173 lines
18 KiB
XML
173 lines
18 KiB
XML
<document id="3CDBA1346B5E62FCF1B37B308A58437D" ID-DOI="10.11646/zootaxa.4052.2.9" ID-GBIF-Dataset="c2fc312c-097b-4aaa-911f-3a0785f6f01c" ID-ISSN="1175-5326" ID-Zenodo-Dep="234196" ID-ZooBank="22AEEFF3-0419-4549-BF29-E7A6AE1D77E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461350448683" checkinUser="plazi" docAuthor="Balashov, Igor & Griffiths, Owen" docDate="2015" docId="9526643BB169FFD5FF2BF87D30CC33A1" docLanguage="en" docName="zt04052p240.pdf" docOrigin="Zootaxa 4052 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Boucardicus monchenkoi Balashov & Griffiths, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="238" masterDocId="691F1C43B169FFD4FFBCFFF234303740" masterDocTitle="Two new species of minute land snails from Madagascar: Boucardicus monchenkoi sp. nov. and B. ambindaensis sp. nov. (Caenogastropoda: Cyclophoridae)" masterLastPageNumber="240" masterPageNumber="237" pageNumber="237" updateTime="1698621729662" updateUser="plazi">
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<mods:mods id="6417FEFCA1D3154C0AFE51FAD02B7E8A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="537041CF8FB7E160EC3F9C87A7B219F7">
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<mods:title id="22A8332975F7DCA19895F1FAC666B9C6">Two new species of minute land snails from Madagascar: Boucardicus monchenkoi sp. nov. and B. ambindaensis sp. nov. (Caenogastropoda: Cyclophoridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="933C8B8D1B5305B89CE5864671B5A0CA" type="personal">
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<mods:roleTerm id="14EBC1B622A1F2E14E282DB8BA374BB5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9D45FD8D402C4DAA812EFBFC11B0449F">Balashov, Igor</mods:namePart>
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</mods:name>
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<mods:name id="C4E6847C0D2BF3174D20D927FB1C287F" type="personal">
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<mods:roleTerm id="9EC247CF8DCD732124F716D4829FB1B5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="641C63F7B6BCF5895B75C64971591B4D">Griffiths, Owen</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="F0B16516660F6D0CED95990C87F00524">text</mods:typeOfResource>
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<mods:titleInfo id="8BD648C7E1842B5DE2858867E529F721">
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<mods:title id="D0AF5E9A607925F381459130EC2C68BC">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="069E7CEA36488DD5B7071D227390BB8C">
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<mods:date id="9C264F410E4A0D7D254D34798CCEFF45">2015</mods:date>
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<mods:detail id="76039177378D38B35C5F90AC473A1D9C" type="volume">
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<mods:number id="51227305065E3A646D7A026996AB8DFE">4052</mods:number>
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</mods:detail>
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<mods:detail id="42C423A2073D57722F51B84646F17F6C" type="issue">
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<mods:number id="92D5479EBA5ACADF055E2CEEE8AE920E">2</mods:number>
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<mods:start id="74E6D4838945B456A93AA999CA6B8F96">237</mods:start>
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<mods:end id="6535C3A7EE6C640FB5F78BC9C13638E7">240</mods:end>
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<mods:classification id="3C6D5FCFB7B6BE52F95E798361C96209">journal article</mods:classification>
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<mods:identifier id="D99C1400491B2B565EFCB1400EF663FC" type="DOI">10.11646/zootaxa.4052.2.9</mods:identifier>
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<mods:identifier id="96605D801CA3586F263EEDD1126C93CC" type="GBIF-Dataset">c2fc312c-097b-4aaa-911f-3a0785f6f01c</mods:identifier>
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<mods:identifier id="26EBDDC0A563B77C1DA9E3313550BB22" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="887BFA24CFB614560A3C59E0E4071613" type="Zenodo-Dep">234196</mods:identifier>
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<mods:identifier id="4777EC92C4FA22E0ACC4D7CD6D25F212" type="ZooBank">22AEEFF3-0419-4549-BF29-E7A6AE1D77E5</mods:identifier>
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</mods:mods>
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<treatment id="9526643BB169FFD5FF2BF87D30CC33A1" ID-DOI="http://doi.org/10.5281/zenodo.6103091" ID-GBIF-Taxon="119646534" ID-Zenodo-Dep="6103091" LSID="urn:lsid:plazi:treatment:9526643BB169FFD5FF2BF87D30CC33A1" httpUri="http://treatment.plazi.org/id/9526643BB169FFD5FF2BF87D30CC33A1" lastPageId="1" lastPageNumber="238" pageId="0" pageNumber="237">
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<subSubSection id="559586A6B169FFD4FF2BF87D36A530E9" box="[151,661,1935,1961]" pageId="0" pageNumber="237" type="nomenclature">
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<paragraph id="1D30D52DB169FFD4FF2BF87D36A530E9" blockId="0.[151,1435,1935,2027]" box="[151,661,1935,1961]" pageId="0" pageNumber="237">
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<heading id="46786241B169FFD4FF2BF87D36A530E9" bold="true" box="[151,661,1935,1961]" fontSize="11" level="1" pageId="0" pageNumber="237" reason="1">
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<emphasis id="2FFB093FB169FFD4FF2BF87D36A530E9" bold="true" box="[151,661,1935,1961]" pageId="0" pageNumber="237">
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<taxonomicName id="DA8FAEAEB169FFD4FF2BF87D35FD30E9" ID-CoL="5WQMW" box="[151,461,1935,1961]" class="Gastropoda" family="Cyclophoridae" genus="Boucardicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="237" phylum="Mollusca" rank="species" species="monchenkoi" status="sp. nov.">
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<emphasis id="2FFB093FB169FFD4FF2BF87D35FD30E9" bold="true" box="[151,461,1935,1961]" italics="true" pageId="0" pageNumber="237">Boucardicus monchenkoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="34C8B444B169FFD4FE68F87D360230E9" box="[468,562,1935,1961]" pageId="0" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="85B4C9A8B169FFD4FD85F87D364F30E9" box="[569,639,1935,1961]" captionStart="FIGURE 1" captionStartId="1.[151,244,1975,1997]" captionTargetBox="[226,1360,1303,1951]" captionTargetId="figure@1.[226,1360,1303,1951]" captionTargetPageId="1" captionText="FIGURE 1. Holotypes of Boucardicus monchenkoi sp. nov. (a) and B. ambindaensis sp. nov. (b)." httpUri="https://zenodo.org/record/234197/files/figure.png" pageId="0" pageNumber="237">Fig. 1</figureCitation>
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<emphasis id="2FFB093FB169FFD4FDC3F87D36BE30E9" bold="true" box="[639,654,1935,1961]" italics="true" pageId="0" pageNumber="237">a</emphasis>
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.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="559586A6B169FFD5FF2BF840369E3657" lastPageId="1" lastPageNumber="238" pageId="0" pageNumber="237" type="materials_examined">
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<paragraph id="1D30D52DB169FFD4FF2BF840354030AB" blockId="0.[151,1435,1935,2027]" pageId="0" pageNumber="237">
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<emphasis id="2FFB093FB169FFD4FF2BF84035183089" bold="true" box="[151,296,1970,1993]" pageId="0" pageNumber="237">
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<typeStatus id="C2346B8FB169FFD4FF2BF84034FF3089" box="[151,207,1970,1993]" pageId="0" pageNumber="237">Type</typeStatus>
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locality
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</emphasis>
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. Coastal grassland near Hotel Dauphin (Avenue Gallien), Tôlanaro city (formerly Fort Dauphin), southeastern
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<collectingCountry id="659895BDB169FFD4FF55F826355C30AB" box="[233,364,2004,2027]" name="Madagascar" pageId="0" pageNumber="237">Madagascar</collectingCountry>
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.
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</paragraph>
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<paragraph id="1D30D52DB168FFD5FF7BFF6A369E3657" blockId="1.[151,1436,152,1249]" pageId="1" pageNumber="238">
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<emphasis id="2FFB093FB168FFD5FF7BFF6A351A37EF" bold="true" box="[199,298,152,175]" pageId="1" pageNumber="238">Material</emphasis>
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.
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<typeStatus id="C2346B8FB168FFD5FE86FF6A35AD37EF" box="[314,413,152,175]" pageId="1" pageNumber="238" type="holotype">Holotype</typeStatus>
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(
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<collectionCode id="7B9E4DE8B168FFD5FE11FF6A35DE37EF" box="[429,494,152,175]" pageId="1" pageNumber="238">IZAN</collectionCode>
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GT 4629) and 10
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<typeStatus id="C2346B8FB168FFD5FD01FF6A371337EF" box="[701,803,152,175]" pageId="1" pageNumber="238" type="paratype">paratypes</typeStatus>
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(
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<quantity id="DA7778C8B168FFD5FC89FF6A375037EF" box="[821,864,152,175]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="1" pageNumber="238" unit="in" value="7.0">7 in</quantity>
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<collectionCode id="7B9E4DE8B168FFD5FCD5FF6A379B37EF" box="[873,939,152,175]" pageId="1" pageNumber="238">IZAN</collectionCode>
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, GT 4630, 3 in AMS, C.483540) are directly from the
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<typeStatus id="C2346B8FB168FFD5FF44FF4835163791" box="[248,294,186,209]" pageId="1" pageNumber="238">type</typeStatus>
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locality (collected by
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<collectingCountry id="659895BDB168FFD5FDAFFF4836193791" box="[531,553,186,209]" name="Iceland" pageId="1" pageNumber="238">O</collectingCountry>
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. Griffiths in
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<date id="6931F3EDB168FFD5FD0CFF4837063791" box="[688,822,186,209]" pageId="1" pageNumber="238" value="2001-03">March 2001</date>
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), 1 more
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<typeStatus id="C2346B8FB168FFD5FC25FF4837C53791" box="[921,1013,186,209]" pageId="1" pageNumber="238" type="paratype">paratype</typeStatus>
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(
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<collectionCode id="7B9E4DE8B168FFD5FBBEFF4830733791" box="[1026,1091,186,209]" pageId="1" pageNumber="238">IZAN</collectionCode>
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GT 4563) was collected "on the sandy hills with succulents
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<quantity id="DA7778C8B168FFD5FE74FF2F367037B4" box="[456,576,221,244]" metricMagnitude="2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="1" pageNumber="238" unit="m" value="110.0" valueMax="120.0" valueMin="100.0">100-120 m</quantity>
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from the ocean near Fort Dauphin" by V. I. Monchenko in February of 1991 together with some shells of the larger land snails.
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</paragraph>
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</subSubSection>
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<subSubSection id="559586A6B168FFD5FF7BFED03760363F" pageId="1" pageNumber="238" type="etymology">
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<paragraph id="1D30D52DB168FFD5FF7BFED03760363F" blockId="1.[151,1436,152,1249]" pageId="1" pageNumber="238">
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<emphasis id="2FFB093FB168FFD5FF7BFED035703679" bold="true" box="[199,320,290,313]" pageId="1" pageNumber="238">Etymology</emphasis>
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. The specific name,
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<taxonomicName id="DA8FAEAEB168FFD5FD8EFED336833679" box="[562,691,289,313]" class="Gastropoda" family="Cyclophoridae" genus="Boucardicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="238" phylum="Mollusca" rank="species" species="monchenkoi">
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<emphasis id="2FFB093FB168FFD5FD8EFED336833679" box="[562,691,289,313]" italics="true" pageId="1" pageNumber="238">monchenkoi</emphasis>
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</taxonomicName>
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, honors collector of the first known specimen—Prof. Vladislav I. Monchenko, Academician of National Academy of Sciences of
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<collectingCountry id="659895BDB168FFD5FCDCFEB73789361C" box="[864,953,325,348]" name="Ukraine" pageId="1" pageNumber="238">Ukraine</collectingCountry>
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, founder of the Department of Invertebrate Fauna and Systematics of I.I. Schmalhausen Institute of Zoology.
