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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.430.8094" ID-PMC="PMC4141162" ID-Pensoft-Pub="1313-2970-430-1" ID-Pensoft-UUID="FFE6FF8CFFD43B66335AD2203774DB49" ID-PubMed="25152663" ID-Zenodo-Dep="578441" ID-ZooBank="86F760563B8B49FB9C86FAD0DB0CBE8C" ModsDocID="1313-2970-430-1" checkinTime="1451245551152" checkinUser="pensoft" docAuthor="Erwin, Terry L. &amp; Zamorano, Laura S." docDate="2014" docId="94E6498E127A059008C6C673214BB971" docLanguage="en" docName="ZooKeys 430: 1-108" docOrigin="ZooKeys 430" docPubDate="2014-08-01" docSource="http://dx.doi.org/10.3897/zookeys.430.8094" docTitle="Pseudophorticus Erwin 2004" docType="treatment" docVersion="5" id="FFE6FF8CFFD43B66335AD2203774DB49" lastPageNumber="21" masterDocId="FFE6FF8CFFD43B66335AD2203774DB49" masterDocTitle="A synopsis of the tribe Lachnophorini, with a new genus of Neotropical distribution and a revision of the Neotropical genus Asklepia Liebke, 1938 (Insecta, Coleoptera, Carabidae)" masterLastPageNumber="108" masterPageNumber="1" pageNumber="21" updateTime="1668159067277" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A synopsis of the tribe Lachnophorini, with a new genus of Neotropical distribution and a revision of the Neotropical genus Asklepia Liebke, 1938 (Insecta, Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Erwin, Terry L.</mods:namePart>
<mods:affiliation>Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
<mods:nameIdentifier type="email">erwint@si.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamorano, Laura S.</mods:namePart>
<mods:affiliation>Research Student, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA &amp; Laboratorio de Zoologia Acuatica LAZOEA, Departamento de Ciencias Biologicas, Universidad de los Andes, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-08-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>430</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>108</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.430.8094</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.430.8094</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-430-1</mods:identifier>
<mods:identifier type="ZooBank">86F760563B8B49FB9C86FAD0DB0CBE8C</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFE6FF8CFFD43B66335AD2203774DB49</mods:identifier>
<mods:identifier type="Zenodo-Dep">578441</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152054744" LSID="urn:lsid:plazi:treatment:94E6498E127A059008C6C673214BB971" httpUri="http://treatment.plazi.org/id/94E6498E127A059008C6C673214BB971" lastPageNumber="21" pageId="20" pageNumber="21">
<subSubSection pageId="20" pageNumber="21" type="nomenclature">
<paragraph pageId="20" pageNumber="21">
<taxonomicName LSID="94E6498E-127A-0590-08C6-C673214BB971" authority="Erwin, 2004" authorityName="Erwin" authorityYear="2004" class="Insecta" family="Carabidae" genus="Pseudophorticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophorticus" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Pseudophorticus Erwin, 2004</taxonomicName>
False beauty-bearing beetles
<figureCitation captionStart="Figure 1922" captionStartId="F7" captionText="Figure 19 - 22. 19 Pseudophorticus sp. (all species in South America are either undescribed or placed in Euphorticus G. Horn). Digital Photo-illustration. Habitus, dorsal aspect, based on specimen ADP 132538 from Pakitza, Peru 20 Selina westermanni Motschulsky. Digital Photo-illustration. Habitus, dorsal aspect, based on specimen ADP 132536 from China Bay, Sri Lanka 21 Stenocheila lacordairei Laporte de Castelnau. Digital Photo-illustration. Habitus, dorsal aspect, based on specimen ADP 133801 from Chapada dos Guimares, Brazil 22 Quammenis spectabilis Erwin. Digital Photo-illustration. Habitus, dorsal aspect, based on specimen ADP 100513 from Estacion Zurqui, Costa Rica." httpUri="https://binary.pensoft.net/fig/30637" pageId="20" pageNumber="21">Figs 19</figureCitation>
<figureCitation captionStart="Figures 2328" captionStartId="F8" captionText="Figures 23 - 28. Mouthparts of adults. 23 Eucaerus sp. Mouthparts, ventral aspect, based on specimen ADP 132552 from Pakitza, Peru 24 Euphorticus sp. Mouthparts, ventral aspect, based on specimen ADP 132558 from Tapilula, Mexico 25 Guatemalteca virgen Erwin. Mouthparts, ventral aspect, based on specimen ADP 132546 from nr. La Virgen, Costa Rica 26 Lachnaces sp. Mouthparts, ventral aspect, based on specimen ADP 132578 from Tambopata Reserved Zone, Explorer's Inn, Peru 27 Lachnophorus sp. Mouthparts, ventral aspect, based on specimen ADP 132570 from Pakitza, Peru 28 Pseudophorticus sp. Mouthparts, ventral aspect, based on specimen ADP 132538 from Pakitza, Peru. Legend: a 3 Antennomere 3; mt Mentum tooth; m Mentum; lp 3 Labial palpomere 3; mp 4 Maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/30638" pageId="20" pageNumber="21">, 28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="reference_group">
<paragraph pageId="20" pageNumber="21">
<taxonomicName authorityName="Erwin" authorityYear="2004" class="Insecta" family="Carabidae" genus="Pseudophorticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Pseudophorticus</taxonomicName>
Erwin, 2004:7
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="type species">
<paragraph pageId="20" pageNumber="21">Type species.</paragraph>
<paragraph pageId="20" pageNumber="21">
<taxonomicName authorityName="Erwin" authorityYear="2004" class="Insecta" family="Carabidae" genus="Pseudophorticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis italics="true" pageId="20" pageNumber="21">Pseudophorticus puncticollis</emphasis>
</taxonomicName>
Erwin, 2004:8
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="way of life">
<paragraph pageId="20" pageNumber="21">Way of life.</paragraph>
<paragraph pageId="20" pageNumber="21">Size range - 4.7 mm to 6.2 mm; these small beetles occur on the ground in rainforests; they are diurnal and run in clearings and on trails in open spots. Nothing is known about their way of life.</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="distribution">
<paragraph pageId="20" pageNumber="21">Distribution.</paragraph>
<paragraph pageId="20" pageNumber="21">
Costa Rica south to
<normalizedToken originalValue="Perú">Peru</normalizedToken>
and southeastern Brazil.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="notes">
<paragraph pageId="20" pageNumber="21">Notes.</paragraph>
<paragraph pageId="20" pageNumber="21">
Currently, one described species is assigned to this genus. Many undescribed species are represented in collections, misidentified as either
<taxonomicName authorityName="Horn" authorityYear="1881" class="Insecta" family="Carabidae" genus="Euphorticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Euphorticus</emphasis>
</taxonomicName>
or
<taxonomicName authorityName="Dejean" authorityYear="1831" class="Insecta" family="Carabidae" genus="Lachnophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Lachnophorus</emphasis>
</taxonomicName>
; the genus is in need of a taxonomic revision. In order to do such a revision, one would need to study all the primary types of both
<taxonomicName authorityName="Horn" authorityYear="1881" class="Insecta" family="Carabidae" genus="Euphorticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Euphorticus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Dejean" authorityYear="1831" class="Insecta" family="Carabidae" genus="Lachnophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Lachnophorus</emphasis>
</taxonomicName>
to discover their correct generic assignments.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="references">
<paragraph pageId="20" pageNumber="21">References.</paragraph>
<paragraph pageId="20" pageNumber="21">
<bibRefCitation author="Erwin, TL" journalOrPublisher="Revista Peruana de Entomologia" pageId="79" pageNumber="80" pagination="1 - 85" refId="B22" refString="Erwin, TL, 1991. Natural history of the carabid beetles at the BIOLAT Rio Manu Biological Station, Pakitza, Peru. Revista Peruana de Entomologia 33: 1 - 85" title="Natural history of the carabid beetles at the BIOLAT Rio Manu Biological Station, Pakitza, Peru." volume="33" year="1991">Erwin (1991)</bibRefCitation>
as
<emphasis italics="true" pageId="20" pageNumber="21">incertae sedis</emphasis>
;
<bibRefCitation author="Erwin, TL" journalOrPublisher="Zootaxa" pageId="79" pageNumber="80" pagination="1 - 18" refId="B24" refString="Erwin, TL, 2004. The Beetle Family Carabidae of Costa Rica and Panama: Descriptions of four new genera and six new species with notes on their way of life (Insecta: Coleoptera). Zootaxa 537: 1 - 18" title="The Beetle Family Carabidae of Costa Rica and Panama: Descriptions of four new genera and six new species with notes on their way of life (Insecta: Coleoptera)." volume="537" year="2004">Erwin (2004)</bibRefCitation>
;
<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="78" pageNumber="79" pagination="1 - 1722" publicationUrl="10.3897/zookeys.245.3416" refId="B11" refString="Bousquet, Y, 2012. Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico (in three parts). ZooKeys 245: 1 - 1722, DOI: 10.3897/zookeys.245.3416" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico (in three parts)." url="10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet (2012)</bibRefCitation>
;
<bibRefCitation author="Erwin, TL" journalOrPublisher="Entomologische Blaetter fuer Biologie und Systematik der Kaefer (2010) 106" pageId="79" pageNumber="80" refId="B33" refString="Erwin, TL, Micheli, CJ, Hevel, GF, 2012. The Beetle Family Carabidae of Guyane (French Guyana): The Tribes, Genera, and number of species known, with notes on their ways of life. (Insecta: Coleoptera: Adephaga). Le Coleopteriste 5: 1-88. [ACOREP, Paris]" title="The Beetle Family Carabidae of Guyane (French Guyana): The Tribes, Genera, and number of species known, with notes on their ways of life. (Insecta: Coleoptera: Adephaga). Le Coleopteriste 5: 1 - 88. [ACOREP, Paris]" year="2012">Erwin et al. (2012)</bibRefCitation>
.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/30637" pageId="20" pageNumber="21" start="Figure 1922" startId="F7">
<paragraph pageId="20" pageNumber="21">
<emphasis bold="true" pageId="20" pageNumber="21">Figure 19-22.</emphasis>
<emphasis bold="true" pageId="20" pageNumber="21">19</emphasis>
<taxonomicName authorityName="Erwin" authorityYear="2004" class="Insecta" family="Carabidae" genus="Pseudophorticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Pseudophorticus</emphasis>
</taxonomicName>
sp. (all species in South America are either undescribed or placed in
<taxonomicName authorityName="Horn" authorityYear="1881" class="Insecta" family="Carabidae" genus="Euphorticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Euphorticus</emphasis>
</taxonomicName>
G. Horn). Digital Photo-illustration. Habitus, dorsal aspect, based on specimen ADP132538 from Pakitza,
<normalizedToken originalValue="Perú">Peru</normalizedToken>
<emphasis bold="true" pageId="20" pageNumber="21">20</emphasis>
<taxonomicName authorityName="Motschulsky" authorityYear="1858" class="Insecta" family="Carabidae" genus="Selina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis italics="true" pageId="20" pageNumber="21">Selina westermanni</emphasis>
</taxonomicName>
Motschulsky. Digital Photo-illustration. Habitus, dorsal aspect, based on specimen ADP132536 from China Bay, Sri Lanka
<emphasis bold="true" pageId="20" pageNumber="21">21</emphasis>
<taxonomicName authorityName="Laporte" authorityYear="1832" class="Insecta" family="Carabidae" genus="Stenocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="lacordairei">
<emphasis italics="true" pageId="20" pageNumber="21">Stenocheila lacordairei</emphasis>
</taxonomicName>
Laporte de Castelnau. Digital Photo-illustration. Habitus, dorsal aspect, based on specimen ADP133801 from Chapada dos
<normalizedToken originalValue="Guimãres">Guimares</normalizedToken>
, Brazil
<emphasis bold="true" pageId="20" pageNumber="21">22</emphasis>
<taxonomicName authorityName="Erwin" authorityYear="2000" class="Insecta" family="Carabidae" genus="Quammenis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis italics="true" pageId="20" pageNumber="21">Quammenis spectabilis</emphasis>
</taxonomicName>
Erwin. Digital Photo-illustration. Habitus, dorsal aspect, based on specimen ADP100513 from
<normalizedToken originalValue="Estacíon">Estacion</normalizedToken>
Zurqui, Costa Rica.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/30638" pageId="20" pageNumber="21" start="Figures 2328" startId="F8">
<paragraph pageId="20" pageNumber="21">
<emphasis bold="true" pageId="20" pageNumber="21">Figures 23-28.</emphasis>
Mouthparts of adults.
<emphasis bold="true" pageId="20" pageNumber="21">23</emphasis>
<taxonomicName authorityName="LeConte" authorityYear="1853" class="Insecta" family="Carabidae" genus="Eucaerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Eucaerus</emphasis>
</taxonomicName>
sp. Mouthparts, ventral aspect, based on specimen ADP132552 from Pakitza,
<normalizedToken originalValue="Perú">Peru</normalizedToken>
<emphasis bold="true" pageId="20" pageNumber="21">24</emphasis>
<taxonomicName authorityName="Horn" authorityYear="1881" class="Insecta" family="Carabidae" genus="Euphorticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Euphorticus</emphasis>
</taxonomicName>
sp. Mouthparts, ventral aspect, based on specimen ADP132558 from Tapilula,
<normalizedToken originalValue="México">Mexico</normalizedToken>
<emphasis bold="true" pageId="20" pageNumber="21">25</emphasis>
<taxonomicName authorityName="Erwin" authorityYear="2004" class="Insecta" family="Carabidae" genus="Guatemalteca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="virgen">
<emphasis italics="true" pageId="20" pageNumber="21">Guatemalteca virgen</emphasis>
</taxonomicName>
Erwin. Mouthparts, ventral aspect, based on specimen ADP132546 from nr. La Virgen, Costa Rica
<emphasis bold="true" pageId="20" pageNumber="21">26</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1872" class="Insecta" family="Carabidae" genus="Lachnaces" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Lachnaces</emphasis>
</taxonomicName>
sp. Mouthparts, ventral aspect, based on specimen ADP132578 from Tambopata Reserved Zone,
<normalizedToken originalValue="Explorers">Explorer's</normalizedToken>
Inn,
<normalizedToken originalValue="Perú">Peru</normalizedToken>
<emphasis bold="true" pageId="20" pageNumber="21">27</emphasis>
<taxonomicName authorityName="Dejean" authorityYear="1831" class="Insecta" family="Carabidae" genus="Lachnophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Lachnophorus</emphasis>
</taxonomicName>
sp. Mouthparts, ventral aspect, based on specimen ADP132570 from Pakitza,
<normalizedToken originalValue="Perú">Peru</normalizedToken>
<emphasis bold="true" pageId="20" pageNumber="21">28</emphasis>
<taxonomicName authorityName="Erwin" authorityYear="2004" class="Insecta" family="Carabidae" genus="Pseudophorticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Pseudophorticus</emphasis>
</taxonomicName>
sp. Mouthparts, ventral aspect, based on specimen ADP132538 from Pakitza,
<normalizedToken originalValue="Perú">Peru</normalizedToken>
. Legend:
<emphasis bold="true" pageId="20" pageNumber="21">a3</emphasis>
Antennomere 3;
<emphasis bold="true" pageId="20" pageNumber="21">mt</emphasis>
Mentum tooth;
<emphasis bold="true" pageId="20" pageNumber="21">m</emphasis>
Mentum;
<emphasis bold="true" pageId="20" pageNumber="21">lp3</emphasis>
Labial palpomere 3;
<emphasis bold="true" pageId="20" pageNumber="21">mp4</emphasis>
Maxillary palpomere 4.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>