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<document id="92D70B92AB98508E6644E7BBD1A8C98C" ID-CLB-Dataset="7497" ID-DOI="10.11646/zootaxa.4949.1.5" ID-GBIF-Dataset="137cdaec-de44-48ae-be6b-fc9982803086" ID-ISSN="1175-5326" ID-Zenodo-Dep="4635941" ID-ZooBank="2C821F67-4B27-48D1-B5F4-2DD2A35BA3FA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616662070952" checkinUser="plazi" docAuthor="Park, Kyu-Tek &amp; Bae, Yang-Seop" docDate="2021" docId="9610DF5C7B51FFDA4E99FF6FFCA22258" docLanguage="en" docName="zootaxa.4949.1.5.pdf" docOrigin="Zootaxa 4949 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Torodora spathiana Park 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="107" masterDocId="6A29A7247B55FFDF4E0EFFF8FFD02161" masterDocTitle="Five new species of the genus Torodora Meyrick (Lepidoptera, Lecithoceridae Torodorinae) from Cambodia, with a tentative check-list of the genus" masterLastPageNumber="114" masterPageNumber="102" pageNumber="106" updateTime="1698934944201" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="8EEF929E8F6BB4C703882660D540B3DD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="56B116B8F9607B1BAAEE8668036502D8">
<mods:title id="C15FA37B0C89E9D06E66735279F48100">Five new species of the genus Torodora Meyrick (Lepidoptera, Lecithoceridae Torodorinae) from Cambodia, with a tentative check-list of the genus</mods:title>
</mods:titleInfo>
<mods:name id="493C5736D49AD99DA01A0EF14071E9F5" type="personal">
<mods:role id="9A1A474444F4EDD9ACD5A7E7BF8847B5">
<mods:roleTerm id="CDD18E858F8B981AE9EC0C163FB5873C">Author</mods:roleTerm>
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<mods:namePart id="7EA38E45A3A8DE53B552C17BE6D95AD6">Park, Kyu-Tek</mods:namePart>
<mods:affiliation id="0EE1700ECDEFEAC9B1CA5A0229B45481">The Korean Academy of Science and Technology, Seongnam, Gyeonggi Prov., 463 - 808 Korea.</mods:affiliation>
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<mods:roleTerm id="735DD7CC34913A880AEB3E4892910100">Author</mods:roleTerm>
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<mods:namePart id="DBADD140D6DB9F5C35CAA069FAE57471">Bae, Yang-Seop</mods:namePart>
<mods:nameIdentifier id="E9D54C0D1F745C689C332C977B684574" type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
<mods:affiliation id="AEFDB787F699F506B6F774DD9910F63D">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012 Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012 Korea. Corresponding author. baeys @ inu. ac. kr; https: // orcid. org / 0000 - 0001 - 7356 - 5633</mods:affiliation>
<mods:nameIdentifier id="92FC44C0FCF5BF5266DDB438BF86BDB5" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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<mods:title id="32C6780D65A825D3415DE9D37345452D">Zootaxa</mods:title>
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<mods:part id="64425E17D952C9D1AF413EAD9A5EF80D">
<mods:date id="3BC83396EC88FABCC8F7DA092EBE6E26">2021</mods:date>
<mods:detail id="483D437789CC3996293FB17BDDEB31D2" type="pubDate">
<mods:number id="6C29485B36F945C7901A8D52567FAF13">2021-03-24</mods:number>
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<mods:detail id="D458BE1C5D0819F7A0CDCEB5F8B907E2" type="volume">
<mods:number id="64DEEF280F58ED68A5B657CDC2CB146B">4949</mods:number>
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<mods:classification id="2EDAFD5E3F3606CB6CDCB221E09F39BC">journal article</mods:classification>
<mods:identifier id="A0B8ADC0DC5015D894E70F028CFC9BE0" type="CLB-Dataset">7497</mods:identifier>
<mods:identifier id="ECE2EC54DD81B6CD4C0357D0447A7B6C" type="DOI">10.11646/zootaxa.4949.1.5</mods:identifier>
<mods:identifier id="ED0C5C6B200C73F56401F74EBCDC3B6E" type="GBIF-Dataset">137cdaec-de44-48ae-be6b-fc9982803086</mods:identifier>
<mods:identifier id="2A5251ABA154371F93CFFD032E530E12" type="ISSN">1175-5326</mods:identifier>
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<treatment id="9610DF5C7B51FFDA4E99FF6FFCA22258" ID-DOI="http://doi.org/10.5281/zenodo.4635961" ID-GBIF-Taxon="180875407" ID-Zenodo-Dep="4635961" LSID="urn:lsid:plazi:treatment:9610DF5C7B51FFDA4E99FF6FFCA22258" httpUri="http://treatment.plazi.org/id/9610DF5C7B51FFDA4E99FF6FFCA22258" lastPageId="5" lastPageNumber="107" pageId="4" pageNumber="106">
<subSubSection id="56A33DC17B51FFDB4E99FF6FFDE621D0" box="[151,566,151,178]" pageId="4" pageNumber="106" type="nomenclature">
<paragraph id="1E066E4A7B51FFDB4E99FF6FFDE621D0" blockId="4.[151,566,151,213]" box="[151,566,151,178]" pageId="4" pageNumber="106">
<heading id="454ED9267B51FFDB4E99FF6FFDE621D0" bold="true" box="[151,566,151,178]" fontSize="11" level="1" pageId="4" pageNumber="106" reason="1">
<emphasis id="2CCDB2587B51FFDB4E99FF6FFDE621D0" bold="true" box="[151,566,151,178]" pageId="4" pageNumber="106">
<taxonomicName id="D9B915C97B51FFDB4E99FF6FFE1C21D0" ID-CoL="9FPVV" authority="Park" authorityName="Park" authorityYear="2021" box="[151,460,151,178]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="106" phylum="Arthropoda" rank="species" species="spathiana" status="sp. nov.">
<emphasis id="2CCDB2587B51FFDB4E99FF6FFE5521D3" bold="true" box="[151,389,151,178]" italics="true" pageId="4" pageNumber="106">Torodora spathiana</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="37FE0F237B51FFDB4FD7FF6FFDE621D0" box="[473,566,151,177]" pageId="4" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B51FFDB4E99FF43FEF821B4" box="[151,296,187,213]" pageId="4" pageNumber="106" type="description">
<paragraph id="1E066E4A7B51FFDB4E99FF43FEF821B4" blockId="4.[151,566,151,213]" box="[151,296,187,213]" pageId="4" pageNumber="106">
(
<figureCitation id="868272CF7B51FFDB4E91FF43FEF021B4" box="[159,288,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1784,1808]" captionTargetBox="[176,1412,657,1740]" captionTargetId="figure-163@4.[151,1436,633,1761]" captionTargetPageId="4" captionText="FIGURE 2. Torodora spathiana Park, sp. nov.: A, adult, holotype; B, labial palpus, in lateral view; C, hind tibia; D, male genitalia, gen. slide no. CIS-6063; E, aedeagus; F, abdominal segments VIII; G, genitalia of T. paratidosa (Wu), gen. slide no. CIS-6689 from Cambodia. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4635945" httpUri="https://zenodo.org/record/4635945/files/figure.png" pageId="4" pageNumber="106">Figs. 2A–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B51FFDB4E99FEFBFE782020" pageId="4" pageNumber="106" type="materials_examined">
<paragraph id="1E066E4A7B51FFDB4E99FEFBFE782020" blockId="4.[151,1437,259,610]" pageId="4" pageNumber="106">
<emphasis id="2CCDB2587B51FFDB4E99FEFBFF09207C" bold="true" box="[151,217,259,285]" pageId="4" pageNumber="106">Type.</emphasis>
<materialsCitation id="AED164177B51FFDB4ED1FEFBFE782020" ID-GBIF-Occurrence="3068362309" collectingDate="2002-07-03" collectingDateMax="2002-07-08" collectingDateMin="2002-07-03" collectorName="Bae &amp; Chae" country="Cambodia" location="Seima Biodiversity Conservation area" pageId="4" pageNumber="106" specimenCount="1" specimenCount-male="1" stateProvince="Mondolkiri" typeStatus="holotype">
<typeStatus id="C102D0E87B51FFDB4ED1FEFBFE9C207C" box="[223,332,259,285]" pageId="4" pageNumber="106" type="holotype">Holotype</typeStatus>
:
<specimenCount id="08BFA5C37B51FFDB4F58FEFCFEBD207F" box="[342,365,260,286]" pageId="4" pageNumber="106" type="male">♂</specimenCount>
,
<emphasis id="2CCDB2587B51FFDB4F76FEFBFE24207C" bold="true" box="[376,500,259,285]" pageId="4" pageNumber="106">
<collectingCountry id="66AE2EDA7B51FFDB4F76FEFBFE24207C" box="[376,500,259,285]" name="Cambodia" pageId="4" pageNumber="106">Cambodia</collectingCountry>
</emphasis>
,
<collectingRegion id="DC7DA0A87B51FFDB4C0CFEFBFD55207C" box="[514,645,259,285]" country="Cambodia" name="Mondolkiri" pageId="4" pageNumber="106">Mondulkiri</collectingRegion>
,
<location id="1B6638917B51FFDB4C81FEFBFBE9207F" LSID="urn:lsid:plazi:treatment:9610DF5C7B51FFDA4E99FF6FFCA22258:1B6638917B51FFDB4C81FEFBFBE9207F" box="[655,1081,259,286]" country="Cambodia" name="Seima Biodiversity Conservation area" pageId="4" pageNumber="106" stateProvince="Mondolkiri">Seima Biodiversity Conservation area</location>
,
<date id="6A07488A7B51FFDB4A4AFEFBFB03207C" box="[1092,1235,259,285]" pageId="4" pageNumber="106" value="2002-07-03" valueMax="2002-07-08" valueMin="2002-07-03">
<collectingDate id="7A43B1627B51FFDB4A4AFEFBFB03207C" box="[1092,1235,259,285]" pageId="4" pageNumber="106" value="2002-07-03" valueMax="2002-07-08" valueMin="2002-07-03">3–8 vii 2002</collectingDate>
</date>
, leg.
<collectorName id="B34C0B9C7B51FFDB4B03FEFCFAE9207F" box="[1293,1337,260,286]" pageId="4" pageNumber="106">Bae</collectorName>
&amp;
<collectorName id="B34C0B9C7B51FFDB4B55FEFBFA48207C" box="[1371,1432,259,285]" pageId="4" pageNumber="106">Chae</collectorName>
; gen. slide no. CIS-6063.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B51FFDB4EC9FEB3FAE520B0" pageId="4" pageNumber="106" type="diagnosis">
<paragraph id="1E066E4A7B51FFDB4EC9FEB3FAE520B0" blockId="4.[151,1437,259,610]" pageId="4" pageNumber="106">
<emphasis id="2CCDB2587B51FFDB4EC9FEB3FE922004" bold="true" box="[199,322,331,357]" pageId="4" pageNumber="106">Diagnosis.</emphasis>
The male genitalia of the new species are similar to those of
<taxonomicName id="D9B915C97B51FFDB4A0EFEB4FAF12004" authority="(Wu, 1994)" baseAuthorityName="Wu" baseAuthorityYear="1994" box="[1024,1313,331,357]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="106" phylum="Arthropoda" rank="species" species="parotidosa">
<emphasis id="2CCDB2587B51FFDB4A0EFEB4FB442004" box="[1024,1172,331,357]" italics="true" pageId="4" pageNumber="106">T. parotidosa</emphasis>
(
<bibRefCitation id="7A2813BB7B51FFDB4AABFEB3FAC92004" author="Wu, C. &amp; Liu, Y." box="[1189,1305,331,357]" pageId="4" pageNumber="106" pagination="55 - 173" refId="ref6370" refString="Wu, C. &amp; Liu, Y. (1994) A study of Chinese Torodora Meyrick, 1894 and description of new species. Sinzoologica, 11, 55 - 173." type="journal article" year="1994">Wu, 1994</bibRefCitation>
)
</taxonomicName>
, which are known in
<collectingCountry id="66AE2EDA7B51FFDB4F06FE97FE9B20E8" box="[264,331,367,393]" name="China" pageId="4" pageNumber="106">China</collectingCountry>
and
<collectingCountry id="66AE2EDA7B51FFDB4F8FFE97FE3720E8" box="[385,487,367,393]" name="Thailand" pageId="4" pageNumber="106">Thailand</collectingCountry>
, but it can be distinguished by the valva extremely narrowed beyond the rectangular basal part, and the juxta with asymmetrical latero-caudal lobes. The new species is also differentiated from the latter by the more or less slender 2
<superScript id="E9CCC3027B51FFDB4FD7FE4FFE3920A4" attach="left" box="[473,489,439,453]" fontSize="6" pageId="4" pageNumber="106">nd</superScript>
segment of labial palpus, whereas
<taxonomicName id="D9B915C97B51FFDB4D7CFE40FBD620B0" baseAuthorityName="Wu" baseAuthorityYear="1994" box="[882,1030,439,465]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="106" phylum="Arthropoda" rank="species" species="parotidosa">
<emphasis id="2CCDB2587B51FFDB4D7CFE40FBD620B0" box="[882,1030,439,465]" italics="true" pageId="4" pageNumber="106">T. parotidosa</emphasis>
</taxonomicName>
has rough scales ventrally.
</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B51FFDA4EC9FE23FE4123AC" lastPageId="5" lastPageNumber="107" pageId="4" pageNumber="106" type="description">
<paragraph id="1E066E4A7B51FFDB4EC9FE23FCF92097" blockId="4.[151,1437,259,610]" box="[199,809,475,502]" pageId="4" pageNumber="106">
<emphasis id="2CCDB2587B51FFDB4EC9FE23FE892094" bold="true" box="[199,345,475,501]" pageId="4" pageNumber="106">Description.</emphasis>
Adult (
<figureCitation id="868272CF7B51FFDB4FA3FE23FDE42094" box="[429,564,475,501]" captionStart="FIGURE 2" captionStartId="4.[151,250,1784,1808]" captionTargetBox="[176,1412,657,1740]" captionTargetId="figure-163@4.[151,1436,633,1761]" captionTargetPageId="4" captionText="FIGURE 2. Torodora spathiana Park, sp. nov.: A, adult, holotype; B, labial palpus, in lateral view; C, hind tibia; D, male genitalia, gen. slide no. CIS-6063; E, aedeagus; F, abdominal segments VIII; G, genitalia of T. paratidosa (Wu), gen. slide no. CIS-6689 from Cambodia. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4635945" httpUri="https://zenodo.org/record/4635945/files/figure.png" pageId="4" pageNumber="106">Figs. 2A–C</figureCitation>
). Wingspan 19.0 mm.
</paragraph>
<paragraph id="1E066E4A7B51FFDB4EC9FE07FE672300" blockId="4.[151,1437,259,610]" pageId="4" pageNumber="106">
<emphasis id="2CCDB2587B51FFDB4EC9FE07FEDD2378" box="[199,269,511,537]" italics="true" pageId="4" pageNumber="106">Head:</emphasis>
Yellowish brown dorsally. Antenna slightly longer than forewing; scape elongated, not dilated apically, grayish orange; flagellum grayish orange, lacking annulations, not ciliate. Second segment of labial palpus thickened; 3
<superScript id="E9CCC3027B51FFDB4EE7FDBFFF272334" attach="left" box="[233,247,583,597]" fontSize="6" pageId="4" pageNumber="106">rd</superScript>
segment broken.
</paragraph>
<caption id="4AC63EC27B51FFDB4E99F900FCAC2639" ID-DOI="http://doi.org/10.5281/zenodo.4635945" ID-Zenodo-Dep="4635945" httpUri="https://zenodo.org/record/4635945/files/figure.png" pageId="4" pageNumber="106" startId="4.[151,250,1784,1808]" targetBox="[176,1412,657,1740]" targetPageId="4">
<paragraph id="1E066E4A7B51FFDB4E99F900FCAC2639" blockId="4.[151,1437,1784,1881]" pageId="4" pageNumber="106">
<emphasis id="2CCDB2587B51FFDB4E99F900FEC52671" bold="true" box="[151,277,1784,1808]" pageId="4" pageNumber="106">FIGURE 2.</emphasis>
<taxonomicName id="D9B915C97B51FFDB4F10F900FDF32671" authority="Park" authorityName="Park" authorityYear="2021" box="[286,547,1784,1808]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="106" phylum="Arthropoda" rank="species" species="spathiana" status="sp. nov.">
<emphasis id="2CCDB2587B51FFDB4F10F900FE392671" box="[286,489,1784,1808]" italics="true" pageId="4" pageNumber="106">Torodora spathiana</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="37FE0F237B51FFDB4C21F901FD552671" box="[559,645,1785,1809]" pageId="4" pageNumber="106" rank="species">
<emphasis id="2CCDB2587B51FFDB4C21F901FDAE2670" bold="true" box="[559,638,1785,1809]" pageId="4" pageNumber="106">sp. nov</emphasis>
.
</taxonomicNameLabel>
: A, adult, holotype; B, labial palpus, in lateral view; C, hind tibia; D, male genitalia, gen. slide no. CIS-6063; E, aedeagus; F, abdominal segments VIII; G, genitalia of
<taxonomicName id="D9B915C97B51FFDB4A4BF8E5FAD92654" authority="(Wu)" baseAuthorityName="Wu" box="[1093,1289,1820,1845]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="106" phylum="Arthropoda" rank="species" species="paratidosa">
<emphasis id="2CCDB2587B51FFDB4A4BF8E5FB1C2655" box="[1093,1228,1820,1844]" italics="true" pageId="4" pageNumber="106">T. paratidosa</emphasis>
(Wu)
</taxonomicName>
, gen. slide no. CIS-6689 from Cambodia. Scale bar for genitalia and aedeagus: 1.0 mm.
</paragraph>
</caption>
<paragraph id="1E066E4A7B51FFDA4EC9F87FFC8A2020" blockId="4.[151,1436,1927,2025]" lastBlockId="5.[151,1437,151,825]" lastPageId="5" lastPageNumber="107" pageId="4" pageNumber="106">
<emphasis id="2CCDB2587B51FFDB4EC9F87FFEC826C0" box="[199,280,1927,1953]" italics="true" pageId="4" pageNumber="106">Thorax</emphasis>
: Tegula and thorax yellowish brown dorsally. Hind tibia with rough scales above from base to near apex; apex with white scales. Forewing ground color grayish orange to yellowish brown, without distinct markings; costa slightly arched around basal 1/3 and beyond 2/3; apex obtuse; termen slightly concave; fringe concolorous with ground color; venation with R
<subScript id="823D6C0F7B50FFDA4C26FF51FDE021D6" attach="left" box="[552,560,169,183]" fontSize="6" pageId="5" pageNumber="107">1</subScript>
arising from beyond middle; distance between origins of R
<subScript id="823D6C0F7B50FFDA4AD8FF51FB0E21D6" attach="left" box="[1238,1246,169,183]" fontSize="6" pageId="5" pageNumber="107">2</subScript>
and R
<subScript id="823D6C0F7B50FFDA4B27FF51FAE121D6" attach="left" box="[1321,1329,169,183]" fontSize="6" pageId="5" pageNumber="107">3</subScript>
less than 1/2 length of R
<subScript id="823D6C0F7B50FFDA4F4DFF35FE9B21BA" attach="left" box="[323,331,205,219]" fontSize="6" pageId="5" pageNumber="107">1</subScript>
and R
<subScript id="823D6C0F7B50FFDA4F98FF35FE4E21BA" attach="left" box="[406,414,205,219]" fontSize="6" pageId="5" pageNumber="107">2</subScript>
; R
<subScript id="823D6C0F7B50FFDA4FCFFF35FE1921BA" attach="left" box="[449,457,205,219]" fontSize="6" pageId="5" pageNumber="107">3</subScript>
stalked with R
<subScript id="823D6C0F7B50FFDA4C7FFF35FD5B21BA" attach="left" box="[625,651,205,219]" fontSize="6" pageId="5" pageNumber="107">4+5</subScript>
for basal 1/3 length; R
<subScript id="823D6C0F7B50FFDA4D9FFF35FC4921BA" attach="left" box="[913,921,205,219]" fontSize="6" pageId="5" pageNumber="107">4</subScript>
and R
<subScript id="823D6C0F7B50FFDA4DEBFF35FC3D21BA" attach="left" box="[997,1005,205,219]" fontSize="6" pageId="5" pageNumber="107">5</subScript>
stalked for more than basal 2/3; R
<subScript id="823D6C0F7B50FFDA4B78FF35FAAE21BA" attach="left" box="[1398,1406,205,219]" fontSize="6" pageId="5" pageNumber="107">5</subScript>
to termen; M
<subScript id="823D6C0F7B50FFDA4F02FF09FEC4219E" attach="left" box="[268,276,241,255]" fontSize="6" pageId="5" pageNumber="107">1</subScript>
remote from R
<subScript id="823D6C0F7B50FFDA4FB0FF09FE07219E" attach="left" box="[446,471,241,255]" fontSize="6" pageId="5" pageNumber="107">3+4</subScript>
; M
<subScript id="823D6C0F7B50FFDA4FF0FF09FDD6219E" attach="left" box="[510,518,241,255]" fontSize="6" pageId="5" pageNumber="107">2</subScript>
nearly parallel to M
<subScript id="823D6C0F7B50FFDA4CE6FF09FD20219E" attach="left" box="[744,752,241,255]" fontSize="6" pageId="5" pageNumber="107">1</subScript>
; CuA
<subScript id="823D6C0F7B50FFDA4D3DFF09FCEB219E" attach="both" box="[819,827,241,255]" fontSize="6" pageId="5" pageNumber="107">1</subScript>
and CuA
<subScript id="823D6C0F7B50FFDA4D90FF09FC76219E" attach="left" box="[926,934,241,255]" fontSize="6" pageId="5" pageNumber="107">2</subScript>
stalked for basal half; cell closed. Hind wing paler and broader than forewing; venation with M
<subScript id="823D6C0F7B50FFDA4CC9FEEDFD1F2042" attach="left" box="[711,719,277,291]" fontSize="6" pageId="5" pageNumber="107">2</subScript>
; close to M
<subScript id="823D6C0F7B50FFDA4D5BFEEDFC8D2042" attach="left" box="[853,861,277,291]" fontSize="6" pageId="5" pageNumber="107">2</subScript>
; M
<subScript id="823D6C0F7B50FFDA4D8BFEEDFC5D2042" attach="left" box="[901,909,277,291]" fontSize="6" pageId="5" pageNumber="107">3</subScript>
and CuA
<subScript id="823D6C0F7B50FFDA4DF7FEEDFBD12042" attach="left" box="[1017,1025,277,291]" fontSize="6" pageId="5" pageNumber="107">1</subScript>
nearly connate at base; apex obtuse; termen slightly concave; fringe concolorous with ground color.
</paragraph>
<paragraph id="1E066E4A7B50FFDA4EC9FEB3FBBA2004" blockId="5.[151,1437,151,825]" box="[199,1130,331,357]" pageId="5" pageNumber="107">
<emphasis id="2CCDB2587B50FFDA4EC9FEB3FEE12004" box="[199,305,331,357]" italics="true" pageId="5" pageNumber="107">Abdomen</emphasis>
(
<figureCitation id="868272CF7B50FFDA4F4EFEB3FE462004" box="[320,406,331,357]" captionStart="FIGURE 3" captionStartId="6.[151,250,1824,1848]" captionTargetBox="[175,1410,205,1764]" captionTargetId="figure-19@6.[151,1436,181,1800]" captionTargetPageId="6" captionText="FIGURE 3. Torodora alterniella Park, sp. nov.: A, adult, paratype from Laos; B, labial palpus and antenna; C, male genitalia, gen. slide no. CIS-8097, paratype from Laos; D, ditto, aedeagus; E, aedeagus of holotype, gen. slide no. CIS-6097; F, abdominal segments V–VIII, holotype; G, female genitalia., gen. slide no. CIS-8082 from Cambodia. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4635947" httpUri="https://zenodo.org/record/4635947/files/figure.png" pageId="5" pageNumber="107">Fig. 3F</figureCitation>
): Spinous zones on dorsal surface nearly triangularly developed.
</paragraph>
<paragraph id="1E066E4A7B50FFDA4EC9FE97FBBA23C8" blockId="5.[151,1437,151,825]" pageId="5" pageNumber="107">
<emphasis id="2CCDB2587B50FFDA4EC9FE97FEBD20E8" box="[199,365,367,393]" italics="true" pageId="5" pageNumber="107">Male genitalia</emphasis>
(
<figureCitation id="868272CF7B50FFDA4F72FE97FDD320E8" box="[380,515,367,393]" captionStart="FIGURE 3" captionStartId="6.[151,250,1824,1848]" captionTargetBox="[175,1410,205,1764]" captionTargetId="figure-19@6.[151,1436,181,1800]" captionTargetPageId="6" captionText="FIGURE 3. Torodora alterniella Park, sp. nov.: A, adult, paratype from Laos; B, labial palpus and antenna; C, male genitalia, gen. slide no. CIS-8097, paratype from Laos; D, ditto, aedeagus; E, aedeagus of holotype, gen. slide no. CIS-6097; F, abdominal segments V–VIII, holotype; G, female genitalia., gen. slide no. CIS-8082 from Cambodia. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4635947" httpUri="https://zenodo.org/record/4635947/files/figure.png" pageId="5" pageNumber="107">Figs. 3D, E</figureCitation>
). Uncus elongated, heavily sclerotized, bent downward, much exceeded posterior margin of the basal plate of gnathos. Median process of gnathos narrowed toward apex, strongly bent beyond 2/3, sharply pointed apically. Tegumen weakly sclerotized, concave in inverted V-shape on anterior margin. Valva broad at base, quadrate in basal part, abruptly narrowed medially; cucullus broadly expanded into spoon-shaped, with round outer margin; sacculus not developed. Vinculum banded, sclerotized. Juxta shield-shaped; latero-caudal processes asymmetrical, left one longer and right one about 1/2 length of the left, with median ridge in anterior half. Aedeagus very stout basally; width of basal half about 1.5 times than the width of basal part of valva, as long as valva; cornuti consisting of a heavily sclerotized, crescent plate with wrinkled tube basally beyond middle, a long, narrow sac containing numerous conic spines from near base to 2/3 length of aedeagus.
</paragraph>
<paragraph id="1E066E4A7B50FFDA4EC9FD4BFE4123AC" blockId="5.[151,1437,151,825]" box="[199,401,691,717]" pageId="5" pageNumber="107">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B50FFDA4EC9FD2EFD6C2390" box="[199,700,726,753]" pageId="5" pageNumber="107" type="distribution">
<paragraph id="1E066E4A7B50FFDA4EC9FD2EFD6C2390" blockId="5.[151,1437,151,825]" box="[199,700,726,753]" pageId="5" pageNumber="107">
<emphasis id="2CCDB2587B50FFDA4EC9FD2EFEB12391" bold="true" box="[199,353,726,752]" pageId="5" pageNumber="107">Distribution.</emphasis>
<collectingCountry id="66AE2EDA7B50FFDA4F69FD2FFE0A2390" box="[359,474,727,753]" name="Cambodia" pageId="5" pageNumber="107">Cambodia</collectingCountry>
(
<collectingRegion id="DC7DA0A87B50FFDA4FE7FD2FFD612390" box="[489,689,727,753]" country="Cambodia" name="Mondolkiri" pageId="5" pageNumber="107">Mondulkiri Prov.</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B50FFDA4EC9FD03FCA22258" pageId="5" pageNumber="107" type="etymology">
<paragraph id="1E066E4A7B50FFDA4EC9FD03FCA22258" blockId="5.[151,1437,151,825]" pageId="5" pageNumber="107">
<emphasis id="2CCDB2587B50FFDA4EC9FD03FE9E2274" bold="true" box="[199,334,763,789]" pageId="5" pageNumber="107">Etymology.</emphasis>
The species name is derived from the Latin,
<emphasis id="2CCDB2587B50FFDA4D47FD03FC442274" box="[841,916,763,789]" italics="true" pageId="5" pageNumber="107">spatha</emphasis>
(= spoon-shaped, paddle), with a Latin suffix, -
<emphasis id="2CCDB2587B50FFDA4EAFFCE7FF0C2259" box="[161,220,799,824]" italics="true" pageId="5" pageNumber="107">anus,</emphasis>
referring /to the paddle-shaped valva in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>
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