397 lines
46 KiB
XML
397 lines
46 KiB
XML
<document id="C1A906E2F78807F4D0637DD90F5661E3" ID-CLB-Dataset="51752" ID-DOI="http://dx.doi.org/10.11646/zootaxa.4029.1.1" ID-GBIF-Dataset="85cd7166-7b23-496d-8b10-7f1dcf2c4d60" ID-ISSN="1175-5326" ID-ZooBank="A625A5A9-EE80-45E0-A9BE-7A183B0996B1" checkinTime="1447441568197" checkinUser="donat" docAuthor="Robert A. Johnson" docDate="2015" docId="971D8786FFC2FF8965D417E2D5792753" docLanguage="en" docName="29902.pdf.imf" docOrigin="Zootaxa 4029 (1)" docTitle="Pogonomyrmex cunicularius" docType="treatment" docVersion="7" lastPageNumber="50" masterDocId="6B24FFFEFFE9FFB865431775D1342613" masterDocTitle="A taxonomic revision of South American species of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae). Part I" masterLastPageNumber="142" masterPageNumber="1" pageNumber="44" updateTime="1732675905917" updateUser="ExternalLinkService">
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<mods:title id="3E10D98CE7499732C2537B02AE918E83">A taxonomic revision of South American species of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae). Part I</mods:title>
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<mods:roleTerm id="55BE7C1A67CA1291C54D701622815781">Author</mods:roleTerm>
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<mods:namePart id="98593F7B43D08405CBFA764A99BB91B5">Robert A. Johnson</mods:namePart>
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<mods:title id="F72EAAB7A10FDB441162F64A1BAFAA50">Zootaxa</mods:title>
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<mods:date id="6B97C8921479DDEF007C46560ADEDBCF">2015</mods:date>
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<mods:number id="5C5D114B2D0CD716FA982BBD40B2C8B1">4029</mods:number>
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<mods:identifier id="6EF55539AE69383DCD31B8809C66E872" type="DOI">http://dx.doi.org/10.11646/zootaxa.4029.1.1</mods:identifier>
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<mods:identifier id="879EE4C5380ED3204791ACE36A366EA7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="971D8786FFC2FF8965D417E2D5792753" ID-DOI="http://doi.org/10.5281/zenodo.6115572" ID-GBIF-Taxon="114920420" ID-Zenodo-Dep="6115572" LSID="urn:lsid:plazi:treatment:971D8786FFC2FF8965D417E2D5792753" httpUri="http://treatment.plazi.org/id/971D8786FFC2FF8965D417E2D5792753" lastPageId="49" lastPageNumber="50" pageId="43" pageNumber="44" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="57AE651BFFC2FF9365D417E2D0C226A2" box="[151,502,151,177]" pageId="43" pageNumber="44" type="nomenclature">
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<paragraph id="1F0B3690FFC2FF9365D417E2D0C226A2" box="[151,502,151,177]" pageId="43" pageNumber="44">
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<heading id="444381FCFFC2FF9365D417E2D0C226A2" bold="true" box="[151,502,151,177]" fontSize="11" level="1" pageId="43" pageNumber="44" reason="1">
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<taxonomicName id="D8B44D13FFC2FF9365D417E2D0C226A2" ID-CoL="4KVV6" box="[151,502,151,177]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="cunicularius">
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<emphasis id="2DC0EA82FFC2FF9365D417E2D0C226A2" bold="true" box="[151,502,151,177]" italics="true" pageId="43" pageNumber="44">Pogonomyrmex cunicularius</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="57AE651BFFC2FF9365D417CFD07E26C0" box="[151,330,186,211]" pageId="43" pageNumber="44" type="description">
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<paragraph id="1F0B3690FFC2FF9365D417CFD07E26C0" box="[151,330,186,211]" pageId="43" pageNumber="44">
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(
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<figureCitation id="878F2A15FFC2FF9365DC17CFD07626C0" box="[159,322,186,211]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStart-2="FIGURE 20" captionStartId-0="44.[151,249,1978,2000]" captionStartId-1="45.[151,249,1978,2000]" captionStartId-2="46.[151,249,1802,1824]" captionTargetBox-0="[450,1132,193,1956]" captionTargetBox-1="[412,1170,193,1955]" captionTargetBox-2="[437,1141,193,1780]" captionTargetId-0="figure@44.[450,1136,193,1956]" captionTargetId-1="figure@45.[412,1175,193,1956]" captionTargetId-2="figure@46.[437,1167,193,1780]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionTargetPageId-2="46" captionText-0="FIGURE 18. Photograph of Pogonomyrmex cunicularius Mayr worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0103052). Photographs by April Nobile from www. AntWeb. org." captionText-1="FIGURE 19. Photograph of Pogonomyrmex cunicularius Mayr ergatoid queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0173109). Photographs by April Nobile from www. AntWeb. org." captionText-2="FIGURE 20. Photograph of Pogonomyrmex cunicularius Mayr male: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0235298). Photographs by Will Ericson from www. AntWeb. org." pageId="43" pageNumber="44">Figures 18–20</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="57AE651BFFC2FF9365D41676D0AE27FC" pageId="43" pageNumber="44" type="reference_group">
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<paragraph id="1F0B3690FFC2FF9365D41676D2572766" pageId="43" pageNumber="44">
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<taxonomicName id="D8B44D13FFC2FF9365D41676D347270B" authority="Mayr, 1887: 613" authorityName="Mayr" authorityPageNumber="613" authorityYear="1887" box="[151,627,258,280]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="cunicularius">
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<emphasis id="2DC0EA82FFC2FF9365D41676D089270B" box="[151,445,258,280]" italics="true" pageId="43" pageNumber="44">Pogonomyrmex cunicularius</emphasis>
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Mayr, 1887: 613
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</taxonomicName>
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(worker, male). Syntypes examined: 3 workers [MSNG], 2 workers [NMW], URUGUAY, no location; 1 male [NMW], URUGUAY, Montevideo: Montevideo; 2 workers [NMW], ARGENTINA, Buenos Aires (Prof. Berg leg.). See also Gallardo, 1932: 120, fig. 13; Johnson, 2010: 169, fig. 2. NMW worker from URUGUAY here designated
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<emphasis id="2DC0EA82FFC2FF9364AD162BD3BE2766" bold="true" box="[494,650,350,373]" pageId="43" pageNumber="44">LECTOTYPE</emphasis>
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[CASENT0173372].
