373 lines
48 KiB
XML
373 lines
48 KiB
XML
<document id="B0E17449D354DDF676D4250BDDBB54FC" ID-DOI="10.1080/00222933.2012.763053" ID-ISSN="1464-5262" ID-Zenodo-Dep="10091554" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628873311945" checkinUser="felipe" docAuthor="Asenjo, Angelico & Ribeiro-Costa, Cibele S." docDate="2013" docId="953B87C00E562D29FEF5A265BCFADF72" docLanguage="en" docName="JNatHist.47.19-20.1257-1285.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 47 (19 - 20)" docSource="https://www.mendeley.com/catalogue/70469a15-3e13-3578-a8ef-927a79c5ef50/" docStyle="DocumentStyle:CD936EBDDF5274E0936345450CC2DAE4.5:JNatHist.2013-2015.journal_article.1cover" docStyleId="CD936EBDDF5274E0936345450CC2DAE4" docStyleName="JNatHist.2013-2015.journal_article.1cover" docStyleVersion="5" docTitle="Leptochirus (Tropiochirus) pachoensis Bernhauer 1917" docType="treatment" docVersion="3" lastPageNumber="1274" masterDocId="6902FFB80E462D3BFFCAA12BBE73DE75" masterDocTitle="Revision of the neotropical subgenus Tropiochirus of the genus Leptochirus Germar 1824 (Coleoptera: Staphylinidae: Osoriinae)" masterLastPageNumber="1285" masterPageNumber="1257" pageNumber="1272" updateTime="1699542363201" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C9AB1CE950C550A8A850A8845503EE72">Revision of the neotropical subgenus Tropiochirus of the genus Leptochirus Germar 1824 (Coleoptera: Staphylinidae: Osoriinae)</mods:title>
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<mods:namePart id="135B9384A6F5674ACB34A46CAFFD246C">Asenjo, Angelico</mods:namePart>
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<mods:namePart id="3492F7259A275ECAD1A4D0A602C8D4F9">Ribeiro-Costa, Cibele S.</mods:namePart>
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<mods:title id="E259430F51538CAFA228825FDE5CA9B0">Journal of Natural History</mods:title>
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<mods:part id="199033CCF84BE180EF19354F7F481729">
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<mods:date id="21E94AABEB446552BC8C222CA362DC92">2013</mods:date>
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<mods:title id="1CE0795DD5B4DC481E88A643F738FD7A">J. Nat. Hist.</mods:title>
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<mods:number id="97132487FEBBA94ACA735325F4A00FB1">2013-03-22</mods:number>
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<mods:number id="F5412294C35A28CD43B15FBA0D166F62">47</mods:number>
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<mods:number id="7DBEAA972EA571E17413FA4DBEA1DCA9">19 - 20</mods:number>
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<mods:classification id="7A19E09EA35AFD2B721271099C14A9F4">journal article</mods:classification>
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<treatment id="953B87C00E562D29FEF5A265BCFADF72" LSID="urn:lsid:plazi:treatment:953B87C00E562D29FEF5A265BCFADF72" httpUri="http://treatment.plazi.org/id/953B87C00E562D29FEF5A265BCFADF72" lastPageId="18" lastPageNumber="1274" pageId="16" pageNumber="1272">
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<subSubSection id="5588655D0E562D2BFEF5A265BDECDD13" box="[319,927,846,870]" pageId="16" pageNumber="1272" type="nomenclature">
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<paragraph id="1D2D36D60E562D2BFEF5A265BDECDD13" blockId="16.[319,927,846,902]" box="[319,927,846,870]" pageId="16" pageNumber="1272">
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<heading id="466581BA0E562D2BFEF5A265BDECDD13" box="[319,927,846,870]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="1272" reason="2">
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<taxonomicName id="DA924D550E562D2BFEF5A265BDECDD13" ID-CoL="3TLRP" authority="Bernhauer, 1917" authorityName="Bernhauer" authorityYear="1917" box="[319,927,846,870]" class="Insecta" family="Staphylinidae" genus="Leptochirus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1272" phylum="Arthropoda" rank="species" species="pachoensis" subGenus="Tropiochirus">
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<emphasis id="2FE6EAC40E562D2BFEF5A265BC92DD13" bold="true" box="[319,737,846,870]" pageId="16" pageNumber="1272">
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<emphasis id="2FE6EAC40E562D2BFEF5A265BFB1DD13" bold="true" box="[319,450,846,870]" italics="true" pageId="16" pageNumber="1272">Leptochirus</emphasis>
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(
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<emphasis id="2FE6EAC40E562D2BFE18A265BC2EDD13" bold="true" box="[466,605,846,870]" italics="true" pageId="16" pageNumber="1272">Tropiochirus</emphasis>
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)
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<emphasis id="2FE6EAC40E562D2BFDA6A265BC92DD13" bold="true" box="[620,737,846,870]" italics="true" pageId="16" pageNumber="1272">pachoensis</emphasis>
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</emphasis>
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<bibRefCitation id="79034B270E562D2BFD22A265BDECDD13" author="Bernhauer M" box="[744,927,846,870]" pageId="16" pageNumber="1272" pagination="49 - 53" refId="ref12682" refString="Bernhauer M. 1917. Neue Arten der Gattungen Piestus, Leptochirus und Conosoma aus Sudamerika. Neue Beitr Syst Insektenk. 1: 45 - 48, 49 - 53." type="journal article" year="1917">Bernhauer, 1917</bibRefCitation>
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</taxonomicName>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="5588655D0E562D2BFE17A245BD72DDF3" box="[477,769,878,902]" pageId="16" pageNumber="1272" type="description">
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<paragraph id="1D2D36D60E562D2BFE17A245BD72DDF3" blockId="16.[319,927,846,902]" box="[477,769,878,902]" pageId="16" pageNumber="1272">
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(
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<figureCitation id="85A92A530E562D2BFE2FA245BC10DDF3" box="[485,611,878,902]" captionStart="Figure 2" captionStartId="5.[211,278,1629,1651]" captionTargetBox="[238,1145,161,1595]" captionTargetId="figure-5@5.[238,1145,161,1596]" captionTargetPageId="5" captionText="Figure 2. Head and protorax of Leptochirus (Tropiochirus). (A) L. (T.) helleri; (B) L. (T.) marina sp. nov. holotype; (C) L. (T.) mery sp. nov. holotype; (D) L. (T.) milton sp. nov. holotype; (E) L. (T.) pachoensis; (F) L. (T.) proteus; (G) L. (T.) venezuelanus; (H) L. (T.) yunganus; (I) L. (T.) zischkaianus. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10091558" httpUri="https://zenodo.org/record/10091558/files/figure.png" pageId="16" pageNumber="1272">Figures 2E</figureCitation>
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,
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<figureCitation id="85A92A530E562D2BFDA4A245BCE3DDF3" box="[622,656,878,902]" captionStart="Figure 3" captionStartId="6.[143,210,1078,1100]" captionTargetBox="[145,1102,167,1045]" captionTargetId="figure-6@6.[145,1102,167,1045]" captionTargetPageId="6" captionText="Figure 3. Left and right mandible, dorsal view. (A) Leptochirus (Tropiochirus) helleri; (B) L. (T.) marina sp. nov. holotype; (C) L. (T.) mery sp. nov. holotype; (D) L. (T.) milton sp. nov. holotype; (E) L. (T.) pachoensis; (F) L. (T.) proteus; (G) L. (T.) venezuelanus; (H) L. (T.) yunganus; (I) L. (T.) zischkaianus. Scale bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10091560" httpUri="https://zenodo.org/record/10091560/files/figure.png" pageId="16" pageNumber="1272">3E</figureCitation>
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,
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||
<figureCitation id="85A92A530E562D2BFD51A245BCD9DDF3" box="[667,682,878,902]" captionStart="Figure 5" captionStartId="8.[143,210,1486,1508]" captionTargetBox="[145,1102,169,1452]" captionTargetId="figure-6@8.[145,1102,169,1453]" captionTargetPageId="8" captionText="Figure 5. Distribution map of Leptochirus (Tropiochirus) helleri, L. (T.) marina sp. nov., L. (T.) mery sp. nov., L. (T.) milton sp. nov., and L. (T.) pachoensis." figureDoi="http://doi.org/10.5281/zenodo.10091562" httpUri="https://zenodo.org/record/10091562/files/figure.png" pageId="16" pageNumber="1272">5</figureCitation>
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,
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<figureCitation id="85A92A530E562D2BFD7CA245BC8ADDF3" box="[694,761,878,902]" captionStart="Figure 6" captionStartId="15.[211,278,1458,1480]" captionText="Figure 6. Leptochirus (Tropiochirus) milton sp. nov. holotype: (A) aedeagus left lateral view; (B) aedeagus dorsal view; (C) aedeagus right lateral view; (D) tergum X [T10] and tergite IX [T9] of male; (E) sternite IX of male; (F) right gonocoxites of female. L. (T.) pachoensis. (G) aedeagus left lateral view; (H) aedeagus dorsal view; (I) aedeagus right lateral view; (J) tergum X [T10] and tergite IX [T9] of male; (K) sternite IX of male; (L) right gonocoxites of female. Leptochirus (T.) proteus. (M) Aedeagus left lateral view; (N) aedeagus dorsal view; (O) aedeagus right lateral view; (P) tergum X [T10] and tergite IX [T9] of male; (Q) sternite IX of male; (R) right gonocoxites of female. Scale bars = 0.25 mm." pageId="16" pageNumber="1272">6G–L</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="5588655D0E562D2BFF45A285BFCFDAF0" pageId="16" pageNumber="1272" type="reference_group">
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<paragraph id="1D2D36D60E562D2BFF45A285BDEDDA73" blockId="16.[143,1103,942,1030]" pageId="16" pageNumber="1272">
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<treatmentCitationGroup id="3D8211F80E562D2BFF45A285BDEDDA73" pageId="16" pageNumber="1272">
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<treatmentCitation id="9C3310C70E562D2BFF45A285BD6FDDB3" author="Bernhauer M" box="[143,796,942,966]" page="47" pageId="16" pageNumber="1272" year="1917">
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<taxonomicName id="DA924D550E562D2BFF45A285BD6FDDB3" ID-CoL="3TLRP" authority="Bernhauer, 1917: 47" authorityName="Bernhauer" authorityPageNumber="47" authorityYear="1917" box="[143,796,942,966]" class="Insecta" family="Staphylinidae" genus="Leptochirus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1272" phylum="Arthropoda" rank="species" species="pachoensis" subGenus="Tropiochirus">
|
||
<emphasis id="2FE6EAC40E562D2BFF45A285BF62DDB3" box="[143,273,942,966]" italics="true" pageId="16" pageNumber="1272">Leptochirus</emphasis>
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||
(
|
||
<emphasis id="2FE6EAC40E562D2BFEE8A285BFDEDDB3" box="[290,429,942,966]" italics="true" pageId="16" pageNumber="1272">Tropiochirus</emphasis>
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)
|
||
<emphasis id="2FE6EAC40E562D2BFE77A285BC41DDB3" box="[445,562,942,966]" italics="true" pageId="16" pageNumber="1272">pachoensis</emphasis>
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||
<bibRefCitation id="79034B270E562D2BFDF0A285BD6FDDB3" author="Bernhauer M" box="[570,796,942,966]" pageId="16" pageNumber="1272" pagination="49 - 53" refId="ref12682" refString="Bernhauer M. 1917. Neue Arten der Gattungen Piestus, Leptochirus und Conosoma aus Sudamerika. Neue Beitr Syst Insektenk. 1: 45 - 48, 49 - 53." type="journal article" year="1917">Bernhauer, 1917: 47</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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[original description, comparison with
|
||
<taxonomicName id="DA924D550E562D2BFEE2A2E4BC1DDD93" ID-CoL="3TLRQ" authority="Fauvel, 1864" authorityName="Fauvel" authorityYear="1864" box="[296,622,974,998]" class="Insecta" family="Staphylinidae" genus="Leptochirus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1277" phylum="Arthropoda" rank="species" species="proteus" subGenus="Tropiochirus">
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||
<emphasis id="2FE6EAC40E562D2BFEE2A2E4BF49DD93" box="[296,314,975,998]" italics="true" pageId="16" pageNumber="1272">L</emphasis>
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||
. (
|
||
<emphasis id="2FE6EAC40E562D2BFE99A2E4BF16DD93" box="[339,357,975,998]" italics="true" pageId="16" pageNumber="1272">T</emphasis>
|
||
.)
|
||
<emphasis id="2FE6EAC40E562D2BFE4AA2E4BFA3DD93" box="[384,464,975,998]" italics="true" pageId="16" pageNumber="1272">proteus</emphasis>
|
||
<bibRefCitation id="79034B270E562D2BFE13A2E5BC1DDD93" author="Fauvel A" box="[473,622,974,998]" pageId="16" pageNumber="1272" pagination="3 - 62" refId="ref12900" refString="Fauvel A. 1864. Etudes sur les Staphylinides de l'Amerique centrale, principalement du Mexique. Notices Entomol. 2: 3 - 62." type="journal article" year="1864">Fauvel, 1864</bibRefCitation>
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||
</taxonomicName>
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||
.
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||
<typeStatus id="C22988740E562D2BFDB1A2E4BCC1DD92" box="[635,690,975,999]" pageId="16" pageNumber="1272">Type</typeStatus>
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||
locality: “Westkolumbien: Pacho in den Ostkordilleren,
|
||
<quantity id="DA6A9B330E562D2BFEA4A2C5BFB7DA73" box="[366,452,1006,1030]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="1272" unit="m" value="2000.0">2000 m</quantity>
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”];
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||
<treatmentCitation id="9C3310C70E562D2BFE28A2C5BC9BDA73" author="Scheerpeltz O" box="[482,744,1006,1030]" page="1000" pageId="16" pageNumber="1272" year="1933">
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||
<bibRefCitation id="79034B270E562D2BFE28A2C5BC9BDA73" author="Scheerpeltz O" box="[482,744,1006,1030]" pageId="16" pageNumber="1272" pagination="989 - 1500" refId="ref13135" refString="Scheerpeltz O. 1933. Coleopterorum Catalogus 6 (129). Berlin (Germany): S. Schenkling, Staphylinidae; p. 989 - 1500." type="book chapter" year="1933">Scheerpeltz, 1933: 1000</bibRefCitation>
|
||
</treatmentCitation>
|
||
[world catalog].
|
||
</treatmentCitationGroup>
|
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</paragraph>
|
||
<paragraph id="1D2D36D60E562D2BFF45A505BFCFDAF0" blockId="16.[143,1104,1070,1157]" pageId="16" pageNumber="1272">
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<treatmentCitationGroup id="3D8211F80E562D2BFF45A505BFCFDAF0" pageId="16" pageNumber="1272">
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<treatmentCitation id="9C3310C70E562D2BFF45A505BCD5DA33" author="Blackwelder RE" box="[143,678,1070,1094]" page="110" pageId="16" pageNumber="1272" year="1944">
|
||
<taxonomicName id="DA924D550E562D2BFF45A505BCD5DA33" ID-CoL="3TLRP" authority="Blackwelder, 1944: 110" authorityName="Blackwelder" authorityPageNumber="110" authorityYear="1944" box="[143,678,1070,1094]" class="Insecta" family="Staphylinidae" genus="Leptochirus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1272" phylum="Arthropoda" rank="species" species="pachoensis">
|
||
<emphasis id="2FE6EAC40E562D2BFF45A505BFE2DA33" box="[143,401,1070,1094]" italics="true" pageId="16" pageNumber="1272">Leptochirus pachoensis</emphasis>
|
||
<bibRefCitation id="79034B270E562D2BFE56A505BCD5DA33" author="Blackwelder RE" box="[412,678,1070,1094]" pageId="16" pageNumber="1272" pagination="1 - 188" refId="ref12770" refString="Blackwelder RE. 1944. Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America, Part 1. USNM Bull. 185: 1 - 188." type="journal article" year="1944">Blackwelder, 1944: 110</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
[checklist from Neotropical region];
|
||
<treatmentCitation id="9C3310C70E562D2BFF45A565BFF2DA10" author="Herman LH" box="[143,385,1101,1125]" page="1105" pageId="16" pageNumber="1272" year="2001">
|
||
<bibRefCitation id="79034B270E562D2BFF45A565BFF2DA10" author="Herman LH" box="[143,385,1101,1125]" pageId="16" pageNumber="1272" pagination="1067 - 1806" refId="ref12990" refString="Herman LH. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. III. Oxyteline group. Bull Am Mus Nat Hist. 265: 1067 - 1806." type="journal article" year="2001">Herman, 2001: 1105</bibRefCitation>
|
||
</treatmentCitation>
|
||
[world catalog];
|
||
<treatmentCitation id="9C3310C70E562D2BFD9AA565BA23DA10" author="Newton AF & Gutierrez Chacon C & Chandler D" box="[592,1104,1101,1125]" page="30" pageId="16" pageNumber="1272" year="2005">
|
||
<bibRefCitation id="79034B270E562D2BFD9AA565BA23DA10" author="Newton AF & Gutierrez Chacon C & Chandler D" box="[592,1104,1101,1125]" pageId="16" pageNumber="1272" pagination="1 - 72" refId="ref13102" refString="Newton AF, Gutierrez Chacon C, Chandler D. 2005. Checklist of the Staphylinidae (Coleoptera) of Colombia. Biota Colombiana. 6; 1: 1 - 72." type="journal article" year="2005">Newton, Gutiérrez and Chandler, 2005: 30</bibRefCitation>
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</treatmentCitation>
|
||
[checklist from
|
||
<collectingCountry id="658576460E562D2BFEF6A546BFC2DAF0" box="[316,433,1133,1157]" name="Colombia" pageId="16" pageNumber="1272">Colombia</collectingCountry>
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].
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||
</treatmentCitationGroup>
|
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5588655D0E562D2BFF45A585BFC1D83D" pageId="16" pageNumber="1272" type="materials_examined">
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<paragraph id="1D2D36D60E562D2BFF45A585BC0BDB11" blockId="16.[143,1104,1197,1380]" pageId="16" pageNumber="1272">
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||
Type material.
|
||
<materialsCitation id="ADFA3C8B0E562D2BFE9CA586BC0BDB11" collectionCode="FMNH" collectorName="M. Bernhauer" country="Colombia" elevation="2000" location="Colombia" pageId="16" pageNumber="1272" specimenCount="1" typeStatus="holotype">
|
||
<taxonomicName id="DA924D550E562D2BFE9CA586BCD6DAB0" authorityName="Blackwelder" authorityYear="1944" box="[342,677,1197,1221]" class="Insecta" family="Staphylinidae" genus="Leptochirus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1257" phylum="Arthropoda" rank="species" species="pachoensis" subGenus="Tropiochirus">
|
||
<emphasis id="2FE6EAC40E562D2BFE9CA586BFABDAB0" box="[342,472,1197,1221]" italics="true" pageId="16" pageNumber="1272">Leptochirus</emphasis>
|
||
(
|
||
<emphasis id="2FE6EAC40E562D2BFE3CA585BC7BDAB0" box="[502,520,1198,1221]" italics="true" pageId="16" pageNumber="1272">T</emphasis>
|
||
.)
|
||
<emphasis id="2FE6EAC40E562D2BFDFAA586BCD6DAB0" box="[560,677,1197,1221]" italics="true" pageId="16" pageNumber="1272">pachoensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="2FE6EAC40E562D2BFD0BA586BD5BDAB0" bold="true" box="[705,808,1197,1221]" pageId="16" pageNumber="1272">
|
||
<typeStatus id="C22988740E562D2BFD0BA586BD5BDAB0" box="[705,808,1197,1221]" pageId="16" pageNumber="1272" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(female): “Pacho Col. [
|
||
<collectingCountry id="658576460E562D2BFF5CA5E6BF7ADA90" box="[150,265,1229,1253]" name="Colombia" pageId="16" pageNumber="1272">Colombia</collectingCountry>
|
||
]
|
||
<emphasis id="2FE6EAC40E562D2BFEC4A5E6BF69DA90" box="[270,282,1229,1253]" italics="true" pageId="16" pageNumber="1272">/</emphasis>
|
||
dh.es will
|
||
<emphasis id="2FE6EAC40E562D2BFE50A5E6BFD5DA90" box="[410,422,1229,1253]" italics="true" pageId="16" pageNumber="1272">/</emphasis>
|
||
<quantity id="DA6A9B330E562D2BFE6CA5E6BC7EDA90" box="[422,525,1229,1253]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="1272" unit="m" value="2000.0">
|
||
<elevation id="96BFD1E50E562D2BFE6CA5E6BC7EDA90" box="[422,525,1229,1253]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="1272" unit="m" value="2000.0">2000 m</elevation>
|
||
</quantity>
|
||
oklis” “Columbia occ.[western]
|
||
<emphasis id="2FE6EAC40E562D2BFC78A5E6BDCDDA90" box="[946,958,1229,1253]" italics="true" pageId="16" pageNumber="1272">/</emphasis>
|
||
cali, Fassl” “pachoensis
|
||
<emphasis id="2FE6EAC40E562D2BFEDDA5C6BF50DB70" box="[279,291,1261,1285]" italics="true" pageId="16" pageNumber="1272">/</emphasis>
|
||
Bernh. Typus unic” “Chicago NHMus
|
||
<emphasis id="2FE6EAC40E562D2BFCD6A5C6BD5BDB70" box="[796,808,1261,1285]" italics="true" pageId="16" pageNumber="1272">/</emphasis>
|
||
<collectorName id="B06753000E562D2BFCE2A5C6BDB3DB70" box="[808,960,1261,1285]" pageId="16" pageNumber="1272">M.Bernhauer</collectorName>
|
||
<emphasis id="2FE6EAC40E562D2BFC0BA5C6BDBEDB70" box="[961,973,1261,1285]" italics="true" pageId="16" pageNumber="1272">/</emphasis>
|
||
Collection” (
|
||
<collectionCode id="7B83AE130E562D2BFF52A426BE87DB50" box="[152,244,1293,1317]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="1272" type="Museum">FMNH</collectionCode>
|
||
).
|
||
<emphasis id="2FE6EAC40E562D2BFECCA426BF30DB50" bold="true" box="[262,323,1293,1317]" pageId="16" pageNumber="1272">Note:</emphasis>
|
||
In the original description,
|
||
<bibRefCitation id="79034B270E562D2BFDB0A426BD4DDB50" author="Bernhauer M" box="[634,830,1293,1317]" pageId="16" pageNumber="1272" pagination="49 - 53" refId="ref12682" refString="Bernhauer M. 1917. Neue Arten der Gattungen Piestus, Leptochirus und Conosoma aus Sudamerika. Neue Beitr Syst Insektenk. 1: 45 - 48, 49 - 53." type="journal article" year="1917">Bernhauer (1917)</bibRefCitation>
|
||
specified that he studied
|
||
<specimenCount id="0B94FD5F0E562D2BFF45A406BF59DB30" box="[143,298,1325,1349]" pageId="16" pageNumber="1272" type="generic">one specimen</specimenCount>
|
||
. We received from the
|
||
<collectionCode id="7B83AE130E562D2BFDE8A406BC09DB31" box="[546,634,1325,1348]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="1272" type="Museum">FMNH</collectionCode>
|
||
a female with the label “Typus”, therefore the specimen is the
|
||
<typeStatus id="C22988740E562D2BFEA3A467BFBEDB11" box="[361,461,1356,1380]" pageId="16" pageNumber="1272" type="holotype">holotype</typeStatus>
|
||
for the species.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="1D2D36D60E562D2BFF45A482BF0BDBB4" blockId="16.[143,1103,1449,1608]" box="[143,376,1449,1473]" pageId="16" pageNumber="1272">
|
||
<heading id="466581BA0E562D2BFF45A482BF0BDBB4" box="[143,376,1449,1473]" fontSize="10" level="3" pageId="16" pageNumber="1272" reason="5">
|
||
<emphasis id="2FE6EAC40E562D2BFF45A482BF0BDBB4" bold="true" box="[143,376,1449,1473]" italics="true" pageId="16" pageNumber="1272">Additional specimens</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="1D2D36D60E562D2BFF45A4FABFC1D83D" blockId="16.[143,1103,1449,1608]" pageId="16" pageNumber="1272">
|
||
<materialsCitation id="ADFA3C8B0E562D2BFF45A4FABCE7D87C" collectingDate="1971-03-04" collectingDateMax="1971-03-05" collectingDateMin="1971-03-04" collectionCode="CNC" collectorName="J. M. Campbell" country="Ecuador" elevation="1500" location="Baeza" municipality="Oritoyacu" pageId="16" pageNumber="1272" specimenCount="1" specimenCount-male="1" stateProvince="Napo">
|
||
<emphasis id="2FE6EAC40E562D2BFF45A4FABF1ADB9C" bold="true" box="[143,361,1489,1513]" pageId="16" pageNumber="1272">
|
||
<collectingCountry id="658576460E562D2BFF45A4FABF69DB9C" box="[143,282,1489,1513]" name="Ecuador" pageId="16" pageNumber="1272">ECUADOR</collectingCountry>
|
||
.
|
||
<emphasis id="2FE6EAC40E562D2BFEECA4FABF11DB9D" bold="true" box="[294,354,1489,1512]" italics="true" pageId="16" pageNumber="1272">
|
||
<collectingRegion id="DF56F8340E562D2BFEECA4FABF11DB9D" box="[294,354,1489,1512]" country="Ecuador" name="Napo" pageId="16" pageNumber="1272">Napo</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
</emphasis>
|
||
<quantity id="DA6A9B330E562D2BFEBAA4FABFDADB9C" box="[368,425,1489,1513]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="1272" unit="km" value="2.0">2 km</quantity>
|
||
south from
|
||
<collectingMunicipality id="FD49ACAC0E562D2BFDF9A4FABCD7DB9C" box="[563,676,1489,1513]" pageId="16" pageNumber="1272">Oritoyacu</collectingMunicipality>
|
||
and
|
||
<quantity id="DA6A9B330E562D2BFD16A4FABD69DB9C" box="[732,794,1489,1513]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="16" pageNumber="1272" unit="km" value="22.0">22km</quantity>
|
||
south from
|
||
<location id="184D600D0E562D2BFC6FA4F9BD9FDB9C" LSID="urn:lsid:plazi:treatment:953B87C00E562D29FEF5A265BCFADF72:184D600D0E562D2BFC6FA4F9BD9FDB9C" box="[933,1004,1490,1513]" country="Ecuador" municipality="Oritoyacu" name="Baeza" pageId="16" pageNumber="1272" stateProvince="Napo">Baeza</location>
|
||
,
|
||
<quantity id="DA6A9B330E562D2BFC3CA4FABA38DB9C" box="[1014,1099,1489,1513]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="1272" unit="m" value="1500.0">
|
||
<elevation id="96BFD1E50E562D2BFC3CA4FABA38DB9C" box="[1014,1099,1489,1513]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="1272" unit="m" value="1500.0">1500 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="692C10160E562D2BFF45A4DABF63D87C" box="[143,272,1521,1545]" pageId="16" pageNumber="1272" value="1971-03-04" valueMax="1971-03-05" valueMin="1971-03-04">
|
||
<collectingDate id="7968E9FE0E562D2BFF45A4DABF63D87C" box="[143,272,1521,1545]" pageId="16" pageNumber="1272" value="1971-03-04" valueMax="1971-03-05" valueMin="1971-03-04">4-5-iii.1971</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="B06753000E562D2BFEEBA4D9BFA9D87C" box="[289,474,1521,1545]" pageId="16" pageNumber="1272">J. M. Campbell</collectorName>
|
||
(
|
||
<specimenCount id="0B94FD5F0E562D2BFE3BA4DABC33D87C" box="[497,576,1521,1545]" pageId="16" pageNumber="1272" type="male">1 male</specimenCount>
|
||
<collectionCode id="7B83AE130E562D2BFD86A4DABCFFD87C" box="[588,652,1521,1545]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="16" pageNumber="1272">CNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="ADFA3C8B0E562D2BFD6FA4DABA38D87C" box="[677,1099,1521,1545]" collectingDate="1976-03-04" collectingDateMax="1976-03-05" collectingDateMin="1976-03-04" collectionCode="ZMHB" collectorName="J. M. Campbell" country="Ecuador" elevation="1500" location="Baeza" municipality="Oritoyacu" pageId="16" pageNumber="1272" specimenCount="1" specimenCount-female="1" stateProvince="Napo">
|
||
<emphasis id="2FE6EAC40E562D2BFD6FA4DABCA9D87C" box="[677,730,1521,1545]" italics="true" pageId="16" pageNumber="1272">idem</emphasis>
|
||
<date id="692C10160E562D2BFD2DA4DABD16D87C" box="[743,869,1521,1545]" pageId="16" pageNumber="1272" value="1976-03-04" valueMax="1976-03-05" valueMin="1976-03-04">
|
||
<collectingDate id="7968E9FE0E562D2BFD2DA4DABD16D87C" box="[743,869,1521,1545]" pageId="16" pageNumber="1272" value="1976-03-04" valueMax="1976-03-05" valueMin="1976-03-04">4-5-iii.1976</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="0B94FD5F0E562D2BFCB6A4DABDADD87C" box="[892,990,1521,1545]" pageId="16" pageNumber="1272" type="female">1 female</specimenCount>
|
||
<collectionCode id="7B83AE130E562D2BFC26A4DABA30D87C" box="[1004,1091,1521,1545]" pageId="16" pageNumber="1272">ZMHB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="ADFA3C8B0E562D2BFF45A73ABD58D85C" box="[143,811,1553,1577]" collectionCode="ZMHB" country="Colombia" location="Distrito Capital" pageId="16" pageNumber="1272" specimenCount="1" specimenCount-female="1" stateProvince="Bogota">
|
||
<emphasis id="2FE6EAC40E562D2BFF45A73ABF87D85C" bold="true" box="[143,500,1553,1577]" pageId="16" pageNumber="1272">
|
||
<collectingCountry id="658576460E562D2BFF45A73ABF58D85C" box="[143,299,1553,1577]" name="Colombia" pageId="16" pageNumber="1272">COLOMBIA</collectingCountry>
|
||
.
|
||
<emphasis id="2FE6EAC40E562D2BFEF3A73ABF87D85C" bold="true" box="[313,500,1553,1577]" italics="true" pageId="16" pageNumber="1272">
|
||
<location id="184D600D0E562D2BFEF3A73ABF9ED85C" LSID="urn:lsid:plazi:treatment:953B87C00E562D29FEF5A265BCFADF72:184D600D0E562D2BFEF3A73ABF9ED85C" box="[313,493,1553,1577]" country="Colombia" name="Distrito Capital" pageId="16" pageNumber="1272" stateProvince="Bogota">Distrito Capital</location>
|
||
:
|
||
</emphasis>
|
||
</emphasis>
|
||
<collectingRegion id="DF56F8340E562D2BFE37A73ABC3CD85C" box="[509,591,1553,1577]" country="Colombia" name="Bogota" pageId="16" pageNumber="1272">Bogotá</collectingRegion>
|
||
(
|
||
<specimenCount id="0B94FD5F0E562D2BFDA9A73ABCB1D85C" box="[611,706,1553,1577]" pageId="16" pageNumber="1272" type="female">1 female</specimenCount>
|
||
<collectionCode id="7B83AE130E562D2BFD01A73ABD51D85C" box="[715,802,1553,1577]" pageId="16" pageNumber="1272">ZMHB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="ADFA3C8B0E562D2BFCF2A73ABFDDD83D" collectionCode="DZPU, ZMHB" country="Colombia" location="Nueva Granada" pageId="16" pageNumber="1272" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<collectingCountry id="658576460E562D2BFCF2A73ABD82D85C" box="[824,1009,1553,1577]" name="Colombia" pageId="16" pageNumber="1272">Nueva Granada</collectingCountry>
|
||
(
|
||
<specimenCount id="0B94FD5F0E562D2BFBCEA73ABA3CD85C" box="[1028,1103,1553,1577]" pageId="16" pageNumber="1272" type="male">1 male</specimenCount>
|
||
<collectionCode id="7B83AE130E562D2BFF45A71ABEADD83D" box="[143,222,1585,1608]" pageId="16" pageNumber="1272">DZPU</collectionCode>
|
||
,
|
||
<specimenCount id="0B94FD5F0E562D2BFF23A71BBF34D83D" box="[233,327,1584,1608]" pageId="16" pageNumber="1272" type="female">1 female</specimenCount>
|
||
<collectionCode id="7B83AE130E562D2BFE84A71BBFD5D83D" box="[334,422,1584,1608]" pageId="16" pageNumber="1272">ZMHB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5588655D0E572D2AFF19A183BAE0DF52" pageId="17" pageNumber="1273" type="diagnosis">
|
||
<paragraph id="1D2D36D60E572D2AFF19A183BF4CDECA" blockId="17.[211,1172,168,295]" box="[211,319,168,191]" pageId="17" pageNumber="1273">
|
||
<heading id="466581BA0E572D2AFF19A183BF4CDECA" box="[211,319,168,191]" fontSize="10" level="3" pageId="17" pageNumber="1273" reason="5">
|
||
<emphasis id="2FE6EAC40E572D2AFF19A183BF4CDECA" box="[211,319,168,191]" italics="true" pageId="17" pageNumber="1273">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="1D2D36D60E572D2AFF19A1E4BAE0DF52" blockId="17.[211,1172,168,295]" pageId="17" pageNumber="1273">
|
||
<taxonomicName id="DA924D550E572D2AFF19A1E4BC7FDE92" authorityName="Blackwelder" authorityYear="1944" box="[211,524,207,231]" class="Insecta" family="Staphylinidae" genus="Leptochirus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1257" phylum="Arthropoda" rank="species" species="pachoensis" subGenus="Tropiochirus">
|
||
<emphasis id="2FE6EAC40E572D2AFF19A1E4BF26DE92" box="[211,341,207,231]" italics="true" pageId="17" pageNumber="1273">Leptochirus</emphasis>
|
||
(
|
||
<emphasis id="2FE6EAC40E572D2AFEA3A1FBBF08DE92" box="[361,379,208,231]" italics="true" pageId="17" pageNumber="1273">T</emphasis>
|
||
.)
|
||
<emphasis id="2FE6EAC40E572D2AFE5DA1E4BC7FDE92" box="[407,524,207,231]" italics="true" pageId="17" pageNumber="1273">pachoensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="2FE6EAC40E572D2AFD86A1FBBC16DE92" box="[588,613,208,231]" italics="true" pageId="17" pageNumber="1273">L.</emphasis>
|
||
<taxonomicName id="DA924D550E572D2AFDBAA1E4BC98DE92" authorityName="Bernhauer" authorityYear="1903" box="[624,747,207,231]" class="Insecta" family="Staphylinidae" genus="Leptochirus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1273" phylum="Arthropoda" rank="species" species="helleri" subGenus="Tropiochirus">
|
||
(
|
||
<emphasis id="2FE6EAC40E572D2AFDB2A1FBBCF9DE92" box="[632,650,208,231]" italics="true" pageId="17" pageNumber="1273">T</emphasis>
|
||
.)
|
||
<emphasis id="2FE6EAC40E572D2AFD6DA1E4BC98DE92" box="[679,747,207,231]" italics="true" pageId="17" pageNumber="1273">helleri</emphasis>
|
||
</taxonomicName>
|
||
have the accessory tooth of the left mandible far from the lateral longitudinal ridge (
|
||
<figureCitation id="85A92A530E572D2AFD3FA1C4BDF8DF72" box="[757,907,239,263]" captionStart="Figure 3" captionStartId="6.[143,210,1078,1100]" captionTargetBox="[145,1102,167,1045]" captionTargetId="figure-6@6.[145,1102,167,1045]" captionTargetPageId="6" captionText="Figure 3. Left and right mandible, dorsal view. (A) Leptochirus (Tropiochirus) helleri; (B) L. (T.) marina sp. nov. holotype; (C) L. (T.) mery sp. nov. holotype; (D) L. (T.) milton sp. nov. holotype; (E) L. (T.) pachoensis; (F) L. (T.) proteus; (G) L. (T.) venezuelanus; (H) L. (T.) yunganus; (I) L. (T.) zischkaianus. Scale bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10091560" httpUri="https://zenodo.org/record/10091560/files/figure.png" pageId="17" pageNumber="1273">Figure 3A, E</figureCitation>
|
||
), but
|
||
<emphasis id="2FE6EAC40E572D2AFC06A1DBBDADDF72" box="[972,990,240,263]" italics="true" pageId="17" pageNumber="1273">L</emphasis>
|
||
.
|
||
<taxonomicName id="DA924D550E572D2AFC21A1C4BAE0DF72" authorityName="Blackwelder" authorityYear="1944" box="[1003,1171,239,263]" class="Insecta" family="Staphylinidae" genus="Leptochirus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1273" phylum="Arthropoda" rank="species" species="pachoensis" subGenus="Tropiochirus">
|
||
(
|
||
<emphasis id="2FE6EAC40E572D2AFC3EA1DBBA75DF72" box="[1012,1030,240,263]" italics="true" pageId="17" pageNumber="1273">T</emphasis>
|
||
.)
|
||
<emphasis id="2FE6EAC40E572D2AFBD4A1C4BAE0DF72" box="[1054,1171,239,263]" italics="true" pageId="17" pageNumber="1273">pachoensis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by the first dorsal tooth of the left mandible bifurcate (
|
||
<figureCitation id="85A92A530E572D2AFBD9A024BAF4DF52" box="[1043,1159,271,295]" captionStart="Figure 3" captionStartId="6.[143,210,1078,1100]" captionTargetBox="[145,1102,167,1045]" captionTargetId="figure-6@6.[145,1102,167,1045]" captionTargetPageId="6" captionText="Figure 3. Left and right mandible, dorsal view. (A) Leptochirus (Tropiochirus) helleri; (B) L. (T.) marina sp. nov. holotype; (C) L. (T.) mery sp. nov. holotype; (D) L. (T.) milton sp. nov. holotype; (E) L. (T.) pachoensis; (F) L. (T.) proteus; (G) L. (T.) venezuelanus; (H) L. (T.) yunganus; (I) L. (T.) zischkaianus. Scale bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10091560" httpUri="https://zenodo.org/record/10091560/files/figure.png" pageId="17" pageNumber="1273">Figure 3E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5588655D0E572D2AFF19A047BF9FD8D3" pageId="17" pageNumber="1273" type="description">
|
||
<paragraph id="1D2D36D60E572D2AFF19A047BF21DFF6" blockId="17.[211,438,364,427]" box="[211,338,364,387]" pageId="17" pageNumber="1273">
|
||
<heading id="466581BA0E572D2AFF19A047BF21DFF6" box="[211,338,364,387]" fontSize="10" level="3" pageId="17" pageNumber="1273" reason="5">
|
||
<emphasis id="2FE6EAC40E572D2AFF19A047BF21DFF6" box="[211,338,364,387]" italics="true" pageId="17" pageNumber="1273">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="1D2D36D60E572D2AFF19A0BFBFC5DFDE" blockId="17.[211,438,364,427]" box="[211,438,403,427]" pageId="17" pageNumber="1273">BL: 14.73; BW: 2.46</paragraph>
|
||
<paragraph id="1D2D36D60E572D2AFF19A0F8BDEEDF9E" blockId="17.[211,925,467,491]" box="[211,925,467,491]" pageId="17" pageNumber="1273">Body strongly depressed. Body all black except tarsi red-brown.</paragraph>
|
||
<paragraph id="1D2D36D60E572D2AFF19A338BFB2DD3F" blockId="17.[211,1172,531,842]" pageId="17" pageNumber="1273">
|
||
<emphasis id="2FE6EAC40E572D2AFF19A338BF64DC5E" bold="true" box="[211,279,531,555]" pageId="17" pageNumber="1273">Head.</emphasis>
|
||
Transverse (
|
||
<figureCitation id="85A92A530E572D2AFE6EA338BC69DC5E" box="[420,538,531,555]" captionStart="Figure 2" captionStartId="5.[211,278,1629,1651]" captionTargetBox="[238,1145,161,1595]" captionTargetId="figure-5@5.[238,1145,161,1596]" captionTargetPageId="5" captionText="Figure 2. Head and protorax of Leptochirus (Tropiochirus). (A) L. (T.) helleri; (B) L. (T.) marina sp. nov. holotype; (C) L. (T.) mery sp. nov. holotype; (D) L. (T.) milton sp. nov. holotype; (E) L. (T.) pachoensis; (F) L. (T.) proteus; (G) L. (T.) venezuelanus; (H) L. (T.) yunganus; (I) L. (T.) zischkaianus. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10091558" httpUri="https://zenodo.org/record/10091558/files/figure.png" pageId="17" pageNumber="1273">Figure 2E</figureCitation>
|
||
). Inner lateral teeth separated by a depression extending to base. Inner and outer lateral teeth fused and resembling a septum; median sulcus deep, sides of median sulcus parallel; laterofrontal depressions oval. Labrum with outer edge strongly angled and lateral-apical edge concave. Mandibles (
|
||
<figureCitation id="85A92A530E572D2AFC2FA359BA12DCFF" box="[997,1121,626,650]" captionStart="Figure 2" captionStartId="5.[211,278,1629,1651]" captionTargetBox="[238,1145,161,1595]" captionTargetId="figure-5@5.[238,1145,161,1596]" captionTargetPageId="5" captionText="Figure 2. Head and protorax of Leptochirus (Tropiochirus). (A) L. (T.) helleri; (B) L. (T.) marina sp. nov. holotype; (C) L. (T.) mery sp. nov. holotype; (D) L. (T.) milton sp. nov. holotype; (E) L. (T.) pachoensis; (F) L. (T.) proteus; (G) L. (T.) venezuelanus; (H) L. (T.) yunganus; (I) L. (T.) zischkaianus. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10091558" httpUri="https://zenodo.org/record/10091558/files/figure.png" pageId="17" pageNumber="1273">Figures 2E</figureCitation>
|
||
,
|
||
<figureCitation id="85A92A530E572D2AFBA1A359BAF8DCFF" box="[1131,1163,626,650]" captionStart="Figure 3" captionStartId="6.[143,210,1078,1100]" captionTargetBox="[145,1102,167,1045]" captionTargetId="figure-6@6.[145,1102,167,1045]" captionTargetPageId="6" captionText="Figure 3. Left and right mandible, dorsal view. (A) Leptochirus (Tropiochirus) helleri; (B) L. (T.) marina sp. nov. holotype; (C) L. (T.) mery sp. nov. holotype; (D) L. (T.) milton sp. nov. holotype; (E) L. (T.) pachoensis; (F) L. (T.) proteus; (G) L. (T.) venezuelanus; (H) L. (T.) yunganus; (I) L. (T.) zischkaianus. Scale bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10091560" httpUri="https://zenodo.org/record/10091560/files/figure.png" pageId="17" pageNumber="1273">3E</figureCitation>
|
||
) with lateral longitudinal ridge occupying less than two-thirds of apical region; external margin strongly developed, attenuated anteriorly. Left mandible (
|
||
<figureCitation id="85A92A530E572D2AFC64A399BA53DCBF" box="[942,1056,690,714]" captionStart="Figure 3" captionStartId="6.[143,210,1078,1100]" captionTargetBox="[145,1102,167,1045]" captionTargetId="figure-6@6.[145,1102,167,1045]" captionTargetPageId="6" captionText="Figure 3. Left and right mandible, dorsal view. (A) Leptochirus (Tropiochirus) helleri; (B) L. (T.) marina sp. nov. holotype; (C) L. (T.) mery sp. nov. holotype; (D) L. (T.) milton sp. nov. holotype; (E) L. (T.) pachoensis; (F) L. (T.) proteus; (G) L. (T.) venezuelanus; (H) L. (T.) yunganus; (I) L. (T.) zischkaianus. Scale bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10091560" httpUri="https://zenodo.org/record/10091560/files/figure.png" pageId="17" pageNumber="1273">Figure 3E</figureCitation>
|
||
) with first dorsal tooth developed, straight and bifurcate, accessory tooth far from lateral longitudinal ridge; second dorsal tooth little developed and resembling a small hill; first ventral tooth with tooth accessory pointed. Submentum with setae arranged in row on anterolateral margin.
|
||
</paragraph>
|
||
<paragraph id="1D2D36D60E572D2AFF19A25ABFFBDADD" blockId="17.[211,1172,881,1192]" pageId="17" pageNumber="1273">
|
||
<emphasis id="2FE6EAC40E572D2AFF19A25ABF5EDDFC" bold="true" box="[211,301,881,905]" pageId="17" pageNumber="1273">Thorax.</emphasis>
|
||
Pronotum (
|
||
<figureCitation id="85A92A530E572D2AFE7CA25ABC5BDDFC" box="[438,552,881,905]" captionStart="Figure 2" captionStartId="5.[211,278,1629,1651]" captionTargetBox="[238,1145,161,1595]" captionTargetId="figure-5@5.[238,1145,161,1596]" captionTargetPageId="5" captionText="Figure 2. Head and protorax of Leptochirus (Tropiochirus). (A) L. (T.) helleri; (B) L. (T.) marina sp. nov. holotype; (C) L. (T.) mery sp. nov. holotype; (D) L. (T.) milton sp. nov. holotype; (E) L. (T.) pachoensis; (F) L. (T.) proteus; (G) L. (T.) venezuelanus; (H) L. (T.) yunganus; (I) L. (T.) zischkaianus. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10091558" httpUri="https://zenodo.org/record/10091558/files/figure.png" pageId="17" pageNumber="1273">Figure 2E</figureCitation>
|
||
) a little wider than long (PL: 2.54; PW: 2.98); superior marginal line of hypomeron not interrupted, not bent to dorsal region at anterior margin and bent to slightly dorsal region at posterior margin; without pair of small depressions near basal constriction. Inferior marginal line of hypomeron reaching anterior margin. Apex of prosternal process strongly expanded and resembling a circular plate, extending considerably beyond the lateral process of hypomeron. Metaventrite with microsculpture longitudinally undulate in median region and with small longitudinal median depression near posterior edge. Mesoscutellum with apex rounded. Elytra longer than broad and without pair of small depressions on posterior-lateral edge (EL: 3.27, EW: 2.95).
|
||
</paragraph>
|
||
<paragraph id="1D2D36D60E572D2AFF19A5FBBF9CDB3D" blockId="17.[211,1172,1232,1352]" pageId="17" pageNumber="1273">
|
||
<emphasis id="2FE6EAC40E572D2AFF19A5FBBF31DA9D" bold="true" box="[211,322,1232,1256]" pageId="17" pageNumber="1273">Abdomen.</emphasis>
|
||
Cylindrical. Sternite III with keel of apex rounded. Tergite IX (
|
||
<figureCitation id="85A92A530E572D2AFBD7A5FBBAF8DA9D" box="[1053,1163,1232,1256]" captionStart="Figure 6" captionStartId="15.[211,278,1458,1480]" captionText="Figure 6. Leptochirus (Tropiochirus) milton sp. nov. holotype: (A) aedeagus left lateral view; (B) aedeagus dorsal view; (C) aedeagus right lateral view; (D) tergum X [T10] and tergite IX [T9] of male; (E) sternite IX of male; (F) right gonocoxites of female. L. (T.) pachoensis. (G) aedeagus left lateral view; (H) aedeagus dorsal view; (I) aedeagus right lateral view; (J) tergum X [T10] and tergite IX [T9] of male; (K) sternite IX of male; (L) right gonocoxites of female. Leptochirus (T.) proteus. (M) Aedeagus left lateral view; (N) aedeagus dorsal view; (O) aedeagus right lateral view; (P) tergum X [T10] and tergite IX [T9] of male; (Q) sternite IX of male; (R) right gonocoxites of female. Scale bars = 0.25 mm." pageId="17" pageNumber="1273">Figure 6J</figureCitation>
|
||
) sclerotized with left and right plates separated, each with six or seven setae of various lengths. Tergite X (
|
||
<figureCitation id="85A92A530E572D2AFE6CA43BBC61DB5D" box="[422,530,1296,1320]" captionStart="Figure 6" captionStartId="15.[211,278,1458,1480]" captionText="Figure 6. Leptochirus (Tropiochirus) milton sp. nov. holotype: (A) aedeagus left lateral view; (B) aedeagus dorsal view; (C) aedeagus right lateral view; (D) tergum X [T10] and tergite IX [T9] of male; (E) sternite IX of male; (F) right gonocoxites of female. L. (T.) pachoensis. (G) aedeagus left lateral view; (H) aedeagus dorsal view; (I) aedeagus right lateral view; (J) tergum X [T10] and tergite IX [T9] of male; (K) sternite IX of male; (L) right gonocoxites of female. Leptochirus (T.) proteus. (M) Aedeagus left lateral view; (N) aedeagus dorsal view; (O) aedeagus right lateral view; (P) tergum X [T10] and tergite IX [T9] of male; (Q) sternite IX of male; (R) right gonocoxites of female. Scale bars = 0.25 mm." pageId="17" pageNumber="1273">Figure 6J</figureCitation>
|
||
) membranous except for sclerotized apical part, with two pairs of long apical setae.
|
||
</paragraph>
|
||
<paragraph id="1D2D36D60E572D2AFF19A45BBD9AD852" blockId="17.[211,1172,1392,1575]" pageId="17" pageNumber="1273">
|
||
<emphasis id="2FE6EAC40E572D2AFF19A45BBF67DBFD" box="[211,276,1392,1416]" italics="true" pageId="17" pageNumber="1273">Male.</emphasis>
|
||
Sternite IX (
|
||
<figureCitation id="85A92A530E572D2AFE63A45BBC53DBFD" box="[425,544,1392,1416]" captionStart="Figure 6" captionStartId="15.[211,278,1458,1480]" captionText="Figure 6. Leptochirus (Tropiochirus) milton sp. nov. holotype: (A) aedeagus left lateral view; (B) aedeagus dorsal view; (C) aedeagus right lateral view; (D) tergum X [T10] and tergite IX [T9] of male; (E) sternite IX of male; (F) right gonocoxites of female. L. (T.) pachoensis. (G) aedeagus left lateral view; (H) aedeagus dorsal view; (I) aedeagus right lateral view; (J) tergum X [T10] and tergite IX [T9] of male; (K) sternite IX of male; (L) right gonocoxites of female. Leptochirus (T.) proteus. (M) Aedeagus left lateral view; (N) aedeagus dorsal view; (O) aedeagus right lateral view; (P) tergum X [T10] and tergite IX [T9] of male; (Q) sternite IX of male; (R) right gonocoxites of female. Scale bars = 0.25 mm." pageId="17" pageNumber="1273">Figure 6K</figureCitation>
|
||
) composed of two plates, distal plate sclerotized except for membranous apex, anterior margin rounded, posterior margin sinuated and with two long setae; proximal plate less sclerotized, greater than apical, broadened in middle and with a projecting angle at posterior border. Aedeagus (
|
||
<figureCitation id="85A92A530E572D2AFCA0A4E4BD8BDB92" box="[874,1016,1487,1511]" captionStart="Figure 6" captionStartId="15.[211,278,1458,1480]" captionText="Figure 6. Leptochirus (Tropiochirus) milton sp. nov. holotype: (A) aedeagus left lateral view; (B) aedeagus dorsal view; (C) aedeagus right lateral view; (D) tergum X [T10] and tergite IX [T9] of male; (E) sternite IX of male; (F) right gonocoxites of female. L. (T.) pachoensis. (G) aedeagus left lateral view; (H) aedeagus dorsal view; (I) aedeagus right lateral view; (J) tergum X [T10] and tergite IX [T9] of male; (K) sternite IX of male; (L) right gonocoxites of female. Leptochirus (T.) proteus. (M) Aedeagus left lateral view; (N) aedeagus dorsal view; (O) aedeagus right lateral view; (P) tergum X [T10] and tergite IX [T9] of male; (Q) sternite IX of male; (R) right gonocoxites of female. Scale bars = 0.25 mm." pageId="17" pageNumber="1273">Figure 6G–I</figureCitation>
|
||
) with median lobe slightly bulbous at base, pointed at apex, weakly sclerotized on dorsal side and a small plate sclerotized on ventral side; parameres short, fused dorsally.
|
||
</paragraph>
|
||
<paragraph id="1D2D36D60E572D2AFF19A764BF9FD8D3" blockId="17.[211,1172,1615,1702]" pageId="17" pageNumber="1273">
|
||
<emphasis id="2FE6EAC40E572D2AFF19A764BF5AD812" box="[211,297,1615,1639]" italics="true" pageId="17" pageNumber="1273">Female.</emphasis>
|
||
Similar to male, except for the gonocoxites divided longitudinally (
|
||
<figureCitation id="85A92A530E572D2AFBD9A764BAF4D812" box="[1043,1159,1615,1639]" captionStart="Figure 6" captionStartId="15.[211,278,1458,1480]" captionText="Figure 6. Leptochirus (Tropiochirus) milton sp. nov. holotype: (A) aedeagus left lateral view; (B) aedeagus dorsal view; (C) aedeagus right lateral view; (D) tergum X [T10] and tergite IX [T9] of male; (E) sternite IX of male; (F) right gonocoxites of female. L. (T.) pachoensis. (G) aedeagus left lateral view; (H) aedeagus dorsal view; (I) aedeagus right lateral view; (J) tergum X [T10] and tergite IX [T9] of male; (K) sternite IX of male; (L) right gonocoxites of female. Leptochirus (T.) proteus. (M) Aedeagus left lateral view; (N) aedeagus dorsal view; (O) aedeagus right lateral view; (P) tergum X [T10] and tergite IX [T9] of male; (Q) sternite IX of male; (R) right gonocoxites of female. Scale bars = 0.25 mm." pageId="17" pageNumber="1273">Figure 6L</figureCitation>
|
||
). The distal plate with apex membranous and long seta; proximal plate with the posterior region pointed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5588655D0E542D29FF45A183BCFADF72" pageId="18" pageNumber="1274" type="distribution">
|
||
<paragraph id="1D2D36D60E542D29FF45A183BF67DEB5" blockId="18.[143,1103,168,263]" box="[143,276,168,192]" pageId="18" pageNumber="1274">
|
||
<heading id="466581BA0E542D29FF45A183BF67DEB5" box="[143,276,168,192]" fontSize="10" level="3" pageId="18" pageNumber="1274" reason="5">
|
||
<emphasis id="2FE6EAC40E542D29FF45A183BF67DEB5" box="[143,276,168,192]" italics="true" pageId="18" pageNumber="1274">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="1D2D36D60E542D29FF45A1E4BCFADF72" blockId="18.[143,1103,168,263]" pageId="18" pageNumber="1274">
|
||
<taxonomicName id="DA924D550E542D29FF45A1E4BFB6DE92" authorityName="Blackwelder" authorityYear="1944" box="[143,453,207,231]" class="Insecta" family="Staphylinidae" genus="Leptochirus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1257" phylum="Arthropoda" rank="species" species="pachoensis" subGenus="Tropiochirus">
|
||
<emphasis id="2FE6EAC40E542D29FF45A1E4BF62DE92" box="[143,273,207,231]" italics="true" pageId="18" pageNumber="1274">Leptochirus</emphasis>
|
||
(
|
||
<emphasis id="2FE6EAC40E542D29FEE9A1FBBF46DE92" box="[291,309,208,231]" italics="true" pageId="18" pageNumber="1274">T</emphasis>
|
||
.)
|
||
<emphasis id="2FE6EAC40E542D29FE9AA1E4BFB6DE92" box="[336,453,207,231]" italics="true" pageId="18" pageNumber="1274">pachoensis</emphasis>
|
||
</taxonomicName>
|
||
is currently known from
|
||
<collectingCountry id="658576460E542D29FD20A1E4BD2DDE92" box="[746,862,207,231]" name="Colombia" pageId="18" pageNumber="1274">Colombia</collectingCountry>
|
||
. In the current study this species is listed from
|
||
<collectingCountry id="658576460E542D29FE61A1C4BC7FDF72" box="[427,524,239,263]" name="Ecuador" pageId="18" pageNumber="1274">Ecuador</collectingCountry>
|
||
(
|
||
<figureCitation id="85A92A530E542D29FDD0A1C4BC08DF72" box="[538,635,239,263]" captionStart="Figure 5" captionStartId="8.[143,210,1486,1508]" captionTargetBox="[145,1102,169,1452]" captionTargetId="figure-6@8.[145,1102,169,1453]" captionTargetPageId="8" captionText="Figure 5. Distribution map of Leptochirus (Tropiochirus) helleri, L. (T.) marina sp. nov., L. (T.) mery sp. nov., L. (T.) milton sp. nov., and L. (T.) pachoensis." figureDoi="http://doi.org/10.5281/zenodo.10091562" httpUri="https://zenodo.org/record/10091562/files/figure.png" pageId="18" pageNumber="1274">Figure 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |