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<document id="ED2FF37BFDE0B2680D934AEF1DE32DF0" ID-DOI="10.5281/zenodo.7666260" ID-ISSN="2305-2562" ID-Zenodo-Dep="7666260" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677090874998" checkinUser="felipe" docAuthor="Plisko, Jadwiga Danuta" docDate="2003" docId="967287D60602AE74FE78FA343A19FB58" docLanguage="en" docName="AfrInvertebr.44.2.279325.pdf" docOrigin="African Invertebrates 44 (2)" docStyle="DocumentStyle:C20B10C002986E51083406A4EACF003A.3:AfrInvertebr.2001-2007.journal_article" docStyleId="C20B10C002986E51083406A4EACF003A" docStyleName="AfrInvertebr.2001-2007.journal_article" docStyleVersion="3" docTitle="Microchaetus distasmosus Plisko 2003, sp.n." docType="treatment" docVersion="3" lastPageNumber="284" masterDocId="6A4BFFAE0601AE71FFA3FFAE3E7DFF87" masterDocTitle="Eleven new South African earthworms (Oligochaeta: Microchaetidae) with new information on some known species, and an inventory of the microchaetids of KwaZulu-Natal" masterLastPageNumber="325" masterPageNumber="279" pageNumber="282" updateTime="1699455353883" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="D66FA9941970C33AE560F99F9273BE97" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="90FAE676824A81537EDE0BAD74768584">
<mods:title id="2258F6064C628A898C78A7D816B0147D">Eleven new South African earthworms (Oligochaeta: Microchaetidae) with new information on some known species, and an inventory of the microchaetids of KwaZulu-Natal</mods:title>
</mods:titleInfo>
<mods:name id="C72390E1CC16582B9AF33598C1925889" type="personal">
<mods:role id="D3161B65EEBDCFBD7CC5BAB7A59FDF17">
<mods:roleTerm id="0FF286FEE7F8429C5620B48F9105ACCE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="99E07E8FF01EF99EA6E9D83AD4BF8352">Plisko, Jadwiga Danuta</mods:namePart>
</mods:name>
<mods:typeOfResource id="46B0CF0017D63471B1FD627A4C970E57">text</mods:typeOfResource>
<mods:relatedItem id="4B674657E52D0F996B533D49F1546F72" type="host">
<mods:titleInfo id="01DE74F4E6B57738E8FF465318E7478F">
<mods:title id="667CD5C2A99028032D146ECE8C6EABFA">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="2448E3868763C32100DF426BE8526C62">
<mods:date id="CBD38A93919724C78417A0614EEDAE42">2003</mods:date>
<mods:detail id="1FC23796776980CB04E016BE525D2992" type="pubDate">
<mods:number id="37F37D31A4AC4391AFC35D8A1E9041C2">2003-12-31</mods:number>
</mods:detail>
<mods:detail id="9A86EBA9FF3D5901A905E45F589FCD3D" type="volume">
<mods:number id="50453EDB998D46682AF64291DD029D16">44</mods:number>
</mods:detail>
<mods:detail id="6835782968C07776571E9EF38238BD94" type="issue">
<mods:number id="5449D3348D4639BE826CD1E849EFF7B0">2</mods:number>
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<mods:extent id="077EC57604F32567E6C21945F6C352CF" unit="page">
<mods:start id="C3EA6221ADEC1CF2CE8AB329D11B3541">279</mods:start>
<mods:end id="15678CD359FEA13F812B99EA81CE494C">325</mods:end>
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<mods:classification id="8BFF5EFF5226F2477189E8A04D7D7BD4">journal article</mods:classification>
<mods:identifier id="673B05221271582271F124FDA7DE40E4" type="DOI">10.5281/zenodo.7666260</mods:identifier>
<mods:identifier id="89EA2225366DFB53FB09C8740A61C9A1" type="ISSN">2305-2562</mods:identifier>
<mods:identifier id="BC1D787486CFE679E5B4E7EA35E9FE78" type="Zenodo-Dep">7666260</mods:identifier>
</mods:mods>
<treatment id="967287D60602AE74FE78FA343A19FB58" ID-DOI="http://doi.org/10.5281/zenodo.7666915" ID-Zenodo-Dep="7666915" LSID="urn:lsid:plazi:treatment:967287D60602AE74FE78FA343A19FB58" httpUri="http://treatment.plazi.org/id/967287D60602AE74FE78FA343A19FB58" lastPageId="5" lastPageNumber="284" pageId="3" pageNumber="282">
<subSubSection id="56C1654B0602AE72FE78FA343D2DFA34" box="[475,848,1434,1459]" pageId="3" pageNumber="282" type="nomenclature">
<paragraph id="1E6436C00602AE72FE78FA343D2DFA34" blockId="3.[181,1142,1434,1670]" box="[475,848,1434,1459]" pageId="3" pageNumber="282">
<heading id="452C81AC0602AE72FE78FA343D2DFA34" box="[475,848,1434,1459]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="282" reason="2">
<taxonomicName id="D9DB4D430602AE72FE78FA343D69FA34" ID-CoL="853G4" authority="Plisko, 2003" authorityName="Plisko" authorityYear="2003" box="[475,788,1434,1459]" class="Clitellata" family="Microchaetidae" genus="Microchaetus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="3" pageNumber="282" phylum="Annelida" rank="species" species="distasmosus" status="sp. nov.">
<emphasis id="2CAFEAD20602AE72FE78FA343D69FA34" bold="true" box="[475,788,1434,1459]" pageId="3" pageNumber="282">Microchaetus distasmosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="379C57A90602AE72FCB8FA343D2DFA34" box="[795,848,1434,1459]" pageId="3" pageNumber="282" rank="species">sp.n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="56C1654B0602AE74FDF8FA113A19FB58" lastPageId="5" lastPageNumber="284" pageId="3" pageNumber="282" type="description">
<paragraph id="1E6436C00602AE72FDF8FA113CB2FA5F" blockId="3.[181,1142,1434,1670]" box="[603,719,1471,1496]" pageId="3" pageNumber="282">
(
<figureCitation id="86E02A450602AE72FDC0FA113CB5FA5F" box="[611,712,1471,1496]" captionStart="Figs 23" captionStartId="4.[144,183,459,479]" captionTargetBox="[146,1105,215,436]" captionTargetId="figure-386@4.[144,1105,281,361]" captionTargetPageId="4" captionText="Figs 23. Microchaetus distasmosus sp. n. 2. Holotype, anterior part of body, ventral view with clitellum and tubercula pubertatis. 3. Paratype, clitellar region, enlarged. [C = clitellum, T = tuberculum pubertatis]." figureDoi="http://doi.org/10.5281/zenodo.7666264" httpUri="https://zenodo.org/record/7666264/files/figure.png" pageId="3" pageNumber="282">Figs 23</figureCitation>
)
</paragraph>
<paragraph id="1E6436C00602AE72FF16FA4B3C42F999" blockId="3.[181,1142,1434,1670]" pageId="3" pageNumber="282">
Etymology: G.
<emphasis id="2CAFEAD20602AE72FEC0FA4B3FACFA7A" box="[355,465,1509,1533]" italics="true" pageId="3" pageNumber="282">distasmos</emphasis>
= uncertainty. Refers to the unusual combination of internal characters observed in this species.
</paragraph>
<paragraph id="1E6436C00602AE75FF16F9833A39FD1C" blockId="3.[181,1142,1434,1670]" lastBlockId="4.[144,1104,578,1667]" lastPageId="4" lastPageNumber="283" pageId="3" pageNumber="282">
<materialsCitation id="AEB33C9D0602AE75FF16F9833A43FD1C" collectingDate="1990-02-24" collectingDateMax="1996-11-26" collectingDateMin="1990-02-24" collectionCode="NMSA, JDP" county="Material" lastPageId="4" lastPageNumber="283" latitude="-28.366667" location="Umfolozi Game Reserve" longLatPrecision="1232" longitude="31.866667" municipality="Mapelane Nature Reserve" pageId="3" pageNumber="282" specimenCount="2" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<collectingCounty id="F7054E4C0602AE72FF16F9833F69F9C1" box="[181,276,1581,1606]" pageId="3" pageNumber="282">Material</collectingCounty>
examined:
<emphasis id="2CAFEAD20602AE72FE3BF9823C29F9C2" bold="true" box="[408,596,1580,1605]" pageId="3" pageNumber="282">
<collectingRegion id="DC1FF8220602AE72FE3BF9823C29F9C2" box="[408,596,1580,1605]" country="South Africa" name="KwaZulu-Natal" pageId="3" pageNumber="282">KwaZulu­Natal</collectingRegion>
</emphasis>
:
<typeStatus id="C16088620602AE72FDC7F9833CB1F9C1" box="[612,716,1581,1606]" pageId="3" pageNumber="282" type="holotype">Holotype</typeStatus>
<collectionCode id="78CAAE050602AE72FD77F9833D5AF9C1" box="[724,807,1581,1606]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="282" type="Museum">NMSA</collectionCode>
/Olig.02532
<collectingMunicipality id="FE00ACBA0602AE72FC10F9833F72F9E1" pageId="3" pageNumber="282">Mapelane Nature Reserve</collectingMunicipality>
(
<geoCoordinate id="7BEF50070602AE72FEBCF9E33F05F9E1" box="[287,376,1613,1638]" degrees="28" direction="south" minutes="54" orientation="latitude" pageId="3" pageNumber="282" precision="925" value="-28.9">28º54'S</geoCoordinate>
:
<geoCoordinate id="7BEF50070602AE72FED8F9E33FA8F9E1" box="[379,469,1613,1638]" degrees="31" direction="east" minutes="55" orientation="longitude" pageId="3" pageNumber="282" precision="925" value="31.916666">31º55'E</geoCoordinate>
) at the edge of forest and grassland, from moist, sandy soil,
<date id="6A6510000602AE72FF16F9C33FEFF901" box="[181,402,1645,1670]" pageId="3" pageNumber="282" value="1996-11-26">
<collectingDate id="7A21E9E80602AE72FF16F9C33FEFF901" box="[181,402,1645,1670]" pageId="3" pageNumber="282" value="1996-11-26">26 November 1996</collectingDate>
</date>
,
<collectionCode id="78CAAE050602AE72FE3EF9C33FB1F901" box="[413,460,1645,1670]" pageId="3" pageNumber="282">JDP</collectionCode>
, TL.
<typeStatus id="C16088620602AE72FDABF9C33C04F901" box="[520,633,1645,1670]" pageId="3" pageNumber="282" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="78CAAE050602AE72FD27F9C33CA5F901" box="[644,728,1645,1670]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="282" type="Museum">NMSA</collectionCode>
/Olig.02377, 6 not fully mature, with tubercula pubertatis, collected together with holotype;
<collectionCode id="78CAAE050605AE75FD5AFDEC3D31FDDC" box="[761,844,578,603]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="4" pageNumber="283" type="Museum">NMSA</collectionCode>
/Olig.00626a
<location id="1B04601B0605AE75FC47FDEC3F4AFDFC" LSID="urn:lsid:plazi:treatment:967287D60602AE74FE78FA343A19FB58:1B04601B0605AE75FC47FDEC3F4AFDFC" county="Material" latitude="-28.366667" longLatPrecision="1232" longitude="31.866667" municipality="Mapelane Nature Reserve" name="Umfolozi Game Reserve" pageId="4" pageNumber="283" stateProvince="KwaZulu-Natal">Umfolozi Game Reserve</location>
(
<geoCoordinate id="7BEF50070605AE75FEEAFDCC3FDEFDFC" box="[329,419,610,635]" degrees="28" direction="south" minutes="22" orientation="latitude" pageId="4" pageNumber="283" precision="925" value="-28.366667">28º22'S</geoCoordinate>
:
<geoCoordinate id="7BEF50070605AE75FE12FDCC3C77FDFC" box="[433,522,610,635]" degrees="31" direction="east" minutes="52" orientation="longitude" pageId="4" pageNumber="283" precision="925" value="31.866667">31º52'E</geoCoordinate>
) on high bank of
<location id="1B04601B0605AE75FD7DFDCC3DACFDFC" LSID="urn:lsid:plazi:treatment:967287D60602AE74FE78FA343A19FB58:1B04601B0605AE75FD7DFDCC3DACFDFC" box="[734,977,610,635]" county="Material" latitude="-28.366667" longLatPrecision="1232" longitude="31.866667" municipality="Mapelane Nature Reserve" name="White Mfolozi River" pageId="4" pageNumber="283" stateProvince="KwaZulu-Natal">White Mfolozi River</location>
, grassland, sandy moist soil,
<date id="6A6510000605AE75FEF7FD2C3C63FD1C" box="[340,542,642,667]" pageId="4" pageNumber="283" value="1990-02-24">
<collectingDate id="7A21E9E80605AE75FEF7FD2C3C63FD1C" box="[340,542,642,667]" pageId="4" pageNumber="283" value="1990-02-24">24 February 1990</collectingDate>
</date>
, 20 cl + 10 juv,
<collectionCode id="78CAAE050605AE75FD72FD2C3C82FD1C" box="[721,767,642,667]" pageId="4" pageNumber="283">JDP</collectionCode>
[associated with
<taxonomicName id="D9DB4D430605AE75FC61FD2C3A49FD1D" baseAuthorityName="Kinberg" baseAuthorityYear="1867" box="[962,1076,642,666]" class="Clitellata" family="Microchaetidae" genus="Microchaetus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="4" pageNumber="283" phylum="Annelida" rank="species" species="natalensis">
<emphasis id="2CAFEAD20605AE75FC61FD2C3A49FD1D" box="[962,1076,642,666]" italics="true" pageId="4" pageNumber="283">natalensis</emphasis>
</taxonomicName>
]
</materialsCitation>
.
</paragraph>
<caption id="4AA466480605AE75FF33FE653F25FD88" ID-DOI="http://doi.org/10.5281/zenodo.7666264" ID-Zenodo-Dep="7666264" httpUri="https://zenodo.org/record/7666264/files/figure.png" pageId="4" pageNumber="283" startId="4.[144,183,459,479]" targetBox="[146,1105,215,436]" targetPageId="4">
<paragraph id="1E6436C00605AE75FF33FE653F25FD88" blockId="4.[144,1104,459,527]" pageId="4" pageNumber="283">
Figs 23.
<taxonomicName id="D9DB4D430605AE75FF48FE653FA6FE58" authority="Plisko, 2003" authorityName="Plisko" authorityYear="2003" box="[235,475,459,479]" class="Clitellata" family="Microchaetidae" genus="Microchaetus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="4" pageNumber="283" phylum="Annelida" rank="species" species="distasmosus" status="sp. nov.">
<emphasis id="2CAFEAD20605AE75FF48FE653FA6FE58" box="[235,475,459,479]" italics="true" pageId="4" pageNumber="283">Microchaetus distasmosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="379C57A90605AE75FE41FE653C69FE58" box="[482,532,459,479]" pageId="4" pageNumber="283" rank="species">sp. n.</taxonomicNameLabel>
2. Holotype, anterior part of body, ventral view with clitellum and tubercula pubertatis. 3. Paratype, clitellar region, enlarged. [C = clitellum, T = tuberculum pubertatis].
</paragraph>
</caption>
<paragraph id="1E6436C00605AE75FF33FD043CF3FD44" blockId="4.[144,1104,578,1667]" box="[144,654,682,707]" pageId="4" pageNumber="283">
Description based on
<typeStatus id="C16088620605AE75FE20FD043F98FD44" box="[387,485,682,707]" pageId="4" pageNumber="283" type="holotype">holotype</typeStatus>
and
<typeStatus id="C16088620605AE75FDB8FD043CF7FD44" box="[539,650,682,707]" pageId="4" pageNumber="283" type="paratype">paratypes</typeStatus>
.
</paragraph>
<paragraph id="1E6436C00605AE75FF33FD7C3F12FD6C" blockId="4.[144,1104,578,1667]" box="[144,367,722,747]" pageId="4" pageNumber="283">External characters:</paragraph>
<paragraph id="1E6436C00605AE75FF33FD543E81FB34" blockId="4.[144,1104,578,1667]" pageId="4" pageNumber="283">
<emphasis id="2CAFEAD20605AE75FF33FD543E97FC95" box="[144,234,762,786]" italics="true" pageId="4" pageNumber="283">General</emphasis>
: body cylindrical, extended during preservation.
<emphasis id="2CAFEAD20605AE75FD59FD543D35FC95" box="[762,840,762,786]" italics="true" pageId="4" pageNumber="283">Colour</emphasis>
:
<typeStatus id="C16088620605AE75FCF6FD543DC8FC94" box="[853,949,762,787]" pageId="4" pageNumber="283" type="holotype">holotype</typeStatus>
and
<typeStatus id="C16088620605AE75FC45FD543A32FC94" box="[998,1103,762,787]" pageId="4" pageNumber="283" type="paratype">paratypes</typeStatus>
preserved in alcohol for over six years do not reveal pigmentation; grey.
<emphasis id="2CAFEAD20605AE75FC60FCB43A3AFCB5" box="[963,1095,794,818]" italics="true" pageId="4" pageNumber="283">Dimensions</emphasis>
:
<typeStatus id="C16088620605AE75FF33FC943E8FFCD4" box="[144,242,826,851]" pageId="4" pageNumber="283" type="holotype">holotype</typeStatus>
preserved, much extended,
<quantity id="D9239B250605AE75FD91FC943CFFFCD4" box="[562,642,826,851]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="283" unit="mm" value="75.0">75 mm</quantity>
long,
<quantity id="D9239B250605AE75FD6FFC943D73FCD4" box="[716,782,826,851]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="283" unit="mm" value="2.0">2 mm</quantity>
wide at 10, 2.
<quantity id="D9239B250605AE75FC12FC943D88FCD4" box="[945,1013,826,851]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="283" unit="mm" value="5.0">5 mm</quantity>
wide at tubercula pubertatis;
<typeStatus id="C16088620605AE75FED9FCF43F98FCF4" box="[378,485,858,883]" pageId="4" pageNumber="283" type="paratype">paratypes</typeStatus>
<quantity id="D9239B250605AE75FE48FCF43C1EFCF4" box="[491,611,858,883]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="8.0" metricValueMin="5.5" pageId="4" pageNumber="283" unit="mm" value="67.5" valueMax="80.0" valueMin="55.0">5580 mm</quantity>
long,
<quantity id="D9239B250605AE75FD04FCF43C9AFCF4" box="[679,743,858,883]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="283" unit="mm" value="2.0">2 mm</quantity>
wide at 10,
<quantity id="D9239B250605AE75FCCFFCF43DB5FCF4" box="[876,968,858,883]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="283" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
at tubercula pubertatis.
<emphasis id="2CAFEAD20605AE75FEA9FCD43FBEFC15" box="[266,451,890,914]" italics="true" pageId="4" pageNumber="283">Segment number</emphasis>
:
<typeStatus id="C16088620605AE75FE72FCD43C4EFC14" box="[465,563,890,915]" pageId="4" pageNumber="283" type="holotype">holotype</typeStatus>
197+ (complete number is uncertain, due to poor preservation);
<typeStatus id="C16088620605AE75FE90FC343FEEFC34" box="[307,403,922,947]" pageId="4" pageNumber="283" type="paratype">paratype</typeStatus>
163.
<emphasis id="2CAFEAD20605AE75FE77FC343C21FC35" box="[468,604,922,946]" italics="true" pageId="4" pageNumber="283">Prostomium</emphasis>
: prolobous, small.
<emphasis id="2CAFEAD20605AE75FC95FC343DB3FC35" box="[822,974,922,946]" italics="true" pageId="4" pageNumber="283">Segmentation</emphasis>
: secondary annulation present on preclitellar segments; 1 and 2 short, simple, with obvious intersegmental furrow 1/2, both segments with irregular longitudinal grooves; 3 simple; 48 with two ringlets, first longer than second; 9 with two ringlets, second short; 10 and the following simple, postclitellarly randomly demarcated.
<emphasis id="2CAFEAD20605AE75FCFAFBB43DEBFBB5" box="[857,918,1050,1074]" italics="true" pageId="4" pageNumber="283">Setae</emphasis>
: minute, closely paired.
<emphasis id="2CAFEAD20605AE75FF40FB943FDDFBD5" box="[227,416,1082,1106]" italics="true" pageId="4" pageNumber="283">Nephridial pores</emphasis>
: not observed.
<emphasis id="2CAFEAD20605AE75FDEFFB943C9EFBD5" box="[588,739,1082,1106]" italics="true" pageId="4" pageNumber="283">Female pores</emphasis>
: obvious on
<typeStatus id="C16088620605AE75FCD7FB943D9FFBD4" box="[884,994,1082,1107]" pageId="4" pageNumber="283" type="paratype">paratypes</typeStatus>
, paired in 14 below
<emphasis id="2CAFEAD20605AE75FF5AFBF43F78FBF5" box="[249,261,1114,1138]" italics="true" pageId="4" pageNumber="283">c</emphasis>
setae.
<emphasis id="2CAFEAD20605AE75FEEEFBF43FBAFBF5" box="[333,455,1114,1138]" italics="true" pageId="4" pageNumber="283">Male pores</emphasis>
: externally not observed; possibly in intersegmental furrow 18/19, as
<taxonomicName id="D9DB4D430605AE75FEA2FBD43FD0FB14" box="[257,429,1146,1171]" class="Aves" genus="Vasa" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="303" phylum="Chordata" rank="species" species="deferentia">vasa deferentia</taxonomicName>
enter body wall in 18.
<emphasis id="2CAFEAD20605AE75FD63FBD43DDDFB15" box="[704,928,1146,1170]" italics="true" pageId="4" pageNumber="283">Spermathecal pores</emphasis>
: externally not observed.
</paragraph>
<paragraph id="1E6436C00605AE75FF09FB143C71FA14" blockId="4.[144,1104,578,1667]" pageId="4" pageNumber="283">
Clitellar region (
<figureCitation id="86E02A450605AE75FED2FB143FA2FB54" box="[369,479,1210,1235]" captionStart="Figs 23" captionStartId="4.[144,183,459,479]" captionTargetBox="[146,1105,215,436]" captionTargetId="figure-386@4.[144,1105,281,361]" captionTargetPageId="4" captionText="Figs 23. Microchaetus distasmosus sp. n. 2. Holotype, anterior part of body, ventral view with clitellum and tubercula pubertatis. 3. Paratype, clitellar region, enlarged. [C = clitellum, T = tuberculum pubertatis]." figureDoi="http://doi.org/10.5281/zenodo.7666264" httpUri="https://zenodo.org/record/7666264/files/figure.png" pageId="4" pageNumber="283">Figs 23</figureCitation>
):
<emphasis id="2CAFEAD20605AE75FE54FB143C1FFB55" box="[503,610,1210,1234]" italics="true" pageId="4" pageNumber="283">Clitellum</emphasis>
: saddle­shaped, white, elongated, clearly segmented, on 1/n11,1228,29; anterior and posterior borders not clearly marked; anteriorly ventral borders extend to line of papillae located on
<emphasis id="2CAFEAD20605AE75FCD1FB543DF3FA95" box="[882,910,1274,1298]" italics="true" pageId="4" pageNumber="283">ab</emphasis>
setae; on 1827 bordering tubercula pubertatis.
<emphasis id="2CAFEAD20605AE75FE50FAB43CA0FAB5" box="[499,733,1306,1330]" italics="true" pageId="4" pageNumber="283">Tubercula pubertatis</emphasis>
: on
<typeStatus id="C16088620605AE75FCB2FAB43D0EFAB4" box="[785,883,1306,1331]" pageId="4" pageNumber="283" type="holotype">holotype</typeStatus>
and
<typeStatus id="C16088620605AE75FC0FFAB43A6AFAB4" box="[940,1047,1306,1331]" pageId="4" pageNumber="283" type="paratype">paratypes</typeStatus>
well developed glandular tubercles, segmented, on 18,1927,28; ventrally extending to
<emphasis id="2CAFEAD20605AE75FB90FA943A32FAD5" box="[1075,1103,1338,1362]" italics="true" pageId="4" pageNumber="283">ab</emphasis>
setae.
<emphasis id="2CAFEAD20605AE75FF70FAF43F4CFAF5" box="[211,305,1370,1394]" italics="true" pageId="4" pageNumber="283">Papillae</emphasis>
: different sizes and shapes, glandular, encircling
<emphasis id="2CAFEAD20605AE75FCE8FAF43D1AFAF5" box="[843,871,1370,1394]" italics="true" pageId="4" pageNumber="283">ab</emphasis>
setae on first ringlets of 57 and also on 1118 (
<figureCitation id="86E02A450605AE75FE19FAD43F83FA14" box="[442,510,1402,1427]" captionStart="Figs 23" captionStartId="4.[144,183,459,479]" captionTargetBox="[146,1105,215,436]" captionTargetId="figure-386@4.[144,1105,281,361]" captionTargetPageId="4" captionText="Figs 23. Microchaetus distasmosus sp. n. 2. Holotype, anterior part of body, ventral view with clitellum and tubercula pubertatis. 3. Paratype, clitellar region, enlarged. [C = clitellum, T = tuberculum pubertatis]." figureDoi="http://doi.org/10.5281/zenodo.7666264" httpUri="https://zenodo.org/record/7666264/files/figure.png" pageId="4" pageNumber="283">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="1E6436C00605AE75FF33FA0C3F15FA3C" blockId="4.[144,1104,578,1667]" box="[144,360,1442,1467]" pageId="4" pageNumber="283">Internal characters:</paragraph>
<paragraph id="1E6436C00605AE74FF33FA643DA1FE60" blockId="4.[144,1104,578,1667]" lastBlockId="5.[181,1143,206,1247]" lastPageId="5" lastPageNumber="284" pageId="4" pageNumber="283">
<emphasis id="2CAFEAD20605AE75FF33FA643EB2FA65" box="[144,207,1482,1506]" italics="true" pageId="4" pageNumber="283">Septa</emphasis>
: 4/5 thin, 5/6 and 6/7 most thickened; 7/8 and 8/9 little thickened.
<emphasis id="2CAFEAD20605AE75FC4CFA643A3AFA65" box="[1007,1095,1482,1506]" italics="true" pageId="4" pageNumber="283">Gizzard</emphasis>
: cylindrical, large, muscular, in 7.
<emphasis id="2CAFEAD20605AE75FDB2FA443C9AF985" box="[529,743,1514,1538]" italics="true" pageId="4" pageNumber="283">Calciferous glands</emphasis>
: in 10 large, separated dorsally and ventrally.
<emphasis id="2CAFEAD20605AE75FE8FF9A43FF1F9A5" box="[300,396,1546,1570]" italics="true" pageId="4" pageNumber="283">Intestine</emphasis>
: commences in 13.
<emphasis id="2CAFEAD20605AE75FDCBF9A43C9CF9A5" box="[616,737,1546,1570]" italics="true" pageId="4" pageNumber="283">Typhlosole</emphasis>
: in
<typeStatus id="C16088620605AE75FCA9F9A43D11F9A4" box="[778,876,1546,1571]" pageId="4" pageNumber="283" type="holotype">holotype</typeStatus>
commences in 20 as thin tube, slightly enlarging backwards, terminating in segment 75.
<emphasis id="2CAFEAD20605AE75FCCCF9843A35F9C5" box="[879,1096,1578,1602]" italics="true" pageId="4" pageNumber="283">Dorsal blood vessel</emphasis>
: 48 double, separated; 9 double, close, cordiform.
<emphasis id="2CAFEAD20605AE75FD7AF9E43A68F9E5" box="[729,1045,1610,1634]" italics="true" pageId="4" pageNumber="283">Paired dorsoventral vessels</emphasis>
: 48 enlarged vessels; 911 moniliform.
<taxonomicName id="D9DB4D430605AE75FD83F9C43D37F904" authority=": meganephridia" authorityName="Meganephridia" box="[544,842,1642,1667]" class="Insecta" family="Crabronidae" genus="Nephridia" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="2CAFEAD20605AE75FD83F9C43CEFF905" box="[544,658,1642,1666]" italics="true" pageId="4" pageNumber="283">Nephridia</emphasis>
: meganephridia
</taxonomicName>
; little coiled loops with thin, extended caeca.
<emphasis id="2CAFEAD20604AE74FE12FF603CDFFF61" box="[433,674,206,230]" italics="true" pageId="5" pageNumber="284">Spermiductal funnels</emphasis>
: holandric arrangement; two pairs, each closely connected posteriorly with anterior parts of seminal vesicles, in 10 and 11 respectively.
<taxonomicName id="D9DB4D430604AE74FEF1FEA03C7AFEA1" box="[338,519,270,294]" class="Aves" genus="Vasa" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="284" phylum="Chordata" rank="species" species="deferentia">
<emphasis id="2CAFEAD20604AE74FEF1FEA03C7AFEA1" box="[338,519,270,294]" italics="true" pageId="5" pageNumber="284">Vasa deferentia</emphasis>
</taxonomicName>
: difficult to observe due to softened body wall and decomposed tissues; probably enter body wall at posterior part of segment 18.
<emphasis id="2CAFEAD20604AE74FBB8FE803F71FEE1" italics="true" pageId="5" pageNumber="284">Seminal vesicles</emphasis>
: two pairs of large sacs commencing at septa 10/11 and 11/12 respectively.
<emphasis id="2CAFEAD20604AE74FF16FEC03F20FE01" box="[181,349,366,390]" italics="true" pageId="5" pageNumber="284">Spermathecae</emphasis>
: small, elongated thecae in 8 and 9; 12 at each side, close to intersegmental furrow 8/9 and 9/10 respectively; ectal parts probably in 8/9 and 9/10, although anterior not confirmed.
<emphasis id="2CAFEAD20604AE74FDBAFE003C0DFE41" box="[537,624,430,454]" italics="true" pageId="5" pageNumber="284">Ovaries</emphasis>
: not observed.
<emphasis id="2CAFEAD20604AE74FCB0FE003DCEFE41" box="[787,947,430,454]" italics="true" pageId="5" pageNumber="284">Genital glands</emphasis>
: glandular tissues in 15, and 1827 associated with internal glands of tubercula pubertatis.
</paragraph>
<paragraph id="1E6436C00604AE74FF16FE583F5CFD68" blockId="5.[181,1143,206,1247]" pageId="5" pageNumber="284">
Biological notes: Mapelane Nature Reserve comprises coastal dune forests extending southwards from the Mfolozi riverbank with a variety of woodland patches and grasslands.
<typeStatus id="C16088620604AE74FE99FD983FDFFDC8" box="[314,418,566,591]" pageId="5" pageNumber="284" type="holotype">Holotype</typeStatus>
and some
<typeStatus id="C16088620604AE74FDBCFD983CF7FDC8" box="[543,650,566,591]" pageId="5" pageNumber="284" type="paratype">paratypes</typeStatus>
were found at the edge of large patches of grassland, about
<quantity id="D9239B250604AE74FED3FDF83FCBFDE8" box="[368,438,598,623]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="284" unit="km" value="10.0">10 km</quantity>
south from the river mouth. The other
<typeStatus id="C16088620604AE74FCC0FDF83DB3FDE8" box="[867,974,598,623]" pageId="5" pageNumber="284" type="paratype">paratypes</typeStatus>
were collected north­west of the
<typeStatus id="C16088620604AE74FED5FDD83FA5FD08" box="[374,472,630,655]" pageId="5" pageNumber="284" type="holotype">holotype</typeStatus>
site on the bank of the White Mfolozi River, covered by high indigenous grasses, and were associated with
<taxonomicName id="D9DB4D430604AE74FD14FD383D54FD29" baseAuthorityName="Kinberg" baseAuthorityYear="1867" box="[695,809,662,686]" class="Clitellata" family="Microchaetidae" genus="Microchaetus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="natalensis">
<emphasis id="2CAFEAD20604AE74FD14FD383D54FD29" box="[695,809,662,686]" italics="true" pageId="5" pageNumber="284">natalensis</emphasis>
</taxonomicName>
. Both localities are similar in vegetation and soil. Transportation of cocoons or individuals by water or in mud can be expected.
</paragraph>
<paragraph id="1E6436C00604AE74FF16FD503DF3FC90" blockId="5.[181,1143,206,1247]" box="[181,910,766,791]" pageId="5" pageNumber="284">
Distribution: Known only from the Mfolozi River banks (
<figureCitation id="86E02A450604AE74FC98FD503D00FC90" box="[827,893,766,791]" captionStart="Fig" captionStartId="3.[182,215,1291,1311]" captionTargetBox="[181,1141,203,1270]" captionTargetId="figure-141@3.[181,1141,203,1270]" captionTargetPageId="3" captionText="Fig. 1. The distribution of species of Microchaetus and Michalakus in KZN. Microchaetus: 1 = ambitus; 2 = caementerii; 3 = distasmosus; 4 = griphus; 5 = herberti; 6 = idechoritus; 7 = mkuzi; 8 = natalensis; 9 = papillatus; 10 = parvus; 11 = pondoanus; 12 = quaerus; 13 = tuberosus; 14 = vernoni; 15 = zaloumisi; 16 = Michalakus initus." figureDoi="http://doi.org/10.5281/zenodo.7666262" httpUri="https://zenodo.org/record/7666262/files/figure.png" pageId="5" pageNumber="284">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="1E6436C00604AE74FF16FC883D47FBF8" blockId="5.[181,1143,206,1247]" pageId="5" pageNumber="284">
Discussion: A distinctive species although similar and probably related to
<taxonomicName id="D9DB4D430604AE74FBB8FC883A0BFCB9" authorityName="Plisko" authorityYear="2003" box="[1051,1142,806,830]" class="Clitellata" family="Microchaetidae" genus="Microchaetus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="quaerus">
<emphasis id="2CAFEAD20604AE74FBB8FC883A0BFCB9" box="[1051,1142,806,830]" italics="true" pageId="5" pageNumber="284">quaerus</emphasis>
</taxonomicName>
described below. Both species have spermathecae in segments 8 and 9. The location of spermiductal pores in intersegmental furrow 8/9, however, was not confirmed in
<taxonomicName id="D9DB4D430604AE74FF16FC283F41FC19" authorityName="Plisko" authorityYear="2003" box="[181,316,902,926]" class="Clitellata" family="Microchaetidae" genus="Microchaetus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="distasmosus">
<emphasis id="2CAFEAD20604AE74FF16FC283F41FC19" box="[181,316,902,926]" italics="true" pageId="5" pageNumber="284">distasmosus</emphasis>
</taxonomicName>
.
<taxonomicName id="D9DB4D430604AE74FEE9FC283F88FC19" authorityName="Plisko" authorityYear="2003" box="[330,501,902,926]" class="Clitellata" family="Microchaetidae" genus="Microchaetus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="distasmosus">
<emphasis id="2CAFEAD20604AE74FEE9FC283F88FC19" box="[330,501,902,926]" italics="true" pageId="5" pageNumber="284">M. distasmosus</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="D9DB4D430604AE74FD1AFC283D6EFC19" authorityName="Plisko" authorityYear="2003" box="[697,787,902,926]" class="Clitellata" family="Microchaetidae" genus="Microchaetus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="quaerus">
<emphasis id="2CAFEAD20604AE74FD1AFC283D6EFC19" box="[697,787,902,926]" italics="true" pageId="5" pageNumber="284">quaerus</emphasis>
</taxonomicName>
in the length and position of the clitellum, shape of tubercula pubertatis, and thickness of preclitellar septa. In the other species of
<taxonomicName id="D9DB4D430604AE74FE8AFC683FBFFC59" authorityName="Rapp" authorityYear="1849" box="[297,450,966,990]" class="Clitellata" family="Microchaetidae" genus="Microchaetus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="genus">
<emphasis id="2CAFEAD20604AE74FE8AFC683FBFFC59" box="[297,450,966,990]" italics="true" pageId="5" pageNumber="284">Microchaetus</emphasis>
</taxonomicName>
the most anterior position of spermathecal pores was observed in
<taxonomicName id="D9DB4D430604AE74FF77FC483F37FC79" authorityName="Michaelsen" authorityYear="1908" box="[212,330,998,1022]" class="Clitellata" family="Microchaetidae" genus="Microchaetus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="namaensis">
<emphasis id="2CAFEAD20604AE74FF77FC483F37FC79" box="[212,330,998,1022]" italics="true" pageId="5" pageNumber="284">namaensis</emphasis>
</taxonomicName>
(known only from the
<collectingRegion id="DC1FF8220604AE74FDFBFC483D7CFC78" box="[600,769,998,1023]" country="South Africa" name="Northern Cape" pageId="5" pageNumber="284">Northern Cape</collectingRegion>
) which has the anterior pores in 9/10 and the posterior in intersegmental furrows 10/11 11/12 respectively. In the other two holandric genera
<taxonomicName id="D9DB4D430604AE74FE1DFB883C4AFBB9" authorityName="Kinberg" authorityYear="1867" box="[446,567,1062,1086]" class="Clitellata" family="Tritogeniidae" genus="Tritogenia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="genus">
<emphasis id="2CAFEAD20604AE74FE1DFB883C4AFBB9" box="[446,567,1062,1086]" italics="true" pageId="5" pageNumber="284">Tritogenia</emphasis>
</taxonomicName>
and
<taxonomicName id="D9DB4D430604AE74FDD4FB883C83FBB9" authorityName="Plisko" authorityYear="1996" box="[631,766,1062,1086]" class="Clitellata" family="Tritogeniidae" genus="Michalakus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="genus">
<emphasis id="2CAFEAD20604AE74FDD4FB883C83FBB9" box="[631,766,1062,1086]" italics="true" pageId="5" pageNumber="284">Michalakus</emphasis>
</taxonomicName>
, the most anterior positions of spermathecal pores are in intersegmental furrows 10/11 11/
<quantity id="D9239B250604AE74FCE9FBE83DFEFBD8" box="[842,899,1094,1119]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="5" pageNumber="284" unit="in" value="12.0">12 in</quantity>
<taxonomicName id="D9DB4D430604AE74FC29FBE83A3BFBD9" authorityName="Plisko &amp; Zicsi" authorityYear="1991" box="[906,1094,1094,1118]" class="Clitellata" family="Tritogeniidae" genus="Tritogenia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="shawi">
<emphasis id="2CAFEAD20604AE74FC29FBE83A3BFBD9" box="[906,1094,1094,1118]" italics="true" pageId="5" pageNumber="284">Tritogenia shawi</emphasis>
</taxonomicName>
and
<taxonomicName id="D9DB4D430604AE74FF16FBC83F5AFBF9" authorityName="Plisko" authorityYear="1997" box="[181,295,1126,1151]" class="Clitellata" family="Tritogeniidae" genus="Tritogenia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="soleata">
<emphasis id="2CAFEAD20604AE74FF16FBC83EB8FBF9" box="[181,197,1126,1150]" italics="true" pageId="5" pageNumber="284">T</emphasis>
.
<emphasis id="2CAFEAD20604AE74FF70FBC83F5AFBF9" box="[211,295,1126,1150]" italics="true" pageId="5" pageNumber="284">soleata</emphasis>
</taxonomicName>
<emphasis id="2CAFEAD20604AE74FE85FBC83F57FBF9" box="[294,298,1126,1150]" italics="true" pageId="5" pageNumber="284">,</emphasis>
and in 10/11 11/12 12/
<quantity id="D9239B250604AE74FD8EFBC83C18FBF8" box="[557,613,1126,1151]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="5" pageNumber="284" unit="in" value="13.0">13 in</quantity>
<taxonomicName id="D9DB4D430604AE74FDC8FBC83D4EFBF9" authorityName="Plisko" authorityYear="1996" box="[619,819,1126,1150]" class="Clitellata" family="Tritogeniidae" genus="Michalakus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="initus">
<emphasis id="2CAFEAD20604AE74FDC8FBC83D4EFBF9" box="[619,819,1126,1150]" italics="true" pageId="5" pageNumber="284">Michalakus initus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1E6436C00604AE74FF73FB283A19FB58" blockId="5.[181,1143,206,1247]" pageId="5" pageNumber="284">
It is noteworthy that in some members of the genus
<taxonomicName id="D9DB4D430604AE74FC97FB283DC2FB19" authorityName="Plisko" authorityYear="1992" box="[820,959,1158,1182]" class="Clitellata" family="Microchaetidae" genus="Proandricus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="genus">
<emphasis id="2CAFEAD20604AE74FC97FB283DC2FB19" box="[820,959,1158,1182]" italics="true" pageId="5" pageNumber="284">Proandricus</emphasis>
</taxonomicName>
assigned to the
<taxonomicName id="D9DB4D430604AE74FF16FB083F43FB39" baseAuthorityName="Reinecke &amp; Ryke" baseAuthorityYear="1969" box="[181,318,1190,1214]" class="Clitellata" family="Microchaetidae" genus="Proandricus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="lesothoensis">
<emphasis id="2CAFEAD20604AE74FF16FB083F43FB39" box="[181,318,1190,1214]" italics="true" pageId="5" pageNumber="284">lesothoensis</emphasis>
</taxonomicName>
­ species group, the utmost anterior spermathecal pores occur in 9/10, or between ringlets in segment 9 (
<taxonomicName id="D9DB4D430604AE74FDB0FB683D7EFB59" authority="Plisko, 2003" authorityName="Plisko" authorityYear="2003" box="[531,771,1222,1246]" class="Clitellata" family="Microchaetidae" genus="Proandricus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="amphius" status="sp. nov.">
<emphasis id="2CAFEAD20604AE74FDB0FB683D7EFB59" box="[531,771,1222,1246]" italics="true" pageId="5" pageNumber="284">Proandricus amphius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="379C57A90604AE74FCA9FB683D3BFB58" box="[778,838,1222,1247]" pageId="5" pageNumber="284" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName id="D9DB4D430604AE74FCDFFB683A6FFB59" authority="Plisko, 2003" authorityName="Plisko" authorityYear="2003" box="[892,1042,1222,1247]" class="Clitellata" family="Microchaetidae" genus="Proandricus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="284" phylum="Annelida" rank="species" species="oresbiosus" status="sp. nov.">
<emphasis id="2CAFEAD20604AE74FCDFFB683DF0FB59" box="[892,909,1222,1246]" italics="true" pageId="5" pageNumber="284">P</emphasis>
.
<emphasis id="2CAFEAD20604AE74FC38FB683A6FFB59" box="[923,1042,1222,1246]" italics="true" pageId="5" pageNumber="284">oresbiosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="379C57A90604AE74FBBBFB683A28FB58" box="[1048,1109,1222,1247]" pageId="5" pageNumber="284" rank="species">sp. n.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
</treatment>
</document>