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<document id="5ECD731C3D68A5A389DAFF9A3249F670" ID-DOI="10.5281/zenodo.10831085" ID-ISSN="1942-1354" ID-Zenodo-Dep="10831085" ID-ZooBank="C1D5B819-A964-4679-B090-84CDBBC59D6A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1688389961601" checkinUser="felipe" docAuthor="Otto, Robert L., Muona, Jyrki &amp; Córdoba-Alfaro, Jim" docDate="2023" docId="954287CDB71DFFA469F0FA3747DA5949" docLanguage="en" docName="InsectaMundi.2023.0991.1-36.pdf" docOrigin="Insecta Mundi 2023 (991)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Onichodon rufus Otto, Muona and Cordoba-Alfaro 2023, new species" docType="treatment" docVersion="4" lastPageNumber="18" masterDocId="697BFFB5B70FFFB76930FFD643695A4D" masterDocTitle="A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama" masterLastPageNumber="36" masterPageNumber="1" pageNumber="17" updateTime="1710774778166" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="644BF948221E6F35445F89E341F1AE7F">A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama</mods:title>
</mods:titleInfo>
<mods:name id="8709EC8547304252DE5BD9791BCC13BA" type="personal">
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<mods:namePart id="B932B72F68FD82BE2FDF5589195D827F">Otto, Robert L.</mods:namePart>
<mods:affiliation id="D96C7B71E9E0C886AAD671C7936C0DA5">W 4806 Chrissie Circle, Shawano, WI 54166, U. S. A.</mods:affiliation>
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<mods:namePart id="149EAF08E2FB6461B9389B484EB7CA4A">Muona, Jyrki</mods:namePart>
<mods:affiliation id="2FB4C2411726F4053FE9E5D6439F6B47">Finnish Museum of Natural History, Zoology Unit PO Box 17, University of Helsinki, Finland FIN- 00014</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="DE84E303515731CC4CDF622B19705843">Córdoba-Alfaro, Jim</mods:namePart>
<mods:affiliation id="665B1D351EF3F3AED7E82E4000ED3568">Insectopia Insect Museum, BioSur Foundation Puerto Jiménez, Península de Osa, Costa Rica jim. cordoba @ biosurcorcovado. org</mods:affiliation>
</mods:name>
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<mods:title id="2DBB5679B20E4E4E22E3213013AB467B">Insecta Mundi</mods:title>
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<mods:part id="05DD0048FCFE16277813405813E2760F">
<mods:date id="1EF4E90FA3A368E5A0CDF57CDA31A5EC">2023</mods:date>
<mods:detail id="9C6C48188B83309177BAE16A942991D8" type="pubDate">
<mods:number id="C376544578B9019F7969C1693B366300">2023-05-26</mods:number>
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<mods:detail id="52AF452DD801B1362DB15FB0DB3F170C" type="volume">
<mods:number id="0AEF9AA4D48CBAF6A58359CB8B5772E8">2023</mods:number>
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<mods:detail id="7406246B10A54E0AD129C93D9D84BCCA" type="issue">
<mods:number id="CC52F31A76643AAE5AE63E244D20AE2B">991</mods:number>
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<mods:classification id="D174D334AC4121B768FC28E5ECF11141">journal article</mods:classification>
<mods:identifier id="821FE5B47B323A59662222257136D04B" type="DOI">10.5281/zenodo.10831085</mods:identifier>
<mods:identifier id="F3BEDB0F187EA5D845932EB91A014C1A" type="ISSN">1942-1354</mods:identifier>
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<treatment id="954287CDB71DFFA469F0FA3747DA5949" ID-DOI="http://doi.org/10.5281/zenodo.8109093" ID-Zenodo-Dep="8109093" LSID="urn:lsid:plazi:treatment:954287CDB71DFFA469F0FA3747DA5949" httpUri="http://treatment.plazi.org/id/954287CDB71DFFA469F0FA3747DA5949" lastPageId="19" lastPageNumber="18" pageId="18" pageNumber="17">
<subSubSection id="55F16550B71DFFA569F0FA3740995FB2" box="[192,1008,1505,1535]" pageId="18" pageNumber="17" type="nomenclature">
<paragraph id="1D5436DBB71DFFA569F0FA3740995FB2" blockId="18.[192,1008,1505,1567]" box="[192,1008,1505,1535]" pageId="18" pageNumber="17">
<heading id="461C81B7B71DFFA569F0FA3740995FB2" box="[192,1008,1505,1535]" fontSize="12" level="2" pageId="18" pageNumber="17" reason="2">
<emphasis id="2F9FEAC9B71DFFA569F0FA3740995FB2" bold="true" box="[192,1008,1505,1535]" pageId="18" pageNumber="17">
<taxonomicName id="DAEB4D58B71DFFA569F0FA3740255FB3" authority="Otto, Muona and Cordoba-Alfaro" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,844,1505,1535]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="rufus" status="sp. nov.">
<emphasis id="2F9FEAC9B71DFFA569F0FA3742FD5FB3" bold="true" box="[192,404,1505,1534]" italics="true" pageId="18" pageNumber="17">Onichodon rufus</emphasis>
Otto, Muona and Córdoba-Alfaro
</taxonomicName>
,
<taxonomicNameLabel id="34AC57B2B71DFFA56A6BFA3440995FB2" box="[859,1008,1506,1535]" pageId="18" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B71DFFA569F0F9D342345C52" box="[192,349,1541,1567]" pageId="18" pageNumber="17" type="description">
<paragraph id="1D5436DBB71DFFA569F0F9D342345C52" blockId="18.[192,1008,1505,1567]" box="[192,349,1541,1567]" pageId="18" pageNumber="17">
<figureCitation id="85D02A5EB71DFFA569F0F9D342345C52" box="[192,349,1541,1567]" captionStart="Figures 3134" captionStartId="18.[192,279,656,682]" captionTargetBox="[353,1279,213,626]" captionTargetId="figure-451@18.[353,1279,213,626]" captionTargetPageId="18" captionText="Figures 3134. Costa Rican macraulacine eucnemids. 31) Onichodon confluentus sp. nov., male holotype, ventral habitus. 32) Onichodon confluentus sp. nov., female allotype, dorsal habitus. 33) Onichodon rufus sp. nov., dorsal habitus. 34) Onichodon rufus sp. nov., ventral habitus. (Scale: 3132 = 5.0 mm; 3334 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831099" httpUri="https://zenodo.org/record/10831099/files/figure.png" pageId="18" pageNumber="17">Figures 3334</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B71DFFA569F0F9F946745C27" pageId="18" pageNumber="17" type="diagnosis">
<paragraph id="1D5436DBB71DFFA569F0F9F946745C27" blockId="18.[192,1440,1583,1643]" pageId="18" pageNumber="17">
<emphasis id="2F9FEAC9B71DFFA569F0F9F942555C04" bold="true" box="[192,316,1583,1609]" pageId="18" pageNumber="17">Diagnosis.</emphasis>
The smaller size along with the reddish coloration with its habitus will distinguish the new species from
<taxonomicName id="DAEB4D58B71DFFA569CEF98742F25C27" authority="Otto &amp; Muona &amp; Córdoba-Alfaro, 2023" authorityName="Otto &amp; Muona &amp; Córdoba-Alfaro" authorityYear="2023" box="[254,411,1616,1643]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="confluentus" status="sp. nov.">
<emphasis id="2F9FEAC9B71DFFA569CEF98742F25C27" box="[254,411,1616,1643]" italics="true" pageId="18" pageNumber="17">O. confluentus</emphasis>
</taxonomicName>
<emphasis id="2F9FEAC9B71DFFA56892F98741405C27" bold="true" box="[418,553,1616,1643]" pageId="18" pageNumber="17">
<taxonomicNameLabel id="34AC57B2B71DFFA56892F98741405C27" box="[418,553,1616,1643]" pageId="18" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</emphasis>
and any other misplaced members of
<taxonomicName id="DAEB4D58B71DFFA56AE3F987474B5C27" box="[979,1058,1617,1642]" class="Insecta" family="Eucnemidae" genus="Fornax" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B71DFFA56AE3F987474B5C27" box="[979,1058,1617,1642]" italics="true" pageId="18" pageNumber="17">Fornax</emphasis>
</taxonomicName>
present in
<collectingCountry id="65FC764BB71DFFA56DAEF98746715C27" box="[1182,1304,1616,1643]" name="Costa Rica" pageId="18" pageNumber="17">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B71DFFA569F0F9AC424C5CB4" pageId="18" pageNumber="17" type="materials_examined">
<paragraph id="1D5436DBB71DFFA569F0F9AC424C5CB4" blockId="18.[190,1441,1658,1785]" pageId="18" pageNumber="17">
<emphasis id="2F9FEAC9B71DFFA569F0F9AC42A15CD9" bold="true" box="[192,456,1658,1684]" pageId="18" pageNumber="17">Type material. Female</emphasis>
<materialsCitation id="AD833C86B71DFFA568E1F9AC424C5CB4" collectingDate="1990-04-25" collectionCode="R, INBC" collectorName="H. A. Hespenheide &amp; L. Otto" country="Costa Rica" latitude="10.433333" location="Pto. Viejo" longLatPrecision="1298" longitude="-84.01667" pageId="18" pageNumber="17" specimenCount="1" specimenCount-female="1" stateProvince="Heredia" typeStatus="holotype">
<emphasis id="2F9FEAC9B71DFFA568E1F9AC41515CD9" bold="true" box="[465,568,1658,1684]" pageId="18" pageNumber="17">
<typeStatus id="C2508879B71DFFA568E1F9AC41515CD9" box="[465,568,1658,1684]" pageId="18" pageNumber="17" type="holotype">holotype</typeStatus>
</emphasis>
: “
<collectingCountry id="65FC764BB71DFFA56B61F9AC419E5CD9" box="[593,759,1658,1684]" name="Costa Rica" pageId="18" pageNumber="17">COSTA RICA</collectingCountry>
:
<collectingRegion id="DF2FF839B71DFFA56A33F9AC40365CD9" box="[771,863,1658,1684]" country="Costa Rica" name="Heredia" pageId="18" pageNumber="17">Heredia</collectingRegion>
; pr: La Selva Bio. Sta.;
<quantity id="DA139B3EB71DFFA56D5AF9AC47CC5CD9" box="[1130,1189,1658,1684]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="17" unit="km" value="3.0">3 km</quantity>
S
<location id="18346000B71DFFA56DF5F9AD46505CD9" LSID="urn:lsid:plazi:treatment:954287CDB71DFFA469F0FA3747DA5949:18346000B71DFFA56DF5F9AD46505CD9" box="[1221,1337,1658,1685]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" name="Pto. Viejo" pageId="18" pageNumber="17" stateProvince="Heredia">Pto. Viejo</location>
;
<geoCoordinate id="78DF501CB71DFFA56C76F9AC46C95CD8" box="[1350,1440,1658,1685]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="18" pageNumber="17" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="78DF501CB71DFFA569F0F94A42485CFB" box="[192,289,1692,1718]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="18" pageNumber="17" precision="925" value="-84.01667">84°01W</geoCoordinate>
” / “
<date id="6955101BB71DFFA5687FF94D42A05CFB" box="[335,457,1691,1718]" pageId="18" pageNumber="17" value="1990-04-25">
<collectingDate id="7911E9F3B71DFFA5687FF94D42A05CFB" box="[335,457,1691,1718]" pageId="18" pageNumber="17" value="1990-04-25">25.IV.1990</collectingDate>
</date>
;
<collectorName id="B01E530DB71DFFA568E5F94A41CD5CFB" box="[469,676,1692,1718]" pageId="18" pageNumber="17">H.A. Hespenheide</collectorName>
” (“25”, “IV” and “0” handwritten on label) / “
<emphasis id="2F9FEAC9B71DFFA56D9AF94D462F5CF8" bold="true" box="[1194,1350,1691,1717]" pageId="18" pageNumber="17">
<typeStatus id="C2508879B71DFFA56D9AF94D46285CF8" box="[1194,1345,1691,1717]" pageId="18" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B71DFFA56C64F94A439E5C9A" authorityName="Newman" authorityYear="1838" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B71DFFA56C64F94A439E5C9A" italics="true" pageId="18" pageNumber="17">Onichodon</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B71DFFA56832F96B42555C9A" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[258,316,1725,1751]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="2F9FEAC9B71DFFA56832F96B42555C9A" box="[258,316,1725,1751]" italics="true" pageId="18" pageNumber="17">rufus</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B71DFFA56870F917423C5C9A" box="[320,341,1729,1751]" pageId="18" pageNumber="17" type="female"></specimenCount>
; Otto, Muona &amp; Córdoba-Alfaro; Det.
<collectionCode id="7BFAAE1EB71DFFA56A30F968407D5C95" box="[768,788,1726,1752]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="17">R</collectionCode>
.
<collectorName id="B01E530DB71DFFA56A28F96840005C9A" box="[792,873,1725,1752]" pageId="18" pageNumber="17">L. Otto</collectorName>
; 2019” (red printed label).
<typeStatus id="C2508879B71DFFA56DBCF96B479D5C9A" box="[1164,1268,1725,1751]" pageId="18" pageNumber="17" type="holotype">Holotype</typeStatus>
to be deposited in
<collectionCode id="7BFAAE1EB71DFFA569ECF90942485CB4" box="[220,289,1759,1785]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="18" pageNumber="17">INBC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B71DFFA469F0F8DE40375828" lastPageId="19" lastPageNumber="18" pageId="18" pageNumber="17" type="description">
<paragraph id="1D5436DBB71DFFA469F0F8DE40375828" blockId="18.[192,1440,1800,1928]" lastBlockId="19.[191,1441,213,613]" lastPageId="19" lastPageNumber="18" pageId="18" pageNumber="17">
<emphasis id="2F9FEAC9B71DFFA569F0F8DE41765D6E" bold="true" box="[192,543,1800,1827]" pageId="18" pageNumber="17">
Description. Female
<typeStatus id="C2508879B71DFFA56888F8DF41765D6E" box="[440,543,1801,1827]" pageId="18" pageNumber="17" type="holotype">holotype</typeStatus>
</emphasis>
: Length, 6.0 mm. Width,
<quantity id="DA139B3EB71DFFA56A7DF8DE40C05D6E" box="[845,937,1800,1827]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="17" unit="mm" value="1.5">1.5 mm</quantity>
. Body elongate; dark reddish; antennae and legs dark reddish; head, pronotum and elytra clothed with short, recumbent yellowish setae (
<figureCitation id="85D02A5EB71DFFA56DE7F8FC462E5D09" box="[1239,1351,1834,1860]" captionStart="Figures 3134" captionStartId="18.[192,279,656,682]" captionTargetBox="[353,1279,213,626]" captionTargetId="figure-451@18.[353,1279,213,626]" captionTargetPageId="18" captionText="Figures 3134. Costa Rican macraulacine eucnemids. 31) Onichodon confluentus sp. nov., male holotype, ventral habitus. 32) Onichodon confluentus sp. nov., female allotype, dorsal habitus. 33) Onichodon rufus sp. nov., dorsal habitus. 34) Onichodon rufus sp. nov., ventral habitus. (Scale: 3132 = 5.0 mm; 3334 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831099" httpUri="https://zenodo.org/record/10831099/files/figure.png" pageId="18" pageNumber="17">Figure 33</figureCitation>
).
<emphasis id="2F9FEAC9B71DFFA56C6DF8FC46F35D09" bold="true" box="[1373,1434,1834,1860]" pageId="18" pageNumber="17">Head</emphasis>
: Surface somewhat shiny, subspherical with crowded punctures; frons convex without carina or fovea; interantennal carina present; apical margin of frontoclypeal region weakly trilobed, more than 2 times wider than base; mandibles stout, bidentate, densely punctate.
<emphasis id="2F9FEAC9B71CFFA46BFFFF03405D5AA2" bold="true" box="[719,820,213,239]" pageId="19" pageNumber="18">Antenna</emphasis>
: Weakly serriform from flagellomeres IIX, attaining about 1/3the length of the body, just beyond pronotal hind angle; flagellomere I longer than wide, longer than II; flagellomeres IIVIII each slightly longer than wide, sub-equal; flagellomere IX slightly longer than VIII.
<emphasis id="2F9FEAC9B71CFFA46C7EFECC43995B1B" bold="true" pageId="19" pageNumber="18">Pronotum</emphasis>
: Surface somewhat shiny, with deep, crowded punctures; quadrate, with well-developed, sharp hind angles; sides parallel-sided at basal 2/3, arcuate at apical 1/3; disc convex, without carina or fovea; base sinuous.
<emphasis id="2F9FEAC9B71CFFA46C09FE8B426F5BD4" bold="true" pageId="19" pageNumber="18">Scutellar shield</emphasis>
: Slightly longer than wide, sub-triangular, shallowly punctate, setose and distally rounded.
<emphasis id="2F9FEAC9B71CFFA46C27FEA946345BD4" bold="true" box="[1303,1373,383,409]" pageId="19" pageNumber="18">Elytra</emphasis>
: Very shallow striae present; interstices slightly elevated; surfaces shiny, transversely rugose through much of the elytra.
<emphasis id="2F9FEAC9B71CFFA469DDFE15424B5B90" bold="true" box="[237,290,451,477]" pageId="19" pageNumber="18">Legs</emphasis>
: First tarsomere as long as the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres IIII simple; metatarsomere IV excavated; metatarsomere V short with basally toothed claws.
<emphasis id="2F9FEAC9B71CFFA468C4FDD1412B586C" bold="true" box="[500,578,519,545]" pageId="19" pageNumber="18">Venter</emphasis>
(
<figureCitation id="85D02A5EB71CFFA46B7FFDD141D6586C" box="[591,703,519,545]" captionStart="Figures 3134" captionStartId="18.[192,279,656,682]" captionTargetBox="[353,1279,213,626]" captionTargetId="figure-451@18.[353,1279,213,626]" captionTargetPageId="18" captionText="Figures 3134. Costa Rican macraulacine eucnemids. 31) Onichodon confluentus sp. nov., male holotype, ventral habitus. 32) Onichodon confluentus sp. nov., female allotype, dorsal habitus. 33) Onichodon rufus sp. nov., dorsal habitus. 34) Onichodon rufus sp. nov., ventral habitus. (Scale: 3132 = 5.0 mm; 3334 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831099" httpUri="https://zenodo.org/record/10831099/files/figure.png" pageId="19" pageNumber="18">Figure 34</figureCitation>
): Closely punctate, with short, recumbent yellowish colored setae; hypomeron with basally opened, lateral antennal grooves; metepisterna parallel-sided; elytral epipleura punctate; metacoxal plates medially 3.06.0 times wider than laterally.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B71CFFA469F0FDA2424958FD" pageId="19" pageNumber="18" type="distribution">
<paragraph id="1D5436DBB71CFFA469F0FDA2424958FD" blockId="19.[192,1440,628,688]" pageId="19" pageNumber="18">
<emphasis id="2F9FEAC9B71CFFA469F0FDA2423258C3" bold="true" box="[192,347,628,654]" pageId="19" pageNumber="18">Distribution.</emphasis>
This eucnemid species is known from the
<typeStatus id="C2508879B71CFFA46A78FDA340C358C2" box="[840,938,629,655]" pageId="19" pageNumber="18" type="holotype">holotype</typeStatus>
taken from La Selva within the province of
<collectingRegion id="DF2FF839B71CFFA469F0FD40427558FD" box="[192,284,662,688]" country="Costa Rica" name="Heredia" pageId="19" pageNumber="18">Heredia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B71CFFA469F0FD16466C5897" box="[192,1285,704,731]" pageId="19" pageNumber="18" type="biology_ecology">
<paragraph id="1D5436DBB71CFFA469F0FD16466C5897" blockId="19.[192,1285,704,731]" box="[192,1285,704,731]" pageId="19" pageNumber="18">
<emphasis id="2F9FEAC9B71CFFA469F0FD1642495897" bold="true" box="[192,288,704,730]" pageId="19" pageNumber="18">Biology.</emphasis>
No biological information is known for the new species. Larvae and pupae are unknown.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B71CFFA469F0FD3D47DA5949" box="[192,1203,746,773]" pageId="19" pageNumber="18" type="etymology">
<paragraph id="1D5436DBB71CFFA469F0FD3D47DA5949" blockId="19.[192,1203,746,773]" box="[192,1203,746,773]" pageId="19" pageNumber="18">
<emphasis id="2F9FEAC9B71CFFA469F0FD3D422D5948" bold="true" box="[192,324,747,773]" pageId="19" pageNumber="18">Etymology.</emphasis>
The specific epithet is derived from its overall reddish coloration of its habitus.
</paragraph>
</subSubSection>
</treatment>
</document>