204 lines
33 KiB
XML
204 lines
33 KiB
XML
<document id="454D1CCB36DA80129AA4DD6DA9E3656C" ENCODING="UTF-8" ID-GBIF-Dataset="de6121e1-d370-43ff-8a55-54eff752a6f8" ModsDocID="z00462p001" checkinTime="1247045511124" checkinUser="thomas" docAuthor="James L. Van Tassell & Carole C. Baldwin" docDate="2004" docId="94AA5D55B870EEE0ACB050C3E9144360" docLanguage="en" docName="2004_VanTassell_Baldwin_gg1.xml" docOrigin="Zootaxa 462" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:73EC2E7B-E100-4C34-9485-3BBC53531905" docTitle="Akko Birdsong and Robins" docType="treatment" docVersion="13" lastPageNumber="14" masterDocId="802F49E080BEEDBE85A511D3CC247C7C" masterDocTitle="A review of the gobiid genus Akko (Teleostei: Gobiidae) with description of a new species." masterLastPageNumber="15" masterPageNumber="1" pageNumber="10" updateTime="1701313966396" updateUser="plazi">
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<mods:title id="433EC7145990EE6EC4BBAE144BCD99DB">A review of the gobiid genus Akko (Teleostei: Gobiidae) with description of a new species.</mods:title>
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<mods:namePart id="276155BF39C309BFE74CBED597765873">James L. Van Tassell</mods:namePart>
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<mods:namePart id="106ABB2E0588E552A786759F97C0DEA1">Carole C. Baldwin</mods:namePart>
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<mods:title id="17A5D4065F17347F25132DAA0DB467E0">Zootaxa</mods:title>
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<mods:part id="D1371EA8E817A049419EC951310BA5B2">
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<mods:date id="BE298760440486C28D1DE0799CB65D19">2004</mods:date>
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<treatment id="94AA5D55B870EEE0ACB050C3E9144360" ID-DOI="http://doi.org/10.5281/zenodo.6270528" ID-GBIF-Taxon="100121585" ID-Zenodo-Dep="6270528" LSID="urn:lsid:plazi:treatment:94AA5D55B870EEE0ACB050C3E9144360" httpUri="http://treatment.plazi.org/id/94AA5D55B870EEE0ACB050C3E9144360" lastPageNumber="14" pageNumber="10">
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<subSubSection id="BCA01CA630B94BCBDDC950F4E1716359" type="nomenclature">
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<paragraph id="36E5448BD70395878397E9004025E754">
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[[
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<taxonomicName id="C68C2AD3CF953018A2D35CF9E6A9EAC5" ID-CoL="RB2" ID-ENA="203341" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong & Robins 1995:676" order="Perciformes" pageNumber="10" phylum="Chordata" rank="genus">Akko Birdsong and Robins</taxonomicName>
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]]
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</paragraph>
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</subSubSection>
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<subSubSection id="34FA848E2B28E2CFE90C3817A2842200" type="discussion">
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<paragraph id="B6722BBCDCB80EEF5F5A7253F16A3DF3" pageNumber="10">Discussion</paragraph>
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<paragraph id="CD28F122122010B1680F82B63C93E2B6" pageNumber="10">
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All species of
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<taxonomicName id="53D7CB41287A2BA9D070CF3EE45FB20B" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong & Robins 1995:676" order="Perciformes" pageNumber="10" phylum="Chordata" rank="genus">Akko</taxonomicName>
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are easily recognized by their large teeth, elongate body, small eyes, anterior nostril located within a fleshy tube, and by having the dorsal, caudal, and anal fins connected by membrane. The species can be differentiated by numbers of lateral scales, vertebrae, and fin rays, and by color. In
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<taxonomicName id="98D9F372B543A398FF702B6DB53A2ABA" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong & Robins 1995:676" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="dionaea">A. dionaea</taxonomicName>
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the lateral scales do not overlap on the caudal peduncle, and there are approximately 76 in the series;
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<taxonomicName id="B2C6C2285D3CE29C8A7056799EC45DD7" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
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and
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<taxonomicName id="77935B537219CDC3B903AA28C63EBAE5" LSID-ZBK="urn:lsid:zoobank.org:act:95C59280-1F8E-4DA1-AE07-37ADDE5877AA" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko rossi van Tassell & Baldwin 2004:3" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="rossi">A. rossi</taxonomicName>
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both have overlapping scales on the caudal peduncle, with 53-60 and 155 scales, respectively, in the lateral series. Color patterns are distinctive, even in preserved specimens.
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<taxonomicName id="BF3E7BE58FF1A906C0B16B444D3A73E5" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong & Robins 1995:676" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="dionaea">Akko dionaea</taxonomicName>
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has no trunk or head melanophores, and only the distal two-thirds of the caudal fin and posterior portion of the anal fin have some melanophores;
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<taxonomicName id="DAEEF983B36A2BA4C1CC6CBB658F3A44" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
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possesses melanophores on the upper portion of the trunk, on the interradial membranes of the dorsal fin, and on the caudal fin;
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<taxonomicName id="69A8C55481B7A5E7CE35F6638B9455AC" LSID-ZBK="urn:lsid:zoobank.org:act:95C59280-1F8E-4DA1-AE07-37ADDE5877AA" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko rossi van Tassell & Baldwin 2004:3" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="rossi">A. rossi</taxonomicName>
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has a heavily pigmented dorsal trunk, and all fins are dark brown. Vertebral counts in
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<taxonomicName id="F38082F0BB4D59458337205EFF9A55BC" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong & Robins 1995:676" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="dionaea">A. dionaea</taxonomicName>
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and
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<taxonomicName id="F4480480126663ECFF5C1423D7ADDFA8" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
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are 11 precaudal and 16 caudal, 11+17 in
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<taxonomicName id="3ED43CF5DE40E0AB3D48E42F429C58AD" LSID-ZBK="urn:lsid:zoobank.org:act:95C59280-1F8E-4DA1-AE07-37ADDE5877AA" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko rossi van Tassell & Baldwin 2004:3" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="rossi">A. rossi</taxonomicName>
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; additional specimens of
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<taxonomicName id="E1AD796664231C481C9AEB56D53B6E82" LSID-ZBK="urn:lsid:zoobank.org:act:95C59280-1F8E-4DA1-AE07-37ADDE5877AA" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko rossi van Tassell & Baldwin 2004:3" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="rossi">A. rossi</taxonomicName>
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are needed to determine if this vertebral count is characteristic for the species.
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<taxonomicName id="F65BF29CAF320132A1CDE4B7FAF6BE92" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong & Robins 1995:676" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="dionaea">Akko dionaea</taxonomicName>
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can be separated from the Pacific species by having fewer pectoral-fin rays: 17 in
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<taxonomicName id="042407B32FF11A9C36222D5AE29D1D2F" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong & Robins 1995:676" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="dionaea">A. dionaea</taxonomicName>
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, 19-21 in
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<taxonomicName id="DF5CB2E69C9ECFF8F08DE06B807B3DA1" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
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, and 19 in
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<taxonomicName id="2E2B0986CB43B9D8309FBF1B7EAFBC14" LSID-ZBK="urn:lsid:zoobank.org:act:95C59280-1F8E-4DA1-AE07-37ADDE5877AA" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko rossi van Tassell & Baldwin 2004:3" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="rossi">A. rossi</taxonomicName>
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. The second dorsal and anal fins both have 15 elements in
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<taxonomicName id="FBE7D8067E9631371480152A34C6CA74" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong & Robins 1995:676" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="dionaea">A. dionaea</taxonomicName>
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and
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<taxonomicName id="03E2AB385E907A1B92E20B10FF9D0125" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
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but 16 in
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<taxonomicName id="9CC8D7A795D72AAA41FD16C7928714A3" LSID-ZBK="urn:lsid:zoobank.org:act:95C59280-1F8E-4DA1-AE07-37ADDE5877AA" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko rossi van Tassell & Baldwin 2004:3" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="rossi">A. rossi</taxonomicName>
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. The pelvic-fin rays are branched in the three species, with
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<taxonomicName id="6D7D5018CCE1AF9514927C431AB37108" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong & Robins 1995:676" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="dionaea">A. dionaea</taxonomicName>
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and
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<taxonomicName id="547A5C2C7BDF0EF195EB0E834DDAFC6D" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
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having all rays dichotomously branched. In
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<taxonomicName id="2AB7802DCC694A6C6DE5A24FBD31813A" LSID-ZBK="urn:lsid:zoobank.org:act:95C59280-1F8E-4DA1-AE07-37ADDE5877AA" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko rossi van Tassell & Baldwin 2004:3" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="rossi">A. rossi</taxonomicName>
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, rays 1,2,3 are serially branched, and rays 4,5 are dichotomously branched. Differences that readily separate the three species are given in Table 3.
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</paragraph>
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<paragraph id="4B1326F483F601F035F8E39405988153" pageNumber="11">
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The head pores in all species of
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<taxonomicName id="0742DEFDB010379CE84A7E65B61B732C" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong & Robins 1995:676" order="Perciformes" pageNumber="11" phylum="Chordata" rank="genus">Akko</taxonomicName>
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are very small and difficult to observe. All species of
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<taxonomicName id="733E3B9A0D4C563271C3CCA405A6A9C0" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong & Robins 1995:676" order="Perciformes" pageNumber="11" phylum="Chordata" rank="genus">Akko</taxonomicName>
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have a short cephalic head canal with two pores, B’ and G’. Pore G’ was reported as absent by Birdsong and Robins (1995) in
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<taxonomicName id="303CAB4A620BDAD93022794AB10D686B" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong & Robins 1995:676" order="Perciformes" pageNumber="11" phylum="Chordata" rank="species" species="dionaea">A. dionaea</taxonomicName>
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, and, although it is absent in the holotype, we have observed it in several of the paratypes. The three species also possess small fleshy flaps on the posterior dorsal edges of the upper and lower lips, a character not mentioned in the original description of
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<taxonomicName id="2028D43845FE434C6518CA569A4AE3A9" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong & Robins 1995:676" order="Perciformes" pageNumber="11" phylum="Chordata" rank="species" species="dionaea">A. dionaea</taxonomicName>
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. Birdsong and Robins (1995) stated that row ot in
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<taxonomicName id="2A9E47D70F6B216FE76C825E020C2FBB" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong & Robins 1995:676" order="Perciformes" pageNumber="11" phylum="Chordata" rank="species" species="dionaea">A. dionaea</taxonomicName>
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has a disjunct extension on the branchiostegals. Our examination of the types of
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<taxonomicName id="7C739EAADF34753E9B1A8AC6747C5E2C" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong & Robins 1995:676" order="Perciformes" pageNumber="11" phylum="Chordata" rank="species" species="dionaea">A. dionaea</taxonomicName>
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with 0.5mm fiber optic lighting shows this row to be complete, not disjunct, and similar to our Figure 2 of
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<taxonomicName id="EAB04D2804AEAE2A797CD799C2574B3E" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="11" phylum="Chordata" rank="species" species="brevis">A. brevis</taxonomicName>
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. A large ossified lacrimal is present in all species of
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<taxonomicName id="6228C2CD322AF4D43277360A9F38DC38" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong & Robins 1995:676" order="Perciformes" pageNumber="11" phylum="Chordata" rank="genus">Akko</taxonomicName>
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. In all other Gobiosomatini and most members of the Gobiinae the lacrimal is small.
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<taxonomicName id="8C7AA4453B549E31AFA9F691BE3AAF97" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="11" phylum="Chordata" rank="species" species="brevis">Akko brevis</taxonomicName>
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has the ability to open its mouth in a complete circle equal to the body diameter. The large lacrimal, which is attached to the maxilla by a broad tendinous sheath, may play a role in this movement.
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</paragraph>
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<paragraph id="13AA31C55647E90A029DA3B785CF8029" pageNumber="12">
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<taxonomicName id="2D352A7D187066EC8702367AB3E2BC06" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong & Robins 1995:676" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Akko</taxonomicName>
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belongs within the tribe Gobiosomatini based on the presence of seven spines in the first dorsal fin, a dorsal-pterygiophore formula of 3-221110, and a vertebral count of 11 precaudal and 16 caudal vertebrae (17 in
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<taxonomicName id="55A7BE3AA82DB16E41E879618B3C106E" LSID-ZBK="urn:lsid:zoobank.org:act:95C59280-1F8E-4DA1-AE07-37ADDE5877AA" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko rossi van Tassell & Baldwin 2004:3" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="rossi">rossi</taxonomicName>
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). Although Birdsong and Robins (1995) placed
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<taxonomicName id="65C05E25DC384CE6C6FE8B28632CEB8F" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong & Robins 1995:676" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Akko</taxonomicName>
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within the tribe, they did not comment further on its relationships within the group because of the large number of autapomorphies possessed by
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<taxonomicName id="6A0B7DA141CCD97E2CD6B21DB272352A" LSID-ZBK="urn:lsid:zoobank.org:act:40FD9887-81A7-4035-B4B2-30CB36945D18" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko dionaea Birdsong & Robins 1995:676" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="dionaea">A. dionaea</taxonomicName>
|
||
. Additional information now available from mtDNA places
|
||
<taxonomicName id="E60F945C4299F3ADD92FF112EE75BF1C" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong & Robins 1995:676" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Akko</taxonomicName>
|
||
within the “
|
||
<taxonomicName id="6BF0F276E37365122E3B0F03D7B22ED3" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
|
||
” group of the Gobiosomatini (Birdsong et al. 1988) and provides support for a sister-group relationship with
|
||
<taxonomicName id="B1A137D0D4CA86B661F402530C75609C" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
|
||
(Ruber et al. 2003).
|
||
</paragraph>
|
||
<paragraph id="8C927F84CDE3F5B4086DF0B2FAD985DB" pageNumber="12">
|
||
Further supporting the placement of
|
||
<taxonomicName id="7CEB907A6BDAFB3E1A2A34AC8B057F27" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong & Robins 1995:676" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Akko</taxonomicName>
|
||
within the “
|
||
<taxonomicName id="B140A4240D60490C7F7D19D01AB53349" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
|
||
” group are patterns of sensory papillae. There are two general categories of sensory papillae patterns within the Gobiosomatini: In the “
|
||
<taxonomicName id="ABD294626446EB2C69E14087A2FBE420" family="Gobiidae" genus="Gobiosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Gobiosoma</taxonomicName>
|
||
” group, row n on the dorsal surface of the head is short, the right and left elements never joining at the dorsal midline, and rows x1 and x2 are not connected to form a continuous row (Fig. 5A); in the “
|
||
<taxonomicName id="F24F8181B5C490390463DA8A2156C264" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
|
||
” group row n is generally elongate, the left and right elements frequently joining at the dorsal midline, and rows x1 and x2 are united to form a single row (Fig. 5B). This arrangement of sensory papillae has been observed in most species of
|
||
<taxonomicName id="71174166028A88F2440830F99CF74156" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
|
||
,
|
||
<taxonomicName id="7B0928D88F902039E136251B6160EB4D" LSID-ZBK="urn:lsid:zoobank.org:act:61A54B05-526D-40E6-8B9B-A351D2F3BB99" family="Gobiidae" genus="Bollmannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Bollmannia Jordan 1890:164" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Bollmannia</taxonomicName>
|
||
, and
|
||
<taxonomicName id="8ECCFFF388B12C72C3279CC14735B827" LSID-ZBK="urn:lsid:zoobank.org:act:D0C705A1-F891-4FFF-A5D6-431BE1972A46" family="Gobiidae" genus="Parrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Parrella Ginsburg 1938:116" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Parrella</taxonomicName>
|
||
of the “
|
||
<taxonomicName id="4AA432BAB600FB4A66590B4B245E26A6" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
|
||
” group by one of us (JVT). All species of
|
||
<taxonomicName id="37C68ECF7CEAF8DE07098850BC6299A3" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong & Robins 1995:676" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Akko</taxonomicName>
|
||
possess a papillae pattern similar to that of the “
|
||
<taxonomicName id="56301F51A618B34A98AD24F0CA10BA33" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="12" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
|
||
” group.
|
||
</paragraph>
|
||
<paragraph id="920CF7D8A1171B98D37470C4FB69DFA9" lastPageNumber="14" pageNumber="13">
|
||
Myological features, particularly origin and insertion patterns within the adductor mandibulae complex, also support the proposed relationship between
|
||
<taxonomicName id="8B8BFF9E3EBE2519E0C85F73F8C4BE7C" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong & Robins 1995:676" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Akko</taxonomicName>
|
||
and other species of the “
|
||
<taxonomicName id="5A4E0DC41B6124196FD8FFEFD04ADF12" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
|
||
” group. In the “
|
||
<taxonomicName id="3551E7BE31C48CFF3B21B6505EA29D6E" family="Gobiidae" genus="Gobiosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Gobiosoma</taxonomicName>
|
||
” group (Fig. 6A) the adductor mandibulae1 complex (A1) originates along the dorsal half of the preopercle and along the lateral edges of the pterotic and sphenotic. The A1 complex separates into A1 beta and A1 alpha sections. An additional separation of the A1, the A1 gamma, occurs in all genera of the “
|
||
<taxonomicName id="2853F2E64FCCF4CE4802E9588DEBB2C8" family="Gobiidae" genus="Gobiosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Gobiosoma</taxonomicName>
|
||
” group except
|
||
<taxonomicName id="3FCFAA2C660074096948549883FE89FD" LSID-ZBK="urn:lsid:zoobank.org:act:FE19BC2A-27FA-4504-8BC8-E76CB54AFFF8" family="Gobiidae" genus="Aruma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Aruma Gobiosoma Ginsburg 1933:16" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Aruma</taxonomicName>
|
||
,
|
||
<taxonomicName id="D46C28EFEAC2CCFECDCE703B18E8DFF1" LSID-ZBK="urn:lsid:zoobank.org:act:E932D49D-30E2-4BDA-BACA-DDD642A785E5" family="Gobiidae" genus="Barbulifer" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Barbulifer Eigenmann & Eigenmann 1888:70" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Barbulifer</taxonomicName>
|
||
,
|
||
<taxonomicName id="DEF85BFECA59F66EA4129F732EB070C8" LSID-ZBK="urn:lsid:zoobank.org:act:C44B38E8-1E20-4D43-AE80-6D8ECCCA343E" family="Gobiidae" genus="Gymneleotris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gymneleotris Bleeker 1874:304" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Gymneleotris</taxonomicName>
|
||
,
|
||
<taxonomicName id="7DE7A9B09CAEEBA71BD8168CC364B30E" LSID-ZBK="urn:lsid:zoobank.org:act:A4D1874E-65F1-4C53-B61A-FCF53B467632" family="Gobiidae" genus="Chriolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chriolepis Gilbert 1892:557" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Chriolepis</taxonomicName>
|
||
,
|
||
<taxonomicName id="1552222F19DEE2F04ECEA9A3D660E1B9" LSID-ZBK="urn:lsid:zoobank.org:act:F997D77F-C2D8-4D88-AF48-4E998B7F1179" family="Gobiidae" genus="Eleotrica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Eleotrica Ginsburg 1933:10" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Eleotrica</taxonomicName>
|
||
,
|
||
<taxonomicName id="B27F88DC06864B67D930F4FC79E0D61A" LSID-ZBK="urn:lsid:zoobank.org:act:C47CF494-51C4-43BC-A5FA-83A9745D26EF" family="Gobiidae" genus="Pycnomma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pycnomma Rutter 1904:252" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Pycnomma</taxonomicName>
|
||
,
|
||
<taxonomicName id="9BEAC7DB10D4333D2B779006A9D972A7" LSID-ZBK="urn:lsid:zoobank.org:act:F0B4BCE2-2987-4E8E-97F0-B55F7723F974" family="Gobiidae" genus="Gobulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gobulus Ginsburg 1933:12" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Gobulus</taxonomicName>
|
||
,
|
||
<taxonomicName id="36D29EC75762F758DE093BF6C10A9FB1" LSID-ZBK="urn:lsid:zoobank.org:act:F80CA779-0C02-4749-A133-712189880C55" family="Gobiidae" genus="Nes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Nes Gobiosoma Ginsburg 1933:25" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Nes</taxonomicName>
|
||
(Van Tassell, 1998). The A1 gamma inserts on the maxilla and on the A2 gamma, the A1 beta inserts on the maxilla, and the A1 alpha inserts on the coronoid process of the dentary. The adductor mandibulae 2 (A2) originates on the ventral half of the preopercle and on the quadrate. The A2 separates into two sections, one inserting along with the A1 alpha on the coronoid process and the other inserting on the maxilla and A1 gamma, near or at the insertion of the primordial ligament.
|
||
</paragraph>
|
||
<paragraph id="AB489D8392CB5D548FE83F15A55E1AFA" pageNumber="14">
|
||
In the “
|
||
<taxonomicName id="EFCCBC301B0D78F3196F5DA4048C9B2F" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="14" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
|
||
” group the A1 complex does not originate along the sphenotic, and the A1 gamma is absent. In
|
||
<taxonomicName id="255E214F41ABFDE41C09216D39F9084A" LSID-ZBK="urn:lsid:zoobank.org:act:61A54B05-526D-40E6-8B9B-A351D2F3BB99" family="Gobiidae" genus="Bollmannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Bollmannia Jordan 1890:164" order="Perciformes" pageNumber="14" phylum="Chordata" rank="genus">Bollmannia</taxonomicName>
|
||
(Fig. 6B) the A1 originates along the dorsal half of the preopercle and on the pterotic; it inserts via two heads, one on the maxilla, in conjunction with the A1 beta, and the second via a tendon on the coronoid process of the dentary. An A1 beta is present medial to the large A1 alpha, originating on the hyomandibular and inserting with the A1 alpha on the maxilla. In
|
||
<taxonomicName id="6D21E6FBB995DEF737F460F1B6E3C9D2" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="14" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
|
||
there is a partial separation of an A1 beta from A1 alpha anteriorly, just prior to the insertion on the maxilla; otherwise, the A1 remains a single muscle mass. The adductor mandibulae 2 is a single mass originating along the ventral half of the preopercle and on the quadrate. It inserts, along with the A1 alpha, via a tendon onto the dentary. The cheek myology pattern of
|
||
<taxonomicName id="A07BF05005CF5BB07104374605CE6C04" LSID-ZBK="urn:lsid:zoobank.org:act:1654F3F7-4BF7-44EE-BD21-80A538552441" family="Gobiidae" genus="Akko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Akko Birdsong & Robins 1995:676" order="Perciformes" pageNumber="14" phylum="Chordata" rank="genus">Akko</taxonomicName>
|
||
is most similar to that of the “
|
||
<taxonomicName id="94BA1FF527E1DD438B68DD8AC2301CDE" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="14" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
|
||
” group (Fig. 4). Cheek myology of
|
||
<taxonomicName id="D33E80C2070C43EF697817021D6DC20B" LSID-ZBK="urn:lsid:zoobank.org:act:D0C705A1-F891-4FFF-A5D6-431BE1972A46" family="Gobiidae" genus="Parrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Parrella Ginsburg 1938:116" order="Perciformes" pageNumber="14" phylum="Chordata" rank="genus">Parrella</taxonomicName>
|
||
and
|
||
<taxonomicName id="EF273198493E85BFB38520DFCA08A8BB" LSID-ZBK="urn:lsid:zoobank.org:act:DAC9635F-D6BA-49D4-832A-BFABA5C2581D" family="Gobiidae" genus="Palatogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Palatogobius Gilbert 1971:30" order="Perciformes" pageNumber="14" phylum="Chordata" rank="genus">Palatogobius</taxonomicName>
|
||
in the “
|
||
<taxonomicName id="E187254E96CCBFB052A956A366241057" LSID-ZBK="urn:lsid:zoobank.org:act:44046A17-7637-4AF9-9121-F993672528FB" family="Gobiidae" genus="Microgobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Microgobius Bleeker 1931:101" order="Perciformes" pageNumber="14" phylum="Chordata" rank="genus">Microgobius</taxonomicName>
|
||
” group has not been examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |