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<document id="929F8ECAB5DFAE4C2926F683BAC6FA79" ID-CLB-Dataset="53488" ID-DOI="10.11646/phytotaxa.585.3.2" ID-GBIF-Dataset="e75aae80-461f-406e-ad01-67d5755a59a9" ID-ISSN="1179-3163" ID-Zenodo-Dep="7690757" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1677742952481" checkinUser="plazi" docAuthor="Boon, Richard G. C. &amp; Van Wyk, Abraham E." docDate="2023" docId="944287ABED12FF97FF11FE69F3B3B21A" docLanguage="en" docName="phytotaxa.585.3.2.pdf.imd" docOrigin="Phytotaxa 585 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.585.3.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Phyllogeiton trachybasis R. G. C. Boon &amp; A. E. van Wyk 2023, sp. nov." docType="treatment" docVersion="10" lastPageNumber="206" masterDocId="687BFFD3ED10FF9AFF99FFDDF618B666" masterDocTitle="A new species of Phyllogeiton (Rhamnaceae: Rhamneae) from Maputaland, South Africa" masterLastPageNumber="209" masterPageNumber="193" pageNumber="195" updateTime="1699475978720" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D266AAFAD7F17E089D32830BD3E5D6C6">A new species of Phyllogeiton (Rhamnaceae: Rhamneae) from Maputaland, South Africa</mods:title>
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<mods:namePart id="68B3A61E8E589DF9D9DBD8ADE68662C5">Boon, Richard G. C.</mods:namePart>
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<mods:affiliation id="3358628702C1D39DDA9A3AFFD18F0D66">Biological Sciences, School of Life Sciences, University of KwaZulu-Natal, Private Bag X 5400, Durban, 4000 South Africa &amp; kzntrees @ gmail. com; https: // orcid. org / 0000 - 0003 - 3109 - 3687</mods:affiliation>
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<mods:namePart id="33F61F4958A810BCD75A508572FA5041">Van Wyk, Abraham E.</mods:namePart>
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<mods:affiliation id="B11AE79E48219128D2360799B25543BD">H. G. W. J. Schweickerdt Herbarium, Department of Plant and Soil Sciences, University of Pretoria, Pretoria, 0002 South Africa National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria, 0001 South Africa &amp; braamvanwyk @ gmail. com; https: // orcid. org / 0000 - 0002 - 0437 - 3272</mods:affiliation>
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<mods:date id="EECD5E6F20DECA7B1164F6E22938C904">2023</mods:date>
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<treatment id="944287ABED12FF97FF11FE69F3B3B21A" ID-DOI="http://doi.org/10.5281/zenodo.7693326" ID-GBIF-Taxon="209385652" ID-Zenodo-Dep="7693326" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A" httpUri="http://treatment.plazi.org/id/944287ABED12FF97FF11FE69F3B3B21A" lastPageId="13" lastPageNumber="206" pageId="2" pageNumber="195">
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<taxonomicName id="DBEB4D3EED12FF98FF11FE69F4EBB7A8" ID-CoL="B8QZ7" authority="R.G.C.Boon &amp; A.E.van Wyk" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[136,755,436,463]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="species" species="trachybasis" status="sp. nov.">
<emphasis id="2E9FEAAFED12FF98FF11FE69F7BCB7A9" bold="true" box="[136,420,436,463]" italics="true" pageId="2" pageNumber="195">Phyllogeiton trachybasis</emphasis>
R.G.C.Boon &amp; A.E.van Wyk
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,
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<taxonomicNameLabel id="35AC57D4ED12FF98FD64FE68F54AB7A8" box="[765,850,437,462]" pageId="2" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="84D02A38ED12FF98FCF8FE69F5DCB7A8" box="[865,964,436,462]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="4.[136,229,1991,2013]" captionStartId-1="5.[136,229,1955,1977]" captionStartId-2="6.[136,229,1953,1975]" captionStartId-3="7.[136,229,1953,1975]" captionTargetBox-0="[151,1435,187,1964]" captionTargetBox-1="[164,1423,190,1931]" captionTargetBox-2="[151,1435,190,1929]" captionTargetBox-3="[136,1450,190,1932]" captionTargetId-0="figure-19@4.[151,1435,187,1964]" captionTargetId-1="figure-18@5.[164,1423,190,1931]" captionTargetId-2="figure-19@6.[151,1435,190,1929]" captionTargetId-3="figure-18@7.[151,1436,190,1928]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Phyllogeiton trachybasis, habit and habitat. Mature tree in Sand Forest at Bonamanzi Game Reserve, Hluhluwe, KwaZuluNatal, South Africa. Photograph: R.G.C. Boon." captionText-1="FIGURE 2. Phyllogeiton trachybasis, bark morphology illustrating variation (A–D) and colour of the wood (E). A. Bark on trunk of old, mature tree. B. Bark on trunk of relatively young tree (sunny habitat). C. Bark on trunk of relatively young tree (shady habitat). D. Exceedingly rough and flaking bark on trunk of mature tree. E. Finely sanded, untreated wood in transverse section; from dead branch ± 130 mm in diameter. Photographs: R.G.C. Boon (A, D), F. du Randt (B, C), &amp; D. Bishop (E)." captionText-2="FIGURE 3. Phyllogeiton trachybasis, leaf, flower and fruit morphology. A. Branchlet with leaves, the latter fresh green, without a bloom and ca. concolorous. B. Flowers, open and in bud. C. Almost ripe fruit. D. Mainly ripe fallen fruits underneath a tree, some with stone removed or otherwise damaged by wildlife. E. Cleaned stones. Photographs: R.G.C. Boon (A, C, D), G. Nichols (B), &amp; E. Douwes (E)." captionText-3="FIGURE 4. Phyllogeiton trachybasis (A–E), with fruit of P. discolor (F) and P. zeyheri (G). A. Fruiting branchlet (fruit almost mature). B. Flower. C. Petal. D. Stamen. E–G. Comparison of the fruit (bottom) of the three species of Phyllogeiton, with corresponding enlarged persistent receptacle and fruit stalk (top). Scale bar = 10 mm (A), 1 mm (B–D), or 4 mm (E–G). Drawings based on photographs and liquid-preserved fresh material deposited in Herb. PRU. Artist: Daleen Roodt." figureDoi-0="http://doi.org/10.5281/zenodo.7690759" figureDoi-1="http://doi.org/10.5281/zenodo.7690761" figureDoi-2="http://doi.org/10.5281/zenodo.7690763" figureDoi-3="http://doi.org/10.5281/zenodo.7702415" httpUri-0="https://zenodo.org/record/7690759/files/figure.png" httpUri-1="https://zenodo.org/record/7690761/files/figure.png" httpUri-2="https://zenodo.org/record/7690763/files/figure.png" httpUri-3="https://zenodo.org/record/7702415/files/figure.png" pageId="2" pageNumber="195">Figs 1–4</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED12FF98FF11FE22F6E0B4C0" pageId="2" pageNumber="195" type="diagnosis">
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<emphasis id="2E9FEAAFED12FF98FF11FE22F713B473" bold="true" box="[136,267,511,533]" pageId="2" pageNumber="195">Diagnosis:—</emphasis>
Morphologically most similar to
<taxonomicName id="DBEB4D3EED12FF98FDDDFDDDF50BB473" box="[580,787,512,533]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED12FF98FDDDFDDDF50BB473" box="[580,787,512,533]" italics="true" pageId="2" pageNumber="195">Phyllogeiton discolor</emphasis>
</taxonomicName>
from which it differs particularly by the following characters: bark on the bole of mature trees almost black, very rough, exfoliating in large pieces; leaves fresh green and without a bloom, ± concolorous, apex acuminate; petals spreading to somewhat ascending at anthesis; fruit oblong-ellipsoid, up to 25 ×
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, exocarp often indistinctly longitudinally ribbed, bright orange-yellow; disc-shaped receptacle at point of fruit stalk’s attachment to base of fruit plane.
</paragraph>
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<emphasis id="2E9FEAAFED12FF98FF11FD69F6C2B4AF" bold="true" box="[136,218,692,713]" pageId="2" pageNumber="195">Type:—</emphasis>
<materialsCitation id="AC833CE0ED12FF98FF43FD69F4A0B48B" ID-GBIF-Occurrence="4041537325" collectingDate="1996-03-05" collectionCode="PRU" collectorName="Abbott" country="South Africa" location="Bonamanzi Game Park" municipality="Hluhluwe" pageId="2" pageNumber="195" specimenCode="081091" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<collectingCountry id="64FC762DED12FF98FF43FD69F791B4AF" box="[218,393,692,713]" name="South Africa" pageId="2" pageNumber="195">SOUTH AFRICA</collectingCountry>
.
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:
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,
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[Reserve], game lodge car park, (2832AA/AB) [2832AB],
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<collectingDate id="7811E995ED12FF98FF24FD05F75EB488" box="[189,326,728,750]" pageId="2" pageNumber="195" value="1996-03-05">5 March 1996</collectingDate>
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,
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<collectorName id="B11E536BED12FF98FEC9FD05F788B48B" box="[336,400,728,749]" pageId="2" pageNumber="195">Abbott</collectorName>
6929
</emphasis>
(
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:
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;
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<materialsCitation id="AC833CE0ED12FF98FD27FD05F564B48B" ID-GBIF-Occurrence="4041537314" box="[702,892,728,750]" collectingDate="1996-03-05" collectionCode="NH, PCE" country="South Africa" location="Bonamanzi Game Park" municipality="Hluhluwe" pageId="2" pageNumber="195" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="isotype">
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,
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).
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</paragraph>
</subSubSection>
<subSubSection id="54F16536ED12FF98FF11FD21F30CB51B" pageId="2" pageNumber="195" type="reference_group">
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“
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<taxonomicName id="DBEB4D3EED12FF98FF0AFD21F6E2B577" ID-CoL="39M4" authority="Boon &amp; Van Wyk, 2023" authorityName="Boon &amp; Van Wyk" authorityYear="2023" box="[147,250,764,785]" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="genus" status="sp. nov.">Berchemia</taxonomicName>
<taxonomicNameLabel id="35AC57D4ED12FF98FE9AFD21F757B577" box="[259,335,764,785]" pageId="2" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
” in Coates
<treatmentCitation id="9D4A10ACED12FF98FE5FFD21F481B574" author="Palgrave, M." box="[454,665,764,786]" page="669" pageId="2" pageNumber="195" year="2002">
<bibRefCitation id="787A4B4CED12FF98FE5FFD21F481B574" author="Palgrave, M." box="[454,665,764,786]" pageId="2" pageNumber="195" refId="ref9035" refString="Palgrave, M. (2002) Keith Coates Palgrave trees of southern Africa, 3 rd ed. Struik, Cape Town, 1212 pp." type="book" year="2002">Palgrave (2002: 669)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="9D4A10ACED12FF98FD3FFD21F541B577" author="Boon, R." box="[678,857,764,785]" page="332" pageId="2" pageNumber="195" year="2010">
<bibRefCitation id="787A4B4CED12FF98FD3FFD21F541B577" author="Boon, R." box="[678,857,764,785]" pageId="2" pageNumber="195" refId="ref8873" refString="Boon, R. [R. G. C.] (2010) Pooley's trees of eastern South Africa. A complete guide. Flora and Fauna Publications Trust, Durban, 626 pp." type="book" year="2010">Boon (2010: 332)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="9D4A10ACED12FF98FCFCFD21F2A5B577" author="Van Wyk, B. &amp; A. E &amp; Van Wyk, P." box="[869,1213,764,786]" page="410" pageId="2" pageNumber="195" year="2013">
<bibRefCitation id="787A4B4CED12FF98FCFCFD21F2A5B577" author="Van Wyk, B. &amp; A. E &amp; Van Wyk, P." box="[869,1213,764,786]" pageId="2" pageNumber="195" refId="ref11626" refString="Van Wyk, B. [A. E.] &amp; Van Wyk, P. (2013) Field guide to trees of southern Africa, 2 nd ed. Struik Nature, Cape Town, 732 pp." type="book" year="2013">Van Wyk &amp; Van Wyk (2013: 410)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="9D4A10ACED12FF98FB53FD21F3B0B574" author="Du Randt, F." box="[1226,1448,764,786]" page="316" pageId="2" pageNumber="195" year="2018">
<bibRefCitation id="787A4B4CED12FF98FB53FD21F3B0B574" author="Du Randt, F." box="[1226,1448,764,786]" pageId="2" pageNumber="195" refId="ref9248" refString="Du Randt, F. (2018) The sand forest of Maputaland. South African National Biodiversity Institute, Pretoria, 416 pp." type="book" year="2018">Du Randt (2018: 316)</bibRefCitation>
</treatmentCitation>
. “
<taxonomicName id="DBEB4D3EED12FF98FF51FCFDF770B553" ID-CoL="LKG8" box="[200,360,800,821]" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="species" species="undefined-a">
<emphasis id="2E9FEAAFED12FF98FF51FCFDF737B553" box="[200,303,800,821]" italics="true" pageId="2" pageNumber="195">Berchemia</emphasis>
sp. A
</taxonomicName>
” in
<treatmentCitation id="9D4A10ACED12FF98FE09FCFDF441B550" author="Schmidt, E." box="[400,601,800,822]" page="546" pageId="2" pageNumber="195" year="2018">
<bibRefCitation id="787A4B4CED12FF98FE09FCFDF441B550" author="Schmidt, E." box="[400,601,800,822]" pageId="2" pageNumber="195" pagination="544 - 552" refId="ref10878" refString="Schmidt, E. (2018) Rhamnaceae. In: Burrows, J. E., Burrows, S. M., Lotter, M. C. &amp; Schmidt, E. (Eds.) Trees and shrubs Mozambique. Publishing Print Matters, Noordhoek, Cape Town, pp. 544 - 552." type="book chapter" year="2018">Schmidt (2018: 546)</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="1C5436BDED12FF98FF11FC99F30CB51B" blockId="2.[136,1452,511,894]" pageId="2" pageNumber="195">
Illustrations:
<bibRefCitation id="787A4B4CED12FF98FE9EFC99F7B3B53F" author="Boon, R." box="[263,427,836,857]" pageId="2" pageNumber="195" refId="ref8873" refString="Boon, R. [R. G. C.] (2010) Pooley's trees of eastern South Africa. A complete guide. Flora and Fauna Publications Trust, Durban, 626 pp." type="book" year="2010">Boon (2010: 333</bibRefCitation>
,
<emphasis id="2E9FEAAFED12FF98FE2CFC99F472B53F" box="[437,618,836,857]" italics="true" pageId="2" pageNumber="195">
<taxonomicName id="DBEB4D3EED12FF98FE2CFC99F404B53F" ID-CoL="39M4" authority="Boon &amp; Van Wyk, 2023" authorityName="Boon &amp; Van Wyk" authorityYear="2023" box="[437,540,836,857]" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="genus" status="sp. nov.">Berchemia</taxonomicName>
<taxonomicNameLabel id="35AC57D4ED12FF98FDB8FC99F472B53F" box="[545,618,836,857]" pageId="2" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
one photograph in second row from top, two in third row from the top);
<bibRefCitation id="787A4B4CED12FF98FA87FC99F73FB518" author="Van Wyk, B. &amp; A. E &amp; Van den Berg, E. &amp; Coates Palgrave, M. &amp; Jordaan, M." pageId="2" pageNumber="195" refId="ref11517" refString="Van Wyk, B. [A. E.], Van den Berg, E., Coates Palgrave, M. &amp; Jordaan, M. (2011) Dictionary of names for southern African trees. Scientific names of indigenous trees, shrubs and climbers with common names from 30 languages. Briza Academic Books, Briza Publications, Pretoria, 956 pp." type="book" year="2011">
Van Wyk
<emphasis id="2E9FEAAFED12FF98FAE5FC99F3B4B53F" box="[1404,1452,836,857]" italics="true" pageId="2" pageNumber="195">et al.</emphasis>
(2011: 125
</bibRefCitation>
,
<emphasis id="2E9FEAAFED12FF98FEA8FCB5F7F0B51B" box="[305,488,872,893]" italics="true" pageId="2" pageNumber="195">
<taxonomicName id="DBEB4D3EED12FF98FEA8FCB5F780B51B" ID-CoL="39M4" authority="Boon &amp; Van Wyk, 2023" authorityName="Boon &amp; Van Wyk" authorityYear="2023" box="[305,408,872,893]" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="genus" status="sp. nov.">Berchemia</taxonomicName>
<taxonomicNameLabel id="35AC57D4ED12FF98FE07FCB5F7F0B51B" box="[414,488,872,893]" pageId="2" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
);
<bibRefCitation id="787A4B4CED12FF98FE62FCB5F4D0B51B" author="Du Randt, F." box="[507,712,872,894]" pageId="2" pageNumber="195" refId="ref9248" refString="Du Randt, F. (2018) The sand forest of Maputaland. South African National Biodiversity Institute, Pretoria, 416 pp." type="book" year="2018">Du Randt (2018: 317</bibRefCitation>
,
<emphasis id="2E9FEAAFED12FF98FD4BFCB5F591B51B" box="[722,905,872,893]" italics="true" pageId="2" pageNumber="195">
<taxonomicName id="DBEB4D3EED12FF98FD4BFCB5F521B51B" ID-CoL="39M4" authority="Boon &amp; Van Wyk, 2023" authorityName="Boon &amp; Van Wyk" authorityYear="2023" box="[722,825,872,893]" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="genus" status="sp. nov.">Berchemia</taxonomicName>
<taxonomicNameLabel id="35AC57D4ED12FF98FCD9FCB5F591B51B" box="[832,905,872,893]" pageId="2" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
);
<bibRefCitation id="787A4B4CED12FF98FC05FCB5F245B518" author="Schmidt, E." box="[924,1117,872,894]" pageId="2" pageNumber="195" pagination="544 - 552" refId="ref10878" refString="Schmidt, E. (2018) Rhamnaceae. In: Burrows, J. E., Burrows, S. M., Lotter, M. C. &amp; Schmidt, E. (Eds.) Trees and shrubs Mozambique. Publishing Print Matters, Noordhoek, Cape Town, pp. 544 - 552." type="book chapter" year="2018">Schmidt (2018: 546</bibRefCitation>
,
<taxonomicName id="DBEB4D3EED12FF98FBFEFCB5F310B51B" ID-CoL="LKG8" box="[1127,1288,872,893]" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="species" species="undefined-a">
<emphasis id="2E9FEAAFED12FF98FBFEFCB5F2D6B51B" box="[1127,1230,872,893]" italics="true" pageId="2" pageNumber="195">Berchemia</emphasis>
sp. A
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED12FF99FF11FC71F77FB7F6" lastPageId="3" lastPageNumber="196" pageId="2" pageNumber="195" type="description">
<paragraph id="1C5436BDED12FF98FF11FC71F2FEB141" blockId="2.[136,1452,940,2083]" pageId="2" pageNumber="195">
Semi-deciduous, erect tree up to 12(–15) m tall.
<emphasis id="2E9FEAAFED12FF98FD36FC71F4E8B5A0" box="[687,752,940,966]" italics="true" pageId="2" pageNumber="195">Trunk</emphasis>
single-stemmed and forking below
<quantity id="DB139B58ED12FF98FB10FC71F2D2B5A1" box="[1161,1226,940,967]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="195" unit="m" value="2.5">2.5 m</quantity>
or double-stemmed from near ground level, individual stems up to
<quantity id="DB139B58ED12FF98FD3EFC0DF4EFB58D" box="[679,759,976,1003]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="195" unit="m" value="0.55">0.55 m</quantity>
diam. at breast height (dbh).
<emphasis id="2E9FEAAFED12FF98FBD5FC0DF29AB58C" box="[1100,1154,976,1002]" italics="true" pageId="2" pageNumber="195">Bark</emphasis>
very rough at the base of old trees, flaking in large pieces, flakes very often persistent, dark, almost black, sometimes with pale grey patches of crustose lichen, bark on upper bole and large branches thick, grey, tessellate, revealing dark bark below where peeling, fissured but fairly smooth on young branches.
<emphasis id="2E9FEAAFED12FF98FD0EFBE1F50DB230" box="[663,789,1084,1110]" italics="true" pageId="2" pageNumber="195">Heartwood</emphasis>
(from dead branch ca.
<quantity id="DB139B58ED12FF98FB85FBE1F262B231" box="[1052,1146,1084,1111]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="195" unit="mm" value="130.0">130 mm</quantity>
in diam.) yellowish brown with a reddish or pinkish tinge towards the sapwood.
<emphasis id="2E9FEAAFED12FF98FD77FBBDF57FB21C" box="[750,871,1120,1146]" italics="true" pageId="2" pageNumber="195">Branchlets</emphasis>
pendent, terete to ± longitudinally ribbed, smooth, lenticellate, initially green, becoming dark brown and then grey, glabrous.
<emphasis id="2E9FEAAFED12FF98FC26FB59F200B2F8" box="[959,1048,1156,1182]" italics="true" pageId="2" pageNumber="195">Stipules</emphasis>
intrapetiolar, base broad, apex acute at first but fugacious,
<quantity id="DB139B58ED12FF98FEEAFB75F7D6B2A5" box="[371,462,1192,1219]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="195" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, green at first, basal remains becoming dark brown.
<emphasis id="2E9FEAAFED12FF98FBA3FB74F290B2A4" box="[1082,1160,1193,1218]" italics="true" pageId="2" pageNumber="195">Leaves</emphasis>
opposite, sub-opposite (or occasionally alternate), distichous, simple, ovate to broadly elliptic, (35–)55–100(–115) × (20–)30–40(–50) mm, base rounded, apex broadly acuminate, occasionally acute, very occasionally obtuse, glabrous, firmly chartaceous, glossy, pale green at first, becoming fresh green, slightly paler but without a bloom below and very often with a yellowish tinge, margin entire, undulate, venation craspedodromous, usually slightly impressed above and slightly raised below, yellowish and paler than the lamina, principal lateral veins in (6–)7–9(–10) alternate pairs, tertiary venation minutely reticulate.
<emphasis id="2E9FEAAFED12FF98FF64FA5DF754B3FC" box="[253,332,1408,1434]" italics="true" pageId="2" pageNumber="195">Petiole</emphasis>
canaliculate above, (7–)11–12(–20) mm long, glabrous.
<emphasis id="2E9FEAAFED12FF98FC24FA5DF246B3FC" box="[957,1118,1408,1434]" italics="true" pageId="2" pageNumber="195">Inflorescences</emphasis>
axillary (or terminal), flowers solitary or in 2–5(–7)-flowered sub-fascicles.
<emphasis id="2E9FEAAFED12FF98FD00FA79F4EDB3D8" box="[665,757,1444,1470]" italics="true" pageId="2" pageNumber="195">Flowers</emphasis>
bisexual, actinomorphic, ca.
<quantity id="DB139B58ED12FF98FBD3FA79F297B3D9" box="[1098,1167,1444,1471]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="195" unit="mm" value="8.0">8 mm</quantity>
diameter, pedicel ca.
<quantity id="DB139B58ED12FF98FA16FA79F6ACB385" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="195" unit="mm" value="10.0">10 mm</quantity>
long, receptacle flattish.
<emphasis id="2E9FEAAFED12FF98FE41FA15F439B384" box="[472,545,1480,1506]" italics="true" pageId="2" pageNumber="195">Sepals</emphasis>
1-seriate, 5, free, ascending at anthesis, ovate to deltate, keeled on the adaxial surface, 3.0–3.5 × 2.0–
<quantity id="DB139B58ED12FF98FE16FA31F7F0B061" box="[399,488,1516,1543]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="195" unit="mm" value="2.5">2.5 mm</quantity>
, pale green to yellow-green, glabrous.
<emphasis id="2E9FEAAFED12FF98FC3CFA31F5F3B060" box="[933,1003,1516,1542]" italics="true" pageId="2" pageNumber="195">Petals</emphasis>
1-seriate, 5, free, alternate with sepals, somewhat spreading to ascending at anthesis, narrow and longitudinally folded upwards to clasp the filament, equal in length to sepals or slightly shorter, base unguiculate, apex cucullate, pale green, glabrous.
<emphasis id="2E9FEAAFED12FF98FBE9F9E9F2D5B028" box="[1136,1229,1588,1614]" italics="true" pageId="2" pageNumber="195">Stamens</emphasis>
5, haplostemonous.
<emphasis id="2E9FEAAFED12FF98FF11F985F6E0B014" box="[136,248,1624,1650]" italics="true" pageId="2" pageNumber="195">Filaments</emphasis>
inserted under the disc, ± equal in length to and clasped by the petals, whitish.
<emphasis id="2E9FEAAFED12FF98FBEEF985F2D6B014" box="[1143,1230,1624,1650]" italics="true" pageId="2" pageNumber="195">Anthers</emphasis>
basifixed, extrorse, whitish.
<emphasis id="2E9FEAAFED12FF98FF70F9A1F737B0F0" box="[233,303,1660,1686]" italics="true" pageId="2" pageNumber="195">Ovary</emphasis>
superior, 2-locular, largely enveloped by the swollen, pale yellow disc, style apex 2-fid.
<emphasis id="2E9FEAAFED12FF98FA88F9A0F353B0F0" box="[1297,1355,1661,1686]" italics="true" pageId="2" pageNumber="195">Fruit</emphasis>
a drupe, remains of disc and receptacle prominent at the base, oblong-ellipsoid, often faintly ribbed longitudinally, this being more pronounced in young and dry fruit, 18–25 ×
<quantity id="DB139B58ED12FF98FD27F919F521B0B9" box="[702,825,1732,1759]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="2" pageNumber="195" unit="mm" value="17.5" valueMax="19.0" valueMin="16.0">16–19 mm</quantity>
, but often narrower (from ca.
<quantity id="DB139B58ED12FF98FB12F919F2C4B0B8" box="[1163,1244,1732,1758]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="195" unit="mm" value="12.0">12 mm</quantity>
) in dried material, receptacle at point of fruit stalk’s attachment to base of fruit disc-shaped and plane, exocarp glabrous, green ripening bright orange-yellow, mesocarp fleshy, endocarp woody, stone ellipsoid-ovoid, 15–20 ×
<quantity id="DB139B58ED12FF98FBFFF8D1F2FAB141" box="[1126,1250,1804,1831]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="2" pageNumber="195" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
.
</paragraph>
<paragraph id="1C5436BDED12FF98FF24F8EDF28EB108" blockId="2.[136,1452,940,2083]" pageId="2" pageNumber="195">
<emphasis id="2E9FEAAFED12FF98FF24F8EDF747B12C" bold="true" box="[189,351,1840,1866]" pageId="2" pageNumber="195">Phenology:—</emphasis>
Flowering is mainly in spring from late October to early December at the same time that the new leaves are produced. Fruits are present from November to May and begin ripening in March.
</paragraph>
<paragraph id="1C5436BDED12FF99FF24F8A5F77FB7F6" blockId="2.[136,1452,940,2083]" lastBlockId="3.[136,1452,158,2057]" lastPageId="3" lastPageNumber="196" pageId="2" pageNumber="195">
<emphasis id="2E9FEAAFED12FF98FF24F8A5F40FB1F4" bold="true" box="[189,535,1912,1938]" pageId="2" pageNumber="195">Distinguishing characters:—</emphasis>
The new species is frequently confused with
<taxonomicName id="DBEB4D3EED12FF98FBBAF8A5F30FB1F4" box="[1059,1303,1912,1938]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED12FF98FBBAF8A5F30FB1F4" box="[1059,1303,1912,1938]" italics="true" pageId="2" pageNumber="195">Phyllogeiton discolor</emphasis>
</taxonomicName>
, but the two species’ ranges are allopatric with a gap of ca.
<quantity id="DB139B58ED12FF98FD29F841F510B1D0" box="[688,776,1948,1974]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="195" unit="km" value="300.0">300 km</quantity>
between the closest populations.
<taxonomicName id="DBEB4D3EED12FF98FB08F841F3B3B1D0" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[1169,1451,1948,1974]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED12FF98FB08F841F3B3B1D0" box="[1169,1451,1948,1974]" italics="true" pageId="2" pageNumber="195">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
is easily distinguished from
<taxonomicName id="DBEB4D3EED12FF98FE54F81CF45EB1BC" box="[461,582,1984,2010]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED12FF98FE54F81CF45EB1BC" box="[461,582,1984,2010]" italics="true" pageId="2" pageNumber="195">P. discolor</emphasis>
</taxonomicName>
by usually growing on dry, sandy soils (
<emphasis id="2E9FEAAFED12FF98FB86F81CF225B1BC" box="[1055,1085,1985,2010]" italics="true" pageId="2" pageNumber="195">vs.</emphasis>
usually adjacent to rivers or on termitaria), bark on the bole of mature trees almost black, very rough, exfoliating in large pieces (
<emphasis id="2E9FEAAFED12FF98FB28F838F2D7B198" box="[1201,1231,2021,2046]" italics="true" pageId="2" pageNumber="195">vs.</emphasis>
grey-brown, rough, tessellated), branchlets and leaves glabrous (
<emphasis id="2E9FEAAFED12FF98FD1FF7D4F4BDBE44" box="[646,677,2057,2082]" italics="true" pageId="2" pageNumber="195">vs.</emphasis>
glabrous or minutely and densely pubescent; on leaves pubescence mainly on the abaxial surface, but the adaxial surface may also be minutely pubescent about the midrib), leaf blade fresh green, without a bloom, ± concolorous (
<emphasis id="2E9FEAAFED13FF99FD09FF1EF4B7B6BA" box="[656,687,195,220]" italics="true" pageId="3" pageNumber="196">vs.</emphasis>
dark green, distinctly paler with a greyish bloom below), leaf apex acuminate (
<emphasis id="2E9FEAAFED13FF99FE94FF3AF733B766" box="[269,299,231,256]" italics="true" pageId="3" pageNumber="196">vs.</emphasis>
acute to obtuse), petals spreading to somewhat ascending at anthesis (
<emphasis id="2E9FEAAFED13FF99FBD4FF3AF27CB766" box="[1101,1124,231,256]" italics="true" pageId="3" pageNumber="196">vs</emphasis>
. spreading to reflexed), fruit oblong-ellipsoid (
<emphasis id="2E9FEAAFED13FF99FEC9FED6F776B742" box="[336,366,267,292]" italics="true" pageId="3" pageNumber="196">vs.</emphasis>
slightly or markedly ovoid), up to 25 ×
<quantity id="DB139B58ED13FF99FCB4FED7F564B743" box="[813,892,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="196" unit="mm" value="19.0">19 mm</quantity>
(
<emphasis id="2E9FEAAFED13FF99FC15FED6F5B2B742" box="[908,938,267,292]" italics="true" pageId="3" pageNumber="196">vs.</emphasis>
up to 20 ×
<quantity id="DB139B58ED13FF99FBB0FED7F263B742" box="[1065,1147,266,292]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="196" unit="mm" value="11.0">11 mm</quantity>
), exocarp often indistinctly longitudinally ribbed (
<emphasis id="2E9FEAAFED13FF99FE10FEF2F7BFB72E" box="[393,423,303,328]" italics="true" pageId="3" pageNumber="196">vs.</emphasis>
plane), bright orange-yellow, even when overripe (
<emphasis id="2E9FEAAFED13FF99FC6EFEF2F20EB72E" box="[1015,1046,303,328]" italics="true" pageId="3" pageNumber="196">vs.</emphasis>
yellow but very often pale yellow; often brownish yellow when overripe), and disc-shaped receptacle at point of fruit stalk’s attachment to base of fruit plane (
<emphasis id="2E9FEAAFED13FF99FF4CFEAAF6F4B7F6" box="[213,236,375,400]" italics="true" pageId="3" pageNumber="196">vs</emphasis>
. concave).
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED13FF99FF24FE47F5B6B447" pageId="3" pageNumber="196" type="etymology">
<paragraph id="1C5436BDED13FF99FF24FE47F5B6B447" blockId="3.[136,1452,158,2057]" pageId="3" pageNumber="196">
<emphasis id="2E9FEAAFED13FF99FF24FE47F77CB7D2" bold="true" box="[189,356,410,436]" pageId="3" pageNumber="196">Etymology:—</emphasis>
The specific epithet “
<taxonomicName id="DBEB4D3EED13FF99FDCBFE47F4CBB7D2" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[594,723,410,436]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED13FF99FDCBFE47F4CBB7D2" box="[594,723,410,436]" italics="true" pageId="3" pageNumber="196">trachybasis</emphasis>
</taxonomicName>
” is derived from the Greek τραχυς,
<emphasis id="2E9FEAAFED13FF99FBFAFE47F2ADB7D2" box="[1123,1205,410,436]" italics="true" pageId="3" pageNumber="196">trachys</emphasis>
, rough or shaggy, and βασις,
<emphasis id="2E9FEAAFED13FF99FF4CFE63F717B7BE" box="[213,271,446,472]" italics="true" pageId="3" pageNumber="196">basis</emphasis>
, base or pedestal, referring to the bark towards the base of the trunk in old trees that becomes very thick, rough and flaking in large pieces (
<figureCitation id="84D02A38ED13FF99FD9AFE3FF451B79A" box="[515,585,482,508]" captionStart="FIGURE 1" captionStartId="4.[136,229,1991,2013]" captionTargetBox="[151,1435,187,1964]" captionTargetId="figure-19@4.[151,1435,187,1964]" captionTargetPageId="4" captionText="FIGURE 1. Phyllogeiton trachybasis, habit and habitat. Mature tree in Sand Forest at Bonamanzi Game Reserve, Hluhluwe, KwaZulu- Natal, South Africa. Photograph: R.G.C. Boon." figureDoi="http://doi.org/10.5281/zenodo.7690759" httpUri="https://zenodo.org/record/7690759/files/figure.png" pageId="3" pageNumber="196">Figs 1</figureCitation>
&amp;
<figureCitation id="84D02A38ED13FF99FDF2FE3FF4ACB79A" box="[619,692,482,508]" captionStart="FIGURE 2" captionStartId="5.[136,229,1955,1977]" captionTargetBox="[164,1423,190,1931]" captionTargetId="figure-18@5.[164,1423,190,1931]" captionTargetPageId="5" captionText="FIGURE 2. Phyllogeiton trachybasis, bark morphology illustrating variation (A–D) and colour of the wood (E). A. Bark on trunk of old, mature tree. B. Bark on trunk of relatively young tree (sunny habitat). C. Bark on trunk of relatively young tree (shady habitat). D. Exceedingly rough and flaking bark on trunk of mature tree. E. Finely sanded, untreated wood in transverse section; from dead branch ± 130 mm in diameter. Photographs: R.G.C. Boon (A, D), F. du Randt (B, C), &amp; D. Bishop (E)." figureDoi="http://doi.org/10.5281/zenodo.7690761" httpUri="https://zenodo.org/record/7690761/files/figure.png" pageId="3" pageNumber="196">2A–D</figureCitation>
). The epithet is used here as a noun in apposition: “the shaggy base” and therefore does not necessarily agree in gender with the genus name.
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED13FF99FF24FDF7F5B1B592" pageId="3" pageNumber="196" type="distribution">
<paragraph id="1C5436BDED13FF99FF24FDF7F304B4D6" blockId="3.[136,1452,158,2057]" pageId="3" pageNumber="196">
<emphasis id="2E9FEAAFED13FF99FF24FDF7F76DB422" bold="true" box="[189,373,554,580]" pageId="3" pageNumber="196">Distribution:—</emphasis>
<taxonomicName id="DBEB4D3EED13FF99FEECFDF7F494B422" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[373,652,554,580]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED13FF99FEECFDF7F494B422" box="[373,652,554,580]" italics="true" pageId="3" pageNumber="196">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
is only known with certainty from the southern part of the Maputaland Centre of Endemism, an area rich in restricted-range plants and animals (
<bibRefCitation id="787A4B4CED13FF99FC54FD93F263B40E" author="Van Wyk, A. E." box="[973,1147,590,616]" pageId="3" pageNumber="196" pagination="198 - 207" refId="ref11406" refString="Van Wyk, A. E. (1996) Biodiversity of the Maputaland Centre. In: Van der Maesen, L. J. G., Van der Burgt, X. M. &amp; Van Medenbach de Rooy, J. M. (Eds.) The biodiversity of African plants. Kluwer Academic Publishers, Dordrecht, pp. 198 - 207." type="book chapter" year="1996">Van Wyk 1996</bibRefCitation>
,
<bibRefCitation id="787A4B4CED13FF99FB11FD92F387B40E" author="Van Wyk, A. E. &amp; Smith, G. F." box="[1160,1439,590,617]" pageId="3" pageNumber="196" refId="ref11475" refString="Van Wyk, A. E. &amp; Smith, G. F. (2001) Regions of floristic endemism in southern Africa: a review with emphasis on succulents. Umdaus Press, Hatfield, Pretoria, 199 pp." type="book" year="2001">Van Wyk &amp; Smith 2001</bibRefCitation>
). The Maputaland Centre is at the southern end of the tropics in Africa (
<bibRefCitation id="787A4B4CED13FF99FC3AFDAFF255B4EA" author="Van Wyk, A. E." box="[931,1101,626,652]" pageId="3" pageNumber="196" pagination="198 - 207" refId="ref11406" refString="Van Wyk, A. E. (1996) Biodiversity of the Maputaland Centre. In: Van der Maesen, L. J. G., Van der Burgt, X. M. &amp; Van Medenbach de Rooy, J. M. (Eds.) The biodiversity of African plants. Kluwer Academic Publishers, Dordrecht, pp. 198 - 207." type="book chapter" year="1996">Van Wyk 1996</bibRefCitation>
) and at the northern end of the Maputaland-Pondoland-Albany Hotspot, one of 36 global biodiversity hotspots (
<bibRefCitation id="787A4B4CED13FF99FB96FD4BF308B4D6" author="Steenkamp, Y. &amp; Van Wyk, B. &amp; A. E &amp; Victor, J. &amp; Hoare, D. &amp; Smith, G. &amp; Dold, T. &amp; Cowling, R." box="[1039,1296,662,688]" pageId="3" pageNumber="196" pagination="219 - 228" refId="ref11056" refString="Steenkamp, Y., Van Wyk, B. [A. E.], Victor, J., Hoare, D., Smith, G., Dold, T. &amp; Cowling, R. (2004) Maputaland-Pondoland-Albany. In: Mittermeier, R. A., Robles Gil, P., Hoffmann, M., Pilgrim, J., Brooks, T., Goettsch Mittermeier, C., Lamoreux, J. &amp; da Fonseca, G. A. B. (Eds.) Hotspots revisited: Earth's biologically richest and most threatened terrestrial ecoregions. Cemex &amp; Conservation International, Washington, pp. 219 - 228." type="book chapter" year="2004">
Steenkamp
<emphasis id="2E9FEAAFED13FF99FB0DFD4AF2D4B4D6" box="[1172,1228,662,688]" italics="true" pageId="3" pageNumber="196">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
<paragraph id="1C5436BDED13FF99FF24FD67F540B57A" blockId="3.[136,1452,158,2057]" pageId="3" pageNumber="196">
<materialsCitation id="AC833CE0ED13FF99FF24FD67F540B57A" ID-GBIF-Occurrence="4041537307" county="Lake St Lucia" location="South African" municipality="Ndumo Game Reserve" pageId="3" pageNumber="196" specimenCount="1" stateProvince="KwaZulu-Natal">
Within the Maputaland Centre,
<taxonomicName id="DBEB4D3EED13FF99FDB7FD66F4D6B4B2" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[558,718,698,724]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED13FF99FDB7FD66F4D6B4B2" box="[558,718,698,724]" italics="true" pageId="3" pageNumber="196">P. trachybasis</emphasis>
</taxonomicName>
is known from 10 localities in
<collectingRegion id="DE2FF85FED13FF99FBDCFD67F2EDB4B2" box="[1093,1269,698,724]" country="South Africa" name="KwaZulu-Natal" pageId="3" pageNumber="196">KwaZulu-Natal</collectingRegion>
(
<figureCitation id="84D02A38ED13FF99FA91FD67F356B4B3" box="[1288,1358,698,725]" captionStart="FIGURE 5" captionStartId="8.[136,229,1283,1305]" captionTargetBox="[151,1436,190,1258]" captionTargetId="figure-19@8.[151,1436,190,1258]" captionTargetPageId="8" captionText="FIGURE 5. Topographical map showing the known distribution (black dots) of Phyllogeiton trachybasis in northeastern KwaZulu-Natal, South Africa. Map based on herbarium collections in BNRH, K, NH, NU, PCE, PRE, PRU, and UDW.The insert shows a map of southern Africa with names of countries; the grey rectangle indicates the area depicted by the topographical map." figureDoi="http://doi.org/10.5281/zenodo.7690765" httpUri="https://zenodo.org/record/7690765/files/figure.png" pageId="3" pageNumber="196">Fig. 5</figureCitation>
). These localities range from Hluhluwe and
<collectingCounty id="F5354E31ED13FF99FD8EFD03F4ABB49E" box="[535,691,734,760]" pageId="3" pageNumber="196">Lake St Lucia</collectingCounty>
in the south to Tembe Elephant Park and
<collectingMunicipality id="FC30ACC7ED13FF99FBE6FD03F364B49F" box="[1151,1404,734,761]" pageId="3" pageNumber="196">Ndumo Game Reserve</collectingMunicipality>
just south of the
<location id="19346066ED13FF99FE8AFCDFF7A8B57A" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED13FF99FE8AFCDFF7A8B57A" box="[275,432,770,796]" county="Lake St Lucia" municipality="Ndumo Game Reserve" name="South African" pageId="3" pageNumber="196" stateProvince="KwaZulu-Natal">South African</location>
and Mozambican border in the north.
</materialsCitation>
</paragraph>
<paragraph id="1C5436BDED13FF99FF24FCFBF5B1B592" blockId="3.[136,1452,158,2057]" pageId="3" pageNumber="196">
There are no specimens of
<taxonomicName id="DBEB4D3EED13FF99FE61FCFAF48FB526" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[504,663,806,832]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED13FF99FE61FCFAF48FB526" box="[504,663,806,832]" italics="true" pageId="3" pageNumber="196">P. trachybasis</emphasis>
</taxonomicName>
at the Institute for Agricultural Research of
<collectingCountry id="64FC762DED13FF99FB02FCFBF337B526" box="[1179,1327,806,832]" name="Mozambique" pageId="3" pageNumber="196">Mozambique</collectingCountry>
herbarium (LMA) and its presence in
<collectingCountry id="64FC762DED13FF99FE23FC97F456B502" box="[442,590,842,868]" name="Mozambique" pageId="3" pageNumber="196">Mozambique</collectingCountry>
needs confirmation. The species is, however, almost certainly present in the far south of
<collectingCounty id="F5354E31ED13FF99FE89FCB3F7AEB5EE" box="[272,438,878,904]" pageId="3" pageNumber="196">Mozambique’s</collectingCounty>
<collectingRegion id="DE2FF85FED13FF99FE27FCB2F498B5EE" box="[446,640,878,905]" country="Mozambique" name="Maputo" pageId="3" pageNumber="196">Maputo Province</collectingRegion>
considering that trees of the new species were found in Tembe Elephant Park on the
<collectingMunicipality id="FC30ACC7ED13FF99FE97FC4FF7B2B5CA" box="[270,426,914,940]" pageId="3" pageNumber="196">South African</collectingMunicipality>
side directly next to the border between the two countries. A sight record (
<collectorName id="B11E536BED13FF99FB75FC4FF373B5CA" box="[1260,1387,914,940]" pageId="3" pageNumber="196">E. Schmidt</collectorName>
, pers. comm.) and information supplied by a traditional healer (
<collectorName id="B11E536BED13FF99FC88FC6BF5BBB5B6" box="[785,931,950,976]" pageId="3" pageNumber="196">C. Hanekom</collectorName>
, pers. comm.) suggest that it may occur at the
<collectingRegion id="DE2FF85FED13FF99FF11FC06F6C7B593" box="[136,223,987,1013]" country="Mozambique" name="Maputo" pageId="3" pageNumber="196">Maputo</collectingRegion>
<location id="19346066ED13FF99FF7FFC07F782B593" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED13FF99FF7FFC07F782B593" box="[230,410,986,1013]" country="Mozambique" county="Mozambique's" municipality="South African" name="Special Reserve" pageId="3" pageNumber="196" stateProvince="Maputo">Special Reserve</location>
south of
<collectingRegion id="DE2FF85FED13FF99FD9AFC06F445B593" box="[515,605,987,1013]" country="Mozambique" name="Maputo" pageId="3" pageNumber="196">Maputo</collectingRegion>
, but confirmation is required.
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED13FF99FF24FC22F209B35E" pageId="3" pageNumber="196" type="materials_examined">
<paragraph id="1C5436BDED13FF99FF24FC22F209B35E" blockId="3.[136,1452,158,2057]" pageId="3" pageNumber="196">
Two other possible localities for the new species should be mentioned. The first is ca.
<quantity id="DB139B58ED13FF99FB1BFC23F2D1B27E" box="[1154,1225,1022,1048]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="196" unit="km" value="60.0">60 km</quantity>
from Caia in
<collectingRegion id="DE2FF85FED13FF99FAFDFC23F6F6B25A" country="Mozambique" name="Sofala" pageId="3" pageNumber="196">Sofala Province</collectingRegion>
,
<collectingCountry id="64FC762DED13FF99FF61FBFFF797B25A" box="[248,399,1058,1084]" name="Mozambique" pageId="3" pageNumber="196">Mozambique</collectingCountry>
, on the EN1 route to Gorongosa (grid 1834BA), where a sterile specimen (
<emphasis id="2E9FEAAFED13FF99FB5FFBFEF6D8B206" italics="true" pageId="3" pageNumber="196">
J.E. &amp; S.M. Burrows
<quantity id="DB139B58ED13FF99FF11FB9BF6D8B206" box="[136,192,1094,1120]" metricMagnitude="2" metricUnit="m" metricValue="2.2369779999999997" pageId="3" pageNumber="196" unit="in" value="8807.0">8807</quantity>
</emphasis>
in Herb. BNRH) was collected by botanists familiar with the tree. This location is nearly
<quantity id="DB139B58ED13FF99FB46FB9BF35DB206" box="[1247,1349,1094,1120]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="196" unit="km" value="1000.0">1000 km</quantity>
north of confirmed localities in KwaZulu-Natal. The second locality is near Siteki in Scarp Forest at an elevation of
<quantity id="DB139B58ED13FF99FADEFBB7F396B2E3" box="[1351,1422,1130,1157]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="196" unit="m" value="450.0">450 m</quantity>
in the Lebombo [
<collectingRegion id="DE2FF85FED13FF99FEB2FB53F786B2CE" box="[299,414,1166,1192]" country="Swaziland" name="Lubombo" pageId="3" pageNumber="196">Lubombo</collectingRegion>
] Mountains of
<collectingCountry id="64FC762DED13FF99FDD2FB53F4B5B2CE" box="[587,685,1166,1192]" name="Swaziland" pageId="3" pageNumber="196">Eswatini</collectingCountry>
(formerly
<collectingCountry id="64FC762DED13FF99FCBEFB53F5B8B2CE" box="[807,928,1166,1192]" name="Swaziland" pageId="3" pageNumber="196">Swaziland</collectingCountry>
), about
<quantity id="DB139B58ED13FF99FC63FB53F25AB2CE" box="[1018,1090,1166,1192]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="196" unit="km" value="40.0">40 km</quantity>
north of Ndumo Game Reserve (grid 2631BD). A specimen collected here (
<emphasis id="2E9FEAAFED13FF99FDF3FB6EF52EB2AA" box="[618,822,1202,1229]" italics="true" pageId="3" pageNumber="196">P. &amp; L. Loffler s.n.</emphasis>
in Herb. BNRH) and photographs supplied by L. Loffler appear to match
<taxonomicName id="DBEB4D3EED13FF99FEA6FB0AF7C2B296" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[319,474,1238,1264]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED13FF99FEA6FB0AF7C2B296" box="[319,474,1238,1264]" italics="true" pageId="3" pageNumber="196">P. trachybasis</emphasis>
</taxonomicName>
, but the five trees seen were all sterile.Another collection from this same locality is also sterile (
<emphasis id="2E9FEAAFED13FF99FF45FB27F76DB372" box="[220,373,1274,1300]" italics="true" pageId="3" pageNumber="196">
Schmidt
<quantity id="DB139B58ED13FF99FEA4FB27F76DB372" box="[317,373,1274,1300]" metricMagnitude="2" metricUnit="m" metricValue="1.029716" pageId="3" pageNumber="196" unit="in" value="4054.0">4054</quantity>
</emphasis>
in Ernst Schmidt Private Herbarium). Given that these specimens lack reproductive material, the presence of the species in
<collectingCountry id="64FC762DED13FF99FE34FAC3F459B35E" box="[429,577,1310,1336]" name="Mozambique" pageId="3" pageNumber="196">Mozambique</collectingCountry>
and
<collectingCountry id="64FC762DED13FF99FDEEFAC3F4C1B35E" box="[631,729,1310,1336]" name="Swaziland" pageId="3" pageNumber="196">Eswatini</collectingCountry>
still needs to be confirmed.
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED13FF92FF24FA9FF51FB08B" lastPageId="8" lastPageNumber="201" pageId="3" pageNumber="196" type="biology_ecology">
<paragraph id="1C5436BDED13FF99FF24FA9FF536B077" blockId="3.[136,1452,158,2057]" pageId="3" pageNumber="196">
<emphasis id="2E9FEAAFED13FF99FF24FA9FF759B33A" bold="true" box="[189,321,1346,1372]" pageId="3" pageNumber="196">Ecology:—</emphasis>
<taxonomicName id="DBEB4D3EED13FF99FEDBFA9FF442B33A" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[322,602,1346,1372]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED13FF99FEDBFA9FF442B33A" box="[322,602,1346,1372]" italics="true" pageId="3" pageNumber="196">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
is a sub-canopy or canopy tree found mainly on well-drained, deep sandy soils at low elevations to about
<quantity id="DB139B58ED13FF99FE7EFABBF436B3E7" box="[487,558,1382,1409]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="196" unit="m" value="100.0">100 m</quantity>
above sea level. Trees grow mainly in Tembe Sandy Bushveld (SVl 18) in open to closed woodland (codes of vegetation
<typeStatus id="C350881FED13FF99FDCAFA56F496B3C3" box="[595,654,1419,1445]" pageId="3" pageNumber="196">types</typeStatus>
follow
<bibRefCitation id="787A4B4CED13FF99FD7CFA57F200B3C2" author="Mucina, L. &amp; Rutherford, M. C." box="[741,1048,1418,1444]" pageId="3" pageNumber="196" refId="ref10329" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
and SANBI 2006–2018). They also grow in Sand Forest (FOz 8) patches in a mosaic with Tembe Sandy Bushveld (
<figureCitation id="84D02A38ED13FF99FB9BFA73F25BB3AE" box="[1026,1091,1454,1480]" captionStart="FIGURE 1" captionStartId="4.[136,229,1991,2013]" captionTargetBox="[151,1435,187,1964]" captionTargetId="figure-19@4.[151,1435,187,1964]" captionTargetPageId="4" captionText="FIGURE 1. Phyllogeiton trachybasis, habit and habitat. Mature tree in Sand Forest at Bonamanzi Game Reserve, Hluhluwe, KwaZulu- Natal, South Africa. Photograph: R.G.C. Boon." figureDoi="http://doi.org/10.5281/zenodo.7690759" httpUri="https://zenodo.org/record/7690759/files/figure.png" pageId="3" pageNumber="196">Fig. 1</figureCitation>
). The species may also occur in Western Maputaland Sandy Bushveld (SVI19) at Ndumo Game Reserve, but information recorded on collecting labels is insufficient to confirm its presence in this vegetation
<typeStatus id="C350881FED13FF99FD6EFA2AF533B077" box="[759,811,1527,1553]" pageId="3" pageNumber="196">type</typeStatus>
.
</paragraph>
<paragraph id="1C5436BDED13FF99FF24F9C7F542B01A" blockId="3.[136,1452,158,2057]" pageId="3" pageNumber="196">
<taxonomicName id="DBEB4D3EED13FF99FF24F9C7F7CEB052" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[189,470,1562,1588]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED13FF99FF24F9C7F7CEB052" box="[189,470,1562,1588]" italics="true" pageId="3" pageNumber="196">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
<materialsCitation id="AC833CE0ED13FF99FE46F9C7F54EB01A" ID-GBIF-Occurrence="4041248301" location="Northern Coastal Forest" municipality="Lake Sibaya" pageId="3" pageNumber="196" specimenCount="1" stateProvince="Western Cape">
has also been collected between
<collectingRegion id="DE2FF85FED13FF99FCCAF9C7F595B052" box="[851,909,1562,1588]" country="South Africa" name="Western Cape" pageId="3" pageNumber="196">Cape</collectingRegion>
Vidal and
<collectingMunicipality id="FC30ACC7ED13FF99FB93F9C7F28FB052" box="[1034,1175,1562,1588]" pageId="3" pageNumber="196">Lake Sibaya</collectingMunicipality>
in dune forest, which is included in the broader category
<location id="19346066ED13FF99FE67F9E3F515B03F" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED13FF99FE67F9E3F515B03F" box="[510,781,1598,1625]" municipality="Lake Sibaya" name="Northern Coastal Forest" pageId="3" pageNumber="196" stateProvince="Western Cape">Northern Coastal Forest</location>
(FOz 7). The most recent of these collections was in 1997, but the demographics of the species in dune forest are unknown
</materialsCitation>
.
</paragraph>
<paragraph id="1C5436BDED13FF99FF24F95BF521B11E" blockId="3.[136,1452,158,2057]" pageId="3" pageNumber="196">
Besides its normal sandy habitat, the species was collected once (
<emphasis id="2E9FEAAFED13FF99FC02F95BF217B0C6" box="[923,1039,1670,1696]" italics="true" pageId="3" pageNumber="196">
Moll
<quantity id="DB139B58ED13FF99FC4EF95BF217B0C6" box="[983,1039,1670,1696]" metricMagnitude="2" metricUnit="m" metricValue="1.343914" pageId="3" pageNumber="196" unit="in" value="5291.0">5291</quantity>
</emphasis>
in Herbs K, NH, PRE) in 1971 from riverine forest embedded in Zululand Lowveld (SVl 23). This was on the Mansiya River in the Hluhluwe-iMfolozi Park where the presence of eight trees growing in the river’s floodplain was recently confirmed (F. du Randt, S. Louw, and S. Mabongwa, pers. comm.). The new species also apparently occurs occasionally in Scarp Forest (FOz 5). It was collected in this forest
<typeStatus id="C350881FED13FF99FE58F8CAF7E9B157" box="[449,497,1815,1841]" pageId="3" pageNumber="196">type</typeStatus>
in
<quantity id="DB139B58ED13FF99FD80F8CBF477B156" box="[537,623,1814,1841]" metricMagnitude="1" metricUnit="m" metricValue="4.975859999999999" pageId="3" pageNumber="196" unit="in" value="1959.0">1959 in</quantity>
the Hluhluwe-iMfolozi Park on a steep, southwest-facing rocky slope at an elevation of about
<quantity id="DB139B58ED13FF99FEE3F8E7F7D9B133" box="[378,449,1850,1877]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="196" unit="m" value="500.0">500 m</quantity>
(
<emphasis id="2E9FEAAFED13FF99FE48F8E7F453B132" box="[465,587,1850,1876]" italics="true" pageId="3" pageNumber="196">
Ward
<quantity id="DB139B58ED13FF99FD8AF8E7F453B132" box="[531,587,1850,1876]" metricMagnitude="1" metricUnit="m" metricValue="7.48538" pageId="3" pageNumber="196" unit="in" value="2947.0">2947</quantity>
</emphasis>
in Herb. NU). A visit by F. du Randt, S. Louw, and S. Mabongwa on
<date id="6855107DED13FF99FACCF8E7F6D8B11E" pageId="3" pageNumber="196" value="2021-05-18">18 May 2021</date>
to try and relocate this plant or plants was unsuccessful.
</paragraph>
<paragraph id="1C5436BDED13FF99FF24F85FF796BE6E" blockId="3.[136,1452,158,2057]" pageId="3" pageNumber="196">
Flowers are visited by honey bees. The fruit is eaten by birds, notably Trumpeter Hornbills,
<taxonomicName id="DBEB4D3EED13FF99FB59F85EF753B1A6" authority="Temminck (1824" authorityName="Temminck" authorityYear="1824" class="Aves" family="Bucerotidae" genus="Bycanistes" kingdom="Animalia" order="Bucerotiformes" pageId="3" pageNumber="196" phylum="Chordata" rank="species" species="bucinator">
<emphasis id="2E9FEAAFED13FF99FB59F85EF3B3B1FA" box="[1216,1451,1922,1948]" italics="true" pageId="3" pageNumber="196">Bycanistes bucinator</emphasis>
<bibRefCitation id="787A4B4CED13FF99FF11F87BF753B1A6" author="Temminck, C. J." box="[136,331,1958,1984]" pageId="3" pageNumber="196" pagination="99" refId="ref11203" refString="Temminck, C. J. (1824) Calao trompette Buceros bucinator. In: Nouveau recueil de planches colorieies d'oiseaux: pour servir de suite et de compleiment aux planches enlumineies de Buffon, eidition in-folio et in- 4 r de l'Imprimerie royale, 1770 (1838), vol. 2, livr. 48, pl. 284. Chez Legras Imbert et Comp, Strasbourgh, p. 99. https: // doi. org / 10.5962 / bhl. title. 51468" type="book chapter" year="1824">Temminck (1824</bibRefCitation>
</taxonomicName>
: livr. 48, pl. 284), at Nibela and Vervet Monkeys,
<taxonomicName id="DBEB4D3EED13FF99FCEDF87BF356B1A6" authority="Cuvier (1821: 2)" authorityName="Cuvier" authorityPageNumber="2" authorityYear="1821" box="[884,1358,1958,1984]" class="Mammalia" family="Cercopithecidae" genus="Chlorocebus" kingdom="Animalia" order="Primates" pageId="3" pageNumber="196" phylum="Chordata" rank="species" species="pygerythrus">
<emphasis id="2E9FEAAFED13FF99FCEDF87BF296B1A6" box="[884,1166,1958,1984]" italics="true" pageId="3" pageNumber="196">Chlorocebus pygerythrus</emphasis>
<bibRefCitation id="787A4B4CED13FF99FB0CF87BF356B1A6" author="Cuvier, F." box="[1173,1358,1958,1984]" pageId="3" pageNumber="196" pagination="1 - 2" refId="ref9095" refString="Cuvier, F. (1821) Le vervet. In: Geoffroy-Saint-Hilaire, E. &amp; Cuvier, F. (Eds.) Histoire naturelle des mammiferes 3 [1824], part 24, Chez A. Belin, Paris, pp. 1 - 2. https: // doi. org / 10.5962 / bhl. title. 78766" type="book chapter" year="1821">Cuvier (1821: 2)</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="787A4B4CED13FF99FAC5F87BF6DBB182" author="Pooley, E. S." pageId="3" pageNumber="196" pagination="1 - 54" refId="ref10531" refString="Pooley, E. S. (1978) A checklist of the plants of Ndumu Game Reserve, north-eastern Zululand. Journal of South African Botany 44: 1 - 54. [https: // archive. org / details / journalofsouthaf 44 unse / page / n 11 / mode / 2 up]" type="journal article" year="1978">Pooley 1978</bibRefCitation>
, C. Hanekom, pers. comm.), although at Nibela it is reported that monkeys and people don’t feed on the fruit (G. Linforth, pers. comm.).
</paragraph>
<caption id="48946635ED14FF9EFF11F81AF456BE67" ID-DOI="http://doi.org/10.5281/zenodo.7690759" ID-Zenodo-Dep="7690759" httpUri="https://zenodo.org/record/7690759/files/figure.png" pageId="4" pageNumber="197" startId="4.[136,229,1991,2013]" targetBox="[151,1435,187,1964]" targetPageId="4">
<paragraph id="1C5436BDED14FF9EFF11F81AF456BE67" blockId="4.[136,1452,1991,2049]" pageId="4" pageNumber="197">
<emphasis id="2E9FEAAFED14FF9EFF11F81AF6E6B1BA" bold="true" box="[136,254,1991,2013]" pageId="4" pageNumber="197">FIGURE 1.</emphasis>
<taxonomicName id="DBEB4D3EED14FF9EFE9CF81AF7EDB1BA" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[261,501,1991,2012]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="4" pageNumber="197" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED14FF9EFE9CF81AF7EDB1BA" box="[261,501,1991,2012]" italics="true" pageId="4" pageNumber="197">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
, habit and habitat. Mature tree in Sand Forest at Bonamanzi Game Reserve, Hluhluwe, KwaZuluNatal, South Africa. Photograph: R.G.C. Boon.
</paragraph>
</caption>
<caption id="48946635ED15FF9FFF11F87EF211BE43" ID-DOI="http://doi.org/10.5281/zenodo.7690761" ID-Zenodo-Dep="7690761" httpUri="https://zenodo.org/record/7690761/files/figure.png" pageId="5" pageNumber="198" startId="5.[136,229,1955,1977]" targetBox="[164,1423,190,1931]" targetPageId="5">
<paragraph id="1C5436BDED15FF9FFF11F87EF211BE43" blockId="5.[136,1452,1955,2085]" pageId="5" pageNumber="198">
<emphasis id="2E9FEAAFED15FF9FFF11F87EF6E1B1DE" bold="true" box="[136,249,1955,1977]" pageId="5" pageNumber="198">FIGURE 2</emphasis>
.
<taxonomicName id="DBEB4D3EED15FF9FFE9EF87EF7E1B1DE" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[263,505,1955,1976]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="198" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED15FF9FFE9EF87EF7E1B1DE" box="[263,505,1955,1976]" italics="true" pageId="5" pageNumber="198">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
, bark morphology illustrating variation (A–D) and colour of the wood (E).
<emphasis id="2E9FEAAFED15FF9FFB7EF87EF2E6B1DE" bold="true" box="[1255,1278,1955,1976]" pageId="5" pageNumber="198">A.</emphasis>
Bark on trunk of old, mature tree.
<emphasis id="2E9FEAAFED15FF9FFEB7F815F75CB1BB" bold="true" box="[302,324,1992,2013]" pageId="5" pageNumber="198">B.</emphasis>
Bark on trunk of relatively young tree (sunny habitat).
<emphasis id="2E9FEAAFED15FF9FFCF8F81AF560B1BB" bold="true" box="[865,888,1991,2013]" pageId="5" pageNumber="198">C.</emphasis>
Bark on trunk of relatively young tree (shady habitat).
<emphasis id="2E9FEAAFED15FF9FFA0DF815F3B4B1BB" bold="true" box="[1428,1452,1992,2013]" pageId="5" pageNumber="198">D.</emphasis>
Exceedingly rough and flaking bark on trunk of mature tree.
<emphasis id="2E9FEAAFED15FF9FFD4AF831F4F1BE67" bold="true" box="[723,745,2028,2049]" pageId="5" pageNumber="198">E.</emphasis>
Finely sanded, untreated wood in transverse section; from dead branch ± 130 mm in diameter. Photographs: R.G.C. Boon (A, D), F. du Randt (B, C), &amp; D. Bishop (E).
</paragraph>
</caption>
<caption id="48946635ED16FF9CFF11F87CF387B198" ID-DOI="http://doi.org/10.5281/zenodo.7690763" ID-Zenodo-Dep="7690763" httpUri="https://zenodo.org/record/7690763/files/figure.png" pageId="6" pageNumber="199" startId="6.[136,229,1953,1975]" targetBox="[151,1435,190,1929]" targetPageId="6">
<paragraph id="1C5436BDED16FF9CFF11F87CF387B198" blockId="6.[136,1452,1953,2047]" pageId="6" pageNumber="199">
<emphasis id="2E9FEAAFED16FF9CFF11F87CF6E5B1D0" bold="true" box="[136,253,1953,1975]" pageId="6" pageNumber="199">FIGURE 3.</emphasis>
<taxonomicName id="DBEB4D3EED16FF9CFE9AF87CF7EAB1D0" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[259,498,1953,1974]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="199" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED16FF9CFE9AF87CF7EAB1D0" box="[259,498,1953,1974]" italics="true" pageId="6" pageNumber="199">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
, leaf, flower and fruit morphology.
<emphasis id="2E9FEAAFED16FF9CFCDEF87CF547B1D0" bold="true" box="[839,863,1953,1974]" pageId="6" pageNumber="199">A.</emphasis>
Branchlet with leaves, the latter fresh green, without a bloom and ca. concolorous.
<emphasis id="2E9FEAAFED16FF9CFECEF818F775B1BC" bold="true" box="[343,365,1989,2010]" pageId="6" pageNumber="199">B.</emphasis>
Flowers, open and in bud.
<emphasis id="2E9FEAAFED16FF9CFDE3F818F489B1BD" bold="true" box="[634,657,1989,2011]" pageId="6" pageNumber="199">C.</emphasis>
Almost ripe fruit.
<emphasis id="2E9FEAAFED16FF9CFCD1F818F578B1BC" bold="true" box="[840,864,1989,2010]" pageId="6" pageNumber="199">D.</emphasis>
Mainly ripe fallen fruits underneath a tree, some with stone removed or otherwise damaged by wildlife.
<emphasis id="2E9FEAAFED16FF9CFDA8F834F45FB198" bold="true" box="[561,583,2025,2046]" pageId="6" pageNumber="199">E.</emphasis>
Cleaned stones. Photographs: R.G.C. Boon (A, C, D), G. Nichols (B), &amp; E. Douwes (E).
</paragraph>
</caption>
<caption id="48946635ED17FF9DFF11F87CF56DBE45" ID-DOI="http://doi.org/10.5281/zenodo.7702415" ID-Zenodo-Dep="7702415" httpUri="https://zenodo.org/record/7702415/files/figure.png" pageId="7" pageNumber="200" startId="7.[136,229,1953,1975]" targetBox="[136,1450,190,1932]" targetPageId="7">
<paragraph id="1C5436BDED17FF9DFF11F87CF56DBE45" blockId="7.[136,1452,1953,2083]" pageId="7" pageNumber="200">
<emphasis id="2E9FEAAFED17FF9DFF11F87CF6E6B1D0" bold="true" box="[136,254,1953,1975]" pageId="7" pageNumber="200">FIGURE 4.</emphasis>
<taxonomicName id="DBEB4D3EED17FF9DFE9DF87CF7EDB1D0" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[260,501,1953,1974]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="200" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED17FF9DFE9DF87CF7EDB1D0" box="[260,501,1953,1974]" italics="true" pageId="7" pageNumber="200">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
(A–E), with fruit of
<taxonomicName id="DBEB4D3EED17FF9DFD26F87FF53EB1D0" box="[703,806,1953,1975]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="200" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED17FF9DFD26F87FF53EB1D0" box="[703,806,1953,1975]" italics="true" pageId="7" pageNumber="200">P. discolor</emphasis>
</taxonomicName>
(F) and
<taxonomicName id="DBEB4D3EED17FF9DFCE0F87FF5CEB1D0" authorityName="Grubov" authorityYear="1949" baseAuthorityName="Sonder" baseAuthorityYear="1860" box="[889,982,1953,1975]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="200" phylum="Tracheophyta" rank="species" species="zeyheri">
<emphasis id="2E9FEAAFED17FF9DFCE0F87FF5CEB1D0" box="[889,982,1953,1975]" italics="true" pageId="7" pageNumber="200">P. zeyheri</emphasis>
</taxonomicName>
(G).
<emphasis id="2E9FEAAFED17FF9DFB92F87CF23BB1D0" bold="true" box="[1035,1059,1953,1974]" pageId="7" pageNumber="200">A.</emphasis>
Fruiting branchlet (fruit almost mature).
<emphasis id="2E9FEAAFED17FF9DFF11F818F686B1BC" bold="true" box="[136,158,1989,2010]" pageId="7" pageNumber="200">B.</emphasis>
Flower.
<emphasis id="2E9FEAAFED17FF9DFF6AF818F712B1BD" bold="true" box="[243,266,1989,2011]" pageId="7" pageNumber="200">C.</emphasis>
Petal.
<emphasis id="2E9FEAAFED17FF9DFED5F818F77CB1BC" bold="true" box="[332,356,1989,2010]" pageId="7" pageNumber="200">D.</emphasis>
Stamen.
<emphasis id="2E9FEAAFED17FF9DFE24F818F7EAB1BD" bold="true" box="[445,498,1989,2011]" pageId="7" pageNumber="200">E–G.</emphasis>
Comparison of the fruit (bottom) of the three species of
<taxonomicName id="DBEB4D3EED17FF9DFB8AF818F296B1BC" authorityName="Herzog" authorityYear="1903" baseAuthorityName="Weberbauer" baseAuthorityYear="1895" box="[1043,1166,1989,2010]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="200" phylum="Tracheophyta" rank="genus">
<emphasis id="2E9FEAAFED17FF9DFB8AF818F296B1BC" box="[1043,1166,1989,2010]" italics="true" pageId="7" pageNumber="200">Phyllogeiton</emphasis>
</taxonomicName>
, with corresponding enlarged persistent receptacle and fruit stalk (top). Scale bar = 10 mm (A), 1 mm (B–D), or 4 mm (E–G). Drawings based on photographs and liquid-preserved fresh material deposited in Herb. PRU. Artist: Daleen Roodt.
</paragraph>
</caption>
<caption id="48946635ED18FF92FF11FADEF268B306" ID-DOI="http://doi.org/10.5281/zenodo.7690765" ID-Zenodo-Dep="7690765" httpUri="https://zenodo.org/record/7690765/files/figure.png" pageId="8" pageNumber="201" startId="8.[136,229,1283,1305]" targetBox="[151,1436,190,1258]" targetPageId="8">
<paragraph id="1C5436BDED18FF92FF11FADEF268B306" blockId="8.[136,1452,1283,1377]" pageId="8" pageNumber="201">
<emphasis id="2E9FEAAFED18FF92FF11FADEF6E5B37E" bold="true" box="[136,253,1283,1305]" pageId="8" pageNumber="201">FIGURE 5.</emphasis>
Topographical map showing the known distribution (black dots) of
<taxonomicName id="DBEB4D3EED18FF92FC1FFADEF26DB37E" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[902,1141,1283,1304]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="201" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED18FF92FC1FFADEF26DB37E" box="[902,1141,1283,1304]" italics="true" pageId="8" pageNumber="201">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
in northeastern KwaZulu-Natal, South Africa. Map based on herbarium collections in BNRH, K, NH, NU, PCE, PRE, PRU, and UDW. The insert shows a map of southern Africa with names of countries; the grey rectangle indicates the area depicted by the topographical map.
</paragraph>
</caption>
<paragraph id="1C5436BDED18FF92FF24FA52F50DB073" blockId="8.[136,1452,1423,2062]" pageId="8" pageNumber="201">
African Elephants,
<taxonomicName id="DBEB4D3EED18FF92FE0DFA52F597B3CC" authority="Blumenbach (1797: 125)" authorityName="Blumenbach" authorityPageNumber="125" authorityYear="1797" box="[404,911,1423,1450]" class="Mammalia" family="Elephantidae" genus="Loxodonta" kingdom="Animalia" order="Proboscidea" pageId="8" pageNumber="201" phylum="Chordata" rank="species" species="africana">
<emphasis id="2E9FEAAFED18FF92FE0DFA52F469B3CF" box="[404,625,1423,1449]" italics="true" pageId="8" pageNumber="201">Loxodonta africana</emphasis>
<bibRefCitation id="787A4B4CED18FF92FDEEFA52F597B3CC" author="Blumenbach, J. F." box="[631,911,1423,1450]" pageId="8" pageNumber="201" refId="ref8833" refString="Blumenbach, J. F. (1797) Handbuch der Naturgeschichte, 5 th ed. Johan Christian Dieterich, Gottingen, 749 pp. https: // doi. org / 10.5962 / bhl. title. 79937" type="book" year="1797">Blumenbach (1797: 125)</bibRefCitation>
</taxonomicName>
, damage trees and this has killed at least one tree at Tembe Elephant Park (C. Hanekom, pers. comm.). Nyala antelope,
<taxonomicName id="DBEB4D3EED18FF92FC01FA6EF354B3AB" authority="Angas (1849: 89)" authorityName="Angas" authorityPageNumber="89" authorityYear="1849" box="[920,1356,1459,1485]" class="Mammalia" family="Bovidae" genus="Tragelaphus" kingdom="Animalia" order="Artiodactyla" pageId="8" pageNumber="201" phylum="Chordata" rank="species" species="angasii">
<emphasis id="2E9FEAAFED18FF92FC01FA6EF265B3AB" box="[920,1149,1459,1485]" italics="true" pageId="8" pageNumber="201">Tragelaphus angasii</emphasis>
<bibRefCitation id="787A4B4CED18FF92FB1CFA6EF354B3AB" author="Angas, G. F." box="[1157,1356,1459,1485]" pageId="8" pageNumber="201" pagination="89 - 90" refId="ref8703" refString="Angas, G. F. (1849) Description of Tragelaphus angasii, Gray, with some account of its habits. Proceedings of the Zoological Society of London 1848: 89 - 90. [https: // www. biodiversitylibrary. org / item / 46216 # page / 125 / mode / 1 up]" type="journal article" year="1849">Angas (1849: 89)</bibRefCitation>
</taxonomicName>
, quickly removed fresh leaves from a cut branch at Bonamanzi Game Reserve (D. Bishop, pers. comm.) and are therefore suspected to browse on low-growing plants of the species.
</paragraph>
<paragraph id="1C5436BDED18FF92FF24F9FDF433B0C0" blockId="8.[136,1452,1423,2062]" pageId="8" pageNumber="201">No young plants are known in the wild, but the species is, despite the hard endocarp of the stone, easy to grow without any special treatment besides removal of the fleshy part of the pericarp (G. Nichols, pers. comm.). Possible reasons for the apparent lack of recruits are herbivory, disease or that conditions suitable for germination and establishment on sandy soils are rare.</paragraph>
<paragraph id="1C5436BDED18FF92FF24F972F51FB08B" blockId="8.[136,1452,1423,2062]" pageId="8" pageNumber="201">
The tree is an occasional host to the epiphytic orchids
<taxonomicName id="DBEB4D3EED18FF92FC8DF972F265B0AF" authority="Lindley (1844" authorityName="Lindley" authorityYear="1844" box="[788,1149,1711,1737]" class="Liliopsida" family="Orchidaceae" genus="Ansellia" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="201" phylum="Tracheophyta" rank="species" species="africana">
<emphasis id="2E9FEAAFED18FF92FC8DF972F5CDB0AF" box="[788,981,1711,1737]" italics="true" pageId="8" pageNumber="201">Ansellia africana</emphasis>
<bibRefCitation id="787A4B4CED18FF92FC42F972F265B0AF" author="Lindley, J." box="[987,1149,1711,1737]" pageId="8" pageNumber="201" pagination="12" refId="ref10074" refString="Lindley, J. (1844) Ansellia africana. Edwards's Botanical Register 30: sub t. 12. [https: // www. biodiversitylibrary. org / item / 9067 # page / 48 / mode / 1 up]" type="journal article" year="1844">Lindley (1844</bibRefCitation>
</taxonomicName>
: sub t. 12) and
<taxonomicName id="DBEB4D3EED18FF92FABBF972F51BB08B" authority="(Jacquin 1760: 30) Garay &amp; Sweet (1974: 206)" authorityName="Garay &amp; Sweet" authorityPageNumber="206" authorityYear="1974" baseAuthorityName="Jacquin" baseAuthorityPageNumber="30" baseAuthorityYear="1760" class="Liliopsida" family="Orchidaceae" genus="Polystachya" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="201" phylum="Tracheophyta" rank="species" species="concreta">
<emphasis id="2E9FEAAFED18FF92FABBF972F6F1B08B" italics="true" pageId="8" pageNumber="201">Polystachya concreta</emphasis>
(
<bibRefCitation id="787A4B4CED18FF92FF6EF90EF7A2B08B" author="Jacquin, N. J. von" box="[247,442,1747,1773]" pageId="8" pageNumber="201" refId="ref9883" refString="Jacquin, N. J. von (1760) Enumeratio systematica plantarum, quas in insulis Caribaeis vicinaque Americes continente detexit novas, aut jam cognitas emendavit. Theodorum Haak, Leiden, 41 pp. https: // doi. org / 10.5962 / bhl. title. 737" type="book" year="1760">Jacquin 1760: 30</bibRefCitation>
)
<bibRefCitation id="787A4B4CED18FF92FE50F90EF51BB08B" author="Garay, L. J. &amp; Sweet, H. R." box="[457,771,1747,1773]" pageId="8" pageNumber="201" pagination="200 - 210" refId="ref9385" refString="Garay, L. J. &amp; Sweet, H. R. (1974) Terminilogia de orchidearum Jacquinii. Orquideologia 9: 200 - 210." type="journal article" year="1974">Garay &amp; Sweet (1974: 206)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED18FF93FF24F92AF404B7BE" lastPageId="9" lastPageNumber="202" pageId="8" pageNumber="201" type="vernacular_names">
<paragraph id="1C5436BDED18FF92FF24F92AF356B153" blockId="8.[136,1452,1423,2062]" pageId="8" pageNumber="201">
<emphasis id="2E9FEAAFED18FF92FF24F92AF7BCB177" bold="true" box="[189,420,1783,1809]" pageId="8" pageNumber="201">Common names:—</emphasis>
English and Afrikaans names in use include “sand ivory” and
<emphasis id="2E9FEAAFED18FF92FBC7F92AF2D6B177" box="[1118,1230,1783,1809]" italics="true" pageId="8" pageNumber="201">sandivoor</emphasis>
respectively. These are also the names recommended by the Dendrological Society of
<collectingCountry id="64FC762DED18FF92FCF6F8C6F5E6B153" box="[879,1022,1819,1845]" name="South Africa" pageId="8" pageNumber="201">South Africa</collectingCountry>
(
<bibRefCitation id="787A4B4CED18FF92FB95F8C6F35BB153" author="Von Durckheim, H. &amp; Van Wyk, B. &amp; A. E &amp; Van den Berg, E. &amp; Coates Palgrave, M. &amp; Jordaan, M." box="[1036,1347,1819,1845]" pageId="8" pageNumber="201" refId="ref11782" refString="Von Durckheim, H., Van Wyk, B. [A. E.], Van den Berg, E., Coates Palgrave, M. &amp; Jordaan, M. (2014) Saklys van Suider-Afrikaanse inheemse bome / Pocket list of southern African indigenous trees, 5 th ed. Briza Academic Books, Briza Publications, Pretoria, 634 pp." type="book" year="2014">
Von Dürckheim
<emphasis id="2E9FEAAFED18FF92FB5EF8C1F2E7B153" box="[1223,1279,1819,1845]" italics="true" pageId="8" pageNumber="201">et al.</emphasis>
2014
</bibRefCitation>
).
</paragraph>
<paragraph id="1C5436BDED18FF93FF24F8E2F7D2B6BA" blockId="8.[136,1452,1423,2062]" lastBlockId="9.[136,1452,158,2093]" lastPageId="9" lastPageNumber="202" pageId="8" pageNumber="201">
There are several Zulu names recorded for
<taxonomicName id="DBEB4D3EED18FF92FD3FF8E2F59AB13F" box="[678,898,1855,1881]" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="201" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED18FF92FD3FF8E2F59AB13F" box="[678,898,1855,1881]" italics="true" pageId="8" pageNumber="201">Berchemia discolor</emphasis>
</taxonomicName>
in literature pertinent to
<collectingRegion id="DE2FF85FED18FF92FB04F8E2F355B13F" box="[1181,1357,1855,1881]" country="South Africa" name="KwaZulu-Natal" pageId="8" pageNumber="201">KwaZulu-Natal</collectingRegion>
(
<bibRefCitation id="787A4B4CED18FF92FAC5F8E2F6DBB11B" author="Pooley, E. S." pageId="8" pageNumber="201" pagination="467 - 479" refId="ref10588" refString="Pooley, E. S. (1980) Some notes on the utilization of natural resources by the tribal people of Maputaland. In: Bruton, M. N. &amp; Cooper, K. H. (Eds.) Studies on the ecology of Maputaland. Rhodes University, Grahamstown and the Natal Branch of the Wildlife Society of Southern Africa, Durban, pp. 467 - 479." type="book chapter" year="1980">Pooley 1980</bibRefCitation>
,
<bibRefCitation id="787A4B4CED18FF92FF48F8BEF754B11B" author="Moll, E. J." box="[209,332,1891,1917]" pageId="8" pageNumber="201" refId="ref10302" refString="Moll, E. J. (1981) Trees of Natal. Eco-lab Trust Fund, University of Cape Town, Rondebosch, 570 pp." type="book" year="1981">Moll 1981</bibRefCitation>
,
<bibRefCitation id="787A4B4CED18FF92FEC2F8BEF7F5B11B" author="Pooley, E." box="[347,493,1891,1917]" pageId="8" pageNumber="201" refId="ref10495" refString="Pooley, E. [E. S.] (1993) The complete field guide to trees of Natal, Zululand &amp; Transkei. Natal Flora Publications Trust, Durban, 512 pp." type="book" year="1993">Pooley 1993</bibRefCitation>
,
<bibRefCitation id="787A4B4CED18FF92FE62F8BEF4E1B11B" author="Hutchings, A. &amp; Scott, A. H. &amp; Lewis, G. &amp; Cunningham, A. B." box="[507,761,1891,1918]" pageId="8" pageNumber="201" refId="ref9719" refString="Hutchings, A., Scott, A. H., Lewis, G. &amp; Cunningham, A. B. (1996) Zulu medicinal plants an inventory. University of Natal Press, Pietermaritzburg in association with University of Zululand, KwaDlangeswa and National Botanical Institute, Claremont, 450 pp." type="book" year="1996">
Hutchings
<emphasis id="2E9FEAAFED18FF92FDE1F8B9F4B5B11B" box="[632,685,1891,1917]" italics="true" pageId="8" pageNumber="201">et al</emphasis>
. 1996
</bibRefCitation>
). These names are likely to be applicable to
<taxonomicName id="DBEB4D3EED18FF92FA9CF8B9F3BCB11B" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[1285,1444,1891,1917]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="201" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED18FF92FA9CF8B9F3BCB11B" box="[1285,1444,1891,1917]" italics="true" pageId="8" pageNumber="201">P. trachybasis</emphasis>
</taxonomicName>
. The name
<emphasis id="2E9FEAAFED18FF92FE99F85AF79BB1C7" box="[256,387,1927,1953]" italics="true" pageId="8" pageNumber="201">ubalatsheni</emphasis>
or
<emphasis id="2E9FEAAFED18FF92FE32F85AF434B1C7" box="[427,556,1927,1953]" italics="true" pageId="8" pageNumber="201">ubaletsheni</emphasis>
is recorded in all of these publications and means “marks” or “make marks on the stone or rock” (A. Koopman, pers. comm.). Semantic links to the names
<emphasis id="2E9FEAAFED18FF92FC72F871F225B1A3" box="[1003,1085,1964,1989]" italics="true" pageId="8" pageNumber="201">umuma</emphasis>
and
<emphasis id="2E9FEAAFED18FF92FBECF871F2DFB1A3" box="[1141,1223,1964,1989]" italics="true" pageId="8" pageNumber="201">umumu</emphasis>
(
<bibRefCitation id="787A4B4CED18FF92FB4EF876F349B1A3" author="Moll, E. J." box="[1239,1361,1963,1989]" pageId="8" pageNumber="201" refId="ref10302" refString="Moll, E. J. (1981) Trees of Natal. Eco-lab Trust Fund, University of Cape Town, Rondebosch, 570 pp." type="book" year="1981">Moll 1981</bibRefCitation>
,
<bibRefCitation id="787A4B4CED18FF92FAC7F876F6DBB18F" author="Pooley, E." pageId="8" pageNumber="201" refId="ref10495" refString="Pooley, E. [E. S.] (1993) The complete field guide to trees of Natal, Zululand &amp; Transkei. Natal Flora Publications Trust, Durban, 512 pp." type="book" year="1993">Pooley 1993</bibRefCitation>
,
<bibRefCitation id="787A4B4CED18FF92FF57F812F7DAB18F" author="Hutchings, A. &amp; Scott, A. H. &amp; Lewis, G. &amp; Cunningham, A. B." box="[206,450,1999,2026]" pageId="8" pageNumber="201" refId="ref9719" refString="Hutchings, A., Scott, A. H., Lewis, G. &amp; Cunningham, A. B. (1996) Zulu medicinal plants an inventory. University of Natal Press, Pietermaritzburg in association with University of Zululand, KwaDlangeswa and National Botanical Institute, Claremont, 450 pp." type="book" year="1996">
Hutchings
<emphasis id="2E9FEAAFED18FF92FED1F80DF761B18F" box="[328,377,1999,2025]" italics="true" pageId="8" pageNumber="201">et al</emphasis>
. 1996
</bibRefCitation>
),
<emphasis id="2E9FEAAFED18FF92FE4CF812F400B18F" box="[469,536,1999,2025]" italics="true" pageId="8" pageNumber="201">uvuka</emphasis>
(
<bibRefCitation id="787A4B4CED18FF92FDBFF812F4A0B18F" author="Pooley, E." box="[550,696,1999,2025]" pageId="8" pageNumber="201" refId="ref10495" refString="Pooley, E. [E. S.] (1993) The complete field guide to trees of Natal, Zululand &amp; Transkei. Natal Flora Publications Trust, Durban, 512 pp." type="book" year="1993">Pooley 1993</bibRefCitation>
,
<bibRefCitation id="787A4B4CED18FF92FD5BF812F5AFB18F" author="Hutchings, A. &amp; Scott, A. H. &amp; Lewis, G. &amp; Cunningham, A. B." box="[706,951,1999,2026]" pageId="8" pageNumber="201" refId="ref9719" refString="Hutchings, A., Scott, A. H., Lewis, G. &amp; Cunningham, A. B. (1996) Zulu medicinal plants an inventory. University of Natal Press, Pietermaritzburg in association with University of Zululand, KwaDlangeswa and National Botanical Institute, Claremont, 450 pp." type="book" year="1996">
Hutchings
<emphasis id="2E9FEAAFED18FF92FCA5F80DF575B18F" box="[828,877,1999,2025]" italics="true" pageId="8" pageNumber="201">et al</emphasis>
. 1996
</bibRefCitation>
), and
<emphasis id="2E9FEAAFED18FF92FC60F812F261B18F" box="[1017,1145,1999,2025]" italics="true" pageId="8" pageNumber="201">umbenduza</emphasis>
(
<emphasis id="2E9FEAAFED18FF92FB10F812F317B18F" box="[1161,1295,1999,2025]" italics="true" pageId="8" pageNumber="201">Tinley 416B</emphasis>
in Herbs NH, NU, PRE) are not clear (A. Koopman, pers. comm.). A herbarium specimen collected at Lake Sibaya bears the name
<emphasis id="2E9FEAAFED19FF93FF11FF43F717B6DE" box="[136,271,158,184]" italics="true" pageId="9" pageNumber="202">umadlozane</emphasis>
(
<emphasis id="2E9FEAAFED19FF93FE86FF43F7F7B6DE" box="[287,495,158,184]" italics="true" pageId="9" pageNumber="202">
Cunningham
<quantity id="DB139B58ED19FF93FE2EFF43F7F7B6DE" box="[439,495,158,184]" metricMagnitude="1" metricUnit="m" metricValue="5.587999999999999" pageId="9" pageNumber="202" unit="in" value="2200.0">2200</quantity>
</emphasis>
in Herb. NU), which is derived from the word
<emphasis id="2E9FEAAFED19FF93FC64FF43F27CB6DE" box="[1021,1124,158,184]" italics="true" pageId="9" pageNumber="202">amadlozi</emphasis>
and refers to ancestral spirits (A. Koopman, pers. comm.).
</paragraph>
<paragraph id="1C5436BDED19FF93FF24FF3BF22AB72E" blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="202">
The name
<emphasis id="2E9FEAAFED19FF93FEAAFF3BF783B766" box="[307,411,230,256]" italics="true" pageId="9" pageNumber="202">umhungu</emphasis>
is used at Nibela, which is at the northern end of Lake
<collectingCountry id="64FC762DED19FF93FB9DFF3BF271B766" box="[1028,1129,230,256]" name="Saint Lucia" pageId="9" pageNumber="202">St. Lucia</collectingCountry>
near Hluhluwe (G. Linforth, pers. comm.). This name and the similar
<emphasis id="2E9FEAAFED19FF93FDCDFED7F4CAB742" box="[596,722,266,292]" italics="true" pageId="9" pageNumber="202">umhungulo</emphasis>
(
<emphasis id="2E9FEAAFED19FF93FD7BFED6F59DB742" box="[738,901,266,293]" italics="true" pageId="9" pageNumber="202">
Gerstner
<quantity id="DB139B58ED19FF93FCD4FED7F59DB742" box="[845,901,266,292]" metricMagnitude="2" metricUnit="m" metricValue="1.326388" pageId="9" pageNumber="202" unit="in" value="5222.0">5222</quantity>
</emphasis>
in Herb. PRE) and
<emphasis id="2E9FEAAFED19FF93FBFAFED7F2F1B742" box="[1123,1257,266,292]" italics="true" pageId="9" pageNumber="202">umhlungulo</emphasis>
(
<bibRefCitation id="787A4B4CED19FF93FB6EFED7F6D9B72E" author="Hutchings, A. &amp; Scott, A. H. &amp; Lewis, G. &amp; Cunningham, A. B." pageId="9" pageNumber="202" refId="ref9719" refString="Hutchings, A., Scott, A. H., Lewis, G. &amp; Cunningham, A. B. (1996) Zulu medicinal plants an inventory. University of Natal Press, Pietermaritzburg in association with University of Zululand, KwaDlangeswa and National Botanical Institute, Claremont, 450 pp." type="book" year="1996">
Hutchings
<emphasis id="2E9FEAAFED19FF93FAEAFED6F3BDB742" box="[1395,1445,266,292]" italics="true" pageId="9" pageNumber="202">et al</emphasis>
. 1996
</bibRefCitation>
) probably mean enticing, alluring, and deceiving (A. Koopman, pers. comm.).
</paragraph>
<paragraph id="1C5436BDED19FF93FF24FE8EF404B7BE" blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="202">
In far northern
<collectingRegion id="DE2FF85FED19FF93FEFDFE8FF40CB70A" box="[356,532,338,364]" country="South Africa" name="KwaZulu-Natal" pageId="9" pageNumber="202">KwaZulu-Natal</collectingRegion>
an informant, M. Tembe, reports that the names
<emphasis id="2E9FEAAFED19FF93FBB2FE8FF2FFB70A" box="[1067,1255,338,364]" italics="true" pageId="9" pageNumber="202">vukakwabafileyo</emphasis>
or
<emphasis id="2E9FEAAFED19FF93FA93FE8FF3B4B70A" box="[1290,1452,338,364]" italics="true" pageId="9" pageNumber="202">vukakwabafile</emphasis>
are used (C. Hanekom, pers. comm.). These names mean “to wake up at the place of those who have died” or “arise from the cemetery”, but A. Koopman (pers. comm.) suggests that their correct
<taxonomicName id="DBEB4D3EED19FF93FC75FE47F241B7D3" box="[1004,1113,410,437]" form="may" pageId="9" pageNumber="202" rank="form">form may</taxonomicName>
be
<emphasis id="2E9FEAAFED19FF93FBE6FE47F315B7D2" box="[1151,1293,410,436]" italics="true" pageId="9" pageNumber="202">vusabafileyo</emphasis>
, which means “wake up the dead or unconscious”.
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED19FF93FF24FE3FF409B3AE" pageId="9" pageNumber="202" type="conservation">
<paragraph id="1C5436BDED19FF93FF24FE3FF598B4EA" blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="202">
<emphasis id="2E9FEAAFED19FF93FF24FE3FF7ACB79A" bold="true" box="[189,436,482,508]" pageId="9" pageNumber="202">Conservation status:</emphasis>
—The EOO of
<taxonomicName id="DBEB4D3EED19FF93FDF8FE3EF4E6B79A" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[609,766,482,508]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="9" pageNumber="202" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED19FF93FDF8FE3EF4E6B79A" box="[609,766,482,508]" italics="true" pageId="9" pageNumber="202">P. trachybasis</emphasis>
</taxonomicName>
was estimated at
<quantity id="DB139B58ED19FF93FC5CFE3FF230B79A" box="[965,1064,482,509]" metricMagnitude="6" metricUnit="m" metricValue="5.951" pageId="9" pageNumber="202" unit="km" value="5951.0">5951 km</quantity>
<superScript id="EB9E9BF5ED19FF93FBBEFE3FF237B796" attach="left" box="[1063,1071,482,496]" fontSize="6" pageId="9" pageNumber="202">2</superScript>
. The AOO calculated was
<quantity id="DB139B58ED19FF93FAC5FE3FF3BBB79A" box="[1372,1443,482,508]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="202" unit="km" value="60.0">60 km</quantity>
<superScript id="EB9E9BF5ED19FF93FA3AFE3FF3B3B796" attach="left" box="[1443,1451,482,496]" fontSize="6" pageId="9" pageNumber="202">2</superScript>
for the
<quantity id="DB139B58ED19FF93FF41FDDBF708B446" box="[216,272,518,544]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="202" unit="km" value="2.0">2 km</quantity>
cell width recommended by the
<bibRefCitation id="787A4B4CED19FF93FDE5FDDBF2B9B446" author="IUCN Standards &amp; Petitions Committee" box="[636,1185,518,544]" pageId="9" pageNumber="202" refId="ref9830" refString="IUCN Standards and Petitions Committee (2019) Guidelines for using the IUCN Red List categories and criteria, version 14. Prepared by the Standards and Petitions Committee. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 23 August 2021)" type="book" year="2019">IUCN Standards and Petitions Committee (2019)</bibRefCitation>
and
<quantity id="DB139B58ED19FF93FB4FFDDBF333B446" box="[1238,1323,518,545]" metricMagnitude="5" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="202" unit="km" value="350.0">350 km</quantity>
<superScript id="EB9E9BF5ED19FF93FAB3FDDBF32AB472" attach="left" box="[1322,1330,518,532]" fontSize="6" pageId="9" pageNumber="202">2</superScript>
for a
<quantity id="DB139B58ED19FF93FAEAFDDBF3B4B446" box="[1395,1452,518,545]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="202" unit="km" value="5.0">5 km</quantity>
cell width. The actual AOO probably lies at the lower end of this range, because even the
<quantity id="DB139B58ED19FF93FB12FDF7F2DDB422" box="[1163,1221,554,580]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="202" unit="km" value="2.0">2 km</quantity>
<superScript id="EB9E9BF5ED19FF93FB5CFDF7F2D5B45E" attach="left" box="[1221,1229,554,568]" fontSize="6" pageId="9" pageNumber="202">2</superScript>
cell width includes aquatic environments and unsuitable, degraded or transformed terrestrial habitats. The distinct bark of
<taxonomicName id="DBEB4D3EED19FF93FA97FD92F3B3B40E" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[1294,1451,590,616]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="9" pageNumber="202" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED19FF93FA97FD92F3B3B40E" box="[1294,1451,590,616]" italics="true" pageId="9" pageNumber="202">P. trachybasis</emphasis>
</taxonomicName>
also means that accessible specimens are unlikely to be overlooked.
</paragraph>
<paragraph id="1C5436BDED19FF93FF24FD4BF2ABB49E" blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="202">
The species occurs in 10 severely fragmented populations as defined by the
<bibRefCitation id="787A4B4CED19FF93FC6EFD4BF296B4D6" author="IUCN" box="[1015,1166,662,688]" pageId="9" pageNumber="202" refId="ref9775" refString="IUCN (2012) IUCN Red List categories and criteria, version 3.1. IUCN Species Survival Commission, Gland, Switzerland and Cambridge, UK, iv + 32 pp. Available from: https: // portals. iucn. org / library / node / 10315 (accessed 23 August 2021)" type="book" year="2012">IUCN (2012)</bibRefCitation>
.
<taxonomicName id="DBEB4D3EED19FF93FB0FFD4BF3B3B4D6" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[1174,1451,662,688]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="9" pageNumber="202" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED19FF93FB0FFD4BF3B3B4D6" box="[1174,1451,662,688]" italics="true" pageId="9" pageNumber="202">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
may occur in patches too small to support a viable population in the long-term and sub-populations are separated from other suitable habitat by large distances, thus genetic or demographic exchange seems unlikely.
</paragraph>
<paragraph id="1C5436BDED19FF93FF24FCDFF75BB27E" blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="202">
There are 70 known mature individuals in the six sub-populations surveyed during field work, namely at Bonamanzi Game Reserve (35 trees), Tembe Elephant Park and surrounds (15 trees), Hluhluwe-iMfolozi Park (8 trees), Nibela Peninsula (8 trees), False Bay Park (2 trees), and Phinda Private Game Reserve (2 trees). At least eight of these trees are in poor condition and appear to be dying and no young plants (&lt;10 years old) are known. The smallest (and presumably youngest) known plant is at Bonamanzi Game Reserve and is
<quantity id="DB139B58ED19FF93FC54FC4FF5E1B5CB" box="[973,1017,914,941]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="202" unit="m" value="4.0">4 m</quantity>
tall and has a dbh of
<quantity id="DB139B58ED19FF93FB72FC4FF326B5CB" box="[1259,1342,914,941]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="202" unit="mm" value="70.0">70 mm</quantity>
. The subpopulation sizes at Ndlozi Peninsula, Lake Bhangazi South, Sodwana Bay, and Ndumo Game Reserve are unknown. Even if known trees represent only about 10% of the species’ total population, there are likely to be fewer than 700 trees in the wild.
</paragraph>
<paragraph id="1C5436BDED19FF93FF24FBFFF4A7B372" blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="202">
<taxonomicName id="DBEB4D3EED19FF93FF24FBFFF7CCB25A" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[189,468,1058,1084]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="9" pageNumber="202" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED19FF93FF24FBFFF7CCB25A" box="[189,468,1058,1084]" italics="true" pageId="9" pageNumber="202">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
appears to be almost entirely confined to protected areas. Bark of the few trees growing outside of these areas is heavily used for medicinal purposes, the removal of which kills individuals or limits growth (
<emphasis id="2E9FEAAFED19FF93FF08FBB7F77CB2E2" box="[145,356,1130,1156]" italics="true" pageId="9" pageNumber="202">
Cunningham
<quantity id="DB139B58ED19FF93FEB5FBB7F77CB2E2" box="[300,356,1130,1156]" metricMagnitude="1" metricUnit="m" metricValue="5.587999999999999" pageId="9" pageNumber="202" unit="in" value="2200.0">2200</quantity>
</emphasis>
in Herb. NU; L. Loffler, G. Linforth, C. Hanekom, pers. comm.). Even within some protected areas trees may not be safe from harvesting (
<bibRefCitation id="787A4B4CED19FF93FD10FB53F54FB2CE" author="Groenewald, Y." box="[649,855,1166,1192]" pageId="9" pageNumber="202" refId="ref9413" refString="Groenewald, Y. (2010) Ndumo reserve hit by invasion, crime. Mail and Guardian. Available from: https: // mg. co. za / article / 2010 - 10 - 15 - ndumo-reserve-hit-by-invasion-crime / (accessed 23 August 2021)" type="url" year="2010">Groenewald 2010</bibRefCitation>
) or they may be damaged or killed by elephants (C. Hanekom, pers. comm.). It is probable that the current scarcity of trees resulted, at least in part, from the excessive stripping of bark for use in traditional medicine (locally referred to by the generic term
<emphasis id="2E9FEAAFED19FF93FBE9FB0BF2A6B296" box="[1136,1214,1238,1264]" italics="true" pageId="9" pageNumber="202">umuthi</emphasis>
, also spelled
<emphasis id="2E9FEAAFED19FF93FAC3FB0AF394B296" box="[1370,1420,1239,1264]" italics="true" pageId="9" pageNumber="202">muti</emphasis>
or
<emphasis id="2E9FEAAFED19FF93FF11FB27F6D0B372" box="[136,200,1274,1300]" italics="true" pageId="9" pageNumber="202">muthi</emphasis>
) over many years (see under “Uses” below).
</paragraph>
<paragraph id="1C5436BDED19FF93FF24FAC3F409B3AE" blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="202">
Taking into account that the overall number of mature individuals is small, the known sub-populations are very small and the population is declining due to natural mortality, harvesting and lack of recruitment, application of the IUCN Red List Categories and Criteria (
<bibRefCitation id="787A4B4CED19FF93FDD0FABBF4D7B3E6" author="IUCN" box="[585,719,1382,1408]" pageId="9" pageNumber="202" refId="ref9775" refString="IUCN (2012) IUCN Red List categories and criteria, version 3.1. IUCN Species Survival Commission, Gland, Switzerland and Cambridge, UK, iv + 32 pp. Available from: https: // portals. iucn. org / library / node / 10315 (accessed 23 August 2021)" type="book" year="2012">IUCN 2012</bibRefCitation>
) suggest that the species should be classified as Endangered (EN) B2(v); C2a(i); D. There is an urgent need to survey populations more fully and to try and establish and address the reasons for the lack of recruitment.
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED19FF93FF24FA0FF54DB182" pageId="9" pageNumber="202" type="description">
<paragraph id="1C5436BDED19FF93FF24FA0FF595B11E" blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="202">
<emphasis id="2E9FEAAFED19FF93FF24FA0FF702B38A" bold="true" box="[189,282,1490,1516]" pageId="9" pageNumber="202">Uses:—</emphasis>
On the oldest known herbarium specimens of the species (
<emphasis id="2E9FEAAFED19FF93FC5BFA0EF270B38A" box="[962,1128,1490,1517]" italics="true" pageId="9" pageNumber="202">
Gerstner
<quantity id="DB139B58ED19FF93FBA9FA0FF270B38A" box="[1072,1128,1490,1516]" metricMagnitude="2" metricUnit="m" metricValue="1.326388" pageId="9" pageNumber="202" unit="in" value="5222.0">5222</quantity>
</emphasis>
in Herb. PRE), collected at False Bay Park in 1944, it is recorded that a tree had 90% of its bark removed by traditional healers. The label of another specimen collected in
<quantity id="DB139B58ED19FF93FE40F9C7F435B052" box="[473,557,1562,1588]" metricMagnitude="1" metricUnit="m" metricValue="5.06222" pageId="9" pageNumber="202" unit="in" value="1993.0">1993 in</quantity>
the corridor between Ndumu Game Reserve and Tembe Elephant Park (
<emphasis id="2E9FEAAFED19FF93FAD7F9C6F6FBB03E" italics="true" pageId="9" pageNumber="202">
Van Wyk BSA
<quantity id="DB139B58ED19FF93FF20F9E3F6FBB03E" box="[185,227,1598,1624]" metricMagnitude="1" metricUnit="m" metricValue="1.89992" pageId="9" pageNumber="202" unit="in" value="748.0">748</quantity>
</emphasis>
in Herbs PRE and PRU) reads: “This is a ‘muti’ tree. The old trunk (about
<quantity id="DB139B58ED19FF93FBAFF9E3F279B03F" box="[1078,1121,1598,1625]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="202" unit="m" value="1.0">1 m</quantity>
high) is regularly stripped of its bark—in patches”. A collection from dune forest at Lake Sibaya (
<emphasis id="2E9FEAAFED19FF93FC1DF9BFF24CB01A" box="[900,1108,1634,1660]" italics="true" pageId="9" pageNumber="202">
Cunningham
<quantity id="DB139B58ED19FF93FB85F9BFF24CB01A" box="[1052,1108,1634,1660]" metricMagnitude="1" metricUnit="m" metricValue="5.587999999999999" pageId="9" pageNumber="202" unit="in" value="2200.0">2200</quantity>
</emphasis>
in Herb. NU) also records that the bark is heavily utilised for medicinal purposes. Trees which may belong to this species observed at Siteki,
<collectingCountry id="64FC762DED19FF93FAD0F95BF3B3B0C6" box="[1353,1451,1670,1696]" name="Swaziland" pageId="9" pageNumber="202">Eswatini</collectingCountry>
(see under “Distribution” above) were debarked. Bark and the roots are heavily utilised east of Tembe Elephant Park and according to a local resident are mixed with material from other species to treat chest ailments (C. Hanekom, pers. comm.). There are few surviving trees in the community area surrounding Nibela at Lake
<collectingCountry id="64FC762DED19FF93FBEDF92FF2C2B16A" box="[1140,1242,1778,1804]" name="Saint Lucia" pageId="9" pageNumber="202">St. Lucia</collectingCountry>
because they have all been debarked and the majority have already died (G. Linforth, pers. comm.). At Nibela evil spirits are believed to be extracted by boiling and steaming bark, and fruit is used to treat impotence (G. Linforth, pers. comm.). At nearby False Bay Park, bark is used as a love charm (
<bibRefCitation id="787A4B4CED19FF93FD13F883F599B11E" author="Hutchings, A. &amp; Scott, A. H. &amp; Lewis, G. &amp; Cunningham, A. B." box="[650,897,1886,1913]" pageId="9" pageNumber="202" refId="ref9719" refString="Hutchings, A., Scott, A. H., Lewis, G. &amp; Cunningham, A. B. (1996) Zulu medicinal plants an inventory. University of Natal Press, Pietermaritzburg in association with University of Zululand, KwaDlangeswa and National Botanical Institute, Claremont, 450 pp." type="book" year="1996">
Hutchings
<emphasis id="2E9FEAAFED19FF93FC9FF882F52FB11E" box="[774,823,1886,1912]" italics="true" pageId="9" pageNumber="202">et al</emphasis>
. 1996
</bibRefCitation>
).
