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<document id="5ACCC00B482CA8A6AD2AC59FE0ED4072" ID-CLB-Dataset="306405" ID-DOI="10.3897/subtbiol.50.139914" ID-GBIF-Dataset="e8f46f87-d198-4799-8175-51218575a895" ID-ZooBank="98DE6D7C-2352-480A-8B85-24B6B14F8279" ID-publisher-id="139914" URI-arpha="F4223F22-CF88-5206-89F3-7A759E9CA35E" URI-zoobank="http://zoobank.org/98DE6D7C-2352-480A-8B85-24B6B14F8279" article-type="research-article" checkinTime="1734017366078" checkinUser="pensoft" docAuthor="López-Orozco, Carlos Mario, Borja-Arrieta, Ricardo, Carpio-Díaz, Yesenia Margarita, Gallão, Jonas Eduardo, Campos-Filho, Ivanklin Soares &amp; Bichuette, Maria Elina" docDate="2024" docId="95728AF8FF5C5659B36FA15236D057B6" docLanguage="en" docName="Subterranean Biology 50: 79-104" docOrigin="Subterranean Biology 50" docSource="https://subtbiol.pensoft.net/article/139914/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Xangoniscus jonasi Lopez-Orozco, Bichuette &amp; Campos-Filho 2024, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="104" masterDocId="F4223F22CF88520689F37A759E9CA35E" masterDocTitle="Describing to preserve – three new species of Xangoniscus (Oniscidea, Styloniscidae) of unprotected caves in dry areas from Bahia state, northeastern Brazil" masterLastPageNumber="104" masterPageNumber="79" pageNumber="79" updateTime="1734018335538" updateUser="ExternalLinkService">
<mods:mods id="512B744F8DA2E0D4D18DF39568AEC682" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="FF16E95E955E10401793C375DFC1E371">Describing to preserve – three new species of Xangoniscus (Oniscidea, Styloniscidae) of unprotected caves in dry areas from Bahia state, northeastern Brazil</mods:title>
</mods:titleInfo>
<mods:name id="CC917A942B172CE68669011DDAC6BD5F" type="personal">
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<mods:namePart id="723B50375D17F3BAD5CFBEF24443E074">López-Orozco, Carlos Mario</mods:namePart>
<mods:nameIdentifier id="FC903CE9B04FBE97E51E3345AB8A8BF8" type="ORCID">0000-0002-3251-7739</mods:nameIdentifier>
<mods:affiliation id="ABCC61FB10FB35E6C017096AFECE6AF3">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos, São Carlos, São Paulo, Brazil</mods:affiliation>
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<mods:name id="7B0392EF1B0319C388D5513FF1879B0B" type="personal">
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<mods:roleTerm id="E530C75CE8CC76503ED46889B44646CB">Author</mods:roleTerm>
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<mods:namePart id="E54A45107581193730627ABB848D0F24">Borja-Arrieta, Ricardo</mods:namePart>
<mods:nameIdentifier id="A6F262163EC1BEEA638DF51CFB99A35E" type="ORCID">0000-0002-5064-5080</mods:nameIdentifier>
<mods:affiliation id="0D251994F0A9C4182C8E4959F50E320E">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos, São Carlos, São Paulo, Brazil</mods:affiliation>
</mods:name>
<mods:name id="2379C88431986AC4EA6CB99A0BEBB5C0" type="personal">
<mods:role id="64F5ACD635F748B3E4F8672909331F0A">
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</mods:role>
<mods:namePart id="83AEF48EC92F86EE0B634E575654D65E">Carpio-Díaz, Yesenia Margarita</mods:namePart>
<mods:nameIdentifier id="CD4E11E19E7F8ADF4BC786E0797EA00E" type="ORCID">0000-0001-5116-9736</mods:nameIdentifier>
<mods:affiliation id="B1E258BD916E015184E80242A97EE9BB">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos, São Carlos, São Paulo, Brazil</mods:affiliation>
</mods:name>
<mods:name id="10590506887ED4A8DA6A608173F20E9E" type="personal">
<mods:role id="EE67C089E2CFB17AD91CF4F9F3A674C9">
<mods:roleTerm id="4C1C6E3E4DCA5FF26F0712F909713F8A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="94DECBFD7D282E22EE6383D6E8367B67">Gallão, Jonas Eduardo</mods:namePart>
<mods:nameIdentifier id="B494183ABF3E76AF66EB4C2853B4D44B" type="ORCID">0000-0002-5268-7946</mods:nameIdentifier>
<mods:affiliation id="0B423BADECD38B96C87C1C488ADFA6B7">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos, São Carlos, São Paulo, Brazil &amp; Instituto Brasileiro de Estudos Subterrâneos, São Paulo, Brazil &amp; INPP – Instituto Nacional de Pesquisas do Pantanal, Cuiabá, Mato Grosso, Brazil</mods:affiliation>
</mods:name>
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<mods:roleTerm id="FEB7C47FE6AD9DF0DE7E549B6A516A6F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DD2CE3C2179092C5DB99AD8E812957E5">Campos-Filho, Ivanklin Soares</mods:namePart>
<mods:nameIdentifier id="FAEFDED05A7B667D93EDDB4410806D74" type="ORCID">0000-0001-6139-8241</mods:nameIdentifier>
<mods:affiliation id="57573B27F24EE4DE6498627504F17270">Department of Biological Sciences, University of Cyprus, Lefkosia (Nicosia), Cyprus</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="5C4A34241407437DC632B12FE82E77E0">Bichuette, Maria Elina</mods:namePart>
<mods:nameIdentifier id="22C680B1ACF70815A3283E8B8EA06A29" type="ORCID">0000-0002-9515-4832</mods:nameIdentifier>
<mods:affiliation id="D82DE3B2AB91A15391254B04FBE33818">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos, São Carlos, São Paulo, Brazil &amp; Instituto Brasileiro de Estudos Subterrâneos, São Paulo, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="9A18A3C5E4446B7C3FC21A0E2796CFDE">text</mods:typeOfResource>
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<mods:titleInfo id="79F65AF9A7FEE491F996790A2141C6EA">
<mods:title id="52AF6A5ECCF7B57C68A7B369FC3675F0">Subterranean Biology</mods:title>
</mods:titleInfo>
<mods:part id="EB03C2ADAFA68914008C22B688B3EF1F">
<mods:date id="AEC3C611DADFBA803A67C928ECF5C0DC">2024</mods:date>
<mods:detail id="E8E8A9CE857E27D7665BF15AD1528B17" type="pubDate">
<mods:number id="B2EB56DBF351A730E51E09B59293F1D2">2024-12-12</mods:number>
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<mods:detail id="7EB3DBB956053705381382E6CA8B1DA6" type="volume">
<mods:number id="64C6E8E4E8DA8746C2351F0B9B40C88D">50</mods:number>
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<mods:start id="F0DDEBFE0249CA637EC4235E934FB23F">79</mods:start>
<mods:end id="75DC316B8B68C8ED4C7F06F79967E8A0">104</mods:end>
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<mods:classification id="59914AD3D585B03D0DC11AB609589997">journal article</mods:classification>
<mods:identifier id="EE5FAC4B08151FA2F9E396382EA31EB1" type="CLB-Dataset">306405</mods:identifier>
<mods:identifier id="F2F6BE04BA706D5DC26042A937A9F37C" type="DOI">10.3897/subtbiol.50.139914</mods:identifier>
<mods:identifier id="2E0C369DE02262667A93F58D05CF4E6C" type="GBIF-Dataset">e8f46f87-d198-4799-8175-51218575a895</mods:identifier>
<mods:identifier id="D0A75C2A02E2315F07AB093F300EC199" type="ZooBank">98DE6D7C-2352-480A-8B85-24B6B14F8279</mods:identifier>
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<treatment id="95728AF8FF5C5659B36FA15236D057B6" ID-DOI="http://doi.org/10.5281/zenodo.14418459" ID-Zenodo-Dep="14418459" ID-arpha="95728AF8-FF5C-5659-B36F-A15236D057B6" ID-zoobank="https://zoobank.org/018645EB-FC2A-4CEA-B1B3-9EC60D23F2BA" LSID="urn:lsid:plazi:treatment:95728AF8FF5C5659B36FA15236D057B6" LSID-ZBK="urn:lsid:zoobank.org:act:018645EB-FC2A-4CEA-B1B3-9EC60D23F2BA" httpUri="http://treatment.plazi.org/id/95728AF8FF5C5659B36FA15236D057B6" scope_family="Styloniscidae" scope_subOrder="Oniscidea">
<subSubSection id="5E725C5BA02473A1C5EBF194BAB6F736" type="nomenclature">
<paragraph id="84F37108DCB1DA1B3D0B1D7E9144612F">
<taxonomicName id="4D48ABAF4A21691F0FD7BAFACC3F2B65" authority="Lopez-Orozco, Bichuette &amp; Campos-Filho" authorityName="Lopez-Orozco, Bichuette &amp; Campos-Filho" authorityYear="2024" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="jonasi" status="sp. nov.">
<emphasis id="D68F7F728F7877EEA2072B70C7DC7F61" italics="true">Xangoniscus jonasi</emphasis>
López-Orozco, Bichuette &amp; Campos-Filho
</taxonomicName>
<taxonomicNameLabel id="54AA55DA7644035DD84EFE55838580B5" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D4DF184F8D0FF9BC42F7881789F18BD9" type="description">
<paragraph id="08CAF6FF817E19968963038459FC0CC6">
<figureCitation id="CC41608EE0EA173C9412CC54954CCE5C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the study area and distribution of the Xangoniscus species." httpUri="https://binary.pensoft.net/fig/1197443">Figs 1</figureCitation>
,
<figureCitation id="0AD724520553B2238DF0479429698702" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B habitus, lateral view C cephalon and pereonite 1, frontal view D cephalon and pereonites 1–2, lateral view E pereonite 7, pleon and telson, dorsal view. Scale bar: 1 mm." httpUri="https://binary.pensoft.net/fig/1197449">7</figureCitation>
,
<figureCitation id="1CFE3BE4D0AA02C51321EC893A562174" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8</figureCitation>
,
<figureCitation id="00C783A9BA7C5A10AD2975C471757F6B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A pereopod 1 B pereopod 2 C pereopod 3 D pereopod 4 E pereopod 5 F pereopod 6 G pereopod 7." httpUri="https://binary.pensoft.net/fig/1197451">9</figureCitation>
,
<figureCitation id="A272240FC87353A51D58D7048309AD21" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A pleopod 1 B genital papilla C pleopod 2 D pleopod 3 exopod E pleopod 4 exopod F pleopod 5 exopod." httpUri="https://binary.pensoft.net/fig/1197452">10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EURAE" type="type material">
<paragraph id="25D6C708051A9FF5C5DFDE2AB220D065">
<heading id="A25148AFEF8AD4D305F78BB6A1B3EB88" reason="title">Type material.</heading>
</paragraph>
<paragraph id="328DAAF3E14E8267297BB48B409971A9">
<materialsCitation id="2BC3EBF8DCDDD5E3D5C137F896E70684" collectingDate="2023-10-24" collectionCode="LES" collectorName="JE Gallao." latitude="-13.510573" location="Gruna da Serra Solta cave" longLatPrecision="1" longitude="-43.75207" municipality="Carinhanha" specimenCode="LES 0030097" specimenCount="1" specimenCount-male="1" stateProvince="Bahia" typeStatus="holotype">
<emphasis id="B3766DF502327008D16799181F940293" bold="true" italics="true">
<typeStatus id="A8794D9CA28AE73E7CE30DDE3E677375" type="holotype">Holotype</typeStatus>
</emphasis>
•
<specimenCount id="8CA66BD708B900938C7151326A2D40DF" count="1" type="male">1 male</specimenCount>
(parts in slides) (
<specimenCode id="B8C8A750185C066F0B566E3D985D342C" collectionCode="LES" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13487" lsid="urn:lsid:biocol.org:col:13487" name="Leeds Museums and Galleries" type="Museum">
<abbrev content-type="institution" id="ABBRID0E6RAE" xlink_title="Laboratório de Estudos Subterrâneos">LES</abbrev>
0030097
</specimenCode>
),
<collectingRegion id="F2800166553458A02B419AA2D168BEF2" country="Brazil" name="Bahia">Bahia</collectingRegion>
,
<collectingMunicipality id="E963B1E4AE89D60278059C74EDCEB31A">Carinhanha</collectingMunicipality>
,
<location id="546087FD24FCAA3C1034E47F7AB8C41C" LSID="urn:lsid:plazi:treatment:95728AF8FF5C5659B36FA15236D057B6:546087FD24FCAA3C1034E47F7AB8C41C" latitude="-13.510573" longLatPrecision="1" longitude="-43.75207" municipality="Carinhanha" name="Gruna da Serra Solta cave" stateProvince="Bahia">Gruna da Serra Solta cave</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHSAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-43.752070,-13.510573]}">
<geoCoordinate id="87893A9911B5EC1BD95590BF0D26EBF6" degrees="13.510573" direction="south" orientation="latitude" precision="1" value="-13.510573">- 13.510573</geoCoordinate>
,
<geoCoordinate id="474E0B83A48AA2C8942DD610094B7510" degrees="43.75207" direction="west" orientation="longitude" precision="1" value="-43.75207">- 43.75207</geoCoordinate>
</named-content>
,
<date id="43634B62246E72A696E5418B59E2E169" value="2023-10-24">
<collectingDate id="37F3C7CC6270A985C4A7929FE02A79EB" value="2023-10-24">24 October 2023</collectingDate>
</date>
, leg.
<collectorName id="E027A8A12985FD6D521AF5286AC169DB">JE Gallão.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMSAE" type="description">
<paragraph id="1C1FA80D3D7265BBB4448A1F30721FF9">
<heading id="1238C7FB92D76E5171EE477786594C68" reason="title">Description.</heading>
</paragraph>
<paragraph id="6895F4AC758181FD1967C72FC01E1B4F">
Maximum body length: male
<quantity id="A1527403836DA586A618E4B6FC6AD37E" metricMagnitude="-3" metricUnit="m" metricValue="6.5" unit="mm" value="6.5">6.5 mm</quantity>
. Body outline as in Figs
<figureCitation id="3AB12076FEEB603FBE1ECA4C6A2463AC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B habitus, lateral view C cephalon and pereonite 1, frontal view D cephalon and pereonites 1–2, lateral view E pereonite 7, pleon and telson, dorsal view. Scale bar: 1 mm." httpUri="https://binary.pensoft.net/fig/1197449">7 A, B</figureCitation>
,
<figureCitation id="BA9176BAB3C0CA4AA9238E40CFFB5F83" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 A</figureCitation>
. Colourless (Fig.
<figureCitation id="7472ABB80D2285A569CFA65592030FD9" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B habitus, lateral view C cephalon and pereonite 1, frontal view D cephalon and pereonites 1–2, lateral view E pereonite 7, pleon and telson, dorsal view. Scale bar: 1 mm." httpUri="https://binary.pensoft.net/fig/1197449">7</figureCitation>
). Dorsal surface smooth, with scattered fringed scale setae (Figs
<figureCitation id="80779D4B2D0410E0E5E166D4371CD7CD" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B habitus, lateral view C cephalon and pereonite 1, frontal view D cephalon and pereonites 1–2, lateral view E pereonite 7, pleon and telson, dorsal view. Scale bar: 1 mm." httpUri="https://binary.pensoft.net/fig/1197449">7 A – E</figureCitation>
,
<figureCitation id="434A627BCE93370E4DBFB5C5CC2057C7" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 B</figureCitation>
). Cephalon (Figs
<figureCitation id="D2AB24D516D459071FAD13820149AA0F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B habitus, lateral view C cephalon and pereonite 1, frontal view D cephalon and pereonites 1–2, lateral view E pereonite 7, pleon and telson, dorsal view. Scale bar: 1 mm." httpUri="https://binary.pensoft.net/fig/1197449">7 C, D</figureCitation>
,
<figureCitation id="C4D0AB2ED904634F5C2B5D67801B0D3A" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 C</figureCitation>
) with large quadrangular antennary lobes; vertex with slightly lateral depression to fit antennae when extended backwards, profrons with V-shaped suprantennal line, not surpassing antennule insertion; eyes absent. Pereonite 1 epimera with distal corners not developed frontwards, posterior corners right-angled; pereonites 2–7 epimera gradually directed backwards, posterior corners progressively more acute; pleon narrower than pereon; pleonites 3–5 epimera posterior point not developed (Figs
<figureCitation id="445399D322B38F4D1F09A14D437CC138" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B habitus, lateral view C cephalon and pereonite 1, frontal view D cephalon and pereonites 1–2, lateral view E pereonite 7, pleon and telson, dorsal view. Scale bar: 1 mm." httpUri="https://binary.pensoft.net/fig/1197449">7 A, B, D</figureCitation>
,
<figureCitation id="49CE774871278B0BD1E0E5DBD19B4F0B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 A</figureCitation>
). Telson with concave sides and rounded apex (Fig.
