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<document id="BE22D41383D7C7516D8D2B6A7BB1BEE4" ID-CLB-Dataset="7497" ID-DOI="10.11646/zootaxa.4949.1.5" ID-GBIF-Dataset="137cdaec-de44-48ae-be6b-fc9982803086" ID-ISSN="1175-5326" ID-Zenodo-Dep="4635941" ID-ZooBank="2C821F67-4B27-48D1-B5F4-2DD2A35BA3FA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616662070952" checkinUser="plazi" docAuthor="Park, Kyu-Tek &amp; Bae, Yang-Seop" docDate="2021" docId="9610DF5C7B50FFD84E99FC72FB96223C" docLanguage="en" docName="zootaxa.4949.1.5.pdf" docOrigin="Zootaxa 4949 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Torodora alterniella Park &amp; Bae 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="109" masterDocId="6A29A7247B55FFDF4E0EFFF8FFD02161" masterDocTitle="Five new species of the genus Torodora Meyrick (Lepidoptera, Lecithoceridae Torodorinae) from Cambodia, with a tentative check-list of the genus" masterLastPageNumber="114" masterPageNumber="102" pageNumber="107" updateTime="1698934944201" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="FADE28CBB0FD437D07A370F14A15A0BA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="985D80C74F70FC0D221DE2BC461C3B3D">
<mods:title id="2E8E24EE582F4B63DC03B90E510D3A00">Five new species of the genus Torodora Meyrick (Lepidoptera, Lecithoceridae Torodorinae) from Cambodia, with a tentative check-list of the genus</mods:title>
</mods:titleInfo>
<mods:name id="2161166073049A3777290C4C7DEDE85C" type="personal">
<mods:role id="81148C808E74608CB8240FA284EE4167">
<mods:roleTerm id="AA9603D222F2F9964D8AD0AEFE208D56">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1C48A14153379B38C1D176588A8719E2">Park, Kyu-Tek</mods:namePart>
<mods:affiliation id="96E3DAADC36E11DF63564928CA9BB33F">The Korean Academy of Science and Technology, Seongnam, Gyeonggi Prov., 463 - 808 Korea.</mods:affiliation>
</mods:name>
<mods:name id="6682F013A770F1C9FDD5C36A5D4D69A5" type="personal">
<mods:role id="D26F49E4751F7F5C0D721BD8586426D2">
<mods:roleTerm id="69AB8F1C7F31ADBF3ABB1C3098C45149">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C5D053826257253A267E0202AF5EF72F">Bae, Yang-Seop</mods:namePart>
<mods:nameIdentifier id="3A0DDB2934CBD80D80541B7D927F61C8" type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
<mods:affiliation id="2F3C207965F113D4A28F0B9FBF825D84">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012 Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012 Korea. Corresponding author. baeys @ inu. ac. kr; https: // orcid. org / 0000 - 0001 - 7356 - 5633</mods:affiliation>
<mods:nameIdentifier id="896D18FAEDF34AB6F2F50B66633CB095" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="BC36076D7A86E51ADA08E8AC615E7BEF">text</mods:typeOfResource>
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<mods:titleInfo id="01BDF6A3EC4FE7A78930D91B47A7AB10">
<mods:title id="75F460012FC21EEF82D38F1FD4C15B3A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A401F53ACB0DCFE7CF4FE163EC591F08">
<mods:date id="0DA5012F06474A2CAF50249F21F4DF9B">2021</mods:date>
<mods:detail id="DD9D1BFC5AD531D8201C8CC4CB746AD3" type="pubDate">
<mods:number id="336550A19D96B7C8230C7D9BA5148D4B">2021-03-24</mods:number>
</mods:detail>
<mods:detail id="6699081F7FAAF4268E4D05C829C0F52B" type="volume">
<mods:number id="C61DFABC5C52F94E42521A686C46B993">4949</mods:number>
</mods:detail>
<mods:detail id="E4D329864E0F0F53CF77D818E2C5233F" type="issue">
<mods:number id="2E3FBDD4E531195A24AC350A93617166">1</mods:number>
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<mods:start id="E791F74C034D48FFA53BDCDFD4689519">102</mods:start>
<mods:end id="649129302C1A82C39FBA3D8BC24752DF">114</mods:end>
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<mods:classification id="C30375720B24DAA82CEE71D828639FF0">journal article</mods:classification>
<mods:identifier id="FC6C8A2D45350D1465632FA0827C3AE5" type="CLB-Dataset">7497</mods:identifier>
<mods:identifier id="57BFDC5EFC07B10FB7536F76B2E070BE" type="DOI">10.11646/zootaxa.4949.1.5</mods:identifier>
<mods:identifier id="5FCDEE1A4AFC5B53209D7F2B8022679C" type="GBIF-Dataset">137cdaec-de44-48ae-be6b-fc9982803086</mods:identifier>
<mods:identifier id="7F3E16F44A54A03E9984F19F1C330E34" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0FC966F173FA4BBD76A083CA8FDA7271" type="Zenodo-Dep">4635941</mods:identifier>
<mods:identifier id="FD7A4BC91A7EFB1CC8E6208F69B268CE" type="ZooBank">2C821F67-4B27-48D1-B5F4-2DD2A35BA3FA</mods:identifier>
</mods:mods>
<treatment id="9610DF5C7B50FFD84E99FC72FB96223C" ID-DOI="http://doi.org/10.5281/zenodo.4663328" ID-GBIF-Taxon="180875408" ID-Zenodo-Dep="4663328" LSID="urn:lsid:plazi:treatment:9610DF5C7B50FFD84E99FC72FB96223C" httpUri="http://treatment.plazi.org/id/9610DF5C7B50FFD84E99FC72FB96223C" lastPageId="7" lastPageNumber="109" pageId="5" pageNumber="107">
<subSubSection id="56A33DC17B50FFDA4E99FC72FDEB22C4" box="[151,571,906,933]" pageId="5" pageNumber="107" type="nomenclature">
<paragraph id="1E066E4A7B50FFDA4E99FC72FDEB22C4" blockId="5.[151,571,906,969]" box="[151,571,906,933]" pageId="5" pageNumber="107">
<heading id="454ED9267B50FFDA4E99FC72FDEB22C4" bold="true" box="[151,571,906,933]" fontSize="11" level="1" pageId="5" pageNumber="107" reason="1">
<emphasis id="2CCDB2587B50FFDA4E99FC72FDEB22C4" bold="true" box="[151,571,906,933]" pageId="5" pageNumber="107">
<taxonomicName id="D9B915C97B50FFDA4E99FC72FE0122C4" ID-CoL="9FPVX" authority="Park" authorityName="Park &amp; Bae" authorityYear="2021" box="[151,465,906,933]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="107" phylum="Arthropoda" rank="species" species="alterniella" status="sp. nov.">
<emphasis id="2CCDB2587B50FFDA4E99FC72FE5A22C4" bold="true" box="[151,394,906,933]" italics="true" pageId="5" pageNumber="107">Torodora alterniella</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="37FE0F237B50FFDA4FD0FC73FDEB22C4" box="[478,571,907,933]" pageId="5" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B50FFDA4E99FC57FEF822A8" box="[151,296,943,969]" pageId="5" pageNumber="107" type="description">
<paragraph id="1E066E4A7B50FFDA4E99FC57FEF822A8" blockId="5.[151,571,906,969]" box="[151,296,943,969]" pageId="5" pageNumber="107">
(
<figureCitation id="868272CF7B50FFDA4E91FC57FEF022A8" box="[159,288,943,969]" captionStart="FIGURE 3" captionStartId="6.[151,250,1824,1848]" captionTargetBox="[175,1410,205,1764]" captionTargetId="figure-19@6.[151,1436,181,1800]" captionTargetPageId="6" captionText="FIGURE 3. Torodora alterniella Park, sp. nov.: A, adult, paratype from Laos; B, labial palpus and antenna; C, male genitalia, gen. slide no. CIS-8097, paratype from Laos; D, ditto, aedeagus; E, aedeagus of holotype, gen. slide no. CIS-6097; F, abdominal segments V–VIII, holotype; G, female genitalia., gen. slide no. CIS-8082 from Cambodia. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4635947" httpUri="https://zenodo.org/record/4635947/files/figure.png" pageId="5" pageNumber="107">Figs. 3A–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B50FFDA4E99FC0FFD91251C" pageId="5" pageNumber="107" type="materials_examined">
<paragraph id="1E066E4A7B50FFDA4E99FC0FFD91251C" blockId="5.[151,1437,1014,2013]" pageId="5" pageNumber="107">
<emphasis id="2CCDB2587B50FFDA4E99FC0FFE972571" bold="true" box="[151,327,1014,1041]" pageId="5" pageNumber="107">Type material.</emphasis>
<materialsCitation id="AED164177B50FFDA4F41FC0FFE4A2554" ID-GBIF-Occurrence="3068362306" collectingDate="2011-05-01" collectingDateMax="2011-05-02" collectingDateMin="2011-05-01" collectorName="JB Heppner" country="Cambodia" elevation="40" location="Spring Vy Lodge" pageId="5" pageNumber="107" specimenCount="1" specimenCount-male="1" stateProvince="Kep" typeStatus="holotype">
<typeStatus id="C102D0E87B50FFDA4F41FC0FFE6C2570" box="[335,444,1015,1041]" pageId="5" pageNumber="107" type="holotype">Holotype</typeStatus>
:
<specimenCount id="08BFA5C37B50FFDA4FC6FC0FFE0F2570" box="[456,479,1015,1041]" pageId="5" pageNumber="107" type="male">♂</specimenCount>
,
<emphasis id="2CCDB2587B50FFDA4FE2FC0EFDB82571" bold="true" box="[492,616,1014,1040]" pageId="5" pageNumber="107">
<collectingCountry id="66AE2EDA7B50FFDA4FE2FC0EFDB82571" box="[492,616,1014,1040]" name="Cambodia" pageId="5" pageNumber="107">Cambodia</collectingCountry>
</emphasis>
,
<collectingRegion id="DC7DA0A87B50FFDA4C76FC0FFD3D2570" box="[632,749,1015,1041]" country="Cambodia" name="Kep" pageId="5" pageNumber="107">Kep Prov.</collectingRegion>
,
<collectingRegion id="DC7DA0A87B50FFDA4CF7FC0FFCF82570" box="[761,808,1015,1041]" country="Cambodia" name="Kep" pageId="5" pageNumber="107">Kep</collectingRegion>
<location id="1B6638917B50FFDA4D3FFC0FFC252570" LSID="urn:lsid:plazi:treatment:9610DF5C7B50FFD84E99FC72FB96223C:1B6638917B50FFDA4D3FFC0FFC252570" box="[817,1013,1015,1041]" country="Cambodia" name="Spring Vy Lodge" pageId="5" pageNumber="107" stateProvince="Kep">Spring Vy Lodge</location>
,
<quantity id="D941C3AF7B50FFDA4A0CFC0FFBEF2570" box="[1026,1087,1015,1041]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="107" unit="m" value="40.0">
<elevation id="959489797B50FFDA4A0CFC0FFBEF2570" box="[1026,1087,1015,1041]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="107" unit="m" value="40.0">40 m</elevation>
</quantity>
,
<date id="6A07488A7B50FFDA4A42FC0FFB1F2570" box="[1100,1231,1015,1041]" pageId="5" pageNumber="107" value="2011-05-01" valueMax="2011-05-02" valueMin="2011-05-01">
<collectingDate id="7A43B1627B50FFDA4A42FC0FFB1F2570" box="[1100,1231,1015,1041]" pageId="5" pageNumber="107" value="2011-05-01" valueMax="2011-05-02" valueMin="2011-05-01">1–2 v 2011</collectingDate>
</date>
, leg.
<collectorName id="B34C0B9C7B50FFDA4B00FC0FFA482570" box="[1294,1432,1015,1041]" pageId="5" pageNumber="107">JB Heppner</collectorName>
; gen. side no. CIS-6097
</materialsCitation>
.
<typeStatus id="C102D0E87B50FFDA4FAAFBE3FDC52554" box="[420,533,1051,1077]" pageId="5" pageNumber="107" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="08BFA5C37B50FFDA4C11FBE3FD942554" box="[543,580,1051,1077]" pageId="5" pageNumber="107" type="female">2♀</specimenCount>
,
<materialsCitation id="AED164177B50FFDA4C5EFBE3FAB82554" ID-GBIF-Occurrence="3068362302" box="[592,1384,1051,1077]" collectingDate="2010-10-07" collectorName="Bae &amp; Ju &amp; Vi" country="Cambodia" location="Mondulkiri" pageId="5" pageNumber="107" specimenCount="1" stateProvince="Mondolkiri" typeStatus="paratype">
<collectingRegion id="DC7DA0A87B50FFDA4C5EFBE3FD032554" box="[592,723,1051,1077]" country="Cambodia" name="Mondolkiri" pageId="5" pageNumber="107">Mondulkiri</collectingRegion>
,
<date id="6A07488A7B50FFDA4CD2FBE3FC902554" box="[732,832,1051,1077]" pageId="5" pageNumber="107" value="2010-10-07">
<collectingDate id="7A43B1627B50FFDA4CD2FBE3FC902554" box="[732,832,1051,1077]" pageId="5" pageNumber="107" value="2010-10-07">7 x 2010</collectingDate>
</date>
, leg.
<collectorName id="B34C0B9C7B50FFDA4D74FBE3FC782554" box="[890,936,1051,1077]" pageId="5" pageNumber="107">Bae</collectorName>
,
<collectorName id="B34C0B9C7B50FFDA4DBDFBE3FC1C2554" box="[947,972,1051,1077]" pageId="5" pageNumber="107">Ju</collectorName>
&amp;
<collectorName id="B34C0B9C7B50FFDA4DE0FBE3FBDC2554" box="[1006,1036,1051,1077]" pageId="5" pageNumber="107">Vi</collectorName>
; gen. side no. CIS-8079, -8082
</materialsCitation>
.
<materialsCitation id="AED164177B50FFDA4B7CFBE2FDEC251C" ID-GBIF-Occurrence="3068362307" collectingDate="2019-07-26" collectorName="Bae &amp; Ju &amp; Vi" country="Laos" elevation="470" latitude="8.456388" location="Park" longLatPrecision="21" longitude="103.05139" pageId="5" pageNumber="107" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Bolikhamxai" typeStatus="paratype">
<specimenCount id="08BFA5C37B50FFDA4B7CFBE2FA472554" box="[1394,1431,1050,1077]" pageId="5" pageNumber="107" type="male">
<emphasis id="2CCDB2587B50FFDA4B7CFBE2FA502555" bold="true" box="[1394,1408,1050,1076]" pageId="5" pageNumber="107">3</emphasis>
♂
</specimenCount>
,
<specimenCount id="08BFA5C37B50FFDA4E99FBC7FF6A2538" box="[151,186,1087,1113]" pageId="5" pageNumber="107" type="female">1♀</specimenCount>
,
<emphasis id="2CCDB2587B50FFDA4EC6FBC7FED22538" bold="true" box="[200,258,1087,1113]" pageId="5" pageNumber="107">
<collectingCountry id="66AE2EDA7B50FFDA4EC6FBC7FED22538" box="[200,258,1087,1113]" name="Laos" pageId="5" pageNumber="107">Laos</collectingCountry>
</emphasis>
,
<collectingRegion id="DC7DA0A87B50FFDA4F1EFBC7FE312538" box="[272,481,1087,1113]" country="Laos" name="Bolikhamxai" pageId="5" pageNumber="107">Bolikhamsai Prov.</collectingRegion>
, PKK nat.
<location id="1B6638917B50FFDA4C55FBC7FD412538" LSID="urn:lsid:plazi:treatment:9610DF5C7B50FFD84E99FC72FB96223C:1B6638917B50FFDA4C55FBC7FD412538" box="[603,657,1087,1113]" country="Laos" latitude="8.456388" longLatPrecision="21" longitude="103.05139" name="Park" pageId="5" pageNumber="107" stateProvince="Bolikhamxai">Park</location>
,
<geoCoordinate id="7B8D088D7B50FFDA4C93FBC7FCDE253A" box="[669,782,1087,1115]" degrees="8" direction="north" minutes="27" orientation="latitude" pageId="5" pageNumber="107" precision="15" seconds="23" value="8.456388">
8
<superScript id="E9CCC3027B50FFDA4CA5FBC7FD63252C" attach="none" box="[683,691,1087,1101]" fontSize="6" pageId="5" pageNumber="107">o</superScript>
27′23˝N
</geoCoordinate>
<geoCoordinate id="7B8D088D7B50FFDA4D1BFBC7FC612538" box="[789,945,1086,1115]" degrees="103" direction="east" minutes="03" orientation="longitude" pageId="5" pageNumber="107" precision="15" seconds="05" value="103.05139">
103
<superScript id="E9CCC3027B50FFDA4D31FBC7FC97252C" attach="left" box="[831,839,1087,1101]" fontSize="6" pageId="5" pageNumber="107">o</superScript>
03′ 05˝E
</geoCoordinate>
,
<quantity id="D941C3AF7B50FFDA4DB2FBC7FBD52538" box="[956,1029,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="5" pageNumber="107" unit="m" value="470.0">
<elevation id="959489797B50FFDA4DB2FBC7FBD52538" box="[956,1029,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="5" pageNumber="107" unit="m" value="470.0">470 m</elevation>
</quantity>
,
<date id="6A07488A7B50FFDA4A1FFBC7FB432538" box="[1041,1171,1087,1113]" pageId="5" pageNumber="107" value="2019-07-26">
<collectingDate id="7A43B1627B50FFDA4A1FFBC7FB432538" box="[1041,1171,1087,1113]" pageId="5" pageNumber="107" value="2019-07-26">26 vii 2019</collectingDate>
</date>
, leg.
<collectorName id="B34C0B9C7B50FFDA4AC0FBC7FB2C2538" box="[1230,1276,1087,1113]" pageId="5" pageNumber="107">Bae</collectorName>
,
<collectorName id="B34C0B9C7B50FFDA4B06FBC7FAF12538" box="[1288,1313,1087,1113]" pageId="5" pageNumber="107">Ju</collectorName>
&amp;
<collectorName id="B34C0B9C7B50FFDA4B4AFBC7FAB22538" box="[1348,1378,1087,1113]" pageId="5" pageNumber="107">Vi</collectorName>
; gen. side no. CIS-8097 (
<specimenCount id="08BFA5C37B50FFDA4F7DFB9BFE5A251C" box="[371,394,1123,1149]" pageId="5" pageNumber="107" type="male">♂</specimenCount>
), CIS-8092 (
<specimenCount id="08BFA5C37B50FFDA4C10FB9BFDE1251C" box="[542,561,1123,1149]" pageId="5" pageNumber="107" type="female">♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B50FFDA4EC9FB7EFBC52418" pageId="5" pageNumber="107" type="diagnosis">
<paragraph id="1E066E4A7B50FFDA4EC9FB7EFBC52418" blockId="5.[151,1437,1014,2013]" pageId="5" pageNumber="107">
<emphasis id="2CCDB2587B50FFDA4EC9FB7EFE9225C1" bold="true" box="[199,322,1158,1184]" pageId="5" pageNumber="107">Diagnosis.</emphasis>
The new species, as well as following two new species, are very similar to
<taxonomicName id="D9B915C97B50FFDA4A88FB7FFF0925A4" authority="(Meyrick, 1932)" baseAuthorityName="Meyrick" baseAuthorityYear="1932" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="107" phylum="Arthropoda" rank="species" species="parthenopis">
<emphasis id="2CCDB2587B50FFDA4A88FB7FFAF725C0" box="[1158,1319,1159,1185]" italics="true" pageId="5" pageNumber="107">T. parthenopis</emphasis>
(Meyrick, 1932)
</taxonomicName>
which was described from
<collectingCountry id="66AE2EDA7B50FFDA4C14FB53FDBC25A4" box="[538,620,1195,1221]" name="Taiwan" pageId="5" pageNumber="107">Taiwan</collectingCountry>
in the forewing pattern by sharing following characters: Ground color grayish-orange with sharply produced apex, a distinct black spot at apex surrounded by creamy-white scales below and small blackish dots along termen, and flagellum of antenna with alternative creamy-white and black rings. However,
<taxonomicName id="D9B915C97B50FFDA4F04FAEFFE3E2450" authority="Park &amp; Bae, 2021" authorityName="Park &amp; Bae" authorityYear="2021" box="[266,494,1303,1329]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="107" phylum="Arthropoda" rank="species" species="alterniella" status="sp. nov.">
<emphasis id="2CCDB2587B50FFDA4F04FAEFFE3E2450" box="[266,494,1303,1329]" italics="true" pageId="5" pageNumber="107">Torodora alterniella</emphasis>
</taxonomicName>
<emphasis id="2CCDB2587B50FFDA4FFAFAEFFD9E2450" bold="true" box="[500,590,1303,1329]" pageId="5" pageNumber="107">
<taxonomicNameLabel id="37FE0F237B50FFDA4FFAFAEFFD9E2450" box="[500,590,1303,1329]" pageId="5" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can also be distinguished from its allies by the male genitalia: uncus broadened with sharply produced latero-apical corner; valva broadly dilated in distal part (cucullus); juxta with short, digitate caudal processes and a long, quadrate lobe on anterior margin medially.
</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B50FFD84EC9FA7AFD2C2300" lastPageId="7" lastPageNumber="109" pageId="5" pageNumber="107" type="description">
<paragraph id="1E066E4A7B50FFDA4EC9FA7AFAFB26F4" blockId="5.[151,1437,1014,2013]" pageId="5" pageNumber="107">
<emphasis id="2CCDB2587B50FFDA4EC9FA7AFE8924FD" bold="true" box="[199,345,1410,1436]" pageId="5" pageNumber="107">Description.</emphasis>
Adult (
<figureCitation id="868272CF7B50FFDA4FA2FA7BFDF824FC" box="[428,552,1410,1437]" captionStart="FIGURE 5" captionStartId="9.[151,250,1691,1715]" captionTargetBox="[176,1413,404,1647]" captionTargetId="figure-129@9.[151,1436,380,1667]" captionTargetPageId="9" captionText="FIGURE 5. Torodora cupriella Park, sp. nov.: A, adult, holotype; B, close-up antenna, with arrow indicated the blackish part; C, labial palpus in lateral view; D, female genitalia, gen. slide no. CIS-8081, holotype; E, close-up abdominal sternite VIII-posterior part of ductus bursae; F, close-up signum. Scale bar for genitalia: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4635955" httpUri="https://zenodo.org/record/4635955/files/figure.png" pageId="5" pageNumber="107">Figs 5A, B</figureCitation>
). Wingspan 13.0–14.0 mm.
<emphasis id="2CCDB2587B50FFDA4D6EFA7BFC4D24FC" box="[864,925,1411,1437]" italics="true" pageId="5" pageNumber="107">Head</emphasis>
: Covered with creamy-white scales anteriorly and grayish-orange posteriorly, with erect creamy-white scales laterally. Antenna longer than forewing, about 1.3 times than the length of forewing; scape elongated, creamy white on dorsal and ventral surface, dark brown anteriorly and posteriorly; flagellum with distinct creamy-white and black rings alternatively. Second segment of labial palpus thickened, slightly arched, dark brown on dorsal surface, grayish orange in basal 2/3, and creamy white speckled with brownish scales in distal 1/3 on outer surface, creamy white on inner surface evenly, with creamy-white apex; 3
<superScript id="E9CCC3027B50FFDA4EABF9A2FF632709" attach="left" box="[165,179,1626,1640]" fontSize="6" pageId="5" pageNumber="107">rd</superScript>
segment slender, slightly longer than 2
<superScript id="E9CCC3027B50FFDA4C60F9A2FDAE2709" attach="left" box="[622,638,1626,1640]" fontSize="6" pageId="5" pageNumber="107">nd</superScript>
segment, strongly upturned, blackish ventrally, creamy white laterally.
<emphasis id="2CCDB2587B50FFDA4E99F987FF3827F8" box="[151,232,1663,1689]" italics="true" pageId="5" pageNumber="107">Thorax</emphasis>
: Tegula and thorax grayish orange dorsally. Hind tibia with rough, pale grayish-orange scales above, speckled with dark-brown scales apically; 1st tarsus long, creamy white; 2
<superScript id="E9CCC3027B50FFDA4D96F95AFC7827D1" attach="left" box="[920,936,1698,1712]" fontSize="6" pageId="5" pageNumber="107">nd</superScript>
and 3
<superScript id="E9CCC3027B50FFDA4DE3F95AFC2B27D1" attach="left" box="[1005,1019,1698,1712]" fontSize="6" pageId="5" pageNumber="107">rd</superScript>
black in basal half and creamy white beyond. Forewing ground color grayish orange to yellowish brown, with an obscure white, oblique line, arising from near middle of costa and a more distinct white oblique line from beyond 3/4 of costa toward termen, terminated at veins R
<subScript id="823D6C0F7B50FFDA4F06F8D9FEF2264E" attach="left" box="[264,290,1825,1839]" fontSize="6" pageId="5" pageNumber="107">3+4</subScript>
; a black, distinct rounded spot at apex, with creamy-white scales at inner side, and smaller black dots along termen, forming a line; apex sharply produced; termen oblique, concave before middle; fringe concolorous with ground color. Hindwing slightly broader, more grayish scales than forewing; apex acute; termen slightly sinuate; fringe concolorous; venation with M
<subScript id="823D6C0F7B50FFDA4C6FF875FDB926FA" attach="left" box="[609,617,1933,1947]" fontSize="6" pageId="5" pageNumber="107">2</subScript>
close to M
<subScript id="823D6C0F7B50FFDA4CE9F875FD3F26FA" attach="left" box="[743,751,1933,1947]" fontSize="6" pageId="5" pageNumber="107">3</subScript>
+CuA
<subScript id="823D6C0F7B50FFDA4D3DF875FCEB26FA" attach="left" box="[819,827,1933,1947]" fontSize="6" pageId="5" pageNumber="107">1</subScript>
at base, M
<subScript id="823D6C0F7B50FFDA4DB8F875FC6E26FA" attach="left" box="[950,958,1933,1947]" fontSize="6" pageId="5" pageNumber="107">3</subScript>
and CuA
<subScript id="823D6C0F7B50FFDA4A24F875FBE226FA" attach="left" box="[1066,1074,1933,1947]" fontSize="6" pageId="5" pageNumber="107">1</subScript>
stalked for basal half.
</paragraph>
<paragraph id="1E066E4A7B50FFDA4EC9F867FB0326BC" blockId="5.[151,1437,1014,2013]" pageId="5" pageNumber="107">
<emphasis id="2CCDB2587B50FFDA4EC9F867FEE126D8" box="[199,305,1951,1977]" italics="true" pageId="5" pageNumber="107">Abdomen</emphasis>
(
<figureCitation id="868272CF7B50FFDA4F4CF867FE4A26D8" box="[322,410,1951,1977]" captionStart="FIGURE 2" captionStartId="4.[151,250,1784,1808]" captionTargetBox="[176,1412,657,1740]" captionTargetId="figure-163@4.[151,1436,633,1761]" captionTargetPageId="4" captionText="FIGURE 2. Torodora spathiana Park, sp. nov.: A, adult, holotype; B, labial palpus, in lateral view; C, hind tibia; D, male genitalia, gen. slide no. CIS-6063; E, aedeagus; F, abdominal segments VIII; G, genitalia of T. paratidosa (Wu), gen. slide no. CIS-6689 from Cambodia. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4635945" httpUri="https://zenodo.org/record/4635945/files/figure.png" pageId="5" pageNumber="107">Fig. 2F</figureCitation>
): Spinous zones on tergite in male narrowly developed along posterior margin with strong spines; sternite VIII more or less trifurcated posteriorly, with large, triangular median protrusion.
</paragraph>
<caption id="4AC63EC27B53FFD94E99F8D8FF3C26C2" ID-DOI="http://doi.org/10.5281/zenodo.4635947" ID-Zenodo-Dep="4635947" httpUri="https://zenodo.org/record/4635947/files/figure.png" pageId="6" pageNumber="108" startId="6.[151,250,1824,1848]" targetBox="[175,1410,205,1764]" targetPageId="6">
<paragraph id="1E066E4A7B53FFD94E99F8D8FF3C26C2" blockId="6.[151,1437,1824,1956]" pageId="6" pageNumber="108">
<emphasis id="2CCDB2587B53FFD94E99F8D8FEC32659" bold="true" box="[151,275,1824,1848]" pageId="6" pageNumber="108">FIGURE 3.</emphasis>
<taxonomicName id="D9B915C97B53FFD94F14F8D8FDF02659" authority="Park" authorityName="Park &amp; Bae" authorityYear="2021" box="[282,544,1824,1848]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="108" phylum="Arthropoda" rank="species" species="alterniella" status="sp. nov.">
<emphasis id="2CCDB2587B53FFD94F14F8D8FE392659" box="[282,489,1824,1848]" italics="true" pageId="6" pageNumber="108">Torodora alterniella</emphasis>
Park
</taxonomicName>
,
<emphasis id="2CCDB2587B53FFD94C24F8D8FDAB2659" bold="true" box="[554,635,1824,1848]" pageId="6" pageNumber="108">
<taxonomicNameLabel id="37FE0F237B53FFD94C24F8D8FDAB2659" box="[554,635,1824,1848]" pageId="6" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, adult, paratype from Laos; B, labial palpus and antenna; C, male genitalia, gen. slide no. CIS-8097, paratype from Laos; D,
<emphasis id="2CCDB2587B53FFD94C71F8BCFD7D263D" box="[639,685,1860,1884]" italics="true" pageId="6" pageNumber="108">ditto</emphasis>
, aedeagus; E, aedeagus of holotype, gen. slide no. CIS-6097; F, abdominal segments V–VIII, holotype; G, female genitalia., gen. slide no. CIS-8082 from Cambodia. Scale bar for genitalia and aedeagus: 1.0 mm.
</paragraph>
</caption>
<paragraph id="1E066E4A7B52FFD84EC9FF6FFD5A20E8" blockId="7.[151,1437,151,861]" pageId="7" pageNumber="109">
<emphasis id="2CCDB2587B52FFD84EC9FF6FFEBD21D0" box="[199,365,151,177]" italics="true" pageId="7" pageNumber="109">Male genitalia</emphasis>
(
<figureCitation id="868272CF7B52FFD84F72FF6FFDD421D0" box="[380,516,151,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1824,1848]" captionTargetBox="[175,1410,205,1764]" captionTargetId="figure-19@6.[151,1436,181,1800]" captionTargetPageId="6" captionText="FIGURE 3. Torodora alterniella Park, sp. nov.: A, adult, paratype from Laos; B, labial palpus and antenna; C, male genitalia, gen. slide no. CIS-8097, paratype from Laos; D, ditto, aedeagus; E, aedeagus of holotype, gen. slide no. CIS-6097; F, abdominal segments V–VIII, holotype; G, female genitalia., gen. slide no. CIS-8082 from Cambodia. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4635947" httpUri="https://zenodo.org/record/4635947/files/figure.png" pageId="7" pageNumber="109">Figs. 3C–D</figureCitation>
): Uncus broadened distally, sharply produced at lateral apices. Gnathos relatively small. Tegumen weakly sclerotized, emarginated in inverted V-shaped on anterior margin. Valva elongated, expanded in basal 1/3 anteriorly; costa gently concave; cucullus dilated, densely setose; sacculus broad at base, narrowly terminated before cucullus. Juxta deeply concave on caudal margin; caudal processes short, digitate with round apex; median protrusion on anterior margin narrowly elongate-quadrate. Aedeagus stout basally, broader than basal part of valva, strongly bent medially, narrowed toward apex; narrowly produced at apex; cornuti consisting of two, large sacs with numerous needle-like spines.
</paragraph>
<paragraph id="1E066E4A7B52FFD84EC9FE6BFD2C2300" blockId="7.[151,1437,151,861]" pageId="7" pageNumber="109">
<emphasis id="2CCDB2587B52FFD84EC9FE6BFE5A20CC" box="[199,394,403,429]" italics="true" pageId="7" pageNumber="109">Female genitalia</emphasis>
(
<figureCitation id="868272CF7B52FFD84F93FE6BFE2620CC" box="[413,502,403,429]" captionStart="FIGURE 3" captionStartId="6.[151,250,1824,1848]" captionTargetBox="[175,1410,205,1764]" captionTargetId="figure-19@6.[151,1436,181,1800]" captionTargetPageId="6" captionText="FIGURE 3. Torodora alterniella Park, sp. nov.: A, adult, paratype from Laos; B, labial palpus and antenna; C, male genitalia, gen. slide no. CIS-8097, paratype from Laos; D, ditto, aedeagus; E, aedeagus of holotype, gen. slide no. CIS-6097; F, abdominal segments V–VIII, holotype; G, female genitalia., gen. slide no. CIS-8082 from Cambodia. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4635947" httpUri="https://zenodo.org/record/4635947/files/figure.png" pageId="7" pageNumber="109">Fig. 3F</figureCitation>
). Abdominal sternite VIII deeply emarginated in U-shape. Apophyses anteriores short, about 1/3 the length of apophyses posteriores. Antrum membraneous, funnel-shaped. Ductus bursae with extremely narrow neck beyond antrum, then broadened, about half width of corpus bursae, as long as corpus bursae; ductus seminalis broad as wide as half of ductus bursae, arising from distal part of ductus bursae. Corpus bursae, large, ovate; signum plate vertically elongate-quadrate, slightly emarginated medially on anterior and posterior margin, with numerous, dense conic spines on surface.
</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B52FFD84EC9FD92FD2923E4" box="[199,761,618,645]" pageId="7" pageNumber="109" type="distribution">
<paragraph id="1E066E4A7B52FFD84EC9FD92FD2923E4" blockId="7.[151,1437,151,861]" box="[199,761,618,645]" pageId="7" pageNumber="109">
<emphasis id="2CCDB2587B52FFD84EC9FD92FEB123E5" bold="true" box="[199,353,618,644]" pageId="7" pageNumber="109">Distribution.</emphasis>
<collectingCountry id="66AE2EDA7B52FFD84F69FD93FE0A23E4" box="[359,474,619,645]" name="Cambodia" pageId="7" pageNumber="109">Cambodia</collectingCountry>
(
<collectingRegion id="DC7DA0A87B52FFD84FE4FD93FDCD23E4" box="[490,541,619,645]" country="Cambodia" name="Kep" pageId="7" pageNumber="109">Kep</collectingRegion>
,
<collectingRegion id="DC7DA0A87B52FFD84C26FD93FD3D23E4" box="[552,749,619,645]" country="Cambodia" name="Mondolkiri" pageId="7" pageNumber="109">Mondulkiri Prov.</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B52FFD84EC9FD77FC9423AC" pageId="7" pageNumber="109" type="etymology">
<paragraph id="1E066E4A7B52FFD84EC9FD77FC9423AC" blockId="7.[151,1437,151,861]" pageId="7" pageNumber="109">
<emphasis id="2CCDB2587B52FFD84EC9FD77FE9E23C8" bold="true" box="[199,334,655,681]" pageId="7" pageNumber="109">Etymology.</emphasis>
The species name is derived from the Latin,
<emphasis id="2CCDB2587B52FFD84D54FD77FC6623C8" box="[858,950,655,681]" italics="true" pageId="7" pageNumber="109">alternus</emphasis>
(= alternative), with a Latin suffix, -
<emphasis id="2CCDB2587B52FFD84B6EFD77FA4C23C8" box="[1376,1436,655,681]" italics="true" pageId="7" pageNumber="109">ellus,</emphasis>
referring to the alternative coloration of rings on the antenna.
</paragraph>
</subSubSection>
<subSubSection id="56A33DC17B52FFD84EC9FD2FFB96223C" pageId="7" pageNumber="109" type="discussion">
<paragraph id="1E066E4A7B52FFD84EC9FD2FFB96223C" blockId="7.[151,1437,151,861]" pageId="7" pageNumber="109">
<emphasis id="2CCDB2587B52FFD84EC9FD2FFEEB2390" bold="true" box="[199,315,727,753]" pageId="7" pageNumber="109">Remarks.</emphasis>
Most of the known species of the genus
<taxonomicName id="D9B915C97B52FFD84D05FD2FFC092390" authority="Meyrick" authorityName="Meyrick" authorityYear="1894" box="[779,985,727,753]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="2CCDB2587B52FFD84D05FD2FFCA32390" box="[779,883,727,753]" italics="true" pageId="7" pageNumber="109">Torodora</emphasis>
Meyrick
</taxonomicName>
have an elongated, slender uncus in the male genitalia, but the uncus of this new species and the following species are widely expanded distally. The other examples with this kind of the uncus are also found in
<taxonomicName id="D9B915C97B52FFD84CF5FCE7FB942258" authority="Gozmany, 1978" authorityName="Gozmany" authorityYear="1978" box="[763,1092,799,825]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="109" phylum="Arthropoda" rank="species" species="aenoptera">
<emphasis id="2CCDB2587B52FFD84CF5FCE7FC592259" box="[763,905,799,824]" italics="true" pageId="7" pageNumber="109">T. aenoptera</emphasis>
<bibRefCitation id="7A2813BB7B52FFD84D81FCE7FB942258" author="Gozmany, L." box="[911,1092,799,825]" pageId="7" pageNumber="109" pagination="306" refId="ref6101" refString="Gozmany, L. (1978) Lecithoceridae, In: Amsel, H. G., Gregor, F., Reisser, H. (Eds.), Microlepidoptera Palaearctica. Vol. 5., Georg Fromme &amp; Co., Wien, pp. 306." type="book chapter" year="1978">Gozmány, 1978</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="D9B915C97B52FFD84A41FCE7FA482258" authority="Gozmany, 1978" authorityName="Gozmany" authorityYear="1978" box="[1103,1432,799,825]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="109" phylum="Arthropoda" rank="species" species="flavescens">
<emphasis id="2CCDB2587B52FFD84A41FCE7FB0D2258" box="[1103,1245,799,825]" italics="true" pageId="7" pageNumber="109">T. flavescens</emphasis>
<bibRefCitation id="7A2813BB7B52FFD84AEAFCE7FA482258" author="Gozmany, L." box="[1252,1432,799,825]" pageId="7" pageNumber="109" pagination="306" refId="ref6101" refString="Gozmany, L. (1978) Lecithoceridae, In: Amsel, H. G., Gregor, F., Reisser, H. (Eds.), Microlepidoptera Palaearctica. Vol. 5., Georg Fromme &amp; Co., Wien, pp. 306." type="book chapter" year="1978">Gozmány, 1978</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="D9B915C97B52FFD84EC9FCBBFE74223C" authority="Wu, 1997" authorityName="Wu" authorityYear="1997" box="[199,420,835,861]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="109" phylum="Arthropoda" rank="species" species="sirtalis">
<emphasis id="2CCDB2587B52FFD84EC9FCBBFEE0223C" box="[199,304,835,861]" italics="true" pageId="7" pageNumber="109">T. sirtalis</emphasis>
Wu, 1997
</taxonomicName>
from
<collectingCountry id="66AE2EDA7B52FFD84FE6FCBBFDFE223C" box="[488,558,835,861]" name="China" pageId="7" pageNumber="109">China</collectingCountry>
, and
<taxonomicName id="D9B915C97B52FFD84C66FCBBFC48223C" authority="Park, 2006" authorityName="Park" authorityYear="2006" box="[616,920,835,861]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="109" phylum="Arthropoda" rank="species" species="bachmaensis">
<emphasis id="2CCDB2587B52FFD84C66FCBBFCC5223C" box="[616,789,835,861]" italics="true" pageId="7" pageNumber="109">T. bachmaensis</emphasis>
Park, 2006
</taxonomicName>
, from
<collectingCountry id="66AE2EDA7B52FFD84DEEFCBBFB93223C" box="[992,1091,835,861]" name="Vietnam" pageId="7" pageNumber="109">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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