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</paragraph>
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<paragraph id="1F0B3690FFC2FF9365D4160BD0AE27FC" pageId="43" pageNumber="44">
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<taxonomicName id="D8B44D13FFC2FF9365D4160BD2072780" authority="Santschi, 1931: 275" authorityName="Santschi" authorityPageNumber="275" authorityYear="1931" box="[151,819,381,403]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="variety" species="cunicularis" variety="brevispinus">
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<emphasis id="2DC0EA82FFC2FF9365D4160BD0852780" box="[151,433,381,403]" italics="true" pageId="43" pageNumber="44">Pogonomyrmex cunicularis</emphasis>
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var.
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<emphasis id="2DC0EA82FFC2FF9364AB1608D3682780" box="[488,604,381,403]" italics="true" pageId="43" pageNumber="44">brevispinus</emphasis>
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Santschi, 1931: 275
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</taxonomicName>
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(worker, ergatoid queen). Syntypes examined: 1 worker, 1 ergatoid queen [MACN], ARGENTINA, Entre Ríos: Estación Sosa (Mac Donagh leg.). Kusnezov, 1951: 251 (synonomy under
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<taxonomicName id="D8B44D13FFC2FF93644F16CFD0BF27C3" box="[268,395,442,464]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="cunicularius">
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<emphasis id="2DC0EA82FFC2FF93644F16CFD0BF27C3" box="[268,395,442,464]" italics="true" pageId="43" pageNumber="44">cunicularius</emphasis>
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</taxonomicName>
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; here confirmed). See also Gallardo, 1932: 123. MACN worker here designated
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<emphasis id="2DC0EA82FFC2FF93604316CFD4A827C3" bold="true" box="[1280,1436,442,464]" pageId="43" pageNumber="44">LECTOTYPE</emphasis>
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[CASENT0249048].
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</paragraph>
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</subSubSection>
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<subSubSection id="57AE651BFFC2FF9365D41568D04E24B3" pageId="43" pageNumber="44" type="diagnosis">
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<paragraph id="1F0B3690FFC2FF9365D41568D04E24B3" pageId="43" pageNumber="44">
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<emphasis id="2DC0EA82FFC2FF9365D41568D0492427" bold="true" box="[151,381,540,564]" pageId="43" pageNumber="44">Worker. Diagnosis.</emphasis>
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Within the
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<taxonomicName id="D8B44D13FFC2FF9367481569D3822427" box="[523,694,540,564]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="cunicularius">
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<emphasis id="2DC0EA82FFC2FF9367481569D3822427" box="[523,694,540,564]" italics="true" pageId="43" pageNumber="44">P. cunicularius</emphasis>
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</taxonomicName>
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-group, the combination of: (1) in profile, petiolar node broadly rounded, (2) inferior propodeal spines rounded, (3) superior propodeal spines moderately long, shorter than distance between their bases, and (4) first gastral tergum smooth and strongly shining uniquely characterize this species (
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<figureCitation id="878F2A15FFC2FF9365BB15F2D05E24B3" box="[248,362,647,672]" captionStart="FIGURE 18" captionStartId="44.[151,249,1978,2000]" captionTargetBox="[450,1132,193,1956]" captionTargetId="figure@44.[450,1136,193,1956]" captionTargetPageId="44" captionText="FIGURE 18. Photograph of Pogonomyrmex cunicularius Mayr worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0103052). Photographs by April Nobile from www. AntWeb. org." pageId="43" pageNumber="44">
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<emphasis id="2DC0EA82FFC2FF9365BB15F2D05E24B3" bold="true" box="[248,362,647,672]" pageId="43" pageNumber="44">Figure 18</emphasis>
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</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="57AE651BFFC2FF97658415D8D32227E7" lastPageId="47" lastPageNumber="48" pageId="43" pageNumber="44" type="description">
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<paragraph id="1F0B3690FFC2FF93658415D8D3BA2523" pageId="43" pageNumber="44">
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<emphasis id="2DC0EA82FFC2FF93658415D8D04224D7" bold="true" box="[199,374,685,708]" pageId="43" pageNumber="44">Measurements</emphasis>
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—lectotype (
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<emphasis id="2DC0EA82FFC2FF93674E15D9D32F24D7" box="[525,539,684,708]" italics="true" pageId="43" pageNumber="44">n</emphasis>
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= 16). HL 2.38 (1.85–2.47); HW 2.07 (1.52–2.10); MOD 0.38 (0.31–0.42); OMD 0.62 (0.45–0.69); SL 1.55 (1.44–1.92); PNW 1.44 (1.04–1.47); HFL 2.57 (2.09–2.68); ML 2.83 (2.05–2.87); PW 0.50 (0.40–0.54); PPW 0.72 (0.55–0.76). Indices: SI 74.88 (79.40–100.00); CI 86.97 (81.01–88.21); OI 18.36 (17.79–21.16); HFI 124.15 (119.23–139.38).
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</paragraph>
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<paragraph id="1F0B3690FFC2FF9365841449D1E722F3" pageId="43" pageNumber="44">
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<emphasis id="2DC0EA82FFC2FF9365841449D0402547" bold="true" box="[199,372,828,852]" pageId="43" pageNumber="44">Redescription.</emphasis>
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Head elongate (CI = 81.01–88.21), widest immediately posterior to mandibles, narrowing posterior to eyes; posterior margin flat to weakly convex in full-face view. Cephalic dorsum with weak to moderately strong, wavy or irregular longitudinal rugae to weakly rugoreticulate, rugae often weaker to indistinct near posterior margin; in full-face view, medial rugae diverging weakly toward posterior corners of head. Cephalic interrugae weakly to moderately granulate, weakly shining. Vertex weakly rugose to weakly to moderately granulate, dull to weakly shining. Anterior margin of clypeus flat to weakly convex; dorsal surface with at least several subparallel longitudinal or oblique rugae. Mandibles with six teeth; mandibular dorsum coarsely rugose. Eyes small, MOD ranging from 0.15–0.18x HL. In profile, eyes situated anterior to middle of head, OMD = 1.36– 1.77x MOD. Antennal scapes long (SI = 74.88–100.00), surpassing vertex by less than length of first funicular segment; entire scape with longitudinal striae. Basal flange of scape well-developed with carinate margin. Psammophore poorly-developed, consisting of short to medium-length hairs scattered across ventral surface of head.