<figureCitation id="D4D4CF4313DEF4436013D2C414B7C93E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 D</figureCitation>
). Antennula (Fig.
<figureCitation id="9DC5317727ED8E6A09DC1501A47A2E20" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 E</figureCitation>
) composed of three articles, proximal and distal articles subequal in length, second article short, distal article with eight very long aesthetascs. Antenna (Fig.
<figureCitation id="704735AFC35D98D275A3B5A212938333" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 F</figureCitation>
) short, not surpassing pereonite 3 when extended backwards; flagellum of equal length of fifth article of peduncle, composed of three articles subequal in length. Left mandible (Fig.
<figureCitation id="8BCD58CD3C888D7C02B7E94138C2A873" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 G</figureCitation>
) with two penicils; right mandible (Fig.
<figureCitation id="8AC1D12E980E58E299DC6CA612D79526" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 H</figureCitation>
) with lacinia mobilis and one penicil. Maxillula (Fig.
<figureCitation id="366791680D7A6403B647D4CCFD9D1E89" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 I</figureCitation>
) inner endite with three penicils; outer endite with 5 + 5 teeth, apically simple, and two plumose stalks. Maxilla (Fig.
<figureCitation id="0C623BC8A9EE3705CDF2D35A444AE57C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 J</figureCitation>
) with setose and bilobate apex, outer lobe smaller. Maxilliped (Fig.
<figureCitation id="D30701A98AB13049710AAA3266D1B2D1" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 K</figureCitation>
) basis enlarged on distal portion bearing fringe of fine setae; first article of palp with two tiny setae, distal articles with three tufts of setae; endite rectangular, outer and medial margins setose, apically with two triangular teeth and large rounded penicil. Uropod (Fig.
<figureCitation id="EFFF2F677E1956FCC8882B7E5E6C6FE5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A habitus, dorsal view B dorsal scale-seta C cephalon, frontal view D telson and uropod E antennula F antenna G left mandible H right mandible I maxillula J maxilla K maxilliped, arrow illustrating the endite in caudal view." httpUri="https://binary.pensoft.net/fig/1197450">8 D</figureCitation>
) branches short, endopod inserted proximally, exopod and endopod equal in length. Pereopods 1–7 (Fig.
<figureCitation id="D79476CFD7E339B0CB627480AD80EFC3" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A pereopod 1 B pereopod 2 C pereopod 3 D pereopod 4 E pereopod 5 F pereopod 6 G pereopod 7." httpUri="https://binary.pensoft.net/fig/1197451">9 A – G</figureCitation>
) gradually elongated, with merus, carpus, and propodus bearing sparse setae; dactylus of one claw bearing many setae on inner and outer margins.
</paragraph>
<paragraph id="8FD413141ECF3467075C2D0EEDB8D346">
<emphasis id="959207396AA9869BC8808D42D615A128" bold="true">Male.</emphasis>
Pereopod 1 (Fig.
<figureCitation id="C8BF06DACA213051DD301899D30EF098" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A pereopod 1 B pereopod 2 C pereopod 3 D pereopod 4 E pereopod 5 F pereopod 6 G pereopod 7." httpUri="https://binary.pensoft.net/fig/1197451">9 A</figureCitation>
) carpus with large antennal grooming brush. Pereopod 2–4 (Fig.
<figureCitation id="BD3100DC5E2D8CF115F1A9FEAC814879" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A pereopod 1 B pereopod 2 C pereopod 3 D pereopod 4 E pereopod 5 F pereopod 6 G pereopod 7." httpUri="https://binary.pensoft.net/fig/1197451">9 B – D</figureCitation>
) merus with scales on sternal margin. Pereopods 5 and 7 (Fig.
<figureCitation id="67604B3DD56F338DA814AEC4A616F807" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A pereopod 1 B pereopod 2 C pereopod 3 D pereopod 4 E pereopod 5 F pereopod 6 G pereopod 7." httpUri="https://binary.pensoft.net/fig/1197451">9 E, G</figureCitation>
) without distinct modifications. Pereopod 6 (Fig.
<figureCitation id="1D4C71EAEED75292CD18CB675A89A9DB" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A pereopod 1 B pereopod 2 C pereopod 3 D pereopod 4 E pereopod 5 F pereopod 6 G pereopod 7." httpUri="https://binary.pensoft.net/fig/1197451">9 F</figureCitation>
) carpus slightly grooved on sternal margin, propodus slightly enlarged on median portion. Genital papilla (Fig.
<figureCitation id="2B1EA932A0164BFA1E313305400A5169" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A pleopod 1 B genital papilla C pleopod 2 D pleopod 3 exopod E pleopod 4 exopod F pleopod 5 exopod." httpUri="https://binary.pensoft.net/fig/1197452">10 B</figureCitation>
) lanceolate. Pleopod 1 (Fig.
