treatments-xml/data/95/5A/87/955A8790397C7634FF1D970EFA16FBEA.xml
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<document id="849776D73647A183949F0B7D2674029A" ID-DOI="10.5281/zenodo.5352614" ID-ISSN="1942-1354" ID-Zenodo-Dep="5352614" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630208383417" checkinUser="felipe" docAuthor="Skelley, Paul E." docDate="2009" docId="955A8790397C7634FF1D970EFA16FBEA" docLanguage="en" docName="0085.pdf" docOrigin="Insecta Mundi 2009 (85)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Polyphylla starkae Skelley 2009, new species" docType="treatment" docVersion="4" lastPageNumber="6" masterDocId="6963FFE8397F7633FFDD904BF92EFFD1" masterDocTitle="A new species of Polyphylla Harris from peninsular Florida Coleoptera: Scarabaeidae: Melolonthinae) with a key to species of the pubescens species group" masterLastPageNumber="14" masterPageNumber="1" pageNumber="2" updateTime="1699185867608" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="AB094B616DF278C4FB49686B7FC8B31D">A new species of Polyphylla Harris from peninsular Florida Coleoptera: Scarabaeidae: Melolonthinae) with a key to species of the pubescens species group</mods:title>
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<mods:namePart id="A8674E84E6E275A8101EF95C9DDEB06D">Skelley, Paul E.</mods:namePart>
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<mods:title id="208DB9BE834615898B5159DD0D3147CA">Insecta Mundi</mods:title>
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<mods:date id="228CA03A29EA0A955A8018AE07B2C9D7">2009</mods:date>
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<mods:number id="A2A6848B9F0A0DE1241B1B368ECFF260">2009-07-24</mods:number>
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<mods:number id="5949893E55D7506C14ACBFF289AE01BA">2009</mods:number>
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<treatment id="955A8790397C7634FF1D970EFA16FBEA" ID-DOI="http://doi.org/10.5281/zenodo.5450555" ID-Zenodo-Dep="5450555" LSID="urn:lsid:plazi:treatment:955A8790397C7634FF1D970EFA16FBEA" httpUri="http://treatment.plazi.org/id/955A8790397C7634FF1D970EFA16FBEA" lastPageId="7" lastPageNumber="6" pageId="3" pageNumber="2">
<subSubSection id="55E9650D397C7630FF1D970EFBF5F88A" box="[192,731,1859,1884]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="1D4C3686397C7630FF1D970EFBF5F88A" blockId="3.[192,731,1859,1915]" box="[192,731,1859,1884]" pageId="3" pageNumber="2">
<heading id="460481EA397C7630FF1D970EFBF5F88A" bold="true" box="[192,731,1859,1884]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
<emphasis id="2F87EA94397C7630FF1D970EFBF5F88A" bold="true" box="[192,731,1859,1884]" pageId="3" pageNumber="2">
<taxonomicName id="DAF34D05397C7630FF1D970EFB02F88A" authority="Skelley" authorityName="Skelley" authorityYear="2009" box="[192,556,1859,1884]" class="Insecta" family="Melolonthidae" genus="Polyphylla" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="starkae" status="sp. nov.">
<emphasis id="2F87EA94397C7630FF1D970EF8EEF88D" bold="true" box="[192,448,1861,1884]" italics="true" pageId="3" pageNumber="2">Polyphylla starkae</emphasis>
Skelley
</taxonomicName>
,
<taxonomicNameLabel id="34B457EF397C7630FDE59708FBF5F88A" box="[568,731,1859,1883]" pageId="3" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="55E9650D397C7630FF1D9728FB4EF8AA" box="[192,608,1891,1915]" pageId="3" pageNumber="2" type="description">
<paragraph id="1D4C3686397C7630FF1D9728FB4EF8AA" blockId="3.[192,731,1859,1915]" box="[192,608,1891,1915]" pageId="3" pageNumber="2">
<figureCitation id="85C82A03397C7630FF1D9728F86FF8AA" box="[192,321,1891,1915]" captionStart-0="Figure 1-5" captionStart-1="Figure 6-9" captionStartId-0="4.[192,276,1426,1448]" captionStartId-1="5.[192,276,1151,1173]" captionTargetBox-0="[194,1440,193,1407]" captionTargetBox-1="[191,1448,192,1122]" captionTargetId-0="figure-20@4.[194,1441,193,1407]" captionTargetId-1="figure-321@5.[191,1448,192,1122]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figure 1-5. Polyphylla starkae, new species. 1) Clear cut type locality in April 2009. 2-5) Adult beetles, male on left, female on right, courtesy D. Almquist, FNAI." captionText-1="Figure 6-9. Polyphylla spp. head and pronotum, males on left, females on right. 6-7) Polyphylla starkae. 8-9) Polyphylla pubescens." figureDoi-0="http://doi.org/10.5281/zenodo.5352616" figureDoi-1="http://doi.org/10.5281/zenodo.5352618" httpUri-0="https://zenodo.org/record/5352616/files/figure.png" httpUri-1="https://zenodo.org/record/5352618/files/figure.png" pageId="3" pageNumber="2">Figure 2-7</figureCitation>
, 10, 14, 18, 22-23, 30-32
</paragraph>
</subSubSection>
<caption id="498C660E397B7637FF1D95D9FBC5FA15" ID-DOI="http://doi.org/10.5281/zenodo.5352616" ID-Zenodo-Dep="5352616" httpUri="https://zenodo.org/record/5352616/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,276,1426,1448]" targetBox="[194,1440,193,1407]" targetPageId="4">
<paragraph id="1D4C3686397B7637FF1D95D9FBC5FA15" blockId="4.[192,1440,1426,1477]" pageId="4" pageNumber="3">
<emphasis id="2F87EA94397B7637FF1D95D9F868FA76" bold="true" box="[192,326,1426,1448]" pageId="4" pageNumber="3">Figure 1-5.</emphasis>
<taxonomicName id="DAF34D05397B7637FE9195D9FB32FA76" authority="Skelley, 2009" authorityName="Skelley" authorityYear="2009" box="[332,540,1426,1447]" class="Insecta" family="Melolonthidae" genus="Polyphylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="starkae" status="sp. nov.">
<emphasis id="2F87EA94397B7637FE9195D9FB32FA76" box="[332,540,1426,1447]" italics="true" pageId="4" pageNumber="3">Polyphylla starkae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="34B457EF397B7637FDF495D8FB9EFA79" box="[553,688,1426,1448]" pageId="4" pageNumber="3" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="2F87EA94397B7637FD6795D9FBFEFA79" bold="true" box="[698,720,1426,1448]" pageId="4" pageNumber="3">1)</emphasis>
Clear cut type locality in April 2009.
<emphasis id="2F87EA94397B7637FBAE95D9FD8EFA79" bold="true" box="[1139,1184,1426,1448]" pageId="4" pageNumber="3">2-5)</emphasis>
Adult beetles, male on left, female on right, courtesy D. Almquist, FNAI.
</paragraph>
</caption>
<subSubSection id="55E9650D397B7637FF1D95A5FBF0F977" pageId="4" pageNumber="3" type="diagnosis">
<paragraph id="1D4C3686397B7637FF1D95A5FBF0F977" blockId="4.[192,1440,1518,1702]" pageId="4" pageNumber="3">
<emphasis id="2F87EA94397B7637FF1D95A5F867F9D7" bold="true" box="[192,329,1518,1542]" pageId="4" pageNumber="3">Diagnosis</emphasis>
.
<taxonomicName id="DAF34D05397B7637FE8795A5FB6DF9D7" authorityName="Skelley" authorityYear="2009" box="[346,579,1518,1542]" class="Insecta" family="Melolonthidae" genus="Polyphylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="starkae">
<emphasis id="2F87EA94397B7637FE8795A5FB6DF9D7" box="[346,579,1518,1542]" italics="true" pageId="4" pageNumber="3">Polyphylla starkae</emphasis>
</taxonomicName>
is a member of the pubescens species group and unique with males having elytral setae clustered into groups, some in vague stripes (
<figureCitation id="85C82A03397B7637FC129645FD1EF9F7" box="[975,1072,1550,1574]" captionStart="Figure 1-5" captionStartId="4.[192,276,1426,1448]" captionTargetBox="[194,1440,193,1407]" captionTargetId="figure-20@4.[194,1441,193,1407]" captionTargetPageId="4" captionText="Figure 1-5. Polyphylla starkae, new species. 1) Clear cut type locality in April 2009. 2-5) Adult beetles, male on left, female on right, courtesy D. Almquist, FNAI." figureDoi="http://doi.org/10.5281/zenodo.5352616" httpUri="https://zenodo.org/record/5352616/files/figure.png" pageId="4" pageNumber="3">Fig. 2, 4</figureCitation>
,
<figureCitation id="85C82A03397B7637FBE79645FD75F9F7" box="[1082,1115,1550,1574]" captionStart="Figure 22-29" captionStartId="9.[192,276,1286,1308]" captionTargetBox="[192,1446,186,1259]" captionTargetId="figure-231@9.[192,1446,186,1259]" captionTargetPageId="9" captionText="Figure 22-29. Polyphylla - pubescens species group, male (left) and female (right). 22-23) P. starkae. 24-25) P. donaldsoni. 26-27) P. pubescens. 28-29) P. woodruffi. These images were taken and reproduced at the same magnification." figureDoi="http://doi.org/10.5281/zenodo.5352620" httpUri="https://zenodo.org/record/5352620/files/figure.png" pageId="4" pageNumber="3">22</figureCitation>
). This species is the largest member of the pubescens group and is strongly sexually dimorphic. Females are more robust than the males (
<figureCitation id="85C82A03397B7637FEC69605F8AAF9B7" box="[283,388,1614,1638]" captionStart="Figure 1-5" captionStartId="4.[192,276,1426,1448]" captionTargetBox="[194,1440,193,1407]" captionTargetId="figure-20@4.[194,1441,193,1407]" captionTargetPageId="4" captionText="Figure 1-5. Polyphylla starkae, new species. 1) Clear cut type locality in April 2009. 2-5) Adult beetles, male on left, female on right, courtesy D. Almquist, FNAI." figureDoi="http://doi.org/10.5281/zenodo.5352616" httpUri="https://zenodo.org/record/5352616/files/figure.png" pageId="4" pageNumber="3">Fig. 3, 5</figureCitation>
,
<figureCitation id="85C82A03397B7637FE4F9605F89EF9B7" box="[402,432,1614,1638]" captionStart="Figure 22-29" captionStartId="9.[192,276,1286,1308]" captionTargetBox="[192,1446,186,1259]" captionTargetId="figure-231@9.[192,1446,186,1259]" captionTargetPageId="9" captionText="Figure 22-29. Polyphylla - pubescens species group, male (left) and female (right). 22-23) P. starkae. 24-25) P. donaldsoni. 26-27) P. pubescens. 28-29) P. woodruffi. These images were taken and reproduced at the same magnification." figureDoi="http://doi.org/10.5281/zenodo.5352620" httpUri="https://zenodo.org/record/5352620/files/figure.png" pageId="4" pageNumber="3">23</figureCitation>
) with legs modified for burrowing (compare Fig. 10, 14) and elytral setae nearly absent. Although flight wings appear fully developed, female elytra are tightly connected at the base along the suture, suggesting females are flightless.
</paragraph>
</subSubSection>
<subSubSection id="55E9650D397B7635FF1D9685F812F96A" lastPageId="6" lastPageNumber="5" pageId="4" pageNumber="3" type="description">
<paragraph id="1D4C3686397B7636FF1D9685FD1EFADD" blockId="4.[192,1440,1742,1894]" lastBlockId="5.[192,1441,1236,1900]" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3">
<emphasis id="2F87EA94397B7637FF1D9685FB16F937" bold="true" box="[192,568,1742,1766]" pageId="4" pageNumber="3">
Description. Male
<typeStatus id="C2488824397B7637FE679685FB16F937" box="[442,568,1742,1766]" pageId="4" pageNumber="3" type="holotype">Holotype</typeStatus>
</emphasis>
. Length
<quantity id="DA0B9B63397B7637FD799685FBD7F937" box="[676,761,1742,1766]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="3" unit="mm" value="23.0">23 mm</quantity>
; width
<quantity id="DA0B9B63397B7637FC8C9685FA89F937" box="[849,935,1742,1766]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="3" unit="mm" value="11.0">11 mm</quantity>
. Body reddish-brown to nearly black with overlying vestiture of golden setae generally dense all over. Head between eyes with moderately coarse punctures separated by about 1 x their diameter, punctures of frons with long erect setae. Clypeus transverse, 2 x wider than long, with broadly reflexed margin, anterior angles rounded; anterior margin broadly truncate in dorsal view, weakly sinuate in anterior view; surface punctures slightly larger than those on frons, separated by 0.5-1.0 x their diameter, setae moderately appressed, not erect. Antennae uniformly pale brown; antennal club large, length 1.5 x width of clypeus.
</paragraph>
<caption id="498C660E397A7636FF1D9434F89EFB63" ID-DOI="http://doi.org/10.5281/zenodo.5352618" ID-Zenodo-Dep="5352618" httpUri="https://zenodo.org/record/5352618/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,276,1151,1173]" targetBox="[191,1448,192,1122]" targetPageId="5">
<paragraph id="1D4C3686397A7636FF1D9434F89EFB63" blockId="5.[192,1440,1151,1202]" pageId="5" pageNumber="4">
<emphasis id="2F87EA94397A7636FF1D9434F866FB44" bold="true" box="[192,328,1151,1173]" pageId="5" pageNumber="4">Figure 6-9.</emphasis>
<taxonomicName id="DAF34D05397A7636FE9294CBF8D0FB44" box="[335,510,1152,1173]" class="Insecta" family="Melolonthidae" genus="Polyphylla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2F87EA94397A7636FE9294CBF8E9FB44" box="[335,455,1152,1173]" italics="true" pageId="5" pageNumber="4">Polyphylla</emphasis>
spp.
</taxonomicName>
head and pronotum, males on left, females on right.
<emphasis id="2F87EA94397A7636FB819434FDA7FB44" bold="true" box="[1116,1161,1151,1173]" pageId="5" pageNumber="4">6-7)</emphasis>
<taxonomicName id="DAF34D05397A7636FB4F94CBFC4DFB44" authorityName="Skelley" authorityYear="2009" box="[1170,1379,1152,1173]" class="Insecta" family="Melolonthidae" genus="Polyphylla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="starkae">
<emphasis id="2F87EA94397A7636FB4F94CBFC4DFB44" box="[1170,1379,1152,1173]" italics="true" pageId="5" pageNumber="4">Polyphylla starkae</emphasis>
</taxonomicName>
.
<emphasis id="2F87EA94397A7636FAAE9434FC8EFB44" bold="true" box="[1395,1440,1151,1173]" pageId="5" pageNumber="4">8-9)</emphasis>
<taxonomicName id="DAF34D05397A7636FF1D94D7F887FB60" authorityName="Cartwright" authorityYear="1939" box="[192,425,1180,1201]" class="Insecta" family="Melolonthidae" genus="Polyphylla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="2F87EA94397A7636FF1D94D7F887FB60" box="[192,425,1180,1201]" italics="true" pageId="5" pageNumber="4">Polyphylla pubescens</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="1D4C3686397A7636FF2D955FFC1AFA3D" blockId="5.[192,1441,1236,1900]" pageId="5" pageNumber="4">Pronotal width = 1.8 x length; lateral margin rounded, weakly serrate because of setae bearing punctures; surface glossy between punctures; punctures coarse, same size as on clypeus, separated by less than 0.5 x their diameter near anterior and posterior angles, less dense on disc where variably separated by 1.0-4.0 x their diameter; each puncture bearing an appressed seta, lateral setae distinctly shorter than those on disc. Pronotal hypomeron with scattered coarse punctures bearing long erect setae, except for completely glabrous apicolateral third near pronotal angle. Scutellum with coarse punctures and setae as on disc, setose punctures nearly coalescing along midline, much sparser laterally.</paragraph>
<paragraph id="1D4C3686397A7636FF2D95BFFC48F8DD" blockId="5.[192,1441,1236,1900]" pageId="5" pageNumber="4">Elytral length 3.2 x length of pronotum; lateral margin moderately reflexed, reflexed lateral edge reaching middle of humerus, not attaining base; surface glossy with glabrous areas near base and coarsely punctate areas on apical two thirds between patches of dense fine punctures, all punctures bearing short appressed seta, setal patches vaguely linear in arrangement, setal covering along the sutural, apical and lateral margins complete although variable in width. Flight wings present, fully functional (specimen observed flying). Venter with metathorax densely covered with long erect golden setae. Each abdominal sternite with band of setae bearing punctures on apical half, band wide laterally, narrowed medially, setae appressed; punctures on first three visible sternites fine and dense, punctures on last two visible sternites 3 x diameter of those on basal sternites, all punctures separated by 1-2 x their diameters.</paragraph>
<paragraph id="1D4C3686397A7635FF2D975FFAD8FE4A" blockId="5.[192,1441,1236,1900]" lastBlockId="6.[192,1441,195,1371]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
All femora somewhat flattened. Pro- and mesofemora narrowed, parallel-sided most of length. Metafemora weakly dilated, wider than mesofemora, margins arcuate, length 3.1 x maximal width. Protibia elongate, with apical tooth and second tooth at apical quarter. Mesotibia narrow entire length, not expanded at apex; with weak external tooth at middle of outer edge small; tooth at basal fourth lacking. Metatibia slender, parallel-sided most of length, wider at apex; width at apex = midline length of third visible abdominal sternite; inner tibial margin slightly curved at apex. Metatibial spurs narrowed to acute point. Legs reddish-brown, contrasting with dark pronotum. All tarsi with tooth on posterior claw 0.75 x length of tooth on anterior claw. Genitalia with paramere length 2.5 x basal width in dorsal view, abruptly narrowed at middle; each paramere with dorsal-lateral groove from middle to apical quarter; apex laterally flattened and rounded in lateral view (
<figureCitation id="85C82A0339797635FCB091C8FAC4FE4A" box="[877,1002,387,411]" captionStart="Figure 30-32" captionStartId="10.[192,276,858,880]" captionTargetBox="[191,1443,191,834]" captionTargetId="figure-487@10.[191,1443,191,834]" captionTargetPageId="10" captionText="Figure 30-32. Polyphylla starkae, male genitalia. 30) Caudal view. 31) Lateral view. 32) Dorsal view. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5352622" httpUri="https://zenodo.org/record/5352622/files/figure.png" pageId="6" pageNumber="5">Fig. 30-32</figureCitation>
).
</paragraph>
<paragraph id="1D4C368639797635FF2D91E8FD7CFDAA" blockId="6.[192,1441,195,1371]" pageId="6" pageNumber="5">
<emphasis id="2F87EA9439797635FF2D91E8F8FBFE6A" bold="true" box="[240,469,419,443]" pageId="6" pageNumber="5">
Female
<typeStatus id="C248882439797635FE8791E8F8E1FE6A" box="[346,463,419,443]" pageId="6" pageNumber="5" type="allotype">Allotype</typeStatus>
.
</emphasis>
Length
<quantity id="DA0B9B6339797635FDEB91E8FBA5FE6A" box="[566,651,419,443]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="5" unit="mm" value="25.0">25 mm</quantity>
; width
<quantity id="DA0B9B6339797635FD3C91E8FA19FE6A" box="[737,823,419,443]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="5" unit="mm" value="13.0">13 mm</quantity>
. Body reddish-brown, elytral and abdominal cuticle thin, structures underneath visible when alive, dorsal vestiture of thin setae sparse. Head between eyes with coarse punctures separated by about 1 x their diameter, punctures of frons with long erect setae. Clypeus trapezoidal, sides converging anteriorly, 2 x wider than long, with narrowly reflexed margin, anterior angles rounded; anterior margin broadly truncate in dorsal view, straight in anterior view; surface punctures same size and distribution as on frons, setae moderately appressed, not erect. Antenna unicolorous pale brown; antennal club short, length 0.25 x width of clypeus.
</paragraph>
<paragraph id="1D4C368639797635FF2D92C8FD42FC8A" blockId="6.[192,1441,195,1371]" pageId="6" pageNumber="5">Pronotal width = 1.7 x length; lateral margin rounded; surface glossy between widely scattered coarse punctures, punctures size 1.0-1.5 x those on clypeus, separated by less than 0.5 x their diameter near anterior and posterior angles, less dense on disc where variably separated by 1.0-4.0 x their diameter; each puncture bearing a seta, which vary from somewhat appressed to erect, lateral setae shorter than those on disc. Pronotal hypomeron with scattered coarse punctures bearing long erect setae, except for completely glabrous apical-lateral third near pronotal angle. Scutellum with laterally with coarse punctures and setae as on disc, smaller punctures nearly coalescing along midline.</paragraph>
<paragraph id="1D4C368639797635FF2D9328F841FBCA" blockId="6.[192,1441,195,1371]" pageId="6" pageNumber="5">
Elytral length 3.3 x length of pronotum; lateral margin weakly reflexed, sharp lateral edge reaching middle of humerus, not attaining base; surface glossy and mostly glabrous with widely scattered punctures bearing fine setae; setal covering visible along the sutural, apical and lateral margins complete although variable in width; elytra were tightly connected at basal quarter along suture and they were separated with some difficulty. Flight wings present. Venter with metathorax and abdomen similar to male
<typeStatus id="C248882439797635FEDD9448F844FBCA" box="[256,362,1027,1051]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="1D4C368639797635FF2D9468FC26FA8A" blockId="6.[192,1441,195,1371]" pageId="6" pageNumber="5">All femora with surface somewhat convex. Pro- and mesofemora weakly arcuate most of length. Metafemora distinctly dilated, 1.5 x wider than mesofemora, margins strongly arcuate, length 2.1 x maximal width. Protibia broad, distinctly tridentate, with elongate apical tooth, elongate middle tooth at apical third, triangular basal tooth at basal quarter. Mesotibia slender entire length, not expanded at apex, width 0.5 x mesofemoral width; with small external tooth at middle of outer edge; tooth at basal fourth lacking; apical spurs narrowed to acute apex spatulate, slightly curved downward, upper spur shorter than lower spur. Metatibia parallel-sided along middle half of length; tooth at middle of outer edge weak; abruptly widened near apex; apical spurs curve downward on lower edge; lower spur flattened, gradually narrowing to acute apex, upper spur somewhat spatulate, widest at middle; portion of apical tibial plate below tarsal junction same size as upper part. All tarsi with teeth equal in size.</paragraph>
<paragraph id="1D4C368639797635FF1D95C8FBDEF98A" blockId="6.[192,1440,1411,1723]" pageId="6" pageNumber="5">
<emphasis id="2F87EA9439797635FF1D95C8F86BFA4A" bold="true" box="[192,325,1411,1435]" pageId="6" pageNumber="5">Variation</emphasis>
.
<emphasis id="2F87EA9439797635FE8A95C8F8B5FA4A" bold="true" box="[343,411,1411,1435]" pageId="6" pageNumber="5">Male</emphasis>
. Length
<quantity id="DA0B9B6339797635FDD295C8FBBFFA4A" box="[527,657,1411,1435]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="6" pageNumber="5" unit="mm" value="23.0" valueMax="25.0" valueMin="21.0">21-25 mm</quantity>
; width
<quantity id="DA0B9B6339797635FD2C95C8FA5DFA4A" box="[753,883,1411,1435]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="6" pageNumber="5" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10-12 mm</quantity>
. Anterior clypeal margin on most is weakly sinuate in anterior view, is somewhat raised in the middle on a few. Lateral pronotal margin, evenly arcuate on most, is slightly angulate on a few. Patches of elytral setae vary in size in the specimens available. On some, their somewhat linear arrangement is clearer, but never prominent, on others it is imperceptible. The elytral pattern of radiating lines in reflected light is not visible in most specimens. Two specimens studied had patterning identical to the other members of this group. Body color also varied from light brown to dark brown, almost black.
</paragraph>
<paragraph id="1D4C368639797635FF2D9628F812F96A" blockId="6.[192,1440,1411,1723]" pageId="6" pageNumber="5">
<emphasis id="2F87EA9439797635FF2D9628F87AF9AA" bold="true" box="[240,340,1635,1659]" pageId="6" pageNumber="5">Female</emphasis>
. Length
<quantity id="DA0B9B6339797635FE609628FB19F9AA" box="[445,567,1635,1659]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="6" pageNumber="5" unit="mm" value="24.0" valueMax="25.0" valueMin="23.0">23-25 mm</quantity>
; width
<quantity id="DA0B9B6339797635FD519628FA26F9AA" box="[652,776,1635,1659]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="6" pageNumber="5" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13-14 mm</quantity>
. Body color in life varied from the reddish-brown (
<figureCitation id="85C82A0339797635FA8B9628F9E1F94A" captionStart="Figure 1-5" captionStartId="4.[192,276,1426,1448]" captionTargetBox="[194,1440,193,1407]" captionTargetId="figure-20@4.[194,1441,193,1407]" captionTargetPageId="4" captionText="Figure 1-5. Polyphylla starkae, new species. 1) Clear cut type locality in April 2009. 2-5) Adult beetles, male on left, female on right, courtesy D. Almquist, FNAI." figureDoi="http://doi.org/10.5281/zenodo.5352616" httpUri="https://zenodo.org/record/5352616/files/figure.png" pageId="6" pageNumber="5">Fig. 3, 5</figureCitation>
) to a chocolate brown, similar to the pinned specimen in
<figureCitation id="85C82A0339797635FC7D96C8FD33F94A" box="[928,1053,1667,1691]" captionStart="Figure 22-29" captionStartId="9.[192,276,1286,1308]" captionTargetBox="[192,1446,186,1259]" captionTargetId="figure-231@9.[192,1446,186,1259]" captionTargetPageId="9" captionText="Figure 22-29. Polyphylla - pubescens species group, male (left) and female (right). 22-23) P. starkae. 24-25) P. donaldsoni. 26-27) P. pubescens. 28-29) P. woodruffi. These images were taken and reproduced at the same magnification." figureDoi="http://doi.org/10.5281/zenodo.5352620" httpUri="https://zenodo.org/record/5352620/files/figure.png" pageId="6" pageNumber="5">Figure 23</figureCitation>
. All specimens darkened when preserved.
</paragraph>
</subSubSection>
<subSubSection id="55E9650D39797634FF1D96A8FB67FD0A" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5" type="materials_examined">
<paragraph id="1D4C368639797635FF1D96A8F8B6F8EA" blockId="6.[192,1441,1763,1851]" pageId="6" pageNumber="5">
<emphasis id="2F87EA9439797635FF1D96A8F8A8F92A" bold="true" box="[192,390,1763,1787]" pageId="6" pageNumber="5">Type Locality.</emphasis>
Auburndale,
<collectingRegion id="DF37F86439797635FDF296A8FBA3F92A" box="[559,653,1763,1787]" country="United States of America" name="Florida" pageId="6" pageNumber="5">Florida</collectingRegion>
,
<collectingCountry id="65E4761639797635FD4596A8FBFCF92A" box="[664,722,1763,1787]" name="United States of America" pageId="6" pageNumber="5">USA</collectingCountry>
(
<figureCitation id="85C82A0339797635FD3F96A8FA08F92A" box="[738,806,1763,1787]" captionStart="Figure 1-5" captionStartId="4.[192,276,1426,1448]" captionTargetBox="[194,1440,193,1407]" captionTargetId="figure-20@4.[194,1441,193,1407]" captionTargetPageId="4" captionText="Figure 1-5. Polyphylla starkae, new species. 1) Clear cut type locality in April 2009. 2-5) Adult beetles, male on left, female on right, courtesy D. Almquist, FNAI." figureDoi="http://doi.org/10.5281/zenodo.5352616" httpUri="https://zenodo.org/record/5352616/files/figure.png" pageId="6" pageNumber="5">Fig. 1</figureCitation>
), which is geographically part of the Winter Haven Ridge. For conservation purposes, and until more populations are discovered, the exact coordinates are not provided here.
</paragraph>
<paragraph id="1D4C368639787634FF1D9088FAE2FF2A" blockId="7.[192,1441,195,731]" pageId="7" pageNumber="6">
<emphasis id="2F87EA9439787634FF1D9088F8A3FF0A" bold="true" box="[192,397,195,219]" pageId="7" pageNumber="6">Type Materials.</emphasis>
Male
<typeStatus id="C248882439787634FE0E9088FB16FF0A" box="[467,568,195,219]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
and female
<typeStatus id="C248882439787634FD1F9088FA0EFF0A" box="[706,800,195,219]" pageId="7" pageNumber="6" type="allotype">allotype</typeStatus>
label data as follows:
<collectingRegion id="DF37F86439787634FBC49088FDB6FF0A" box="[1049,1176,195,219]" country="United States of America" name="Florida" pageId="7" pageNumber="6">FLORIDA</collectingRegion>
: Polk Co., Auburndale,
<date id="694D104639787634FF1D90A8F844FF2A" box="[192,362,227,251]" pageId="7" pageNumber="6" value="2009-04-15">15 April 2009</date>
, P. Skelley,
<collectionCode id="7BE2AE4339787634FE2390AFFB3BFF2A" box="[510,533,228,251]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="6">R</collectionCode>
. Morris,
<collectionCode id="7BE2AE4339787634FD5C90AFFBB8FF2A" box="[641,662,228,251]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="6">T</collectionCode>
. Palmer, mating at dusk.
</paragraph>
<paragraph id="1D4C368639787634FF2D914FFC1EFE4A" blockId="7.[192,1441,195,731]" pageId="7" pageNumber="6">
<typeStatus id="C248882439787634FF2D914FF842FECD" box="[240,364,260,284]" pageId="7" pageNumber="6" type="paratype">Paratypes</typeStatus>
(40) were all collected at the same locality by various collectors, here listed in alphabetical order: D. Almquist,
<collectionCode id="7BE2AE4339787634FE6C916FF8E6FEEA" box="[433,456,292,315]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="6">R</collectionCode>
. Morris,
<collectionCode id="7BE2AE4339787634FDEE916FFB66FEEA" box="[563,584,292,315]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="6">T</collectionCode>
. Palmer, P. Skelley, D. Stark. Unless otherwise noted,
<typeStatus id="C248882439787634FB3E916FFC70FEED" box="[1251,1374,292,316]" pageId="7" pageNumber="6" type="paratype">paratypes</typeStatus>
were collected at light on the following dates:
<date id="694D104639787634FD4F9108FA36FE8A" box="[658,792,323,347]" pageId="7" pageNumber="6" value="2007-04-10">10-IV-2007</date>
(3M);
<date id="694D104639787634FCBC9108FAC8FE8A" box="[865,998,323,347]" pageId="7" pageNumber="6" value="2008-04-22">22-IV-2008</date>
(8M);
<date id="694D104639787634FBF29108FDB2FE8A" box="[1071,1180,323,347]" pageId="7" pageNumber="6" value="2008-05-08">8-V-2008</date>
(1M);
<date id="694D104639787634FB389108FC75FE8A" box="[1253,1371,323,347]" pageId="7" pageNumber="6" value="2009-04-04">4-IV-2009</date>
(2M);
<date id="694D104639787634FF1D9128F819FEAA" box="[192,311,355,379]" pageId="7" pageNumber="6" value="2009-04-06">6-IV-2009</date>
(1M);
<date id="694D104639787634FE5D9128F8D7FEAA" box="[384,505,355,379]" pageId="7" pageNumber="6" value="2009-04-06">6-IV-2009</date>
, flying at 4:56 pm (1M);
<date id="694D104639787634FCCD9128FAA9FEAA" box="[784,903,355,379]" pageId="7" pageNumber="6" value="2009-04-09">9-IV-2009</date>
(10M);
<date id="694D104639787634FC039128FD4DFEAA" box="[990,1123,355,379]" pageId="7" pageNumber="6" value="2009-04-11">11-IV-2009</date>
(4M);
<date id="694D104639787634FB719128FC1FFEAA" box="[1196,1329,355,379]" pageId="7" pageNumber="6" value="2009-04-12">12-IV-2009</date>
(1M);
<date id="694D104639787634FAA79128F80BFE4A" pageId="7" pageNumber="6" value="2009-04-14">14- IV-2009</date>
, under ground after dusk (1F);
<date id="694D104639787634FD4691C8FA09FE4A" box="[667,807,387,411]" pageId="7" pageNumber="6" value="2009-04-15">15-IV-2009</date>
, mating at dusk (2F, 4M);
<date id="694D104639787634FBBC91C8FDC4FE4A" box="[1121,1258,387,411]" pageId="7" pageNumber="6" value="2009-04-26">26-IV-2009</date>
(2M).
</paragraph>
<paragraph id="1D4C368639787634FF2D91E8FB67FD0A" blockId="7.[192,1441,195,731]" pageId="7" pageNumber="6">
<materialsCitation id="AD9B3CDB39787634FF2D91E8FBE7FE2A" collectionCode="FSCA, ABSC" collectorName="Gainesville, FL &amp; At" county="Lake Placid" location="The" municipality="Archbold Biological Station" pageId="7" pageNumber="6" specimenCount="1" typeStatus="holotype">
<location id="182C605D39787634FF2D91E8F830FE6A" LSID="urn:lsid:plazi:treatment:955A8790397C7634FF1D970EFA16FBEA:182C605D39787634FF2D91E8F830FE6A" box="[240,286,419,443]" county="Lake Placid" municipality="Archbold Biological Station" name="The" pageId="7" pageNumber="6">The</location>
<typeStatus id="C248882439787634FEFF91E8F8A9FE6A" box="[290,391,419,443]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
,
<typeStatus id="C248882439787634FE5291E8F8C5FE6A" box="[399,491,419,443]" pageId="7" pageNumber="6" type="allotype">allotype</typeStatus>
and a series of
<typeStatus id="C248882439787634FD4F91EFFA28FE6D" box="[658,774,420,444]" pageId="7" pageNumber="6" type="paratype">paratypes</typeStatus>
are deposited in the
<location id="182C605D39787634FC3691E8FCB2FE6A" LSID="urn:lsid:plazi:treatment:955A8790397C7634FF1D970EFA16FBEA:182C605D39787634FC3691E8FCB2FE6A" box="[1003,1436,419,443]" county="Lake Placid" municipality="Archbold Biological Station" name="Florida State Collection of Arthropods" pageId="7" pageNumber="6">Florida State Collection of Arthropods</location>
,
<collectorName id="B006535039787634FF1D9188F85AFE0A" box="[192,372,451,475]" pageId="7" pageNumber="6">Gainesville, FL</collectorName>
(
<collectionCode id="7BE2AE4339787634FE5D9188F8E2FE0A" box="[384,460,451,475]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="7" pageNumber="6" type="Museum">FSCA</collectionCode>
).
<collectorName id="B006535039787634FE069188F8D6FE0A" box="[475,504,451,475]" pageId="7" pageNumber="6">At</collectorName>
least
<specimenCount id="0BF5FD0F39787634FDE7918FFBFDFE0D" box="[570,723,452,476]" pageId="7" pageNumber="6" type="generic" typeStatus="paratype">one paratype</specimenCount>
will be deposited in each of the following collections:
<collectingMunicipality id="FD28ACFC39787634FAE89188F8B5FE2A" pageId="7" pageNumber="6">Archbold Biological Station</collectingMunicipality>
,
<collectingCounty id="F42D4E0A39787634FE7891A8FB19FE2A" box="[421,567,483,507]" pageId="7" pageNumber="6">Lake Placid</collectingCounty>
, FL (
<collectionCode id="7BE2AE4339787634FDAF91A8FBEFFE2A" box="[626,705,483,507]" pageId="7" pageNumber="6">ABSC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD9B3CDB39787634FD0E91A8FABEFDCA" collectionCode="V, AVRC" collectorName="Evans &amp; Richmond, VA" location="Virginia Department of Conservation" municipality="Division of Natural Heritage" pageId="7" pageNumber="6" specimenCount="1" typeStatus="holotype">
A.
<collectionCode id="7BE2AE4339787634FD2E91AFFA24FE2A" box="[755,778,484,507]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="6" type="Museum">V</collectionCode>
.
<collectorName id="B006535039787634FCCE91AFFA4CFE2A" box="[787,866,484,507]" pageId="7" pageNumber="6">Evans</collectorName>
,
<location id="182C605D39787634FCB191A8FC1FFE2A" LSID="urn:lsid:plazi:treatment:955A8790397C7634FF1D970EFA16FBEA:182C605D39787634FCB191A8FC1FFE2A" box="[876,1329,483,507]" municipality="Division of Natural Heritage" name="Virginia Department of Conservation" pageId="7" pageNumber="6">Virginia Department of Conservation</location>
and Recreation,
<collectingMunicipality id="FD28ACFC39787634FEFF9248FB59FDCA" box="[290,631,515,539]" pageId="7" pageNumber="6">Division of Natural Heritage</collectingMunicipality>
,
<collectorName id="B006535039787634FDA29248FA05FDCA" box="[639,811,515,539]" pageId="7" pageNumber="6">Richmond, VA</collectorName>
(
<collectionCode id="7BE2AE4339787634FCEA9248FAA6FDCA" box="[823,904,515,539]" pageId="7" pageNumber="6">AVRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD9B3CDB39787634FC459248FCB2FDCA" box="[920,1436,515,539]" collectionCode="BMNH" location="Natural History Museum" municipality="London" pageId="7" pageNumber="6" specimenCount="1" typeStatus="holotype">
<location id="182C605D39787634FC459248FDE7FDCA" LSID="urn:lsid:plazi:treatment:955A8790397C7634FF1D970EFA16FBEA:182C605D39787634FC459248FDE7FDCA" box="[920,1225,515,539]" municipality="London" name="Natural History Museum" pageId="7" pageNumber="6">Natural History Museum</location>
,
<collectingMunicipality id="FD28ACFC39787634FB0F9248FC02FDCA" box="[1234,1324,515,539]" pageId="7" pageNumber="6">London</collectingMunicipality>
(
<collectionCode id="7BE2AE4339787634FAE49248FCBAFDCA" box="[1337,1428,515,539]" pageId="7" pageNumber="6">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD9B3CDB39787634FF1D9268FA40FDEA" box="[192,878,547,571]" collectionCode="CMNC" country="Canada" location="Canadian Museum of Nature" municipality="Ottawa" pageId="7" pageNumber="6" specimenCount="1" typeStatus="holotype">
<location id="182C605D39787634FF1D9268FB07FDEA" LSID="urn:lsid:plazi:treatment:955A8790397C7634FF1D970EFA16FBEA:182C605D39787634FF1D9268FB07FDEA" box="[192,553,547,571]" country="Canada" municipality="Ottawa" name="Canadian Museum of Nature" pageId="7" pageNumber="6">Canadian Museum of Nature</location>
,
<collectingMunicipality id="FD28ACFC39787634FDEE9268FBBCFDEA" box="[563,658,547,571]" pageId="7" pageNumber="6">Ottawa</collectingMunicipality>
,
<collectingCountry id="65E4761639787634FD409268FBD3FDEA" box="[669,765,547,571]" name="Canada" pageId="7" pageNumber="6">Canada</collectingCountry>
(
<collectionCode id="7BE2AE4339787634FCD69268FA48FDEA" box="[779,870,547,571]" pageId="7" pageNumber="6">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD9B3CDB39787634FCA4926FFA24FD8A" collectionCode="CA, DALC" collectorName="D. A. LaRue" country="Canada" county="Riverside" location="Entomology Research Museum" municipality="University of California" pageId="7" pageNumber="6" specimenCount="1" typeStatus="holotype">
<collectorName id="B006535039787634FCA4926FFD3EFDEA" box="[889,1040,547,571]" pageId="7" pageNumber="6">D. A. LaRue</collectorName>
,
<location id="182C605D39787634FBC69268FCB2FDEA" LSID="urn:lsid:plazi:treatment:955A8790397C7634FF1D970EFA16FBEA:182C605D39787634FBC69268FCB2FDEA" box="[1051,1436,547,571]" country="Canada" county="Riverside" municipality="University of California" name="Entomology Research Museum" pageId="7" pageNumber="6">Entomology Research Museum</location>
,
<collectingMunicipality id="FD28ACFC39787634FF1D9208F8C4FD8A" box="[192,490,579,603]" pageId="7" pageNumber="6">University of California</collectingMunicipality>
,
<collectingCounty id="F42D4E0A39787634FE289208FB40FD8A" box="[501,622,579,603]" pageId="7" pageNumber="6">Riverside</collectingCounty>
,
<collectionCode id="7BE2AE4339787634FDA49208FB8EFD8A" box="[633,672,579,603]" country="USA" name="Chicago Academy of Sciences" pageId="7" pageNumber="6" type="Museum">CA</collectionCode>
(
<collectionCode id="7BE2AE4339787634FD729208FA2CFD8A" box="[687,770,579,603]" pageId="7" pageNumber="6">DALC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD9B3CDB39787634FCC8920FFDBAFD8A" box="[789,1172,579,603]" collectionCode="DSIC" collectorName="D. Stark" country="Canada" location="Lakeland" pageId="7" pageNumber="6" specimenCount="1" typeStatus="holotype">
<collectorName id="B006535039787634FCC8920FFAAFFD8A" box="[789,897,579,603]" pageId="7" pageNumber="6">D. Stark</collectorName>
,
<location id="182C605D39787634FC509208FD28FD8A" LSID="urn:lsid:plazi:treatment:955A8790397C7634FF1D970EFA16FBEA:182C605D39787634FC509208FD28FD8A" box="[909,1030,579,603]" country="Canada" name="Lakeland" pageId="7" pageNumber="6">Lakeland</location>
, FL (
<collectionCode id="7BE2AE4339787634FB999208FDA2FD8A" box="[1092,1164,579,603]" pageId="7" pageNumber="6">DSIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD9B3CDB39787634FB7D920FFB61FDAA" collectionCode="DERC" collectorName="D. E. Russell" country="Canada" location="Miami University" municipality="Oxford" pageId="7" pageNumber="6" specimenCount="1" typeStatus="holotype">
<collectorName id="B006535039787634FB7D920FFC6DFD8A" box="[1184,1347,579,603]" pageId="7" pageNumber="6">D. E. Russell</collectorName>
,
<location id="182C605D39787634FA929208F866FDAA" LSID="urn:lsid:plazi:treatment:955A8790397C7634FF1D970EFA16FBEA:182C605D39787634FA929208F866FDAA" country="Canada" municipality="Oxford" name="Miami University" pageId="7" pageNumber="6">Miami University</location>
,
<collectingMunicipality id="FD28ACFC39787634FE899228F882FDAA" box="[340,428,611,635]" pageId="7" pageNumber="6">Oxford</collectingMunicipality>
, OH (
<collectionCode id="7BE2AE4339787634FE299228FB66FDAA" box="[500,584,611,635]" pageId="7" pageNumber="6">DERC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="AD9B3CDB39787634FD81922FFD6AFDAA" box="[604,1092,611,635]" collectionCode="JZIC" country="Czech Republic" location="Zidek" pageId="7" pageNumber="6" specimenCount="1" stateProvince="Praha" typeStatus="holotype">
J.
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and
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.
</paragraph>
</subSubSection>
<subSubSection id="55E9650D39787634FF1D9348FA16FBEA" pageId="7" pageNumber="6" type="etymology">
<paragraph id="1D4C368639787634FF1D9348FBCEFC8A" blockId="7.[192,1440,771,859]" pageId="7" pageNumber="6">
<emphasis id="2F87EA9439787634FF1D9348F87CFCCA" bold="true" box="[192,338,771,795]" pageId="7" pageNumber="6">Etymology</emphasis>
. Named for Donna Stark, who first collected this beetle, attempted to identify it, and brought it to our attention (see
<bibRefCitation id="79624B7739787634FE3D9368FBADFCEA" author="Palmer, T." box="[480,643,803,827]" pageId="7" pageNumber="6" pagination="5" refId="ref9037" refString="Palmer, T. 2009. A rare find: Amateur collector finds a new beetle species in Polk. The Ledger, June 11, 2009: A 1, A 5. [The Ledger is Polk County's Newspaper, published in Lakeland, FL. This article was also available online through www. theledger. com, viewed June 11, 2009]" type="journal article" year="2009">Palmer 2009</bibRefCitation>
). Survey efforts such as hers are greatly needed to discover and document our indigenous invertebrate fauna.
</paragraph>
<paragraph id="1D4C368639787634FF1D93C8FA16FBEA" blockId="7.[192,1440,899,1083]" pageId="7" pageNumber="6">
<emphasis id="2F87EA9439787634FF1D93C8F887FC4A" bold="true" box="[192,425,899,923]" pageId="7" pageNumber="6">Common Names.</emphasis>
<bibRefCitation id="79624B7739787634FE6C93C8FA3AFC4A" author="Woodruff, R. E. &amp; M. Deyrup" box="[433,788,899,924]" pageId="7" pageNumber="6" pagination="368 - 455" refId="ref9456" refString="Woodruff, R. E., and M. Deyrup. 1994. Order Coleoptera, Family Scarabaeidae, Scarab Beetles. p. 368 - 455. In: M. Deyrup and R. Franz (eds.). Rare and endangered biota of Florida. Volume IV. Invertebrates. University of Florida Press; Gainesville, Florida. 798 p." type="book chapter" year="1994">Woodruff and Deyrup (1994)</bibRefCitation>
called “
<taxonomicName id="DAF34D0539787634FCA593CFFD3AFC4A" authorityName="Cartwright" authorityYear="1939" box="[888,1044,899,923]" class="Insecta" family="Melolonthidae" genus="Polyphylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="2F87EA9439787634FCA593CFFD3AFC4A" box="[888,1044,899,923]" italics="true" pageId="7" pageNumber="6">P. pubescens</emphasis>
</taxonomicName>
” the “panhandle beach scarab.” Unfortunately, this name refers to the species later recognized and named
<taxonomicName id="DAF34D0539787634FB9293EFFCB2FC6A" authority="Skelley (2004)" authorityName="Skelley" authorityYear="2003" box="[1103,1436,931,955]" class="Insecta" family="Melolonthidae" genus="Polyphylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="woodruffi">
<emphasis id="2F87EA9439787634FB9293EFFDCBFC6A" box="[1103,1253,931,955]" italics="true" pageId="7" pageNumber="6">P. woodruffi</emphasis>
<bibRefCitation id="79624B7739787634FB3693E8FCB2FC6A" author="Skelley, P. E." box="[1259,1436,931,955]" pageId="7" pageNumber="6" pagination="129 - 156" refId="ref9221" refString="Skelley, P. E. 2004. Review of the tribe Melolonthini in the southeastern United States (Coleoptera: Scarabaeidae: Melolonthinae). Insecta Mundi 17 (3 - 4) [2003]: 129 - 156." type="journal article" year="2004">Skelley (2004)</bibRefCitation>
</taxonomicName>
. True
<taxonomicName id="DAF34D0539787634FEDD938FF8B6FC0A" authorityName="Cartwright" authorityYear="1939" box="[256,408,963,987]" class="Insecta" family="Melolonthidae" genus="Polyphylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="2F87EA9439787634FEDD938FF8B6FC0A" box="[256,408,963,987]" italics="true" pageId="7" pageNumber="6">P. pubescens</emphasis>
</taxonomicName>
is not known to occur on beaches and is here named the “Eglin uplands scarab”. Given the immediate interest
<taxonomicName id="DAF34D0539787634FE0193AFFB7AFC2A" authorityName="Skelley" authorityYear="2009" box="[476,596,995,1019]" class="Insecta" family="Melolonthidae" genus="Polyphylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="starkae">
<emphasis id="2F87EA9439787634FE0193AFFB7AFC2A" box="[476,596,995,1019]" italics="true" pageId="7" pageNumber="6">P. starkae</emphasis>
</taxonomicName>
has raised among naturalists, conservation groups and other political entities in
<collectingRegion id="DF37F86439787634FE9F9448F88EFBCA" box="[322,416,1027,1051]" country="United States of America" name="Florida" pageId="7" pageNumber="6">Florida</collectingRegion>
, it is fitting to propose a common name for this new species. Thus, I am proposing
<taxonomicName id="DAF34D0539787634FA5A944FF835FBEA" authorityName="Skelley" authorityYear="2009" class="Insecta" family="Melolonthidae" genus="Polyphylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="starkae">
<emphasis id="2F87EA9439787634FA5A944FF835FBEA" italics="true" pageId="7" pageNumber="6">P. starkae</emphasis>
</taxonomicName>
be known as the “Auburndale scrub scarab.”
</paragraph>
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</treatment>
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