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</paragraph>
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<paragraph id="1F0B3690FFC2FF936584139ED5622037" pageId="43" pageNumber="44">Promesonotal profile moderately convex, propodeum flat; all mesosomal surfaces with subparallel, irregular rugae to weakly rugoreticulate. In dorsal view, humeral shoulders of pronotum rounded. Dorsum of promesonotum and sides of pronotum with transverse, oblique to longitudinal, irregular rugae to rugoreticulate; mesopleura with irregular rugae that angle posterodorsally. Superior propodeal spines moderately long, acuminate, shorter than distance between their bases; spines connected by well-defined keel; wavy to irregular transverse rugae on propodeal dorsum traverse ventrally or anteroventrally on sides. Inferior propodeal spines well-developed, triangular, wider than high, tips broadly rounded to bluntly angulate. Propodeal spiracles narrowly ovate facing posterad. Interrugae on mesosoma smooth to weakly granulate, weakly to strongly shining. Legs long (HFL = 2.09–2.68 mm), weakly to moderately coriarious to granulate, dull to weakly shining.</paragraph>
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<paragraph id="1F0B3690FFC2FF936584115AD2AA2133" pageId="43" pageNumber="44">Peduncle of petiole about 0.8x length of petiolar node, anteroventral margin with bluntly angulate to angulate triangular process. In profile, posterior surface of petiolar node flattened; node asymmetrical with anterior surface shorter than posterior surface, apex broadly rounded. In dorsal view, petiolar node about 1.5x longer than wide, widest near anterior margin. Sides and posterior surface of petiolar node with weak to moderately coarse, irregular, transverse rugae, or granulate-punctate. Dorsum of postpetiole convex in profile; in dorsal view, widest near posterior margin, narrowing to anterior margin, maximum width about equal to length, strongly granulate-punctate, dull to weakly shining. First gastral tergum smooth, strongly shining.</paragraph>
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<paragraph id="1F0B3690FFC2FF9365841058D54121C7" pageId="43" pageNumber="44">
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Erect to semi-erect yellowish pilosity moderately abundant on head, variable in length, longest hairs approaching MOD. Moderately abundant suberect yellowish pilosity on scape; abundant decumbent hairs on funicular segments. Legs with moderately abundant subdecumbent to decumbent yellowish setae. Mesosoma, petiolar node, postpetiole, and gastral terga with moderately dense erect setae, mostly similar in length, longest hairs not exceeding MOD. Body concolorous tannish-orange to tannish-red (
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<figureCitation id="878F2A15FFC2FF9366B010C9D55021C7" box="[1011,1124,1980,2004]" captionStart="FIGURE 18" captionStartId="44.[151,249,1978,2000]" captionTargetBox="[450,1132,193,1956]" captionTargetId="figure@44.[450,1136,193,1956]" captionTargetPageId="44" captionText="FIGURE 18. Photograph of Pogonomyrmex cunicularius Mayr worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0103052). Photographs by April Nobile from www. AntWeb. org." pageId="43" pageNumber="44">
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<emphasis id="2DC0EA82FFC2FF9366B010C9D55021C7" bold="true" box="[1011,1124,1980,2004]" pageId="43" pageNumber="44">Figure 18</emphasis>
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</figureCitation>
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).
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</paragraph>
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<caption id="4BCB6618FFC5FF9465D410CFD5BD21FC" pageId="44" pageNumber="45" targetBox="[450,1132,193,1956]" targetPageId="44">
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<paragraph id="1F0B3690FFC5FF9465D410CFD5BD21FC" pageId="44" pageNumber="45">
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<emphasis id="2DC0EA82FFC5FF9465D410CFD02B21C3" bold="true" box="[151,287,1978,2000]" pageId="44" pageNumber="45">FIGURE 18.</emphasis>
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Photograph of
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<taxonomicName id="D8B44D13FFC5FF9464FF10CED22821C3" authority="Mayr" authorityName="Mayr" box="[444,796,1978,2000]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="cunicularius">
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<emphasis id="2DC0EA82FFC5FF9464FF10CED3EB21C3" box="[444,735,1978,2000]" italics="true" pageId="44" pageNumber="45">Pogonomyrmex cunicularius</emphasis>
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Mayr
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</taxonomicName>
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worker: (
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<emphasis id="2DC0EA82FFC5FF9466C310CFD2A621C3" bold="true" box="[896,914,1978,2000]" pageId="44" pageNumber="45">A</emphasis>
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) frontal view of head, (
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<emphasis id="2DC0EA82FFC5FF9461C110CED5A721C3" bold="true" box="[1154,1171,1979,2000]" pageId="44" pageNumber="45">B</emphasis>
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) lateral view of body, and (
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<emphasis id="2DC0EA82FFC5FF9465DC10ACD18521FC" bold="true" box="[159,177,2009,2031]" pageId="44" pageNumber="45">C</emphasis>
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) dorsal view of body (CASENT0103052). Photographs by April Nobile from www.AntWeb.org.
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</paragraph>
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</caption>
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<caption id="4BCB6618FFC4FF9565D410CFD5C621FC" pageId="45" pageNumber="46" targetBox="[412,1170,193,1955]" targetPageId="45">
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<paragraph id="1F0B3690FFC4FF9565D410CFD5C621FC" pageId="45" pageNumber="46">
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<emphasis id="2DC0EA82FFC4FF9565D410CFD01521C3" bold="true" box="[151,289,1978,2000]" pageId="45" pageNumber="46">FIGURE 19.</emphasis>
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Photograph of
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<taxonomicName id="D8B44D13FFC4FF95648110CED21221C3" authority="Mayr" authorityName="Mayr" box="[450,806,1978,2000]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="cunicularius">
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<emphasis id="2DC0EA82FFC4FF95648110CED3D321C3" box="[450,743,1978,2000]" italics="true" pageId="45" pageNumber="46">Pogonomyrmex cunicularius</emphasis>
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Mayr
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</taxonomicName>
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ergatoid queen: (
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<emphasis id="2DC0EA82FFC4FF95669F10CFD2DA21C3" bold="true" box="[988,1006,1978,2000]" pageId="45" pageNumber="46">A</emphasis>
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) frontal view of head, (
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<emphasis id="2DC0EA82FFC4FF9561A610CED5C221C3" bold="true" box="[1253,1270,1979,2000]" pageId="45" pageNumber="46">B</emphasis>
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) lateral view of body, and (
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<emphasis id="2DC0EA82FFC4FF95644B10ACD02E21FC" bold="true" box="[264,282,2009,2031]" pageId="45" pageNumber="46">C</emphasis>
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) dorsal view of body (CASENT0173109). Photographs by April Nobile from www.AntWeb.org.
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</paragraph>
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</caption>
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<caption id="4BCB6618FFC7FF9665D4107FD5BC212D" pageId="46" pageNumber="47" targetBox="[437,1141,193,1780]" targetPageId="46">
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<paragraph id="1F0B3690FFC7FF9665D4107FD5BC212D" pageId="46" pageNumber="47">
|
||
<emphasis id="2DC0EA82FFC7FF9665D4107FD0142133" bold="true" box="[151,288,1802,1824]" pageId="46" pageNumber="47">FIGURE 20.</emphasis>
|
||
Photograph of
|
||
<taxonomicName id="D8B44D13FFC7FF966482107ED2102133" authority="Mayr" authorityName="Mayr" box="[449,804,1802,1824]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFC7FF966482107ED3D12133" box="[449,741,1802,1824]" italics="true" pageId="46" pageNumber="47">Pogonomyrmex cunicularius</emphasis>
|
||
Mayr
|
||
</taxonomicName>
|
||
male: (
|
||
<emphasis id="2DC0EA82FFC7FF966630107FD2B12133" bold="true" box="[883,901,1802,1824]" pageId="46" pageNumber="47">A</emphasis>
|
||
) frontal view of head, (
|
||
<emphasis id="2DC0EA82FFC7FF96613F107ED5B92133" bold="true" box="[1148,1165,1803,1824]" pageId="46" pageNumber="47">B</emphasis>
|
||
) lateral view of body, and (
|
||
<emphasis id="2DC0EA82FFC7FF9665DC105DD185212D" bold="true" box="[159,177,1832,1854]" pageId="46" pageNumber="47">C</emphasis>
|
||
) dorsal view of body (CASENT0235298). Photographs by Will Ericson from www.AntWeb.org.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1F0B3690FFC7FF96658410F1D21E21F6" pageId="46" pageNumber="47">
|
||
<emphasis id="2DC0EA82FFC7FF96658410F1D33A218E" bold="true" box="[199,526,1924,1949]" pageId="46" pageNumber="47">Ergatoid Queen. Diagnosis.</emphasis>
|
||
This caste is diagnosed by: (1) ergatoid, with small ocelli on head, (2) in profile, petiolar node rounded, (3) inferior propodeal spines wider than high, apex broadly rounded, and (4) first gastral tergum smooth and polished, strongly shining (
|
||
<figureCitation id="878F2A15FFC7FF9667EB10B9D22E21F6" box="[680,794,1996,2021]" captionStart="FIGURE 19" captionStartId="45.[151,249,1978,2000]" captionTargetBox="[412,1170,193,1955]" captionTargetId="figure@45.[412,1175,193,1956]" captionTargetPageId="45" captionText="FIGURE 19. Photograph of Pogonomyrmex cunicularius Mayr ergatoid queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0173109). Photographs by April Nobile from www. AntWeb. org." pageId="46" pageNumber="47">
|
||
<emphasis id="2DC0EA82FFC7FF9667EB10B9D22E21F6" bold="true" box="[680,794,1996,2021]" pageId="46" pageNumber="47">Figure 19</emphasis>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1F0B3690FFC6FF97658417EDD3E726EB" pageId="47" pageNumber="48">
|
||
<emphasis id="2DC0EA82FFC6FF97658417EDD04226A3" bold="true" box="[199,374,152,176]" pageId="47" pageNumber="48">Measurements</emphasis>
|
||
—(
|
||
<emphasis id="2DC0EA82FFC6FF9764D817EDD09D26A3" box="[411,425,152,176]" italics="true" pageId="47" pageNumber="48">n</emphasis>
|
||
= 12). HL 2.26–2.66; HW 1.96–2.38; MOD 0.35–0.45; OMD 0.54–0.66; SL 1.53–1.87; PNW 1.23–1.60; HFL 2.27–2.70; ML 2.46–3.20; PW 0.51–0.67; PPW 0.71–0.93. Indices: SI 74.63–87.24; CI 85.38–90.49; OI 17.07–20.74; HFI 111.34–125.37.
|
||
</paragraph>
|
||
<paragraph id="1F0B3690FFC6FF9765841670D312279B" pageId="47" pageNumber="48">
|
||
<emphasis id="2DC0EA82FFC6FF9765841670D0BF270F" bold="true" box="[199,395,260,284]" pageId="47" pageNumber="48">Male. Diagnosis.</emphasis>
|
||
This caste is diagnosed by: (1) first gastral tergum lacking striae, (2) femur long (HFL> 1.95 mm), HFI> 150.0, (3) head weakly elongate (CI <100.0), (4) posterior surface of petiolar node rugose to rugoreticulate, (5) in profile, petiolar node rounded, (6) superior propodeal spines consist of teeth to short spines, and (7) notauli present (
|
||
<figureCitation id="878F2A15FFC6FF9764E7161AD321279B" box="[420,533,367,392]" captionStart="FIGURE 20" captionStartId="46.[151,249,1802,1824]" captionTargetBox="[437,1141,193,1780]" captionTargetId="figure@46.[437,1167,193,1780]" captionTargetPageId="46" captionText="FIGURE 20. Photograph of Pogonomyrmex cunicularius Mayr male: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0235298). Photographs by Will Ericson from www. AntWeb. org." pageId="47" pageNumber="48">
|
||
<emphasis id="2DC0EA82FFC6FF9764E7161AD321279B" bold="true" box="[420,533,367,392]" pageId="47" pageNumber="48">Figure 20</emphasis>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1F0B3690FFC6FF97658416E0D32227E7" pageId="47" pageNumber="48">
|
||
<emphasis id="2DC0EA82FFC6FF97658416E0D04227BF" bold="true" box="[199,374,405,428]" pageId="47" pageNumber="48">Measurements</emphasis>
|
||
—(
|
||
<emphasis id="2DC0EA82FFC6FF9764D816E1D09D27BF" box="[411,425,404,428]" italics="true" pageId="47" pageNumber="48">n</emphasis>
|
||
= 12). HL 1.30–1.77; HW 1.17–1.54; MOD 0.51–0.64; OMD 0.21–0.32; SL 0.38–0.49; HFL 1.98–2.48; ML 2.51–3.25; PW 0.48–0.67; PPW 0.63–0.97. Indices: SI 24.84–35.66; CI 82.89–92.81; OI 39.61–47.86; HFI 154.69–182.05.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="57AE651BFFC6FF976584168AD1E3254F" pageId="47" pageNumber="48" type="materials_examined">
|
||
<paragraph id="1F0B3690FFC6FF976584168AD1E3254F" pageId="47" pageNumber="48">
|
||
<emphasis id="2DC0EA82FFC6FF976584168AD24C240B" bold="true" box="[199,888,511,536]" pageId="47" pageNumber="48">
|
||
Additional material examined. ARGENTINA:
|
||
<emphasis id="2DC0EA82FFC6FF9767BD168AD24C240B" bold="true" box="[766,888,511,536]" italics="true" pageId="47" pageNumber="48">Corrientes</emphasis>
|
||
</emphasis>
|
||
: Rt 14 at km 423, 90 m, Feb 27, 2003 (CASC; RAJC); Mercedes, Feb 1918 (MACN; USNM).
|
||
<emphasis id="2DC0EA82FFC6FF9767871550D277242F" bold="true" box="[708,835,548,572]" italics="true" pageId="47" pageNumber="48">Entre Ríos</emphasis>
|
||
: Pueblo Liebig, 70’, Dec 17, 2005 & Feb 13, 2010 (CSC; MCZ; RAJC); Rt 14 at 4.5 km N San José/Colón exit, 80’, Dec 13, 2006 (RAJC); Rt 14 at 5.1 mi N Gualeguychú, 60’, Dec 17, 2005 (RAJC); Estancia Sosa, no date (MACN; MLPA); Hernandarias, Jun 7, 1951 (FML; MZUSP); El Palmar, Nov 24, 2005 (CSC); Parque Nacional El Palmar, Jan 5, 2006 (RGPC); La Picada, May 1951 (FML).
|
||
<emphasis id="2DC0EA82FFC6FF97643315C1D35424DF" bold="true" box="[368,608,692,716]" pageId="47" pageNumber="48">
|
||
URUGUAY:
|
||
<emphasis id="2DC0EA82FFC6FF97674F15C1D35424DF" bold="true" box="[524,608,692,716]" italics="true" pageId="47" pageNumber="48">Artigas</emphasis>
|
||
</emphasis>
|
||
: Pintado Grande, Feb 22, 1969 (FML; LACM); Grutas de Chiflero, Feb 22, 1961 & no date (FML; LACM); R de Chiflero, Feb 22, 1961 (FML); Rt 30, no date (FML).
|
||
<emphasis id="2DC0EA82FFC6FF97619B15A2D40024E3" bold="true" box="[1240,1332,727,752]" italics="true" pageId="47" pageNumber="48">Colonia</emphasis>
|
||
: Colonia Suissa (= Nueva Helvecia), Mar 11, 1969 (LACM).
|
||
<emphasis id="2DC0EA82FFC6FF9766471589D2BD2507" bold="true" box="[772,905,764,788]" italics="true" pageId="47" pageNumber="48">Maldonado</emphasis>
|
||
: La Sierra, no date (LACM; MCZ; MSNG).
|
||
<emphasis id="2DC0EA82FFC6FF9765D4146AD02A252B" bold="true" box="[151,286,799,824]" italics="true" pageId="47" pageNumber="48">Montevideo</emphasis>
|
||
: Montevideo, no date (MCZ; MLPA; USNM).
|
||
<emphasis id="2DC0EA82FFC6FF97660A146AD28F252B" bold="true" box="[841,955,799,824]" italics="true" pageId="47" pageNumber="48">Paysandú</emphasis>
|
||
: Paysandú, Feb 25, 1961 (FML) (
|
||
<emphasis id="2DC0EA82FFC6FF97600E146AD1F3254F" bold="true" pageId="47" pageNumber="48">
|
||
<figureCitation id="878F2A15FFC6FF97600E146AD186254F" captionStart="FIGURE 21" captionStartId="48.[151,249,1981,2003]" captionTargetBox="[408,1179,195,1959]" captionTargetId="figure@48.[408,1179,195,1959]" captionTargetPageId="48" captionText="FIGURE 21. Geographic distribution of: (A) Pogonomyrmex cunicularius Mayr, (B) Pogonomyrmex pencosensis Forel, and (C) Pogonomyrmex serpens Santschi. The larger black circle in each panel (when present) denotes the type locality." pageId="47" pageNumber="48">Figure 21</figureCitation>
|
||
A
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="57AE651BFFC6FF976584141DD2A325FF" pageId="47" pageNumber="48" type="etymology">
|
||
<paragraph id="1F0B3690FFC6FF976584141DD2A325FF" pageId="47" pageNumber="48">
|
||
<emphasis id="2DC0EA82FFC6FF976584141DD07A2593" bold="true" box="[199,334,872,896]" pageId="47" pageNumber="48">Etymology.</emphasis>
|
||
The specific epithet,
|
||
<taxonomicName id="D8B44D13FFC6FF97677C1412D3FF2593" box="[575,715,871,896]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFC6FF97677C1412D3FF2593" box="[575,715,871,896]" italics="true" pageId="47" pageNumber="48">cunicularius</emphasis>
|
||
</taxonomicName>
|
||
, (from Latin
|
||
<emphasis id="2DC0EA82FFC6FF9766221412D2802593" box="[865,948,871,896]" italics="true" pageId="47" pageNumber="48">cunicul</emphasis>
|
||
-, which indicates a miner or burrower, plus the Latin suffix -
|
||
<emphasis id="2DC0EA82FFC6FF97641A14F9D0A725B7" box="[345,403,908,932]" italics="true" pageId="47" pageNumber="48">arius</emphasis>
|
||
, which denotes belonging to) apparently refers to the soil nests excavated by this species. Mayr did not discuss the naming of this species, but Kusnezov (1949) wrote that
|
||
<taxonomicName id="D8B44D13FFC6FF97611014C5D5C925DB" box="[1107,1277,943,968]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFC6FF97611014C5D5C925DB" box="[1107,1277,943,968]" italics="true" pageId="47" pageNumber="48">P. cunicularius</emphasis>
|
||
</taxonomicName>
|
||
was the only Argentinian species of
|
||
<taxonomicName id="D8B44D13FFC6FF9764DA14A1D37D25FF" box="[409,585,980,1004]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2DC0EA82FFC6FF9764DA14A1D37D25FF" box="[409,585,980,1004]" italics="true" pageId="47" pageNumber="48">Pogonomyrmex</emphasis>
|
||
</taxonomicName>
|
||
that constructed a nest crater.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="57AE651BFFC6FF9765841482D5EA23F7" pageId="47" pageNumber="48" type="discussion">
|
||
<paragraph id="1F0B3690FFC6FF9765841482D43222D7" pageId="47" pageNumber="48">
|
||
<emphasis id="2DC0EA82FFC6FF9765841482D0782203" bold="true" box="[199,332,1015,1040]" pageId="47" pageNumber="48">Discussion.</emphasis>
|
||
<taxonomicName id="D8B44D13FFC6FF97641A148DD3952203" box="[345,673,1015,1040]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFC6FF97641A148DD3952203" box="[345,673,1015,1040]" italics="true" pageId="47" pageNumber="48">Pogonomyrmex cunicularius</emphasis>
|
||
</taxonomicName>
|
||
is not known to co-occur with any other
|
||
<taxonomicName id="D8B44D13FFC6FF9761E7148DD4652203" box="[1188,1361,1015,1040]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFC6FF9761E7148DD4652203" box="[1188,1361,1015,1040]" italics="true" pageId="47" pageNumber="48">P. cunicularius</emphasis>
|
||
</taxonomicName>
|
||
-group species but additional collections may show it co-occurs with both
|
||
<taxonomicName id="D8B44D13FFC6FF9766D31369D5062227" box="[912,1074,1052,1076]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pencosensis">
|
||
<emphasis id="2DC0EA82FFC6FF9766D31369D2AA2227" box="[912,926,1052,1076]" italics="true" pageId="47" pageNumber="48">P</emphasis>
|
||
.
|
||
<emphasis id="2DC0EA82FFC6FF9766EE1369D5062227" box="[941,1074,1052,1076]" italics="true" pageId="47" pageNumber="48">pencosensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D8B44D13FFC6FF9761281369D5E92227" box="[1131,1245,1052,1076]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="serpens">
|
||
<emphasis id="2DC0EA82FFC6FF9761281369D5E92227" box="[1131,1245,1052,1076]" italics="true" pageId="47" pageNumber="48">P. serpens</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="D8B44D13FFC6FF9761AF1369D017224B" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFC6FF9761AF1369D017224B" italics="true" pageId="47" pageNumber="48">Pogonomyrmex cunicularius</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from these two species based on the following characters: (1) in profile, petiolar node broadly rounded, (2) inferior propodeal spines rounded, and (3) first gastral tergum smooth and strongly shining. In
|
||
<taxonomicName id="D8B44D13FFC6FF97645513FDD08C22B3" box="[278,440,1160,1184]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pencosensis">
|
||
<emphasis id="2DC0EA82FFC6FF97645513FDD08C22B3" box="[278,440,1160,1184]" italics="true" pageId="47" pageNumber="48">P. pencosensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D8B44D13FFC6FF9764B313FDD35522B3" box="[496,609,1160,1184]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="serpens">
|
||
<emphasis id="2DC0EA82FFC6FF9764B313FDD35522B3" box="[496,609,1160,1184]" italics="true" pageId="47" pageNumber="48">P. serpens</emphasis>
|
||
</taxonomicName>
|
||
: (1) in profile, the petiolar node is angulate, (2) inferior propodeal spines are acuminate, and (3) first gastral tergum is weakly to moderately coriarious, dull to weakly shining.
|
||
</paragraph>
|
||
<paragraph id="1F0B3690FFC6FF97658413BAD5EA23F7" pageId="47" pageNumber="48">
|
||
Santschi erected
|
||
<taxonomicName id="D8B44D13FFC6FF9764C713A5D3D522FB" box="[388,737,1231,1256]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="variety" species="cunicularius" variety="brevispinus">
|
||
<emphasis id="2DC0EA82FFC6FF9764C713A5D31F22FB" box="[388,555,1231,1256]" italics="true" pageId="47" pageNumber="48">P. cunicularius</emphasis>
|
||
var.
|
||
<emphasis id="2DC0EA82FFC6FF97672213BAD3D522FB" box="[609,737,1231,1256]" italics="true" pageId="47" pageNumber="48">brevispinus</emphasis>
|
||
</taxonomicName>
|
||
because, compared to the type specimen, the propodeal spines were shorter (no longer than maximum eye diameter), the petiolar node was narrower, and the postpetiole was slightly longer than wide along the posterior margin. Kusnezov (1951) synonomized
|
||
<taxonomicName id="D8B44D13FFC6FF9761F1126DD0232347" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="variety" species="cunicularius" variety="brevispinus">
|
||
<emphasis id="2DC0EA82FFC6FF9761F1126DD4572323" box="[1202,1379,1303,1328]" italics="true" pageId="47" pageNumber="48">P. cunicularius</emphasis>
|
||
var.
|
||
<emphasis id="2DC0EA82FFC6FF9765D41249D0232347" box="[151,279,1340,1364]" italics="true" pageId="47" pageNumber="48">brevispinus</emphasis>
|
||
</taxonomicName>
|
||
under
|
||
<taxonomicName id="D8B44D13FFC6FF9764251249D33A2347" box="[358,526,1340,1364]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFC6FF9764251249D33A2347" box="[358,526,1340,1364]" italics="true" pageId="47" pageNumber="48">P. cunicularius</emphasis>
|
||
</taxonomicName>
|
||
without discussion, and I concur with his synonomy. The propodeal spines were slightly shorter in the syntype worker of
|
||
<taxonomicName id="D8B44D13FFC6FF97673A1215D2D5236B" box="[633,993,1375,1400]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="variety" species="cunicularius" variety="brevispinus">
|
||
<emphasis id="2DC0EA82FFC6FF97673A1215D210236B" box="[633,804,1375,1400]" italics="true" pageId="47" pageNumber="48">P. cunicularius</emphasis>
|
||
var.
|
||
<emphasis id="2DC0EA82FFC6FF976622122AD2D5236B" box="[865,993,1375,1400]" italics="true" pageId="47" pageNumber="48">brevispinus</emphasis>
|
||
</taxonomicName>
|
||
that I examined, but their length was within the range of variation observed among workers. The width of the petiolar node and shape of the postpetiole also were within the range of variation observed among workers. Finally, the syntype ergatoid queen of
|
||
<taxonomicName id="D8B44D13FFC6FF9760C412DDD0EE23F7" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="variety" species="cunicularius" variety="brevispinus">
|
||
<emphasis id="2DC0EA82FFC6FF9760C412DDD01723F7" italics="true" pageId="47" pageNumber="48">P. cunicularius</emphasis>
|
||
var.
|
||
<emphasis id="2DC0EA82FFC6FF97641912B9D0EE23F7" box="[346,474,1484,1508]" italics="true" pageId="47" pageNumber="48">brevispinus</emphasis>
|
||
</taxonomicName>
|
||
and queens of
|
||
<taxonomicName id="D8B44D13FFC6FF9767C712B9D21F23F7" box="[644,811,1484,1508]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFC6FF9767C712B9D21F23F7" box="[644,811,1484,1508]" italics="true" pageId="47" pageNumber="48">P. cunicularius</emphasis>
|
||
</taxonomicName>
|
||
did not display noticeable differences.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="57AE651BFFC6FF976584129AD29F2163" pageId="47" pageNumber="48" type="biology_ecology">
|
||
<paragraph id="1F0B3690FFC6FF976584129AD1E1208B" pageId="47" pageNumber="48">
|
||
<emphasis id="2DC0EA82FFC6FF976584129AD012201B" bold="true" box="[199,294,1519,1544]" pageId="47" pageNumber="48">Biology.</emphasis>
|
||
The biology of
|
||
<taxonomicName id="D8B44D13FFC6FF97649C1285D3B3201B" box="[479,647,1519,1544]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFC6FF97649C1285D3B3201B" box="[479,647,1519,1544]" italics="true" pageId="47" pageNumber="48">P. cunicularius</emphasis>
|
||
</taxonomicName>
|
||
is poorly known but it is likely similar to that of
|
||
<taxonomicName id="D8B44D13FFC6FF9761FA1285D46F201B" box="[1209,1371,1520,1544]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pencosensis">
|
||
<emphasis id="2DC0EA82FFC6FF9761FA1285D46F201B" box="[1209,1371,1520,1544]" italics="true" pageId="47" pageNumber="48">P. pencosensis</emphasis>
|
||
</taxonomicName>
|
||
. This species is a solitary forager that can travel>25 m from the nest. Nests of
|
||
<taxonomicName id="D8B44D13FFC6FF9766AC1161D5AD203F" box="[1007,1177,1556,1580]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFC6FF9766AC1161D5AD203F" box="[1007,1177,1556,1580]" italics="true" pageId="47" pageNumber="48">P. cunicularius</emphasis>
|
||
</taxonomicName>
|
||
usually have a pebble tumulus up to 25 cm in diameter with a nest entrance that is up to 2–3 cm in diameter. Colonies of
|
||
<taxonomicName id="D8B44D13FFC6FF9761B7114DD4A82043" box="[1268,1436,1591,1616]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFC6FF9761B7114DD4A82043" box="[1268,1436,1591,1616]" italics="true" pageId="47" pageNumber="48">P. cunicularius</emphasis>
|
||
</taxonomicName>
|
||
probably contain 500 to>1000 workers, and they can produce>100 ergatoid queens and numerous males (pers. obs.).
|
||
</paragraph>
|
||
<paragraph id="1F0B3690FFC6FF97658411D1D29F2163" pageId="47" pageNumber="48">
|
||
Sexuals have been collected from nests from 13 December to 13 February, and one founding queen was excavated on 17 February, indicating the mating flights occur during the austral summer. The large number of ergatoid queens produced by colonies (>100) infers that queens use independent colony founding (see Peeters
|
||
<emphasis id="2DC0EA82FFC6FF9760CB1199D180213B" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
|
||
, 2012), which is an unusual behavior for ergatoid queens (see Johnson, 2010). One excavated queen was haplometrotic, and founding queens are probably semi-claustral (they forage) (see Johnson, 2010). Queens contained 9–12 ovarioles (
|
||
<emphasis id="2DC0EA82FFC6FF976482102DD0FB2163" box="[449,463,1880,1904]" italics="true" pageId="47" pageNumber="48">n</emphasis>
|
||
= 3), compared to four in workers (
|
||
<emphasis id="2DC0EA82FFC6FF976622102DD25B2163" box="[865,879,1880,1904]" italics="true" pageId="47" pageNumber="48">n</emphasis>
|
||
= 6).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="4BCB6618FFD9FF8865D410C8D41F21E1" pageId="48" pageNumber="49" targetBox="[408,1179,195,1959]" targetPageId="48">
|
||
<paragraph id="1F0B3690FFD9FF8865D410C8D41F21E1" pageId="48" pageNumber="49">
|
||
<emphasis id="2DC0EA82FFD9FF8865D410C8D01521C0" bold="true" box="[151,289,1981,2003]" pageId="48" pageNumber="49">FIGURE 21.</emphasis>
|
||
Geographic distribution of: (
|
||
<emphasis id="2DC0EA82FFD9FF88670C10C8D35621C0" bold="true" box="[591,610,1981,2003]" pageId="48" pageNumber="49">A</emphasis>
|
||
)
|
||
<taxonomicName id="D8B44D13FFD9FF88673110CBD2E321C0" authority="Mayr" authorityName="Mayr" box="[626,983,1981,2003]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFD9FF88673110CBD2A321C0" box="[626,919,1981,2003]" italics="true" pageId="48" pageNumber="49">Pogonomyrmex cunicularius</emphasis>
|
||
Mayr
|
||
</taxonomicName>
|
||
, (
|
||
<emphasis id="2DC0EA82FFD9FF8866A810CBD2C821C0" bold="true" box="[1003,1020,1982,2003]" pageId="48" pageNumber="49">B</emphasis>
|
||
)
|
||
<taxonomicName id="D8B44D13FFD9FF88614F10CBD45F21C0" authority="Forel" authorityName="Forel" box="[1036,1387,1981,2003]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="pencosensis">
|
||
<emphasis id="2DC0EA82FFD9FF88614F10CBD41821C0" box="[1036,1324,1982,2003]" italics="true" pageId="48" pageNumber="49">Pogonomyrmex pencosensis</emphasis>
|
||
Forel
|
||
</taxonomicName>
|
||
, and (
|
||
<emphasis id="2DC0EA82FFD9FF8865DC10A9D18521E1" bold="true" box="[159,177,2012,2034]" pageId="48" pageNumber="49">C</emphasis>
|
||
)
|
||
<taxonomicName id="D8B44D13FFD9FF88658310A8D37721E1" authority="Santschi. The" authorityName="Santschi. The" box="[192,579,2012,2034]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="serpens">
|
||
<emphasis id="2DC0EA82FFD9FF88658310A8D08721E1" box="[192,435,2013,2034]" italics="true" pageId="48" pageNumber="49">Pogonomyrmex serpens</emphasis>
|
||
Santschi. The
|
||
</taxonomicName>
|
||
larger black circle in each panel (when present) denotes the type locality.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="57AE651BFFD8FF89658417EDD5792753" pageId="49" pageNumber="50" type="distribution">
|
||
<paragraph id="1F0B3690FFD8FF89658417EDD5792753" pageId="49" pageNumber="50">
|
||
<taxonomicName id="D8B44D13FFD8FF89658417EDD33E26A3" box="[199,522,151,176]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFD8FF89658417EDD33E26A3" box="[199,522,151,176]" italics="true" pageId="49" pageNumber="50">Pogonomyrmex cunicularius</emphasis>
|
||
</taxonomicName>
|
||
is a lowland species that occurs at elevations from 20–90 m. This species is only known to occur east and north of Río Paraná in the Espinal, Uruguayan Savanna, and Humid Pampas ecoregions as defined by Olson
|
||
<emphasis id="2DC0EA82FFD8FF89641E1795D0B926EB" box="[349,397,223,248]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
|
||
. (2001) (
|
||
<emphasis id="2DC0EA82FFD8FF8964B617AAD34E26EB" bold="true" box="[501,634,223,248]" pageId="49" pageNumber="50">
|
||
<figureCitation id="878F2A15FFD8FF8964B617AAD35126EB" box="[501,613,223,248]" captionStart="FIGURE 21" captionStartId="48.[151,249,1981,2003]" captionTargetBox="[408,1179,195,1959]" captionTargetId="figure@48.[408,1179,195,1959]" captionTargetPageId="48" captionText="FIGURE 21. Geographic distribution of: (A) Pogonomyrmex cunicularius Mayr, (B) Pogonomyrmex pencosensis Forel, and (C) Pogonomyrmex serpens Santschi. The larger black circle in each panel (when present) denotes the type locality." pageId="49" pageNumber="50">Figure 21</figureCitation>
|
||
A
|
||
</emphasis>
|
||
). The very wide Río Paraná may act as a barrier to dispersal by ergatoid queens. At present,
|
||
<taxonomicName id="D8B44D13FFD8FF89643C1671D31E270F" box="[383,554,260,284]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="cunicularius">
|
||
<emphasis id="2DC0EA82FFD8FF89643C1671D31E270F" box="[383,554,260,284]" italics="true" pageId="49" pageNumber="50">P. cunicularius</emphasis>
|
||
</taxonomicName>
|
||
appears to be rare with very fragmented populations that likely result from intense and widespread agricultural use over most of its historic range (
|
||
<emphasis id="2DC0EA82FFD8FF8966F41652D5092753" bold="true" box="[951,1085,295,320]" pageId="49" pageNumber="50">
|
||
<figureCitation id="878F2A15FFD8FF8966F41652D51C2753" box="[951,1064,295,320]" captionStart="FIGURE 21" captionStartId="48.[151,249,1981,2003]" captionTargetBox="[408,1179,195,1959]" captionTargetId="figure@48.[408,1179,195,1959]" captionTargetPageId="48" captionText="FIGURE 21. Geographic distribution of: (A) Pogonomyrmex cunicularius Mayr, (B) Pogonomyrmex pencosensis Forel, and (C) Pogonomyrmex serpens Santschi. The larger black circle in each panel (when present) denotes the type locality." pageId="49" pageNumber="50">Figure 21</figureCitation>
|
||
A
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |