treatments-xml/data/94/72/87/947287DFFFB4FF9B7EAA12DBFC2CFA80.xml
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<document id="F57E41880AE80C6C7B97A7ED513078A2" ID-DOI="10.11646/zootaxa.5026.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5300736" ID-ZooBank="44594360-3408-4C4C-A4E6-1025F171A49F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630039285316" checkinUser="plazi" docAuthor="Michie, Laura A., Barnes, R. S. K., Clark, Paul F., Bennett, Wayne A. &amp; Cragg, Simon M." docDate="2021" docId="947287DFFFB4FF9B7EAA12DBFC2CFA80" docLanguage="en" docName="zootaxa.5026.4.2.pdf" docOrigin="Zootaxa 5026 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tubuca dussumieri" docType="treatment" docVersion="4" lastPageNumber="501" masterDocId="684BFFA7FFA1FF8E7E3D124CFF94FFB1" masterDocTitle="Distinguishing ten sympatric species of fiddler crab (Decapoda: Ocypodidae) using a suite of phenotypic characteristics" masterLastPageNumber="506" masterPageNumber="480" pageNumber="501" updateTime="1699139032236" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="BDCF5C514C8A34A325736BC796F3E82E">Distinguishing ten sympatric species of fiddler crab (Decapoda: Ocypodidae) using a suite of phenotypic characteristics</mods:title>
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<mods:namePart id="689FCE9FC052AE3ACD2A0D05CC82952C">Michie, Laura A.</mods:namePart>
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<mods:namePart id="60A67088167F0E034BB34C8864C13B9B">Barnes, R. S. K.</mods:namePart>
<mods:affiliation id="F8F698894831B0C1274BE1EA99917048">Department of Zoology and Conservation Research Institute, University of Cambridge, CB 2 3 EJ, UK.</mods:affiliation>
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<mods:namePart id="48D0C58299193FE337189862F718F8D7">Clark, Paul F.</mods:namePart>
<mods:affiliation id="78F50E68708F0A8A4E40A8416DE3A95A">Department of Life Sciences, The Natural History Museum, Cromwell Rad, London, SW 7 5 BD, UK.</mods:affiliation>
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<mods:namePart id="3E2C81956C47A07ED3D5ADBACEAF5126">Bennett, Wayne A.</mods:namePart>
<mods:affiliation id="FF648D11799D717D6CA095B120D3A44A">University of West Florida, 11000 University Parkway, Pensacola, US.</mods:affiliation>
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<mods:namePart id="9175CEFADD432E129D051F182B597146">Cragg, Simon M.</mods:namePart>
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<mods:affiliation id="E0428A9DD55902E0668790B2959FF1C3">simon. cragg @ port. ac. uk; https: // orcid. org / 0000 - 0003 - 1082 - 7653</mods:affiliation>
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<mods:date id="B8EABFD465F2E6211C55612FE20A1BAA">2021</mods:date>
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<subSubSection id="54C16542FFB4FF9B7EAA12DBFD5FFF00" box="[151,715,151,178]" pageId="21" pageNumber="501" type="nomenclature">
<paragraph id="1C6436C9FFB4FF9B7EAA12DBFD5FFF00" blockId="21.[151,715,151,214]" box="[151,715,151,178]" pageId="21" pageNumber="501">
<heading id="472C81A5FFB4FF9B7EAA12DBFD5FFF00" bold="true" box="[151,715,151,178]" fontSize="11" level="1" pageId="21" pageNumber="501" reason="1">
