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<document id="368F411C28F7481B2F4C0E3B57C09E01" ID-DOI="10.11646/zootaxa.4052.2.9" ID-GBIF-Dataset="c2fc312c-097b-4aaa-911f-3a0785f6f01c" ID-ISSN="1175-5326" ID-Zenodo-Dep="234196" ID-ZooBank="22AEEFF3-0419-4549-BF29-E7A6AE1D77E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461350448683" checkinUser="plazi" docAuthor="Balashov, Igor &amp; Griffiths, Owen" docDate="2015" docId="9526643BB16BFFD6FF2BFF65364932CF" docLanguage="en" docName="zt04052p240.pdf" docOrigin="Zootaxa 4052 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Boucardicus Balashov &amp; Griffiths, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="239" masterDocId="691F1C43B169FFD4FFBCFFF234303740" masterDocTitle="Two new species of minute land snails from Madagascar: Boucardicus monchenkoi sp. nov. and B. ambindaensis sp. nov. (Caenogastropoda: Cyclophoridae)" masterLastPageNumber="240" masterPageNumber="237" pageNumber="239" updateTime="1698621729662" updateUser="plazi">
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<mods:title id="6CFD38DD4EE937182C8C973BCD4BE8FD">Two new species of minute land snails from Madagascar: Boucardicus monchenkoi sp. nov. and B. ambindaensis sp. nov. (Caenogastropoda: Cyclophoridae)</mods:title>
</mods:titleInfo>
<mods:name id="3587EDD11FE707301C2EC53A1B9E5D19" type="personal">
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<mods:roleTerm id="71E053BE57B54B9A2EE96CD0B2C48AAA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1624EAC0DA6BBADA10A4744045B1E9AF">Balashov, Igor</mods:namePart>
</mods:name>
<mods:name id="2A1EDD679F2118E7261CD29B45DFD133" type="personal">
<mods:role id="8B0C4A66CFB3271ED690437FC610B2DE">
<mods:roleTerm id="06A15A52D2478F5DAAC81393E5FD4562">Author</mods:roleTerm>
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<mods:namePart id="4AE13A52798CE191154625F3A3C12773">Griffiths, Owen</mods:namePart>
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<mods:typeOfResource id="9FA31154D26374CCB3B62EC91AA80F9B">text</mods:typeOfResource>
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<mods:title id="AD58586094D2A239FA84FD55D6F4F563">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9A8462C92C06532C9DCA68E9E47DB325">
<mods:date id="0984D03F469616E946C164907391DD71">2015</mods:date>
<mods:detail id="F1819B5BC91592FAD671B987E36C720B" type="volume">
<mods:number id="2DAFF106394EFA40FE7ABD496EC7AF26">4052</mods:number>
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<mods:detail id="7E703E92703310C60E19FEDB39130901" type="issue">
<mods:number id="BF89994BC23E7872A184454BE8BDE3C7">2</mods:number>
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<mods:classification id="39ECD89B2522D388A6786BF0CCE2CE0C">journal article</mods:classification>
<mods:identifier id="C5300D385DAD993EFD35D5101600B1FA" type="DOI">10.11646/zootaxa.4052.2.9</mods:identifier>
<mods:identifier id="AB22F3C5CFD259EB7970C565830361F6" type="GBIF-Dataset">c2fc312c-097b-4aaa-911f-3a0785f6f01c</mods:identifier>
<mods:identifier id="DC8E40A82856700EB0FB8E61A17EC023" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F9EB51BCB9ABF5CFD622E1D9B31F34DA" type="Zenodo-Dep">234196</mods:identifier>
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</mods:mods>
<treatment id="9526643BB16BFFD6FF2BFF65364932CF" ID-DOI="http://doi.org/10.5281/zenodo.6103093" ID-GBIF-Taxon="119646531" ID-Zenodo-Dep="6103093" LSID="urn:lsid:plazi:treatment:9526643BB16BFFD6FF2BFF65364932CF" httpUri="http://treatment.plazi.org/id/9526643BB16BFFD6FF2BFF65364932CF" lastPageNumber="239" pageId="2" pageNumber="239">
<subSubSection id="559586A6B16BFFD6FF2BFF65369A37F2" box="[151,682,151,178]" pageId="2" pageNumber="239" type="nomenclature">
<paragraph id="1D30D52DB16BFFD6FF2BFF65369A37F2" blockId="2.[151,1436,151,1423]" box="[151,682,151,178]" pageId="2" pageNumber="239">
<heading id="46786241B16BFFD6FF2BFF65369A37F2" bold="true" box="[151,682,151,178]" fontSize="11" level="1" pageId="2" pageNumber="239" reason="1">
<emphasis id="2FFB093FB16BFFD6FF2BFF65369A37F2" bold="true" box="[151,682,151,178]" pageId="2" pageNumber="239">
<taxonomicName id="DA8FAEAEB16BFFD6FF2BFF65350037F1" ID-CoL="7NTNR" box="[151,304,151,177]" class="Gastropoda" family="Cyclophoridae" genus="Boucardicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="239" phylum="Mollusca" rank="genus">
<emphasis id="2FFB093FB16BFFD6FF2BFF65350037F1" bold="true" box="[151,304,151,177]" italics="true" pageId="2" pageNumber="239">Boucardicus</emphasis>
</taxonomicName>
<taxonomicName id="DA8FAEAEB16BFFD6FE8BFF6A35D237F2" box="[311,482,152,178]" class="Gastropoda" family="Cyclophoridae" genus="Boucardicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="239" phylum="Mollusca" rank="species" species="ambindaensis" status="sp. nov.">ambindaensis</taxonomicName>
<taxonomicNameLabel id="34C8B444B16BFFD6FE55FF6A367737F2" box="[489,583,152,178]" pageId="2" pageNumber="239" rank="species">sp. nov.</taxonomicNameLabel>
<figureCitation id="85B4C9A8B16BFFD6FDF3FF6A36A437F2" box="[591,660,152,178]" captionStart="FIGURE 1" captionStartId="1.[151,244,1975,1997]" captionTargetBox="[226,1360,1303,1951]" captionTargetId="figure@1.[226,1360,1303,1951]" captionTargetPageId="1" captionText="FIGURE 1. Holotypes of Boucardicus monchenkoi sp. nov. (a) and B. ambindaensis sp. nov. (b)." httpUri="https://zenodo.org/record/234197/files/figure.png" pageId="2" pageNumber="239">Fig. 1</figureCitation>
<emphasis id="2FFB093FB16BFFD6FD28FF65369337F1" bold="true" box="[660,675,151,177]" italics="true" pageId="2" pageNumber="239">b</emphasis>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="559586A6B16BFFD6FF2BFF48309336E1" pageId="2" pageNumber="239" type="materials_examined">
<paragraph id="1D30D52DB16BFFD6FF2BFF4836EC3657" blockId="2.[151,1436,151,1423]" pageId="2" pageNumber="239">
<emphasis id="2FFB093FB16BFFD6FF2BFF48351B3791" bold="true" box="[151,299,186,209]" pageId="2" pageNumber="239">
<typeStatus id="C2346B8FB16BFFD6FF2BFF4834FF3791" box="[151,207,186,209]" pageId="2" pageNumber="239">Type</typeStatus>
locality
</emphasis>
. Dry deciduous forest among the large vertical limestone rocks (tsingy), west side of Tsingy Beanka reserve, east of Ambinda and Belitsaka villages, around
<quantity id="DA7778C8B16BFFD6FD42FF2F377437B4" box="[766,836,221,244]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="239" unit="km" value="50.0">50 km</quantity>
east of Maintirano town, central-western
<collectingCountry id="659895BDB16BFFD6FAA9FF2F31A837B4" box="[1301,1432,221,244]" name="Madagascar" pageId="2" pageNumber="239">Madagascar</collectingCountry>
.
<geoCoordinate id="78BBB3EAB16BFFD6FF2BFEF235293657" box="[151,281,256,279]" direction="north" orientation="latitude" pageId="2" pageNumber="239" precision="1" value="18.04465">18.04465°N</geoCoordinate>
<geoCoordinate id="78BBB3EAB16BFFD6FEA3FEF2359C3657" box="[287,428,256,279]" direction="east" orientation="longitude" pageId="2" pageNumber="239" precision="1" value="44.501118">44.501117°E</geoCoordinate>
,
<quantity id="DA7778C8B16BFFD6FE0BFEF235CA3657" box="[439,506,256,279]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="239" unit="m" value="300.0">300 m</quantity>
altitude (from GPS).
</paragraph>
<paragraph id="1D30D52DB16BFFD6FF7BFED0309336E1" blockId="2.[151,1436,151,1423]" pageId="2" pageNumber="239">
<emphasis id="2FFB093FB16BFFD6FF7BFED0351A3679" bold="true" box="[199,298,290,313]" pageId="2" pageNumber="239">Material</emphasis>
.
<typeStatus id="C2346B8FB16BFFD6FE8BFED035AA3679" box="[311,410,290,313]" pageId="2" pageNumber="239" type="holotype">Holotype</typeStatus>
(
<collectionCode id="7B9E4DE8B16BFFD6FE14FED035D93679" box="[424,489,290,313]" pageId="2" pageNumber="239">IZAN</collectionCode>
GT 4631) and 9
<typeStatus id="C2346B8FB16BFFD6FD23FED037353679" box="[671,773,290,313]" pageId="2" pageNumber="239" type="paratype">paratypes</typeStatus>
(
<quantity id="DA7778C8B16BFFD6FCAFFED0370C3679" box="[787,828,290,313]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="239" unit="in" value="6.0">6 in</quantity>
<collectionCode id="7B9E4DE8B16BFFD6FCFFFED037B53679" box="[835,901,290,313]" pageId="2" pageNumber="239">IZAN</collectionCode>
, GT 4632, 3 in AMS, C.483541) are directly from the
<typeStatus id="C2346B8FB16BFFD6FF02FEB734DC361C" box="[190,236,325,348]" pageId="2" pageNumber="239">type</typeStatus>
locality (collected by R. Randalana &amp; G. Middleton on 2th October of 2006), 3 more
<typeStatus id="C2346B8FB16BFFD6FB39FEB730DB361C" box="[1157,1259,325,348]" pageId="2" pageNumber="239" type="paratype">paratypes</typeStatus>
(
<quantity id="DA7778C8B16BFFD6FB46FEB73114361C" box="[1274,1316,325,348]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="239" unit="in" value="2.0">2 in</quantity>
<collectionCode id="7B9E4DE8B16BFFD6FA97FEB7315D361C" box="[1323,1389,325,348]" pageId="2" pageNumber="239">IZAN</collectionCode>
, GT 4633, 1 in AMS, C.483542) were collected in other locations in south-eastern part of Tsingy Beanka reserve east to Belitsaka village on 3th October of 2006 (from GPS:
<geoCoordinate id="78BBB3EAB16BFFD6FD6DFE78375136E1" box="[721,865,394,417]" direction="north" orientation="latitude" pageId="2" pageNumber="239" precision="1" value="18.054167">18.054167°N</geoCoordinate>
<geoCoordinate id="78BBB3EAB16BFFD6FCDBFE7837C636E1" box="[871,1014,394,417]" direction="east" orientation="longitude" pageId="2" pageNumber="239" precision="1" value="44.533333">44.533333°E</geoCoordinate>
,
<quantity id="DA7778C8B16BFFD6FBBDFE78307436E1" box="[1025,1092,394,417]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="239" unit="m" value="350.0">350 m</quantity>
altitude)
</paragraph>
</subSubSection>
<subSubSection id="559586A6B16BFFD6FF7BFE5F36C83684" box="[199,760,429,452]" pageId="2" pageNumber="239" type="etymology">
<paragraph id="1D30D52DB16BFFD6FF7BFE5F36C83684" blockId="2.[151,1436,151,1423]" box="[199,760,429,452]" pageId="2" pageNumber="239">
<emphasis id="2FFB093FB16BFFD6FF7BFE5F35703684" bold="true" box="[199,320,429,452]" pageId="2" pageNumber="239">Etymology</emphasis>
. From occurrence near Ambinda village.
</paragraph>
</subSubSection>
<subSubSection id="559586A6B16BFFD6FF7BFE2236BC3267" pageId="2" pageNumber="239" type="diagnosis">
<paragraph id="1D30D52DB16BFFD6FF7BFE223562350F" blockId="2.[151,1436,151,1423]" pageId="2" pageNumber="239">
<emphasis id="2FFB093FB16BFFD6FF7BFE22350436A7" bold="true" box="[199,308,464,487]" pageId="2" pageNumber="239">Diagnosis</emphasis>
. Shell is pupilloid-like, almost smooth (with weak radial lines), aperture is almost round, with single columellar lamella relatively deep in aperture that goes through almost whole body-whorl inside it and single palatal lamella in the begging of body-whorl (could be visible through wall of shell). Basal peristome is almost complete, slightly reflected.
</paragraph>
<paragraph id="1D30D52DB16BFFD6FF7BFDA835BF35F7" blockId="2.[151,1436,151,1423]" pageId="2" pageNumber="239">
<emphasis id="2FFB093FB16BFFD6FF7BFDA8357B3531" bold="true" box="[199,331,602,625]" pageId="2" pageNumber="239">Dimensions</emphasis>
. Height of shell
<quantity id="DA7778C8B16BFFD6FDB9FDA836B93531" box="[517,649,602,625]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="2" pageNumber="239" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.42.5 mm</quantity>
, diameter
<quantity id="DA7778C8B16BFFD6FD40FDA8374D3531" box="[764,893,602,625]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="2" pageNumber="239" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.21.3 mm</quantity>
(
<typeStatus id="C2346B8FB16BFFD6FC31FDA837DC3531" box="[909,1004,602,625]" pageId="2" pageNumber="239" type="holotype">holotype</typeStatus>
2.4x
<quantity id="DA7778C8B16BFFD6FB99FDA8304B3531" box="[1061,1147,602,625]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="239" unit="mm" value="1.2">1.2 mm</quantity>
). Aperture height
<quantity id="DA7778C8B16BFFD6FAFDFDA831A73531" box="[1345,1431,602,625]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="239" unit="mm" value="0.7">0.7 mm</quantity>
, aperture width 0.8 mB. Hight of body whorl—1.11.2 mB. Diameter of embryonic whorls (~1.5) 0.30.4 mB. Diameter of 3rd whorl—
<quantity id="DA7778C8B16BFFD6FE84FD5235BC35F7" box="[312,396,672,695]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="239" unit="mm" value="0.8">0.8 mm</quantity>
.
</paragraph>
<paragraph id="1D30D52DB16BFFD6FF7BFD3037CB3317" blockId="2.[151,1436,151,1423]" pageId="2" pageNumber="239">
<emphasis id="2FFB093FB16BFFD6FF7BFD3035303599" bold="true" box="[199,256,706,729]" pageId="2" pageNumber="239">Shell</emphasis>
is pupilloid-like, ovate-pointed, moderately thin-walled, translucent, of 5 moderately convexed whorls (sometimes 5.25). Suture is moderate. Body-whorl periphery round. Coloration is light-brown. Whole shell is smooth (only somewhere weak radial lines are present), except post-constriction swelling that irregularly striated or even sometimes almost ribbed. Body-whorl constriction occurs about 0.6 whorl before the peristome. The post-constriction swelling is moderately weak, about
<quantity id="DA7778C8B16BFFD6FDA5FCBF365B3424" box="[537,619,845,868]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="239" unit="mm" value="0.3">0.3 mm</quantity>
width, placed left to columellar peristome. Aperture is almost round, with a single clear columellar lamella near edge of aperture in the middle of columellar peristome. Columellar lamella is placed perpendicularly to the columellar peristome, goes almost through whole body-whorl on columella (one
<typeStatus id="C2346B8FB16BFFD6FAB0FC60315834E9" box="[1292,1384,914,937]" pageId="2" pageNumber="239" type="paratype">paratype</typeStatus>
was studied inside). Inside body-whorl there is a long palatal lamella that could be visible through wall of the shell in fresh specimens. In
<typeStatus id="C2346B8FB16BFFD6FE8EFC2A35A234AF" box="[306,402,984,1007]" pageId="2" pageNumber="239" type="holotype">holotype</typeStatus>
, as it looks from outside, palatal lamella goes from beginning of body-whorl till almost last 0.5 whorl (length about
<quantity id="DA7778C8B16BFFD6FED2FC0835F43351" box="[366,452,1018,1041]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="239" unit="mm" value="0.7">0.7 mm</quantity>
). In
<typeStatus id="C2346B8FB16BFFD6FE4DFC08367D3351" box="[497,589,1018,1041]" pageId="2" pageNumber="239" type="paratype">paratype</typeStatus>
that was studied inside (old not translucent shell) lamella is short,
<quantity id="DA7778C8B16BFFD6FAB2FC08315F3351" box="[1294,1391,1018,1041]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="239" unit="mm" value="0.25">0.25 mm</quantity>
, but large and high. Peristome is almost complete, slightly interrupted (in upper part in place of connection with penultimate whorl). Basal peristome slightly reflected. Lip moderate. Umbilicus drop-shaped.
</paragraph>
<paragraph id="1D30D52DB16BFFD6FF7BFB90316633DC" blockId="2.[151,1436,151,1423]" pageId="2" pageNumber="239">
<emphasis id="2FFB093FB16BFFD6FF7BFB90350F3339" bold="true" box="[199,319,1122,1145]" pageId="2" pageNumber="239">Variability</emphasis>
. All 13
<typeStatus id="C2346B8FB16BFFD6FE25FB9035F73339" box="[409,455,1122,1145]" pageId="2" pageNumber="239">type</typeStatus>
specimens are very similar in the size, shape, sculpture, swelling and aperture, including
<specimenCount id="0B891EA4B16BFFD6FA32FB90353633DC" pageId="2" pageNumber="239" type="generic">3 specimens</specimenCount>
from second locality. Therefore same to the previous species known variability of the shell is low here.
</paragraph>
<paragraph id="1D30D52DB16BFFD6FF7BFB5A36BC3267" blockId="2.[151,1436,151,1423]" pageId="2" pageNumber="239">
<emphasis id="2FFB093FB16BFFD6FF7BFB5A355133FF" bold="true" box="[199,353,1192,1215]" pageId="2" pageNumber="239">Identification</emphasis>
. From the other pupilloid-like species of the genus except
<taxonomicName id="DA8FAEAEB16BFFD6FC4AFB5A30DB33FF" box="[1014,1259,1191,1215]" class="Gastropoda" family="Cyclophoridae" genus="Boucardicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="239" phylum="Mollusca" rank="species" species="pseudogastrocoptus">
<emphasis id="2FFB093FB16BFFD6FC4AFB5A30DB33FF" box="[1014,1259,1191,1215]" italics="true" pageId="2" pageNumber="239">B. pseudogastrocoptus</emphasis>
</taxonomicName>
it differs by its round aperture with almost complete peristome. From the most—by the well developed both columellar and palatal lamellae. From
<taxonomicName id="DA8FAEAEB16BFFD6FEF8FB1F36093244" box="[324,569,1260,1284]" class="Gastropoda" family="Cyclophoridae" genus="Boucardicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="239" phylum="Mollusca" rank="species" species="pseudogastrocoptus">
<emphasis id="2FFB093FB16BFFD6FEF8FB1F36093244" box="[324,569,1260,1284]" italics="true" pageId="2" pageNumber="239">B. pseudogastrocoptus</emphasis>
</taxonomicName>
it differs by presence of palatal lamella and much more developed columellar lamella, by absence of lip and by smaller shell.
</paragraph>
</subSubSection>
<subSubSection id="559586A6B16BFFD6FF7BFAC0364932CF" pageId="2" pageNumber="239" type="distribution">
<paragraph id="1D30D52DB16BFFD6FF7BFAC0364932CF" blockId="2.[151,1436,151,1423]" pageId="2" pageNumber="239">
<emphasis id="2FFB093FB16BFFD6FF7BFAC035D53209" bold="true" box="[199,485,1330,1353]" pageId="2" pageNumber="239">Distribution and habitats</emphasis>
. It lives in leaflitter in deciduous forests among the large vertical limestone rocks (tsingy) placed on altitude of
<quantity id="DA7778C8B16BFFD6FECBFAA735C1322C" box="[375,497,1365,1388]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="2" pageNumber="239" unit="m" value="325.0" valueMax="350.0" valueMin="300.0">300350 m</quantity>
, in Tsingy Beanka reserve east of Ambinda and Belitsaka villages, around
<quantity id="DA7778C8B16BFFD6FAB0FAA7317F322C" box="[1292,1359,1365,1388]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="239" unit="km" value="50.0">50 km</quantity>
east of Maintirano city, central-western
<collectingCountry id="659895BDB16BFFD6FE4EFA8A364532CF" box="[498,629,1400,1423]" name="Madagascar" pageId="2" pageNumber="239">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>