but obvious; origin of dorsal fin behind gill opening, ending anterior to tip of tail about 1 eye diameter; origin of anal fin slightly behind anus, ending same as dorsal fin; caudal and pectoral fins absent.</paragraph>
<paragraph id="9A86DFC600974BE4481541D2E35E554E">
<emphasis id="BC8A81D4FB2FA88621D9E0882FB37927" bold="true">
<emphasis id="03F68B9E064D5C6705BC753E54CD13FD" bold="true" italics="true">Coloration</emphasis>
.
</emphasis>
No information of fresh coloration. After the preservation by alcohol (ca 60 %) directly, body clearly bicolored, generally darkish brown dorsally and pale yellowish brown ventrally; numerous melanophores present dorsally, the border going down toward tip of tail; lateral-line pores margined by pale spots blank at least anterior to anus, but in tail the pattern gradually faded; head darker dorsally, lower jaw also dusky; dorsal fin dusky but margin pale yellowish white; eye with whitish margin by skin covering eye; anal fin pale yellowish white except dusky tip of tail ca 1 / 2 HL.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJNAG" type="distribution">
<paragraph id="5FA6D6045051DBED35047B3BE3F15478">
<heading id="C7F3B543491029BC4C05FC2314980C9D" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="51E05121E3866512B114CEE3B42D82B0">
Manado,
<collectingRegion id="57CB02FCA9123423E28EA38AA3111CC5" country="Indonesia" name="Sulawesi Utara">Sulawesi</collectingRegion>
,
<collectingCountry id="C29C69116AB9FA96CF27F9B1CAAE6F63" name="Indonesia">Indonesia</collectingCountry>
(
<typeStatus id="31617842CAD363FECAF2F3E60DD4279D" type="holotype">holotype</typeStatus>
) and eastern
<collectingCountry id="D6B92A3D6AF36BF172009E2636A58F10" name="Taiwan">Taiwan</collectingCountry>
. In both places, this species was collected from rivers, but there is no detailed information for the
<typeStatus id="18AE64AB41AF0D889E3A9BC700A6E588" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EONAG" type="Ecological note">
<paragraph id="A046C4A029B3FF8EBD791BC71A5F8394">
<heading id="4567C489FD556ED4E3FFC0EFEA3471F8" reason="title">Ecological note.</heading>
</paragraph>
<paragraph id="31188DA7687FD161FC2AB132BCD2E79F">
<specimenCount id="6CE86FB5F728D5B273D45FC3FCEC8891" count="1" type="generic">One specimen</specimenCount>
was collected from a river mouth together with many
<taxonomicName id="6F9DBAF6B998DA76CE84E5E4DC56B79E" family="Ophichthidae" genus="Lamnostoma" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="1C110510D74897C7D320736DA3BD44A5" italics="true">Lamnostoma</emphasis>
spp.
</taxonomicName>
, a genus which is commonly found in the freshwater environments. However, the species might be rare because only one individual was found among approximately 200 individuals of
<taxonomicName id="D8C00BED1F83EAC667BAF48AF7447DE4" family="Ophichthidae" genus="Lamnostoma" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="genus">
<emphasis id="25F80B412467EF6E0E0AFB326D564518" italics="true">Lamnostoma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECOAG" type="remarks">
<paragraph id="E3402672AF7929AA202703E9C68FAC67">
<heading id="646D7E8E3DDEAA0512E4C5B19306AE34" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="D7D731E7D4782BBB4120F0B491833FD0">
<taxonomicName id="D149AA70306900B7D75F4CD7765F1E62" authorityName="Bleeker" authorityYear="1858" family="Ophichthidae" genus="Sphagebranchus" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">
<emphasis id="E9BA43F9CD1BDDE3226963CE6990793A" italics="true">Sphagebranchus kaupii</emphasis>
</taxonomicName>
is one of the oldest names in the genus
<taxonomicName id="81688964775A8B5E3418C94BA6A251D5" authorityName="Whitley" authorityYear="1940" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="genus">
<emphasis id="0DD03363A8C994C813C8F641EE2819CC" italics="true">Yirrkala</emphasis>
</taxonomicName>
, and it lacks detailed morphological information except for the original description (
<bibRefCitation id="69EB0D77C8B9CD634BF02A456545407F" DOI="10.5962/bhl.title.144153" author="Bleeker" firstAuthor="Bleeker" issue="8" journalOrPublisher="Acta Societatis Regiae Scientiarum Indo-Neêrlandicae" pagination="1-4" refId="B1" refString="Bleeker P (1858) Twaalfde bijdrage tot de kennis der vischfauna van Celebes. Visschen van Manado. Acta Societatis Regiae Scientiarum Indo-Neêrlandicae 5 (8): 14. https://doi.org/10.5962/bhl.title.144153" title="Twaalfde bijdrage tot de kennis der vischfauna van Celebes. Visschen van Manado." volume="5" year="1858">Bleeker 1858</bibRefCitation>
). Based on YHs investigation, the only known Bleeker specimen, deposited in the Natural History Museum (
<named-content id="D572CED32C29BC19B3B059F70E2EE589" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="3260228FED89F898CD267F302B1D2AC9">BMNH</collectionCode>
</named-content>
1867.11. 28.304), should be regarded as the
<typeStatus id="B3C831B3BDB0616B68C1D5477A7DFD1A" type="holotype">holotype</typeStatus>
because its similar length agrees with the original description (e. g.
<quantity id="1183E3332E24B62B9284674484FECA17" metricMagnitude="-1" metricUnit="m" metricValue="3.42" unit="mm" value="342.0">342 mm</quantity>
<abbrev id="ABBRID0ECPAG" xlink_title="Total length">TL</abbrev>
vs
<quantity id="FC7547035142311842BD9C0BEE2525B8" metricMagnitude="-1" metricUnit="m" metricValue="3.62" unit="mm" value="362.0">362 mm</quantity>
<abbrev id="ABBRID0EGPAG" xlink_title="Total length">TL</abbrev>
in the original description) (Fig.
<figureCitation id="F8F53CB36A8856AD12415B97E319AACE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Sphagebranchus kaupii, BMNH 1867.11. 28.304, 342 mm TL A whole body B lateral view of head C dorsal view of head. All photos provided by Natural History Museum." figureDoi="10.3897/zookeys.1220.130885.figure5" httpUri="https://binary.pensoft.net/fig/1194277">5</figureCitation>
).
</paragraph>
<caption id="162089439DE45B018C8DEBB88F0EE93D" ID-DOI="10.3897/zookeys.1220.130885.figure5" ID-arpha="E531405D-6123-5C5C-B1A5-DC188BBD9E34" httpUri="https://binary.pensoft.net/fig/1194277" startId="F5">
<paragraph id="6A546FB1E3C531847B11BEBFCA2DB061">
<label id="37FD4BC49006ED31699F87D9E55031EA">Figure 5.</label>
</paragraph>
<paragraph id="44991531BD2B7FA164A6799224D4BCAC">
Holotype of
<taxonomicName id="13CF7D296BD35F0E92A5B7DF3F67A10E" authority=", BMNH" authorityName="BMNH" family="Ophichthidae" genus="Sphagebranchus" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">
<emphasis id="9AE4808AE1B85B0A51473F329E405F72" italics="true">Sphagebranchus kaupii</emphasis>
,
<named-content id="D64A0B2F268CB0D3EA65F2C18DFA9C49" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">BMNH</named-content>
</taxonomicName>
1867.11. 28.304, 342 mm
<abbrev id="ABBRID0EFQAG" xlink_title="Total length">TL</abbrev>
<emphasis id="7C7FA065347EAD7A2465F8B615187BFF" bold="true">A</emphasis>
whole body
<emphasis id="99224F79FF4170D440EC5556F0C6EF6D" bold="true">B</emphasis>
lateral view of head
<emphasis id="1EB877A05A8C57E04E7141E3FD7BD304" bold="true">C</emphasis>
dorsal view of head. All photos provided by Natural History Museum.
</paragraph>
</caption>
<paragraph id="CBF039C13CD42F05BA722C4202653720">
Our specimens closely match the
<typeStatus id="52B649D24B517957C506C3AD75AC2665" type="holotype">holotype</typeStatus>
in body proportions and vertebral counts in the original description.
<bibRefCitation id="B38CA8DC162747044A2F5201121D027B" DOI="10.5962/bhl.title.144153" author="Bleeker P" firstAuthor="Bleeker" issue="8" journalOrPublisher="Acta Societatis Regiae Scientiarum Indo-Neêrlandicae" pagination="1-4" refId="B1" refString="Bleeker P (1858) Twaalfde bijdrage tot de kennis der vischfauna van Celebes. Visschen van Manado. Acta Societatis Regiae Scientiarum Indo-Neêrlandicae 5 (8): 14. https://doi.org/10.5962/bhl.title.144153" title="Twaalfde bijdrage tot de kennis der vischfauna van Celebes. Visschen van Manado." volume="5" year="1858">Bleeker (1858)</bibRefCitation>
noted that it has drop-shaped markings on the lateral line, which are also found in our specimens. Although
<bibRefCitation id="7872951888EAC5691EAEEC4FE76625FC" author="Günther A" firstAuthor="Gunther" journalOrPublisher="Taylor and Francis, London" refId="B7" refString="Günther A (1870) Catalogue of the Fishes in the British Museum, Vol. 8. Taylor and Francis, London, 549 pp." title="Catalogue of the Fishes in the British Museum, Vol. 8." year="1870">Günther (1870)</bibRefCitation>
mentioned that it has uniform body coloration, it is assumed that
<typeStatus id="360E4ACE4415C5E9997B7EFFCB459606" type="holotype">holotype</typeStatus>
specimen was once dried and bleached beforehand because its very poor present condition had been caused by the drying process. There are several differences in the proportions of the head, snout, eye, and upper-jaw lengths between the original description of
<taxonomicName id="88AAD46E3ADC8C5F7B98E944800FE549" family="Ophichthidae" genus="Sphagebranchus" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">
<emphasis id="80E6504DFA21856E4A9DF96D4AA2796E" italics="true">S. kaupii</emphasis>
</taxonomicName>
and Taiwanese specimens (Table
<tableCitation id="9B0FA5DB2B26BDAB5C570059E0B7ECCE" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Counts and measurements of three Yirrkala species newly collected from Taiwan, with type information." httpUri="http://table.plazi.org/id/BB17082F8F822513CD451454E60E62F5" tableUuid="BB17082F8F822513CD451454E60E62F5">1</tableCitation>
). The Taiwanese population is possibly an undescribed species, but the morphological differences could also be due to intraspecific variation or differences in the method of preservation, or the time in preservative. As we were unable to directly observe the
<typeStatus id="27F14C5C725722997529D2373E48D55E" type="holotype">holotype</typeStatus>
and the number of specimens observed was limited, we refrain from considering it a separate species.
</paragraph>
<paragraph id="E3C151E56D76DA87E0ECF80954B1BE99">
Most of congeners of
<taxonomicName id="E9A115F6571993D35383947693932D42" authorityName="Whitley" authorityYear="1940" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="genus">
<emphasis id="9557A29EFB53C7A578ABFE7395B6CC37" italics="true">Yirrkala</emphasis>
</taxonomicName>
have a dorsal fin that originates around, and usually slightly behind, the gill opening (see above). Only four species,
<taxonomicName id="4181F9A7CFE63FD0F328E45F0515561E" baseAuthorityName="Weber &amp; de Beaufort" baseAuthorityYear="1916" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="gjellerupi">
<emphasis id="ABE38BE2BFB877EEED6EED332CE45503" italics="true">Y. gjellerupi</emphasis>
</taxonomicName>
,
<taxonomicName id="0045CD38B5E94EBE3D3D7423BA2198EE" family="Ophichthidae" genus="Sphagebranchus" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">
<emphasis id="4DC8A35F380C65B189C66399E550C2C1" italics="true">Y. kaupii</emphasis>
</taxonomicName>
,
<taxonomicName id="9FC0C7ACA80FCC1950C01C44991FF33F" authorityName="McCosker" authorityYear="1999" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="insolitus">
<emphasis id="345E9E31EF4AB8FDA031AD4A9D88193D" italics="true">Y. insolitus</emphasis>
</taxonomicName>
, and
<taxonomicName id="C12E5E9893BE914F08ECF0114BC89E8E" baseAuthorityName="Norman" baseAuthorityYear="1939" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="omanensis">
<emphasis id="0D7C37DCDE3CEF451B36E538D93A234C" italics="true">Y. omanensis</emphasis>
</taxonomicName>
, have the origin of the fin located more than half a head length behind the gill opening. Moreover, the latter two species are unique because their dorsal-fin origins are situated far behind, near the anus.
<taxonomicName id="8B2E7634A3A9F8B6D22F49A4D346E6CC" baseAuthorityName="Bleeker" baseAuthorityYear="1858" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">
<emphasis id="80FCBBE8B99637FF27E26731816650D8" italics="true">Yirrkala kaupii</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="23C89AAC3C0FFFA4162FA10AFA8065EE" baseAuthorityName="Weber &amp; de Beaufort" baseAuthorityYear="1916" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="gjellerupi">
<emphasis id="A5887248DE6E4B81C2AC54D14C4CFF59" italics="true">Y. gjellerupi</emphasis>
</taxonomicName>
in the vertebral counts and body proportions, including head length, tail length, and snout length. Both species inhabit rivers far from the river mouth (
<bibRefCitation id="B179083AD9438CB4CA58C89B5BD518D5" DOI="10.1353/psc.2007.0009" author="McCosker" etAl="et al." firstAuthor="McCosker" journalOrPublisher="Pacific Science" pagination="141-144" refId="B23" refString="McCosker JE, Boseto D, Jenkins AP (2007) Redescription of Yirrkala gjellerupi, a poorly known freshwater Indo-Pacific snake eel (Anguilliformes: Ophichthidae). Pacific Science 61: 141144. https://doi.org/10.1353/psc.2007.0009" title="Redescription of Yirrkala gjellerupi, a poorly known freshwater Indo-Pacific snake eel (Anguilliformes: Ophichthidae)." volume="61" year="2007">McCosker et al. 2007</bibRefCitation>
). However,
<taxonomicName id="B796B8FC167F375B160D41C6130D8902" baseAuthorityName="Bleeker" baseAuthorityYear="1858" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="kaupii">
<emphasis id="1C2737C11C4F43ED2B6545AB98D37637" italics="true">Y. kaupii</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="9D28F56EA0D1E91592604BA9BF1744EB" baseAuthorityName="Weber &amp; de Beaufort" baseAuthorityYear="1916" family="Ophichthidae" genus="Yirrkala" kingdom="Animalia" order="Anguilliformes" phylum="Chordata" rank="species" species="gjellerupi">
<emphasis id="59641BF38B8686B79E6149B701E77880" italics="true">Y. gjellerupi</emphasis>
</taxonomicName>
by the position of the rictus (behind a vertical through posterior margin of eye, vs before), and the presence of a small protrusion along around margin of posterior nostril (vs absent).
</paragraph>
</subSubSection>
</treatment>
</document>