<taxonomicName id="DBDB4D4AFFB4FF9B7EAA12DBFD5FFF00" ID-CoL="59JY5" authority="(H. Milne Edwards, 1852)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1852" box="[151,715,151,178]" class="Malacostraca" family="Ocypodidae" genus="Tubuca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="501" phylum="Arthropoda" rank="species" species="dussumieri">
<emphasis id="2EAFEADBFFB4FF9B7EAA12DBFD5FFF00" bold="true" box="[151,715,151,178]" pageId="21" pageNumber="501">
<emphasis id="2EAFEADBFFB4FF9B7EAA12DBFE14FF03" bold="true" box="[151,384,151,178]" italics="true" pageId="21" pageNumber="501">Tubuca dussumieri</emphasis>
(H. Milne Edwards, 1852)
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="54C16542FFB4FF9B7EAA12F7FEACFF67" box="[151,312,187,214]" pageId="21" pageNumber="501" type="description">
<paragraph id="1C6436C9FFB4FF9B7EAA12F7FEACFF67" blockId="21.[151,715,151,214]" box="[151,312,187,214]" pageId="21" pageNumber="501">
(
<figureCitation id="84E02A4CFFB4FF9B7EA212F7FEA4FF67" box="[159,304,187,214]" captionStart="FIGURE 12" captionStartId="20.[151,250,1897,1921]" captionTargetBox="[162,1425,181,1874]" captionTargetId="figure-17@20.[162,1425,181,1874]" captionTargetPageId="20" captionText="FIGURE 12. Tubuca dussumieri (H. Milne Edwards, 1852): ad Sulawesi Tenggara, Indonesia.Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300760" httpUri="https://zenodo.org/record/5300760/files/figure.png" pageId="21" pageNumber="501">Figure 12aj</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFB4FF9B7EAA134FFDECFED4" pageId="21" pageNumber="501" type="materials_examined">
<paragraph id="1C6436C9FFB4FF9B7EAA134FFDECFED4" blockId="21.[151,1437,259,1330]" pageId="21" pageNumber="501">
<emphasis id="2EAFEADBFFB4FF9B7EAA134FFE15FEAC" bold="true" box="[151,385,259,285]" pageId="21" pageNumber="501">Material examined.</emphasis>
<materialsCitation id="ACB33C94FFB4FF9B7FB5134FFDA7FEF0" collectingDate="2014-08-26" collectorName="L. Michie" country="Indonesia" latitude="-5.497222" location="Kaledupa" longLatPrecision="21" longitude="123.755554" pageId="21" pageNumber="501" specimenCount="22" specimenCount-female="13" specimenCount-male="9">
<specimenCount id="0ADDFD40FFB4FF9B7FB5134FFE20FEAF" box="[392,436,259,286]" pageId="21" pageNumber="501" type="male">9 ♂</specimenCount>
,
<specimenCount id="0ADDFD40FFB4FF9B7FFD134FFE6CFEAF" box="[448,504,259,286]" pageId="21" pageNumber="501" type="female">13 ♀</specimenCount>
,
<location id="19046012FFB4FF9B7C3B134FFDE0FEAC" LSID="urn:lsid:plazi:treatment:947287DFFFB4FF9B7EAA12DBFC2CFA80:19046012FFB4FF9B7C3B134FFDE0FEAC" box="[518,628,259,285]" country="Indonesia" latitude="-5.497222" longLatPrecision="21" longitude="123.755554" name="Kaledupa" pageId="21" pageNumber="501">Kaledupa</location>
,
<collectingCountry id="64CC7659FFB4FF9B7C42134FFD7BFEAC" box="[639,751,259,285]" name="Indonesia" pageId="21" pageNumber="501">Indonesia</collectingCountry>
,
<geoCoordinate id="79EF500EFFB4FF9B7CC7134FFC14FEAF" box="[762,896,259,286]" degrees="5" direction="south" minutes="29" orientation="latitude" pageId="21" pageNumber="501" precision="15" seconds="50" value="-5.497222">5º 29 50S</geoCoordinate>
,
<geoCoordinate id="79EF500EFFB4FF9B7DB0134FFBA5FEAC" box="[909,1073,259,286]" degrees="123" direction="east" minutes="45" orientation="longitude" pageId="21" pageNumber="501" precision="15" seconds="20" value="123.755554">123º 45 20E</geoCoordinate>
, mud flat in front of mangroves, under houses,
<collectorName id="B12E531FFFB4FF9B7F0A1364FE33FEF0" box="[311,423,295,322]" pageId="21" pageNumber="501">L. Michie</collectorName>
,
<date id="68651009FFB4FF9B7F8C136BFDA7FEF0" box="[433,563,295,321]" pageId="21" pageNumber="501" value="2014-08-26">
<collectingDate id="7821E9E1FFB4FF9B7F8C136BFDA7FEF0" box="[433,563,295,321]" pageId="21" pageNumber="501" value="2014-08-26">26.08.2014</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="ACB33C94FFB4FF9B7C00136BFDE1FED4" collectingDate="2014-08-28" collectorName="L. Michie" country="Indonesia" latitude="-5.497222" location="Kaledupa" longLatPrecision="21" longitude="123.75584" pageId="21" pageNumber="501" specimenCount="18" specimenCount-male="18">
<specimenCount id="0ADDFD40FFB4FF9B7C00136BFDE1FEF3" box="[573,629,295,322]" pageId="21" pageNumber="501" type="male">11 ♂</specimenCount>
,
<specimenCount id="0ADDFD40FFB4FF9B7CBD1364FD3FFEF3" box="[640,683,296,322]" pageId="21" pageNumber="501" type="male">7 ♂</specimenCount>
,
<location id="19046012FFB4FF9B7C8A136BFCB1FEF0" LSID="urn:lsid:plazi:treatment:947287DFFFB4FF9B7EAA12DBFC2CFA80:19046012FFB4FF9B7C8A136BFCB1FEF0" box="[695,805,295,321]" country="Indonesia" latitude="-5.497222" longLatPrecision="21" longitude="123.75584" name="Kaledupa" pageId="21" pageNumber="501">Kaledupa</location>
,
<collectingCountry id="64CC7659FFB4FF9B7D12136BFC0BFEF0" box="[815,927,295,321]" name="Indonesia" pageId="21" pageNumber="501">Indonesia</collectingCountry>
,
<geoCoordinate id="79EF500EFFB4FF9B7D94136BFBBAFEF3" box="[937,1070,295,322]" degrees="5" direction="south" minutes="29" orientation="latitude" pageId="21" pageNumber="501" precision="15" seconds="50" value="-5.497222">5º 29 50S</geoCoordinate>
,
<geoCoordinate id="79EF500EFFB4FF9B7A04136BFB4FFEF0" box="[1081,1243,295,322]" degrees="123" direction="east" minutes="45" orientation="longitude" pageId="21" pageNumber="501" precision="15" seconds="21" value="123.75584">123º 45 21E</geoCoordinate>
, mud flat in front of mangroves, coll.
<collectorName id="B12E531FFFB4FF9B7F4B1300FE73FED4" box="[374,487,331,358]" pageId="21" pageNumber="501">L. Michie</collectorName>
,
<date id="68651009FFB4FF9B7FCE1307FDE1FED4" box="[499,629,331,357]" pageId="21" pageNumber="501" value="2014-08-28">
<collectingDate id="7821E9E1FFB4FF9B7FCE1307FDE1FED4" box="[499,629,331,357]" pageId="21" pageNumber="501" value="2014-08-28">28.08.2014</collectingDate>
</date>
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.
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFB4FF9B7EFA1323FE59FBA0" pageId="21" pageNumber="501" type="diagnosis">
<paragraph id="1C6436C9FFB4FF9B7EFA1323FB5AFE44" blockId="21.[151,1437,259,1330]" pageId="21" pageNumber="501">
<emphasis id="2EAFEADBFFB4FF9B7EFA1323FED6FE38" bold="true" box="[199,322,367,393]" pageId="21" pageNumber="501">Diagnosis.</emphasis>
Carapace: broader than long, smooth; front narrow and practically straight; anterolateral angles acute, not produced; anterolateral margins short; dorsolateral margins distinct and converging. Major cheliped: elongated dactyl and pollex; pollex with medial tooth; gape between dactyl and pollex is triangular in shape due to distinct angle of dactyl; breadth of dactyl consistent along its entire length and only tapers at tip.
</paragraph>
<paragraph id="1C6436C9FFB4FF9B7EFA13B3FBEDFD18" blockId="21.[151,1437,259,1330]" pageId="21" pageNumber="501">
<emphasis id="2EAFEADBFFB4FF9B7EFA13B3FECCFDA8" bold="true" box="[199,344,511,537]" pageId="21" pageNumber="501">Colouration</emphasis>
(
<figureCitation id="84E02A4CFFB4FF9B7F5813B3FE6FFDAB" box="[357,507,511,538]" captionStart="FIGURE 12" captionStartId="20.[151,250,1897,1921]" captionTargetBox="[162,1425,181,1874]" captionTargetId="figure-17@20.[162,1425,181,1874]" captionTargetPageId="20" captionText="FIGURE 12. Tubuca dussumieri (H. Milne Edwards, 1852): ad Sulawesi Tenggara, Indonesia.Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300760" httpUri="https://zenodo.org/record/5300760/files/figure.png" pageId="21" pageNumber="501">Figure 12ad</figureCitation>
). Adult male. Carapace: black, sometimes with two or three central white markings, anterior ventral surface of carapace and third maxillipeds normally black. Major cheliped: dactyl generally white extending dorsally onto manus, pollex with a white tip and remainder orange which extends ventrally onto manus, usually becoming darker, carpus is a mixture of white, orange and brown, remainder of cheliped usually orange. Minor cheliped &amp; ambulatory legs: generally black, sometimes with last pair blue/white.
</paragraph>
<paragraph id="1C6436C9FFB4FF9B7EFA10FFFE81FD40" blockId="21.[151,1437,259,1330]" pageId="21" pageNumber="501">Adult female. Carapace: similar to male. Ambulatory legs: similar to males, except sometimes all legs are blue/white.</paragraph>
<paragraph id="1C6436C9FFB4FF9B7EFA10B7FD17FC8B" blockId="21.[151,1437,259,1330]" pageId="21" pageNumber="501">Juvenile male. Carapace: most often black with two or three central white markings. Major cheliped: dactyl and pollex normally white, remainder is orange.</paragraph>
<paragraph id="1C6436C9FFB4FF9B7EFA110FFC73FCEC" blockId="21.[151,1437,259,1330]" box="[199,999,835,862]" pageId="21" pageNumber="501">Juvenile female. Similar to adult females. Ambulatory legs: blue/white.</paragraph>
<paragraph id="1C6436C9FFB4FF9B7EFA112BFBE3FC14" blockId="21.[151,1437,259,1330]" pageId="21" pageNumber="501">
<emphasis id="2EAFEADBFFB4FF9B7EFA112BFE5BFC30" bold="true" box="[199,463,871,897]" pageId="21" pageNumber="501">Gonopod morphology</emphasis>
(
<figureCitation id="84E02A4CFFB4FF9B7FDD112BFDEDFC33" box="[480,633,871,898]" captionStart="FIGURE 12" captionStartId="20.[151,250,1897,1921]" captionTargetBox="[162,1425,181,1874]" captionTargetId="figure-17@20.[162,1425,181,1874]" captionTargetPageId="20" captionText="FIGURE 12. Tubuca dussumieri (H. Milne Edwards, 1852): ad Sulawesi Tenggara, Indonesia.Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300760" httpUri="https://zenodo.org/record/5300760/files/figure.png" pageId="21" pageNumber="501">Figure 12eh</figureCitation>
). G1 curved in dorso-lateral direction; large anterior flange, with dark spine at tip of anterior edge; distal protuberance short; pore of sperm channel in midline.
</paragraph>
<paragraph id="1C6436C9FFB4FF9B7EFA11E3FE59FBA0" blockId="21.[151,1437,259,1330]" pageId="21" pageNumber="501">
<emphasis id="2EAFEADBFFB4FF9B7EFA11E3FEC0FC78" bold="true" box="[199,340,943,969]" pageId="21" pageNumber="501">Gastric mill</emphasis>
(
<figureCitation id="84E02A4CFFB4FF9B7F5C11E3FE7EFC7B" box="[353,490,943,970]" captionStart="FIGURE 12" captionStartId="20.[151,250,1897,1921]" captionTargetBox="[162,1425,181,1874]" captionTargetId="figure-17@20.[162,1425,181,1874]" captionTargetPageId="20" captionText="FIGURE 12. Tubuca dussumieri (H. Milne Edwards, 1852): ad Sulawesi Tenggara, Indonesia.Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300760" httpUri="https://zenodo.org/record/5300760/files/figure.png" pageId="21" pageNumber="501">Figure 12i, j</figureCitation>
). Urocardiac ossicles simple, with 5 transverse ridges of median teeth, 14 decreasing in size distally, fifth one larger and attached to plate dorsally; stem region has 1 pair of cusps. Lateral tooth plate with 18 comb-shaped teeth.
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFB4FF9B7EFA1657FB85FBCF" pageId="21" pageNumber="501" type="biology_ecology">
<paragraph id="1C6436C9FFB4FF9B7EFA1657FB85FBCF" blockId="21.[151,1437,259,1330]" pageId="21" pageNumber="501">
<emphasis id="2EAFEADBFFB4FF9B7EFA1657FEB3FB84" bold="true" box="[199,295,1051,1077]" pageId="21" pageNumber="501">Biology.</emphasis>
<taxonomicName id="DBDB4D4AFFB4FF9B7F0C1657FD9CFB84" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1852" box="[305,520,1051,1077]" class="Malacostraca" family="Ocypodidae" genus="Tubuca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="501" phylum="Arthropoda" rank="species" species="dussumieri">
<emphasis id="2EAFEADBFFB4FF9B7F0C1657FD9CFB84" box="[305,520,1051,1077]" italics="true" pageId="21" pageNumber="501">Tubuca dussumieri</emphasis>
</taxonomicName>
is a medium-size species (largest male CL =
<quantity id="DB239B2CFFB4FF9B7A211657FB10FB87" box="[1052,1156,1051,1078]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="21" pageNumber="501" unit="mm" value="14.6">14.6 mm</quantity>
, CB =
<quantity id="DB239B2CFFB4FF9B7AE61657FAD0FB87" box="[1243,1348,1051,1078]" metricMagnitude="-2" metricUnit="m" metricValue="2.94" pageId="21" pageNumber="501" unit="mm" value="29.4">29.4 mm</quantity>
, largest female CL =
<quantity id="DB239B2CFFB4FF9B7F151673FE19FBEB" box="[296,397,1087,1114]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="21" pageNumber="501" unit="mm" value="15.9">15.9 mm</quantity>
, CB =
<quantity id="DB239B2CFFB4FF9B7FE51673FDABFBEB" box="[472,575,1087,1114]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="21" pageNumber="501" unit="mm" value="24.9">24.9 mm</quantity>
); one of the least abundant species on shore, present in mid and high intertidal zones, predominantly associated with shade under buildings or mangrove trees.
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFB4FF9B7EFA16CBFB57FB10" box="[199,1219,1159,1186]" pageId="21" pageNumber="501" type="distribution">
<paragraph id="1C6436C9FFB4FF9B7EFA16CBFB57FB10" blockId="21.[151,1437,259,1330]" box="[199,1219,1159,1186]" pageId="21" pageNumber="501">
<emphasis id="2EAFEADBFFB4FF9B7EFA16CBFEF5FB10" bold="true" box="[199,353,1159,1185]" pageId="21" pageNumber="501">Distribution.</emphasis>
<collectingCountry id="64CC7659FFB4FF9B7F5A16CBFE39FB10" box="[359,429,1159,1185]" name="China" pageId="21" pageNumber="501">China</collectingCountry>
,
<collectingCountry id="64CC7659FFB4FF9B7F8516CBFD99FB10" box="[440,525,1159,1185]" name="Taiwan" pageId="21" pageNumber="501">Taiwan</collectingCountry>
,
<collectingCountry id="64CC7659FFB4FF9B7C2A16CBFDE9FB10" box="[535,637,1159,1185]" name="Thailand" pageId="21" pageNumber="501">Thailand</collectingCountry>
,
<collectingCountry id="64CC7659FFB4FF9B7CB416CBFD6DFB10" box="[649,761,1159,1185]" name="Indonesia" pageId="21" pageNumber="501">Indonesia</collectingCountry>
, Northern
<collectingCountry id="64CC7659FFB4FF9B7D5316CBFC4CFB10" box="[878,984,1159,1185]" name="Australia" pageId="21" pageNumber="501">Australia</collectingCountry>
,
<collectingCountry id="64CC7659FFB4FF9B7DDE16C4FB2BFB10" box="[995,1215,1159,1186]" name="Papua New Guinea" pageId="21" pageNumber="501">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFB4FF9B7EFA16E7FC2CFA80" pageId="21" pageNumber="501" type="discussion">
<paragraph id="1C6436C9FFB4FF9B7EFA16E7FC2CFA80" blockId="21.[151,1437,259,1330]" pageId="21" pageNumber="501">
<emphasis id="2EAFEADBFFB4FF9B7EFA16E7FEAFFB74" bold="true" box="[199,315,1195,1221]" pageId="21" pageNumber="501">Remarks.</emphasis>
<taxonomicName id="DBDB4D4AFFB4FF9B7F7C16E7FD81FB74" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1852" box="[321,533,1195,1221]" class="Malacostraca" family="Ocypodidae" genus="Tubuca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="501" phylum="Arthropoda" rank="species" species="dussumieri">
<emphasis id="2EAFEADBFFB4FF9B7F7C16E7FD81FB74" box="[321,533,1195,1221]" italics="true" pageId="21" pageNumber="501">Tubuca dussumieri</emphasis>
</taxonomicName>
is the largest species at Ambeua which, along with the large anterior flange of the G1, distinguishes it from the other
<taxonomicName id="DBDB4D4AFFB4FF9B7C291683FD50FB58" box="[532,708,1231,1257]" class="Malacostraca" family="Ocypodidae" genus="Tubuca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="501" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2EAFEADBFFB4FF9B7C291683FDF2FB58" box="[532,614,1231,1257]" italics="true" pageId="21" pageNumber="501">Tubuca</emphasis>
species.
</taxonomicName>
The specimens examined agree with the original description. This is however, the first description of the gastric mill for
<taxonomicName id="DBDB4D4AFFB4FF9B7CD616B8FC17FABC" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1852" box="[747,899,1267,1294]" class="Malacostraca" family="Ocypodidae" genus="Tubuca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="501" phylum="Arthropoda" rank="species" species="dussumieri">
<emphasis id="2EAFEADBFFB4FF9B7CD616B8FD6FFABC" box="[747,763,1268,1293]" italics="true" pageId="21" pageNumber="501">T</emphasis>
.
<emphasis id="2EAFEADBFFB4FF9B7D3516BFFC17FABC" box="[776,899,1267,1293]" italics="true" pageId="21" pageNumber="501">dussumieri</emphasis>
</taxonomicName>
and the simple morphology which distinguishes it from other Ambeua genera and is similar to the other
<taxonomicName id="DBDB4D4AFFB4FF9B7D3B175BFC2CFA80" box="[774,952,1303,1329]" class="Malacostraca" family="Ocypodidae" genus="Tubuca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="501" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2EAFEADBFFB4FF9B7D3B175BFCCCFA80" box="[774,856,1303,1329]" italics="true" pageId="21" pageNumber="501">Tubuca</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>