<figureCitation id="E45A4B8F959F607EFE50E15661C28F58" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A pleopod 1 B genital papilla C pleopod 2 D pleopod 3 exopod E pleopod 4 exopod F pleopod 5 exopod." httpUri="https://binary.pensoft.net/fig/1197452">10 A</figureCitation>
) exopod trapezoidal, longer than wide, distal margin slightly convex, inner and outer margins bearing fringe of fine setae; endopod longer than exopod, with narrow basal article and flagelliform distal article; basipod distal margin elongate, longer than exopod, with semicircular apex, with fine and long setae. Pleopod 2 (Fig.
<figureCitation id="09C317DF5729B46F23162F33607D55F6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A pleopod 1 B genital papilla C pleopod 2 D pleopod 3 exopod E pleopod 4 exopod F pleopod 5 exopod." httpUri="https://binary.pensoft.net/fig/1197452">10 C</figureCitation>
) exopod trapezoidal, distal margin slightly convex bearing three setae; endopod of two articles, distal article about three times as long as proximal, with distal margin subquadrangular and apex rounded, with transversal process in V-like on apex in ventral view. Pleopod 3 (Fig.
<figureCitation id="B370DEC9AD4E6B8038E7B09BDEAD3B22" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A pleopod 1 B genital papilla C pleopod 2 D pleopod 3 exopod E pleopod 4 exopod F pleopod 5 exopod." httpUri="https://binary.pensoft.net/fig/1197452">10 D</figureCitation>
) exopod triangular, longer than wide, covering pleopods 1 and 2, fringed with short setae. Pleopod 4 and 5 (Fig.
<figureCitation id="BEDCE9C0214A0E9E15C6BEB2B9393522" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Xangoniscus jonasi López-Orozco, Bichuette &amp; Campos-Filho, sp. nov. Male A pleopod 1 B genital papilla C pleopod 2 D pleopod 3 exopod E pleopod 4 exopod F pleopod 5 exopod." httpUri="https://binary.pensoft.net/fig/1197452">10 E, F</figureCitation>
) exopods trapezoidal, wider than long, with margins bearing several short setae.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGWAE" type="etymology">
<paragraph id="D81D376E1837EC48CCA4A65B2ED289AB">
<heading id="355C082D42829C5DEB6128CBF729AB72" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="A96EBEE2B91AE095580BA49A9AB9A7D2">
The new species is named after Jonas Kahnwald, the protagonist of the “ Dark ” series, who explores caves to travel through time and space. The epithet
<taxonomicName id="E492BE8B6D4E030F8B7ECEE16601051A" authorityName="Lopez-Orozco, Bichuette &amp; Campos-Filho" authorityYear="2024" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="jonasi">
“
<emphasis id="AB16AF6CD4ADD950BD31A3A445DBF4F0" italics="true">jonasi</emphasis>
</taxonomicName>
” is a tribute to the isolation and sense of separation from time and space that caves represent, evoking the central theme of “ Dark ”. The species reflects extreme adaptation to dark depths, just as Jonas adapts to the complexities and paradoxes of time.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWWAE" type="distribution">
<paragraph id="367E2E4330636878EDE0C11B164A2C67">
<heading id="EC9A1B70C02780BAE77992E718218062" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="8AE6C38E3FC4CF9F756E9B1E283C35F5">
Presently known only from Gruna da Serra Solta cave in the Serra do Ramalho karst area, state of
<collectingRegion id="93606B97B21DF514790B14A567B77038" country="Brazil" name="Bahia">Bahia</collectingRegion>
, northeastern
<collectingCountry id="FD5D197412FB898A146BAD5AD8A7D66C" name="Brazil">Brazil</collectingCountry>
(Fig.
<figureCitation id="681D93D65A39FDC254560D8819A26B68" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the study area and distribution of the Xangoniscus species." httpUri="https://binary.pensoft.net/fig/1197443">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAXAE" type="remarks">
<paragraph id="C1839A560A3F7B9E6B58057E2ED6B121">
<heading id="43DF747CF6424B12CE7F8F71D6568483" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="74438A709FFA6CDA8BAD224B9A91ABB1">
<taxonomicName id="3A7FE67C4DB5A3E9BAF5FB7F22C19A80" authorityName="López-Orozco &amp; Borja-Arrieta &amp; Carpio-Díaz &amp; Gallão &amp; Campos-Filho &amp; Bichuette" authorityYear="2024" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="jonasi" status="sp. nov.">
<emphasis id="406791BB10F64A337313EA43915C2ABE" italics="true">Xangoniscus jonasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5661538D3FB46C0E40EBDF446276D29" rank="species">sp. nov.</taxonomicNameLabel>
differs from all other species of the genus in having the antennula with eight long aesthetascs (
<emphasis id="09F521A06284F2018087AF015799BEA5" italics="true">vs.</emphasis>
two in
<taxonomicName id="2409AA89CEB83A3AEB57E5CA36D3A886" authorityName="Campos-Filho, Araujo &amp; Taiti" authorityYear="2014" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="aganju">
<emphasis id="4B6ADE8DF60C33C1C96AC08E923803F3" italics="true">X. aganju</emphasis>
</taxonomicName>
,
<taxonomicName id="352556B453A7323A6D6DA40289D60905" authorityName="Campos-Filho, Gallo &amp; Bichuette" authorityYear="2022" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="lapaensis">
<emphasis id="7B35F59DE8F6C728B8DD7E4D6A508EE4" italics="true">X. lapaensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="E7D2780FB2FCEBBE34233C33CCCC4996" authorityName="Campos-Filho, Bichuette &amp; Taiti" authorityYear="2016" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="odara">
<emphasis id="BFBDFBD41D0B5C26CF3ED678A924E946" italics="true">X. odara</emphasis>
</taxonomicName>
, three in
<taxonomicName id="2A296EF406359E9D4C383D156BC9B2A0" authorityName="Bastos-Pereira, Souza &amp; Ferreira" authorityYear="2017" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="ibiracatuensis">
<emphasis id="3487F20BF11291C4A8BAC56C2437070D" italics="true">X. ibiracatuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="5BFACD68035AD80AD9D1255A0ECF47D0" authorityName="Cardoso, Bastos-Pereira, Souza &amp; Ferreira" authorityYear="2020" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="lundi">
<emphasis id="E5A1402CA2B7350C8B2B64980C8716BE" italics="true">X. lundi</emphasis>
</taxonomicName>
and
<taxonomicName id="75F604E77ACDBC07CAE6FD6F026AB239" authorityName="Cardoso, Bastos-Pereira, Souza &amp; Ferreira" authorityYear="2020" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="santinhoi">
<emphasis id="E38D5B6F781D635C648FA7882C9DDBCF" italics="true">X. santinhoi</emphasis>
</taxonomicName>
, four in
<taxonomicName id="7A875EE503E17193165D4950ADEF3C4F" authorityName="Cardoso, Bastos-Pereira, Souza &amp; Ferreira" authorityYear="2020" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="dagua">
<emphasis id="507548D3C8FDE4E5D1B953B6590F68D0" italics="true">X. dagua</emphasis>
</taxonomicName>
, four or five in
<taxonomicName id="1F42A109442FE6B1E4445678F1F7804A" authorityName="Cardoso, Bastos-Pereira, Souza &amp; Ferreira" authorityYear="2020" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="ceci">
<emphasis id="CB16F4FD549E9AA18030F4F23E206B43" italics="true">X. ceci</emphasis>
</taxonomicName>
, five in
<taxonomicName id="A6B9F9C08E2A501DE316B4B1B15D515A" authorityName="Campos-Filho, Gallao &amp; Bichuette" authorityYear="2022" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="loboi">
<emphasis id="18F8807520C1B66B1CA87E0CBEE71CA1" italics="true">X. loboi</emphasis>
</taxonomicName>
, six in
<taxonomicName id="FD3550489533B15E653259195FE08F6B" authorityName="Bastos-Pereira, Souza &amp; Ferreira" authorityYear="2017" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="itacarambiensis">
<emphasis id="A441A5F9EC744543402B109CDFC1AE22" italics="true">X. itacarambiensis</emphasis>
</taxonomicName>
, twelve in
<taxonomicName id="E9FA51A29156DA0C133BA2313BC3BC09" authorityName="López-Orozco &amp; Borja-Arrieta &amp; Carpio-Díaz &amp; Gallão &amp; Campos-Filho &amp; Bichuette" authorityYear="2024" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="chaimowiczi" status="sp. nov.">
<emphasis id="40A3C30A6CB40478B88B9DF7AEFB3E8C" italics="true">X. chaimowiczi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7660122A649EF46A9A24EE1642A6F6BA" rank="species">sp. nov.</taxonomicNameLabel>
), pleopod 1 basipod distal margin semicircular apex, longer than exopod (
<emphasis id="E922FD3D8B31F0F9B183F28F4297D4B4" italics="true">vs.</emphasis>
simples and shorter than exopod in
<taxonomicName id="E542311B003C9DCD58F7A005543769E2" authorityName="Cardoso, Bastos-Pereira, Souza &amp; Ferreira" authorityYear="2020" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="lundi">
<emphasis id="AB2FAD28403D59BAB35E03118CFC7849" italics="true">X. lundi</emphasis>
</taxonomicName>
,
<taxonomicName id="BD6616442F974F5A8F8238B22B8C4671" authorityName="Cardoso, Bastos-Pereira, Souza &amp; Ferreira" authorityYear="2020" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="dagua">
<emphasis id="AC6844F86624D35A26ECF4B659638FBA" italics="true">X. dagua</emphasis>
</taxonomicName>
,
<taxonomicName id="6FC3816BF7F583B5FE48B26694DB6EB0" authorityName="Cardoso, Bastos-Pereira, Souza &amp; Ferreira" authorityYear="2020" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="ceci">
<emphasis id="EF96AC5509AA50CD0A9EFBF9250A0B4C" italics="true">X. ceci</emphasis>
</taxonomicName>
,
<taxonomicName id="7AB7D2002AB17FD21B5479EF9B58F743" authorityName="Bastos-Pereira, Souza &amp; Ferreira" authorityYear="2017" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="ibiracatuensis">
<emphasis id="A3E0D57579C105185507104CB3AF777D" italics="true">X. ibiracatuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="6BC747E4952C9A124F42B5ADEC5D2201" authorityName="Bastos-Pereira, Souza &amp; Ferreira" authorityYear="2017" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="itacarambiensis">
<emphasis id="FAA2C8CFA7EF2CDB480CDA83715F0274" italics="true">X. itacarambiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="ACBFAE2EA90E4839868C2887015C2B7B" authorityName="Campos-Filho, Gallao &amp; Bichuette" authorityYear="2022" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="loboi">
<emphasis id="6BD6B76C4CD128011CCF1A42236DE75E" italics="true">X. loboi</emphasis>
</taxonomicName>
,
<taxonomicName id="E4486F43A4F6D23805C5619E00B2F806" authorityName="Campos-Filho, Bichuette &amp; Taiti" authorityYear="2016" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="odara">
<emphasis id="866CE98435D0AC08EB02187E85BB8EDF" italics="true">X. odara</emphasis>
</taxonomicName>
,
<taxonomicName id="E09A081E5FF2CF0FA6DAA3445D34FEA2" authorityName="Cardoso, Bastos-Pereira, Souza &amp; Ferreira" authorityYear="2020" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="santinhoi">
<emphasis id="E73F2AC9B222776C393D1990A0F45A30" italics="true">X. santinhoi</emphasis>
</taxonomicName>
, acute and longer than exopod in
<taxonomicName id="4386F567BE6C4AE7ACB7C2D441CAB61B" authorityName="López-Orozco &amp; Borja-Arrieta &amp; Carpio-Díaz &amp; Gallão &amp; Campos-Filho &amp; Bichuette" authorityYear="2024" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="chaimowiczi" status="sp. nov.">
<emphasis id="BF9B1BB57EAD9D385DB2D0A351561FD6" italics="true">X. chaimowiczi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31CA3A403BC5BCC501C7A888073B157E" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="821B9A521233DF874DE3D279572FB301" authorityName="Campos-Filho, Gallo &amp; Bichuette" authorityYear="2022" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="lapaensis">
<emphasis id="2BC53A3062E7FA43B0F0BB5CA07F4B3B" italics="true">X. lapaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="0FD10EB1F7F829DC4B6C4C963D92D176" authorityName="Campos-Filho, Araujo &amp; Taiti" authorityYear="2014" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="aganju">
<emphasis id="893ADF918368B3A3EADEDD8AE81D8231" italics="true">X. aganju</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EK6AE" type="natural history">
<paragraph id="3E9A84D0F552172E2ED22C007A84DF27">
<heading id="8B4A55914D098473519D7B6289CA55D8" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="E364251A5C94E42ACEB88324A19A71B8">
The species showed low abundance (only one individual caught along the cave). The only individual of
<taxonomicName id="3CC2F02BF1249BFD63A3341EBB28A459" authorityName="López-Orozco &amp; Borja-Arrieta &amp; Carpio-Díaz &amp; Gallão &amp; Campos-Filho &amp; Bichuette" authorityYear="2024" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="jonasi" status="sp. nov.">
<emphasis id="E24F7DAD043AFA09DF121536E5EB2483" italics="true">X. jonasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5B1606C2D0975AD5DF6299B008117EBE" rank="species">sp. nov.</taxonomicNameLabel>
captured was found in the aphotic zone on a very wet, muddy substrate, under the cracks formed by the stream current, just a few meters from a pond (Fig.
<figureCitation id="7257518B0CE7B4624E48B0BE4D996649" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the study area and distribution of the Xangoniscus species." httpUri="https://binary.pensoft.net/fig/1197443">1 C</figureCitation>
).
</paragraph>
<caption id="940C23451308252830E78B8C25A20A0A" ID-arpha="BA6EDD3B-EEAE-59C8-85CA-094D88D01422" httpUri="https://binary.pensoft.net/fig/1197449" startId="F7">
<paragraph id="5E9385257DBCBC3949711EE0415DDFF3">
<label id="244B0B84B87114060978C1FB0801FDC6">Figure 7.</label>
</paragraph>
<paragraph id="B95A43C269C3889885A880D4DC6B4877">
<taxonomicName id="CFC71FB351B006AB2B8707ACA4F7A801" authority="Lopez-Orozco, Bichuette &amp; Campos-Filho" authorityName="Lopez-Orozco, Bichuette &amp; Campos-Filho" authorityYear="2024" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="jonasi" status="sp. nov.">
<emphasis id="C2ACFB5B6786670529F02FB8A4A6E7A6" italics="true">Xangoniscus jonasi</emphasis>
López-Orozco, Bichuette &amp; Campos-Filho
</taxonomicName>
,
<taxonomicNameLabel id="10C45EDB0B2EAE790C6588371AC511B7" rank="species">sp. nov.</taxonomicNameLabel>
Male
<emphasis id="F311F5606D570E08EF2E7E21D91E7EF1" bold="true">A</emphasis>
habitus, dorsal view
<emphasis id="2A12F221F7F301EEF10336C04D817C4C" bold="true">B</emphasis>
habitus, lateral view
<emphasis id="14CCCB694926F45769BC2DE97FDF169A" bold="true">C</emphasis>
cephalon and pereonite 1, frontal view
<emphasis id="C369078338ED5E7B5F07EFABD3FAB868" bold="true">D</emphasis>
cephalon and pereonites 1–2, lateral view
<emphasis id="EE149B8E6462D6CC922A9315E6DF7C7E" bold="true">E</emphasis>
pereonite 7, pleon and telson, dorsal view. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="6C4D122B2A235A7E520C625FB77D652B" ID-arpha="A7AE9D76-46AD-509C-9A03-3094D868ED79" httpUri="https://binary.pensoft.net/fig/1197450" startId="F8">
<paragraph id="800FE89F79A19BC8D25475A2B18E8496">
<label id="C650B6EB3B8869D420FAA903B9266104">Figure 8.</label>
</paragraph>
<paragraph id="115C4215ACF899356B09A313E48A1D42">
<taxonomicName id="22B9108A4549D7969BE58BA892E121DA" authority="Lopez-Orozco, Bichuette &amp; Campos-Filho" authorityName="Lopez-Orozco, Bichuette &amp; Campos-Filho" authorityYear="2024" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="jonasi" status="sp. nov.">
<emphasis id="8327C61D2553411F715E3734D106F5EB" italics="true">Xangoniscus jonasi</emphasis>
López-Orozco, Bichuette &amp; Campos-Filho
</taxonomicName>
,
<taxonomicNameLabel id="9D4B056691821F1D3115625B4C34E34F" rank="species">sp. nov.</taxonomicNameLabel>
Male
<emphasis id="C28C22CE94A2B7EA39FE4D9DA1C905EA" bold="true">A</emphasis>
habitus, dorsal view
<emphasis id="5E69E37B7F2EF2ECDF16A6C8355B4612" bold="true">B</emphasis>
dorsal scale-seta
<emphasis id="5E9B626DE28DB4B4F4B6FF32F08ED0A0" bold="true">C</emphasis>
cephalon, frontal view
<emphasis id="2549F397342789C39024D2876230EA12" bold="true">D</emphasis>
telson and uropod
<emphasis id="E7416F9E87569F437566A6F82F74B0B8" bold="true">E</emphasis>
antennula
<emphasis id="D50B9F02FA0417B0E3CAD8BCCEEED167" bold="true">F</emphasis>
antenna
<emphasis id="EC4F3B0807ED754F52AF9CDF74D65FAB" bold="true">G</emphasis>
left mandible
<emphasis id="434C8EA3A35D2569CEF816F2BDAD5CE3" bold="true">H</emphasis>
right mandible
<emphasis id="3C15E6DC929631DE5EE9BFA33BAC0348" bold="true">I</emphasis>
maxillula
<emphasis id="DB207A482BC51E7CAFC77E484B9C308F" bold="true">J</emphasis>
maxilla
<emphasis id="7FDA74821F1535EA6065660E0D6B6F49" bold="true">K</emphasis>
maxilliped, arrow illustrating the endite in caudal view.
</paragraph>
</caption>
<caption id="A5D0B6E77A6C65DF3C49895085776A99" ID-arpha="6432334F-B8AA-5737-9056-941205C9CB1F" httpUri="https://binary.pensoft.net/fig/1197451" startId="F9">
<paragraph id="E97F9D01498E4272251784545763AB5A">
<label id="B4B6D5B47A0605683257736CF61AB7C2">Figure 9.</label>
</paragraph>
<paragraph id="26662B795FB0A31EC4FA26B0D2E7E84D">
<taxonomicName id="A8D4ECE025CA58849A4A08DE13E83A02" authority="Lopez-Orozco, Bichuette &amp; Campos-Filho" authorityName="Lopez-Orozco, Bichuette &amp; Campos-Filho" authorityYear="2024" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="jonasi" status="sp. nov.">
<emphasis id="75612069C6781FA6A538EE2C6F5BBD64" italics="true">Xangoniscus jonasi</emphasis>
López-Orozco, Bichuette &amp; Campos-Filho
</taxonomicName>
,
<taxonomicNameLabel id="3343F0395AD4B0F5DB9E66A0BE39FB67" rank="species">sp. nov.</taxonomicNameLabel>
Male
<emphasis id="A7F46B350EA8058B4178BDA7724FBA8D" bold="true">A</emphasis>
pereopod 1
<emphasis id="B58048D914CFA1D8DEB3C3476BB0C72F" bold="true">B</emphasis>
pereopod 2
<emphasis id="99258972EE2465752D8D5915DEA733AF" bold="true">C</emphasis>
pereopod 3
<emphasis id="2B488BC5787B6E2C4BA6F5D4D47B35A5" bold="true">D</emphasis>
pereopod 4
<emphasis id="574EE500155CF069943ED6595BEBD70B" bold="true">E</emphasis>
pereopod 5
<emphasis id="E6D9B036BDED2F203CC0BB0946C135BD" bold="true">F</emphasis>
pereopod 6
<emphasis id="47852190E20A9E5D7C10BD95D3B34AA0" bold="true">G</emphasis>
pereopod 7.
</paragraph>
</caption>
<caption id="88B19A3D79FAB011A0F03B1A2891D712" ID-arpha="251D7A5D-6354-58EB-9B86-6572D67912A5" httpUri="https://binary.pensoft.net/fig/1197452" startId="F10">
<paragraph id="966D17E7270E2CC2A4CCDC02905532A9">
<label id="AF1BCF36A68E945A8D68F19D9D43DDAB">Figure 10.</label>
</paragraph>
<paragraph id="D042533AC82D0FEC260015A8847230FD">
<taxonomicName id="45F4A845AF633F8D26C2B987F0207F89" authority="Lopez-Orozco, Bichuette &amp; Campos-Filho" authorityName="Lopez-Orozco, Bichuette &amp; Campos-Filho" authorityYear="2024" class="Malacostraca" family="Styloniscidae" genus="Xangoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="jonasi" status="sp. nov.">
<emphasis id="78993508871F94F98CF39BBCA13B6373" italics="true">Xangoniscus jonasi</emphasis>
López-Orozco, Bichuette &amp; Campos-Filho
</taxonomicName>
,
<taxonomicNameLabel id="8E230813531D5E23C34E18524328184D" rank="species">sp. nov.</taxonomicNameLabel>
Male
<emphasis id="9AE54CF618AD12DF0C29669F209ECC9E" bold="true">A</emphasis>
pleopod 1
<emphasis id="17A98D7C1918C1FDC8EDD2E871CA0987" bold="true">B</emphasis>
genital papilla
<emphasis id="F8618014AEFBEC615A7BC90EA199FB53" bold="true">C</emphasis>
pleopod 2
<emphasis id="2DF75958AA8D993FC3252BFDA507A1CE" bold="true">D</emphasis>
pleopod 3 exopod
<emphasis id="D4300FD20A8A95E3CE7C0D145E01B5D0" bold="true">E</emphasis>
pleopod 4 exopod
<emphasis id="529DF7B853A2A6EF9ED36246DCD4C96F" bold="true">F</emphasis>
pleopod 5 exopod.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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