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</paragraph>
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</subSubSection>
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<subSubSection id="559586A6B168FFD5FF7BFE78366933DC" pageId="1" pageNumber="238" type="diagnosis">
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<paragraph id="1D30D52DB168FFD5FF7BFE7835A336A7" blockId="1.[151,1436,152,1249]" pageId="1" pageNumber="238">
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<emphasis id="2FFB093FB168FFD5FF7BFE78350436E1" bold="true" box="[199,308,394,417]" pageId="1" pageNumber="238">Diagnosis</emphasis>
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. Shell is pupilloid-like, clearly ribbed, aperture is ovate-triangular with single well developed columellar lamella that goes from beginning of body-whorl till edge of the aperture and clearly visible in aperture. Peristome is complete, not reflected.
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</paragraph>
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<paragraph id="1D30D52DB168FFD5FF7BFE00354C350F" blockId="1.[151,1436,152,1249]" pageId="1" pageNumber="238">
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<emphasis id="2FFB093FB168FFD5FF7BFE00357B3549" bold="true" box="[199,331,498,521]" pageId="1" pageNumber="238">Dimensions</emphasis>
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. Height of shell
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<quantity id="DA7778C8B168FFD5FE47FE00364D3549" box="[507,637,498,521]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="1" pageNumber="238" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
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, diameter 1.3–1.5 mB.
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<typeStatus id="C2346B8FB168FFD5FCC9FE0037E83549" box="[885,984,498,521]" pageId="1" pageNumber="238" type="holotype">Holotype</typeStatus>
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2.7x1.5 mB. Aperture height
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<quantity id="DA7778C8B168FFD5FAA9FE0031A73549" box="[1301,1431,498,521]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="1" pageNumber="238" unit="mm" value="0.8" valueMax="0.9" valueMin="0.7">0.7–0.9 mm</quantity>
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, aperture width 0.7–0.8 mB. Height of body whorl—1.1–1.2 mB. Diameter of embryonic whorls (~1.5) 0.4 mB. Diameter of 3rd whorl—
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<quantity id="DA7778C8B168FFD5FE80FDCA3547350F" box="[316,375,568,591]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="238" unit="mm" value="1.0">1 mm</quantity>
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.
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</paragraph>
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<paragraph id="1D30D52DB168FFD5FF7BFDA8370534E9" blockId="1.[151,1436,152,1249]" pageId="1" pageNumber="238">
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<emphasis id="2FFB093FB168FFD5FF7BFDA835303531" bold="true" box="[199,256,602,625]" pageId="1" pageNumber="238">Shell</emphasis>
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is pupilloid-like, ovate-pointed, moderately thick-walled, not translucent, of 5–5.5 strongly convexed whorls. Suture is deep. Body-whorl periphery round. Coloration is brown (
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<typeStatus id="C2346B8FB168FFD5FCDEFD8F37AC35D4" box="[866,924,637,660]" pageId="1" pageNumber="238">types</typeStatus>
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are mainly lost their pigmentation). Embryonic whorls (~1.5) are smooth. Pre-constriction sculpture is regularly radially ribbed, on the 3rd whorl there are around 30 ribs in
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<quantity id="DA7778C8B168FFD5FF5EFD3035113599" box="[226,289,706,729]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="238" unit="mm" value="1.0">1 mm</quantity>
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(distance between ribs about
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<quantity id="DA7778C8B168FFD5FDDDFD3036F43599" box="[609,708,706,729]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="238" unit="mm" value="0.04">0.04 mm</quantity>
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), on the 4th whorl—around 22 ribs in
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<quantity id="DA7778C8B168FFD5FBDDFD3030AF3599" box="[1121,1183,706,729]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="238" unit="mm" value="1.0">1 mm</quantity>
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(placed less regularly). First post-constriction ribs are same to 4th whorl, but placed more distantly. On the last 0.25 whorl before the peristome the ribs are much stronger. Body-whorl constriction occurs about 0.6 whorls before the peristome. The post-constriction swelling is moderate, about
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<quantity id="DA7778C8B168FFD5FE7DFCD836223401" box="[449,530,810,833]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="238" unit="mm" value="0.3">0.3 mm</quantity>
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width, placed left to columellar peristome. Aperture is ovate-triangular, with a single clear columellar lamella (peg) near edge of aperture in the middle of columellar peristome. Lamella is placed strictly perpendicularly to the columellar peristome, goes deeply in the aperture. Peristome is complete, not reflected, only slightly near umbilicus. Lip moderate. Umbilicus drop-shaped.
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</paragraph>
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<paragraph id="1D30D52DB168FFD5FF7BFC4734DB3351" blockId="1.[151,1436,152,1249]" pageId="1" pageNumber="238">
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<emphasis id="2FFB093FB168FFD5FF7BFC47350F348C" bold="true" box="[199,319,949,972]" pageId="1" pageNumber="238">Variability</emphasis>
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. All 12
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<typeStatus id="C2346B8FB168FFD5FE1AFC4735E4348C" box="[422,468,949,972]" pageId="1" pageNumber="238">type</typeStatus>
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specimens are very similar in size, shape, sculpture, swelling and aperture. An older specimen from the other locality is almost identical to the
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<typeStatus id="C2346B8FB168FFD5FD41FC2A376E34AF" box="[765,862,984,1007]" pageId="1" pageNumber="238" type="holotype">holotype</typeStatus>
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. Therefore known variability of the shell is low in this species.
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</paragraph>
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<paragraph id="1D30D52DB168FFD5FF7BFBEF366933DC" blockId="1.[151,1436,152,1249]" pageId="1" pageNumber="238">
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<emphasis id="2FFB093FB168FFD5FF7BFBEF35513374" bold="true" box="[199,353,1053,1076]" pageId="1" pageNumber="238">Identification</emphasis>
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. From the all other pupilloid-like species of the genus it differs by its complete aperture, from mostalso by its clearly ribbed shell. From
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<taxonomicName id="DA8FAEAEB168FFD5FDF5FBB2369D3317" box="[585,685,1088,1111]" class="Gastropoda" family="Cyclophoridae" genus="Boucardicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="238" phylum="Mollusca" rank="species" species="menoi">
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<emphasis id="2FFB093FB168FFD5FDF5FBB2369D3317" box="[585,685,1088,1111]" italics="true" pageId="1" pageNumber="238">B. menoi</emphasis>
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</taxonomicName>
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—by much more developed columellar lamella visible in aperture, presence of post-constriction swelling and larger shell. From
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<taxonomicName id="DA8FAEAEB168FFD5FC81FB90378E3339" box="[829,958,1121,1145]" class="Gastropoda" family="Cyclophoridae" genus="Boucardicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="238" phylum="Mollusca" rank="species" species="delicatus">
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<emphasis id="2FFB093FB168FFD5FC81FB90378E3339" box="[829,958,1121,1145]" italics="true" pageId="1" pageNumber="238">B. delicatus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="DA8FAEAEB168FFD5FC44FB90305D3339" box="[1016,1133,1122,1145]" class="Gastropoda" family="Cyclophoridae" genus="Boucardicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="238" phylum="Mollusca" rank="species" species="minutus">
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<emphasis id="2FFB093FB168FFD5FC44FB90305D3339" box="[1016,1133,1122,1145]" italics="true" pageId="1" pageNumber="238">B. minutus</emphasis>
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</taxonomicName>
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—by much larger shell and much more developed columellar lamella.
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</paragraph>
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</subSubSection>
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<subSubSection id="559586A6B168FFD5FF7BFB5A30CC33A1" pageId="1" pageNumber="238" type="distribution">
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<paragraph id="1D30D52DB168FFD5FF7BFB5A30CC33A1" blockId="1.[151,1436,152,1249]" pageId="1" pageNumber="238">
|
||
<emphasis id="2FFB093FB168FFD5FF7BFB5A35D533FF" bold="true" box="[199,485,1192,1215]" pageId="1" pageNumber="238">Distribution and habitats</emphasis>
|
||
. It is only known from coastal grasslands near Tôlanaro city (south-eastern
|
||
<collectingCountry id="659895BDB168FFD5FAB7FB5A31A133FF" box="[1291,1425,1192,1215]" name="Madagascar" pageId="1" pageNumber="238">Madagascar</collectingCountry>
|
||
). This area was once covered in primary forest growing on coastal dunes and has now mostly been cleared.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |