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<document id="63831B9AEE2FEC2745DE073DAEE1C039" ID-DOI="10.5281/zenodo.173822" ID-GBIF-Dataset="21e98840-1260-4cd7-b4e9-aa758c313563" ID-ISSN="1175­5326" ID-Zenodo-Dep="173822" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459785603673" checkinUser="plazi" docAuthor="Barrio-Amorós, César L." docDate="2006" docId="9671FC29FFCE4A39FEFFFE83B4F5E545" docLanguage="en" docName="zt01309p068.pdf" docOrigin="Zootaxa 1309" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Phyllomedusa tarsius" docType="treatment" docVersion="9" lastPageNumber="64" masterDocId="6A488451FFC74A33FFF7FFE5B652E41B" masterDocTitle="A new species of Phyllomedusa (Anura: Hylidae: Phyllomedusinae) from northwestern Venezuela" masterLastPageNumber="68" masterPageNumber="55" pageNumber="64" updateTime="1698218439328" updateUser="plazi">
<mods:mods id="CA5DCF8ED7A051E12AED7F65340F40BA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B2E4E648FEA6C8235D05B60C64C38549">
<mods:title id="D884B59D41F11DDE6CE676118F518BB9">A new species of Phyllomedusa (Anura: Hylidae: Phyllomedusinae) from northwestern Venezuela</mods:title>
</mods:titleInfo>
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<mods:namePart id="4F35D24B0BE565F5B00AEF9AF3602E2F">Barrio-Amorós, César L.</mods:namePart>
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<mods:date id="A2CA929E270A46A0D9412EF812E5C148">2006</mods:date>
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<mods:number id="FE8673EA19B475741C0D88C241669890">1309</mods:number>
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<mods:classification id="79912AC9F57CA79CB8E1790165222315">journal article</mods:classification>
<mods:identifier id="419D6AD948F6C7EB76AC649E566FA944" type="DOI">10.5281/zenodo.173822</mods:identifier>
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<mods:identifier id="4FC5964889E585DA79224A7F50F80812" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="BED4D2203D8A20B02048A9FE3544E404" type="Zenodo-Dep">173822</mods:identifier>
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<treatment id="9671FC29FFCE4A39FEFFFE83B4F5E545" ID-DOI="http://doi.org/10.5281/zenodo.5665360" ID-GBIF-Taxon="124860098" ID-Zenodo-Dep="5665360" LSID="urn:lsid:plazi:treatment:9671FC29FFCE4A39FEFFFE83B4F5E545" httpUri="http://treatment.plazi.org/id/9671FC29FFCE4A39FEFFFE83B4F5E545" lastPageId="10" lastPageNumber="64" pageId="9" pageNumber="64">
<subSubSection id="56C21EB4FFCE4A3AFEFFFE83B272E728" pageId="9" pageNumber="64" type="nomenclature">
<paragraph id="1E674D3FFFCE4A3AFEFFFE83B40FE59B" blockId="9.[264,605,358,384]" box="[264,605,358,384]" pageId="9" pageNumber="64">
<heading id="452FFA53FFCE4A3AFEFFFE83B40FE59B" bold="true" box="[264,605,358,384]" fontSize="11" level="1" pageId="9" pageNumber="64" reason="1">
<emphasis id="2CAC912DFFCE4A3AFEFFFE83B40FE59B" bold="true" box="[264,605,358,384]" pageId="9" pageNumber="64">
<taxonomicName id="D9D836BCFFCE4A3AFEFFFE83B459E59B" box="[264,523,358,384]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tarsius">
<emphasis id="2CAC912DFFCE4A3AFEFFFE83B459E59B" bold="true" box="[264,523,358,384]" italics="true" pageId="9" pageNumber="64">Phyllomedusa tarsius</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
<paragraph id="1E674D3FFFCE4A3AFEFFFE54B272E728" blockId="9.[264,1323,433,819]" pageId="9" pageNumber="64">
<collectingCountry id="66CF0DAFFFCE4A3AFEFFFE54B778E5D0" box="[264,298,433,459]" name="South Georgia and the South Sandwich Islands" pageId="9" pageNumber="64">De</collectingCountry>
la
<bibRefCitation id="7A4930CEFFCE4A3AFEA7FE54B78FE5D0" author="Riva" box="[336,477,433,459]" pageId="9" pageNumber="66" refString="De la Riva, I. (1999) A new Phyllomedusa from southwestern Amazonia (Amphibia: Anura: Hylidae). Revista Espanola de Herpetologia, 13, 123 - 131." type="journal article" year="1999">Riva (1999)</bibRefCitation>
comments on the poorly defined
<taxonomicName id="D9D836BCFFCE4A3AFC9AFE54B5EFE5D1" box="[877,957,433,458]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tarsius">
<emphasis id="2CAC912DFFCE4A3AFC9AFE54B5EFE5D1" box="[877,957,433,458]" italics="true" pageId="9" pageNumber="64">tarsius</emphasis>
</taxonomicName>
group, which to him consisted in the following species:
<taxonomicName id="D9D836BCFFCE4A3AFDB6FE3CB530E5E9" box="[577,866,473,498]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="boliviana">
<emphasis id="2CAC912DFFCE4A3AFDB6FE3CB530E5E9" box="[577,866,473,498]" italics="true" pageId="9" pageNumber="64">Phyllomedusa boliviana</emphasis>
</taxonomicName>
,
<emphasis id="2CAC912DFFCE4A3AFC82FE3CB73FE601" italics="true" pageId="9" pageNumber="64">
<taxonomicName id="D9D836BCFFCE4A3AFC82FE3CB5B5E5E9" box="[885,999,473,498]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="camba">P. camba</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFC00FE3CB2D3E5E9" box="[1015,1153,473,498]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="coelestis">P. coelestis</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFB66FE3CB357E5E9" box="[1169,1285,473,498]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tarsius">P. tarsius</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFAE2FE3CB73FE601" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="trinitatis">P. trinitatis</taxonomicName>
</emphasis>
and
<taxonomicName id="D9D836BCFFCE4A3AFE52FDE4B449E601" box="[421,539,513,538]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="venusta">
<emphasis id="2CAC912DFFCE4A3AFE52FDE4B449E601" box="[421,539,513,538]" italics="true" pageId="9" pageNumber="64">P. venusta</emphasis>
</taxonomicName>
.
<bibRefCitation id="7A4930CEFFCE4A3AFDDFFDE4B560E600" box="[552,818,513,539]" pageId="9" pageNumber="66" refString="Faivovich, J., C. E. B. Haddad, P. Garcia, D. R. Frost, J. A. Campbell, and W. C. Wheeler. (2005) Systematic review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic revision. Bulletin of the American Museum of Natural History, 294, 1 - 240." type="journal article">
Faivovich
<emphasis id="2CAC912DFFCE4A3AFD53FDE4B48CE601" box="[676,734,513,538]" italics="true" pageId="9" pageNumber="64">et al.</emphasis>
(2005)
</bibRefCitation>
only tested (based on 5100 base pairs from four mithochondrial and five nuclear genes)
<emphasis id="2CAC912DFFCE4A3AFCCDFDCCB5FBE659" box="[826,937,553,578]" italics="true" pageId="9" pageNumber="64">P. t a r s i u s</emphasis>
of the group. Creating a new
<taxonomicName id="D9D836BCFFCE4A3AFEFFFDB4B736E670" box="[264,356,593,619]" class="Amphibia" family="Hylidae" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="family">Hylidae</taxonomicName>
taxonomy, they included
<taxonomicName id="D9D836BCFFCE4A3AFD42FDB4B56CE671" box="[693,830,593,618]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="sauvagii">
<emphasis id="2CAC912DFFCE4A3AFD42FDB4B56CE671" box="[693,830,593,618]" italics="true" pageId="9" pageNumber="64">P. sauvagii</emphasis>
</taxonomicName>
in the same group with
<taxonomicName id="D9D836BCFFCE4A3AFB7EFDB4B2ACE671" box="[1161,1278,593,618]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tarsius">
<emphasis id="2CAC912DFFCE4A3AFB7EFDB4B2ACE671" box="[1161,1278,593,618]" italics="true" pageId="9" pageNumber="64">P. tarsius</emphasis>
</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFAE2FDB4B725E689" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="boliviana">
<emphasis id="2CAC912DFFCE4A3AFAE2FDB4B725E689" italics="true" pageId="9" pageNumber="64">P. boliviana</emphasis>
</taxonomicName>
and
<emphasis id="2CAC912DFFCE4A3AFE42FD9CB44DE689" box="[437,543,633,658]" italics="true" pageId="9" pageNumber="64">P. c a m b a</emphasis>
. However,
<taxonomicName id="D9D836BCFFCE4A3AFD58FD9CB56CE689" box="[687,830,633,658]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="boliviana">
<emphasis id="2CAC912DFFCE4A3AFD58FD9CB56CE689" box="[687,830,633,658]" italics="true" pageId="9" pageNumber="64">P. boliviana</emphasis>
</taxonomicName>
and
<emphasis id="2CAC912DFFCE4A3AFC8BFD9CB5B4E689" box="[892,998,633,658]" italics="true" pageId="9" pageNumber="64">P. c a m b a</emphasis>
(which are definitely very similar to each other) differs considerably from
<taxonomicName id="D9D836BCFFCE4A3AFCB3FD44B595E6A1" box="[836,967,673,698]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="coelestis">
<emphasis id="2CAC912DFFCE4A3AFCB3FD44B595E6A1" box="[836,967,673,698]" italics="true" pageId="9" pageNumber="64">P. coelestis</emphasis>
</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFC2FFD44B217E6A1" box="[984,1093,673,698]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tarsius">
<emphasis id="2CAC912DFFCE4A3AFC2FFD44B217E6A1" box="[984,1093,673,698]" italics="true" pageId="9" pageNumber="64">P. tarsius</emphasis>
</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFBA1FD44B28BE6A1" box="[1110,1241,673,698]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="trinitatis">
<emphasis id="2CAC912DFFCE4A3AFBA1FD44B28BE6A1" box="[1110,1241,673,698]" italics="true" pageId="9" pageNumber="64">P. trinitatis</emphasis>
</taxonomicName>
and
<taxonomicName id="D9D836BCFFCE4A3AFAE2FD44B733E6F9" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="venusta">
<emphasis id="2CAC912DFFCE4A3AFAE2FD44B733E6F9" italics="true" pageId="9" pageNumber="64">P. venusta</emphasis>
</taxonomicName>
, especially in appearance of the eye (see below), and
<taxonomicName id="D9D836BCFFCE4A3AFC0AFD2CB2D3E6F9" box="[1021,1153,713,738]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="sauvagii">
<emphasis id="2CAC912DFFCE4A3AFC0AFD2CB2D3E6F9" box="[1021,1153,713,738]" italics="true" pageId="9" pageNumber="64">P. sauvagii</emphasis>
</taxonomicName>
is clearly not assignable to this group (it is one of the most distinctive
<taxonomicName id="D9D836BCFFCE4A3AFC4BFD14B230E711" box="[956,1122,753,778]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="2CAC912DFFCE4A3AFC4BFD14B230E711" box="[956,1122,753,778]" italics="true" pageId="9" pageNumber="64">Phyllomedusa</emphasis>
</taxonomicName>
, and probably it deserves its own species group,
<bibRefCitation id="7A4930CEFFCE4A3AFD77FCFCB511E728" author="Cannatella" box="[640,835,793,819]" pageId="9" pageNumber="66" refString="Cannatella, D. (1980) A review of the Phyllomedusa buckleyi group (Anura: Hylidae). Occasional. Papers of the Museum of Natural History of the University of Kansas, 87, 1 - 40." type="journal article" year="1980">Cannatella 1980</bibRefCitation>
;
<collectingCountry id="66CF0DAFFFCE4A3AFCB8FCFCB523E728" box="[847,881,793,819]" name="South Georgia and the South Sandwich Islands" pageId="9" pageNumber="64">De</collectingCountry>
la
<bibRefCitation id="7A4930CEFFCE4A3AFC61FCFCB246E728" author="Riva" box="[918,1044,793,819]" pageId="9" pageNumber="66" refString="De la Riva, I. (1999) A new Phyllomedusa from southwestern Amazonia (Amphibia: Anura: Hylidae). Revista Espanola de Herpetologia, 13, 123 - 131." type="journal article" year="1999">Riva 1999</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="4AA71DB7FFCE4A3AFEFFFB6EB276E0F3" httpUri="https://zenodo.org/record/173829/files/figure.png" pageId="9" pageNumber="64" targetBox="[264,1323,876,1139]" targetPageId="9">
<paragraph id="1E674D3FFFCE4A3AFEFFFB6EB276E0F3" blockId="9.[264,1323,1163,1256]" pageId="9" pageNumber="64">
<emphasis id="2CAC912DFFCE4A3AFEFFFB6EB7DCE0B8" bold="true" box="[264,398,1163,1187]" pageId="9" pageNumber="64">FIGURE 7.</emphasis>
Audiospectrogram of the recorded call of
<taxonomicName id="D9D836BCFFCE4A3AFC99FB69B21FE0B8" box="[878,1101,1164,1187]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="nieldi" status="sp. nov.">
<emphasis id="2CAC912DFFCE4A3AFC99FB69B21FE0B8" box="[878,1101,1164,1187]" italics="true" pageId="9" pageNumber="64">Phyllomedusa nieldi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="379F2C56FFCE4A3AFBA0FB6EB2E2E0B8" box="[1111,1200,1163,1187]" pageId="9" pageNumber="64" rank="species">
<emphasis id="2CAC912DFFCE4A3AFBA0FB6EB2E2E0B8" bold="true" box="[1111,1200,1163,1187]" pageId="9" pageNumber="64">sp. nov.</emphasis>
</taxonomicNameLabel>
A. Simple note. B. Series of notes. C. Double note. The horizontal bar corresponds to time in seconds, separated by decimals of seconds. The vertical axis is the frequency in Hz.
</paragraph>
</caption>
<subSubSection id="56C21EB4FFCE4A39FECFFAF9B4F5E545" lastPageId="10" lastPageNumber="65" pageId="9" pageNumber="64" type="description">
<paragraph id="1E674D3FFFCE4A3AFECFFAF9B5A8E30D" blockId="9.[264,1324,1308,1894]" pageId="9" pageNumber="64">
I propose the following definition of the
<taxonomicName id="D9D836BCFFCE4A3AFCCBFAF9B5FCE12E" box="[828,942,1308,1333]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tarsius">
<emphasis id="2CAC912DFFCE4A3AFCCBFAF9B5FCE12E" box="[828,942,1308,1333]" italics="true" pageId="9" pageNumber="64">P. tarsius</emphasis>
</taxonomicName>
group (osteological and larval characters not considered: the larvae of
<emphasis id="2CAC912DFFCE4A3AFD20FAA1B59FE146" box="[727,973,1348,1373]" italics="true" pageId="9" pageNumber="64">
<taxonomicName id="D9D836BCFFCE4A3AFD20FAA1B50EE146" box="[727,860,1348,1373]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="coelestis">P. coelestis</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFC91FAA1B59AE146" box="[870,968,1348,1373]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="neildi">P. neildi</taxonomicName>
,
</emphasis>
and
<emphasis id="2CAC912DFFCE4A3AFBF3FAA1B224E146" box="[1028,1142,1348,1373]" italics="true" pageId="9" pageNumber="64">P. v e n u s t a</emphasis>
are unknown): moderate to large phyllomedusines with sexual dimorphism in size (males from
<quantity id="D920E0DAFFCE4A3AFB41FA89B764E1B5" metricMagnitude="-2" metricUnit="m" metricValue="7.515" metricValueMax="9.7" metricValueMin="5.33" pageId="9" pageNumber="64" unit="mm" value="75.15" valueMax="97.0" valueMin="53.3">53.3 to 97 mm</quantity>
­smallest measurement corresponding to
<emphasis id="2CAC912DFFCE4A3AFCC7FA71B5EFE1B6" box="[816,957,1428,1453]" italics="true" pageId="9" pageNumber="64">
<taxonomicName id="D9D836BCFFCE4A3AFCC7FA71B5EAE1B6" box="[816,952,1428,1453]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="coelestis">P. coelestis</taxonomicName>
,
</emphasis>
the biggest to
<emphasis id="2CAC912DFFCE4A3AFB80FA71B2B6E1B6" box="[1143,1252,1428,1453]" italics="true" pageId="9" pageNumber="64">P. t a r s i u s</emphasis>
from Ecuador­; females from
<quantity id="D920E0DAFFCE4A3AFDD2FA59B48FE1CD" box="[549,733,1468,1494]" metricMagnitude="-2" metricUnit="m" metricValue="8.74" metricValueMax="11.18" metricValueMin="6.3" pageId="9" pageNumber="64" unit="mm" value="87.4" valueMax="111.8" valueMin="63.0">63 to 111.8 mm</quantity>
–smallest measurement corresponding to
<emphasis id="2CAC912DFFCE4A3AFB33FA59B34DE1CE" box="[1220,1311,1468,1493]" italics="true" pageId="9" pageNumber="64">P. n e i l d i</emphasis>
, the biggest to
<emphasis id="2CAC912DFFCE4A3AFE36FA01B463E1E6" box="[449,561,1508,1533]" italics="true" pageId="9" pageNumber="64">P. t a r s i u s</emphasis>
­): (1) Foot and hands without webbing; (2) parotoid glands indistinct (in
<emphasis id="2CAC912DFFCE4A3AFE45F9E9B4F3E23E" box="[434,673,1548,1573]" italics="true" pageId="9" pageNumber="64">
<taxonomicName id="D9D836BCFFCE4A3AFE45F9E9B448E23E" box="[434,538,1548,1573]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="neildi">P. neildi</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFDDAF9E9B4F3E23E" box="[557,673,1548,1573]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tarsius">P. tarsius</taxonomicName>
</emphasis>
from
<collectingCountry id="66CF0DAFFFCE4A3AFD02F9E9B53FE23D" box="[757,877,1548,1574]" name="Venezuela" pageId="9" pageNumber="64">Venezuela</collectingCountry>
and some
<taxonomicName id="D9D836BCFFCE4A3AFC08F9E9B2DAE23E" box="[1023,1160,1548,1573]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="trinitatis">
<emphasis id="2CAC912DFFCE4A3AFC08F9E9B2DAE23E" box="[1023,1160,1548,1573]" italics="true" pageId="9" pageNumber="64">P. trinitatis</emphasis>
</taxonomicName>
), moderately developed (
<emphasis id="2CAC912DFFCE4A3AFE66F9D1B4F0E256" box="[401,674,1588,1613]" italics="true" pageId="9" pageNumber="64">
<taxonomicName id="D9D836BCFFCE4A3AFE66F9D1B447E256" box="[401,533,1588,1613]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="coelestis">P. coelestis</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFDD6F9D1B4F0E256" box="[545,674,1588,1613]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="trinitatis">P. trinitatis</taxonomicName>
</emphasis>
) to well developed (some
<emphasis id="2CAC912DFFCE4A3AFC21F9D1B26DE256" box="[982,1087,1588,1613]" italics="true" pageId="9" pageNumber="64">P. t a r s i u s</emphasis>
and
<taxonomicName id="D9D836BCFFCE4A3AFB8DF9D1B2A2E256" box="[1146,1264,1588,1613]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="venusta">
<emphasis id="2CAC912DFFCE4A3AFB8DF9D1B2A2E256" box="[1146,1264,1588,1613]" italics="true" pageId="9" pageNumber="64">P. venusta</emphasis>
</taxonomicName>
); (3) dorsal skin smooth with fine and scattered small tubercles (
<emphasis id="2CAC912DFFCE4A3AFC36F9B9B2E0E26E" box="[961,1202,1628,1653]" italics="true" pageId="9" pageNumber="64">
<taxonomicName id="D9D836BCFFCE4A3AFC36F9B9B271E26E" box="[961,1059,1628,1653]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="neildi">P. neildi</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFBC7F9B9B2E0E26E" box="[1072,1202,1628,1653]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="trinitatis">P. trinitatis</taxonomicName>
</emphasis>
), pustules (
<taxonomicName id="D9D836BCFFCE4A3AFEE5F961B7DBE286" box="[274,393,1668,1693]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="venusta">
<emphasis id="2CAC912DFFCE4A3AFEE5F961B7DBE286" box="[274,393,1668,1693]" italics="true" pageId="9" pageNumber="64">P. venusta</emphasis>
</taxonomicName>
), shagreen (
<taxonomicName id="D9D836BCFFCE4A3AFDEAF961B4D9E286" box="[541,651,1668,1693]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tarsius">
<emphasis id="2CAC912DFFCE4A3AFDEAF961B4D9E286" box="[541,651,1668,1693]" italics="true" pageId="9" pageNumber="64">P. tarsius</emphasis>
</taxonomicName>
), or posteriorly tuberculate (
<taxonomicName id="D9D836BCFFCE4A3AFC15F961B237E286" box="[994,1125,1668,1693]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="coelestis">
<emphasis id="2CAC912DFFCE4A3AFC15F961B237E286" box="[994,1125,1668,1693]" italics="true" pageId="9" pageNumber="64">P. coelestis</emphasis>
</taxonomicName>
); (4) males with vocal slits (
<taxonomicName id="D9D836BCFFCE4A3AFE67F949B440E2DE" box="[400,530,1708,1733]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="coelestis">
<emphasis id="2CAC912DFFCE4A3AFE67F949B440E2DE" box="[400,530,1708,1733]" italics="true" pageId="9" pageNumber="64">P. coelestis</emphasis>
</taxonomicName>
) or without (
<emphasis id="2CAC912DFFCE4A3AFD58F949B24FE2DE" box="[687,1053,1708,1733]" italics="true" pageId="9" pageNumber="64">
P. n e i l d i,
<taxonomicName id="D9D836BCFFCE4A3AFCE9F949B5DDE2DE" box="[798,911,1708,1733]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tarsius">P. tarsius</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFC6CF949B24FE2DE" box="[923,1053,1708,1733]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="trinitatis">P. trinitatis</taxonomicName>
</emphasis>
); (5) Toe I longer than II; (6) iris golden, orange or reddish, with black fine reticulations (
<emphasis id="2CAC912DFFCE4A3AFBE1F931B7BAE30E" italics="true" pageId="9" pageNumber="64">
<taxonomicName id="D9D836BCFFCE4A3AFBE1F931B2C8E2F6" box="[1046,1178,1748,1773]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="coelestis">P. coelestis</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFB50F931B35BE2F6" box="[1191,1289,1748,1773]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="neildi">P. neildi</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFAE2F931B709E30E" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tarsius">P. tarsius</taxonomicName>
,
<taxonomicName id="D9D836BCFFCE4A3AFE91F919B7BAE30E" box="[358,488,1788,1813]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="trinitatis">P. trinitatis</taxonomicName>
</emphasis>
,
<taxonomicName id="D9D836BCFFCE4A3AFE00F919B43FE30E" box="[503,621,1788,1813]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="venusta">
<emphasis id="2CAC912DFFCE4A3AFE00F919B43FE30E" box="[503,621,1788,1813]" italics="true" pageId="9" pageNumber="64">P. venusta</emphasis>
</taxonomicName>
); (7) vomerine processes present.
</paragraph>
<paragraph id="1E674D3FFFCE4A39FECFF8C1B4F5E545" blockId="9.[264,1324,1308,1894]" lastBlockId="10.[264,1323,284,350]" lastPageId="10" lastPageNumber="65" pageId="9" pageNumber="64">
I propose to exclude
<taxonomicName id="D9D836BCFFCE4A3AFDC3F8C1B493E326" box="[564,705,1828,1853]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="boliviana">
<emphasis id="2CAC912DFFCE4A3AFDC3F8C1B493E326" box="[564,705,1828,1853]" italics="true" pageId="9" pageNumber="64">P. boliviana</emphasis>
</taxonomicName>
and
<emphasis id="2CAC912DFFCE4A3AFD0AF8C1B534E326" box="[765,870,1828,1853]" italics="true" pageId="9" pageNumber="64">P. c a m b a</emphasis>
(included in the group by
<collectingCountry id="66CF0DAFFFCE4A3AFB5DF8C1B29EE325" box="[1194,1228,1828,1854]" name="South Georgia and the South Sandwich Islands" pageId="9" pageNumber="64">De</collectingCountry>
la
<bibRefCitation id="7A4930CEFFCE4A3AFB04F8C1B716E37D" author="Riva" pageId="9" pageNumber="66" refString="De la Riva, I. (1999) A new Phyllomedusa from southwestern Amazonia (Amphibia: Anura: Hylidae). Revista Espanola de Herpetologia, 13, 123 - 131." type="journal article" year="1999">Riva 1999</bibRefCitation>
) and
<taxonomicName id="D9D836BCFFCE4A3AFE72F8A9B457E37E" box="[389,517,1868,1893]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="sauvagii">
<emphasis id="2CAC912DFFCE4A3AFE72F8A9B457E37E" box="[389,517,1868,1893]" italics="true" pageId="9" pageNumber="64">P. sauvagii</emphasis>
</taxonomicName>
(included in the group by
<bibRefCitation id="7A4930CEFFCE4A3AFCCDF8A9B266E37D" author="Faivovich" box="[826,1076,1868,1894]" pageId="9" pageNumber="66" refString="Faivovich, J., C. E. B. Haddad, P. Garcia, D. R. Frost, J. A. Campbell, and W. C. Wheeler. (2005) Systematic review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic revision. Bulletin of the American Museum of Natural History, 294, 1 - 240." type="journal article" year="2005">
Faivovich
<emphasis id="2CAC912DFFCE4A3AFC41F8A9B5A3E37E" box="[950,1009,1868,1893]" italics="true" pageId="9" pageNumber="64">et al.</emphasis>
2005
</bibRefCitation>
) because they do not share the most striking character (herein considered as a synapomorphy) of the group, the golden iris with black reticulations.
</paragraph>
</subSubSection>
</treatment>
</document>
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