</paragraph>
<paragraph id="1C5436BDED19FF93FF24F85FF54DB182" blockId="9.[136,1452,158,2093]" pageId="9" pageNumber="202">
There are no records of the fruit being eaten by humans, which is surprising as the fruits of other
<taxonomicName id="DBEB4D3EED19FF93FA85F85FF6C1B1A6" pageId="9" pageNumber="202">
<emphasis id="2E9FEAAFED19FF93FA85F85FF3B3B1FA" box="[1308,1451,1922,1948]" italics="true" pageId="9" pageNumber="202">Phyllogeiton</emphasis>
species
</taxonomicName>
are popular (e.g.
<bibRefCitation id="787A4B4CED19FF93FE3CF87BF42EB1A6" author="Pooley, E. S." box="[421,566,1958,1984]" pageId="9" pageNumber="202" pagination="467 - 479" refId="ref10588" refString="Pooley, E. S. (1980) Some notes on the utilization of natural resources by the tribal people of Maputaland. In: Bruton, M. N. &amp; Cooper, K. H. (Eds.) Studies on the ecology of Maputaland. Rhodes University, Grahamstown and the Natal Branch of the Wildlife Society of Southern Africa, Durban, pp. 467 - 479." type="book chapter" year="1980">Pooley 1980</bibRefCitation>
,
<bibRefCitation id="787A4B4CED19FF93FDDBF87AF577B1A6" author="Van Wyk, B. - E. &amp; Gericke, N." box="[578,879,1958,1985]" pageId="9" pageNumber="202" refId="ref11667" refString="Van Wyk, B. - E. &amp; Gericke, N. (2000) People's plants - a guide to useful plants of southern Africa. Briza Publications, Pretoria, 351 pp." type="book" year="2000">Van Wyk &amp; Gericke 2000</bibRefCitation>
), with fruit of
<taxonomicName id="DBEB4D3EED19FF93FB8EF87AF288B1A6" box="[1047,1168,1958,1984]" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="9" pageNumber="202" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED19FF93FB8EF87AF288B1A6" box="[1047,1168,1958,1984]" italics="true" pageId="9" pageNumber="202">P. discolor</emphasis>
</taxonomicName>
even having potential in terms of commercialisation and domestication (
<bibRefCitation id="787A4B4CED19FF93FD05F817F550B182" author="Lusepani, N. E." box="[668,840,1994,2020]" pageId="9" pageNumber="202" refId="ref10159" refString="Lusepani, N. E. (1999) Reproductive biology and utilisation of Berchemia discolor (Klotzsch) Hemsley (Rhamnaceae). Unpublished MSc thesis, University of Stellenbosch, Stellenbosch, 109 pp." type="book" year="1999">Lusepani 1999</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED19FF90FF24F833F33DB206" lastPageId="10" lastPageNumber="203" pageId="9" pageNumber="202" type="materials_examined">
<paragraph id="1C5436BDED19FF90FF24F833F33DB206" blockId="9.[136,1452,158,2093]" lastBlockId="10.[136,1452,158,2093]" lastPageId="10" lastPageNumber="203" pageId="9" pageNumber="202">
<materialsCitation id="AC833CE0ED19FF93FF24F833F461BE4A" ID-GBIF-Occurrence="4041537310" collectingDate="1956-05-09" collectionCode="NU" collectorName="Hancock" country="South Africa" lastPageId="9" lastPageNumber="203" location="Ndumo" municipality="Ndumu" pageId="9" pageNumber="202" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<emphasis id="2E9FEAAFED19FF93FF24F833F479BE6E" bold="true" box="[189,609,2030,2056]" pageId="9" pageNumber="202">
Additional collections (
<typeStatus id="C350881FED19FF93FE4EF833F44CBE6E" box="[471,596,2030,2056]" pageId="9" pageNumber="202" type="paratype">paratypes</typeStatus>
):
</emphasis>
—
<collectingCountry id="64FC762DED19FF93FDE7F833F556BE6E" box="[638,846,2030,2056]" name="South Africa" pageId="9" pageNumber="202">SOUTH AFRICA</collectingCountry>
.
<emphasis id="2E9FEAAFED19FF93FCC5F833F23ABE6E" bold="true" box="[860,1058,2030,2056]" pageId="9" pageNumber="202">
<collectingRegion id="DE2FF85FED19FF93FCC5F833F205BE6E" box="[860,1053,2030,2056]" country="South Africa" name="KwaZulu-Natal" pageId="9" pageNumber="202">KwaZulu-Natal</collectingRegion>
:
</emphasis>
<collectingMunicipality id="FC30ACC7ED19FF93FBB5F833F298BE6E" box="[1068,1152,2030,2056]" pageId="9" pageNumber="202">Ndumu</collectingMunicipality>
[
<location id="19346066ED19FF93FB0BF833F2F4BE6E" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED19FF93FB0BF833F2F4BE6E" box="[1170,1260,2030,2056]" country="South Africa" municipality="Ndumu" name="Ndumo" pageId="9" pageNumber="202" stateProvince="KwaZulu-Natal">Ndumo</location>
]
<location id="19346066ED19FF93FB64F833F3BFBE6F" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED19FF93FB64F833F3BFBE6F" box="[1277,1447,2030,2057]" country="South Africa" municipality="Ndumu" name="Game Reserve" pageId="9" pageNumber="202" stateProvince="KwaZulu-Natal">Game Reserve</location>
, (2632CD),
<date id="6855107DED19FF93FE93F7CFF782BE4B" box="[266,410,2066,2093]" pageId="9" pageNumber="202" value="1956-05-09">
<collectingDate id="7811E995ED19FF93FE93F7CFF782BE4B" box="[266,410,2066,2093]" pageId="9" pageNumber="202" value="1956-05-09">9 May 1956</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED19FF93FE31F7CFF43BBE4A" box="[424,547,2066,2092]" italics="true" pageId="9" pageNumber="202">
<collectorName id="B11E536BED19FF93FE31F7CFF414BE4A" box="[424,524,2066,2092]" pageId="9" pageNumber="202">Hancock</collectorName>
9
</emphasis>
(
<collectionCode id="7AFAAE78ED19FF93FDACF7CFF47FBE4A" box="[565,615,2066,2092]" pageId="9" pageNumber="202">NU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED19FF90FD1BF7CFF7B4B6DE" ID-GBIF-Occurrence="4041537313" collectingDate="1959-02-23" collectionCode="NH, NU, PRE" collectorName="Tinley" lastPageId="10" lastPageNumber="203" location="Ndumo" municipality="Ndumu" pageId="9" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED19FF93FD1BF7CFF4CEBE4A" box="[642,726,2066,2092]" pageId="9" pageNumber="202">Ndumu</collectingMunicipality>
[
<location id="19346066ED19FF93FD7FF7CFF558BE4A" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED19FF93FD7FF7CFF558BE4A" box="[742,832,2066,2092]" municipality="Ndumu" name="Ndumo" pageId="9" pageNumber="202">Ndumo</location>
]
<location id="19346066ED19FF93FCC9F7CFF5E2BE4B" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED19FF93FCC9F7CFF5E2BE4B" box="[848,1018,2066,2093]" municipality="Ndumu" name="Game Reserve" pageId="9" pageNumber="202">Game Reserve</location>
, (2632CD),
<date id="6855107DED19FF93FB13F7CFF341BE4B" box="[1162,1369,2066,2093]" pageId="9" pageNumber="202" value="1959-02-23">
<collectingDate id="7811E995ED19FF93FB13F7CFF341BE4B" box="[1162,1369,2066,2093]" pageId="9" pageNumber="202" value="1959-02-23">23 February 1959</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED19FF90FAFEF7CFF6DCB6DE" italics="true" lastPageId="10" lastPageNumber="203" pageId="9" pageNumber="202">
<collectorName id="B11E536BED19FF93FAFEF7CFF3B3BE4A" box="[1383,1451,2066,2092]" pageId="9" pageNumber="202">Tinley</collectorName>
416B
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FF4FFF43F71FB6DE" box="[214,263,158,184]" pageId="10" pageNumber="203">NH</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FE83FF43F752B6DE" box="[282,330,158,184]" pageId="10" pageNumber="203">NU</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FEC4FF43F782B6DE" box="[349,410,158,184]" pageId="10" pageNumber="203">PRE</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FE2FFF43F3B4B6DE" ID-GBIF-Occurrence="4041537323" box="[438,1452,158,185]" collectingDate="1968-02-28" collectionCode="PRE" collectorName="Ross &amp; Moll" lastPageId="10" lastPageNumber="203" location="Drift" municipality="Drift" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<locationDeviation id="0DE62F9FED1AFF90FE2FFF43F45EB6DE" bearing="180.0" box="[438,582,158,184]" distance="4828.032000000001" pageId="10" pageNumber="203">
<quantity id="DB139B58ED1AFF90FE2FFF43F413B6DE" box="[438,523,158,184]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="10" pageNumber="203" unit="mi" value="3.0">3 miles</quantity>
S of
</locationDeviation>
Makane’s
<collectingMunicipality id="FC30ACC7ED1AFF90FD51FF43F519B6DE" box="[712,769,158,184]" pageId="10" pageNumber="203">Drift</collectingMunicipality>
, (2632CD),
<date id="6855107DED1AFF90FC0AFF43F27CB6DE" box="[915,1124,158,184]" pageId="10" pageNumber="203" value="1968-02-28">
<collectingDate id="7811E995ED1AFF90FC0AFF43F27CB6DE" box="[915,1124,158,184]" pageId="10" pageNumber="203" value="1968-02-28">28 February 1968</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FBEAFF42F352B6DE" box="[1139,1354,158,185]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FBEAFF42F31FB6DE" box="[1139,1287,158,185]" pageId="10" pageNumber="203">Ross &amp; Moll</collectorName>
1820
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FAC4FF43F383B6DE" box="[1373,1435,158,184]" pageId="10" pageNumber="203">PRE</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FF11FF1EF756B766" ID-GBIF-Occurrence="4041537319" collectingDate="1964-02-08" collectionCode="NH, NU" collectorName="Tinley" lastPageId="10" lastPageNumber="203" location="Ndumo" municipality="Ndumu" pageId="10" pageNumber="202" specimenCount="1" stateProvince="Ingwavuma District" typeStatus="paratype">
<collectingRegion id="DE2FF85FED1AFF90FF11FF1EF775B6BA" box="[136,365,194,221]" pageId="10" pageNumber="203">Ingwavuma District</collectingRegion>
,
<collectingMunicipality id="FC30ACC7ED1AFF90FEE5FF1FF7C8B6BA" box="[380,464,194,220]" pageId="10" pageNumber="203">Ndumu</collectingMunicipality>
[
<location id="19346066ED1AFF90FE79FF1FF422B6BA" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FE79FF1FF422B6BA" box="[480,570,194,220]" municipality="Ndumu" name="Ndumo" pageId="10" pageNumber="203" stateProvince="Ingwavuma District">Ndumo</location>
]
<location id="19346066ED1AFF90FDD2FF1FF4EDB6BB" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FDD2FF1FF4EDB6BB" box="[587,757,194,221]" municipality="Ndumu" name="Game Reserve" pageId="10" pageNumber="203" stateProvince="Ingwavuma District">Game Reserve</location>
,
<location id="19346066ED1AFF90FC9AFF1FF5B9B6BB" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FC9AFF1FF5B9B6BB" box="[771,929,194,221]" municipality="Ndumu" name="Matini Forest" pageId="10" pageNumber="203" stateProvince="Ingwavuma District">Matini Forest</location>
, E. area, (2732AA),
<date id="6855107DED1AFF90FB0FFF1FF341B6BA" box="[1174,1369,194,220]" pageId="10" pageNumber="203" value="1964-02-08">
<collectingDate id="7811E995ED1AFF90FB0FFF1FF341B6BA" box="[1174,1369,194,220]" pageId="10" pageNumber="203" value="1964-02-08">8 February 1964</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FAFEFF1FF6AAB766" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FAFEFF1FF3B3B6BA" box="[1383,1451,194,220]" pageId="10" pageNumber="203">Tinley</collectorName>
943
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FF5FFF3BF6EFB766" box="[198,247,230,256]" pageId="10" pageNumber="203">NH</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FE92FF3BF724B766" box="[267,316,230,256]" pageId="10" pageNumber="203">NU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FEC0FF3BF3B4B766" ID-GBIF-Occurrence="4041537335" box="[345,1452,230,257]" collectingDate="1968-02-26" collectionCode="PRE" collectorName="Venter" lastPageId="10" lastPageNumber="203" location="Makani's" municipality="Makatini Flats" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FEC0FF3BF41BB766" box="[345,515,230,256]" pageId="10" pageNumber="203">Makatini Flats</collectingMunicipality>
,
<location id="19346066ED1AFF90FD8BFF3BF462B766" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FD8BFF3BF462B766" box="[530,634,230,256]" municipality="Makatini Flats" name="Makani's" pageId="10" pageNumber="203">Makani’s</location>
[Makane’s] Pont, (2732AA),
<date id="6855107DED1AFF90FC45FF3BF2B7B766" box="[988,1199,230,256]" pageId="10" pageNumber="203" value="1968-02-26">
<collectingDate id="7811E995ED1AFF90FC45FF3BF2B7B766" box="[988,1199,230,256]" pageId="10" pageNumber="203" value="1968-02-26">26 February 1968</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FB27FF3AF351B766" box="[1214,1353,230,256]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FB27FF3AF31EB766" box="[1214,1286,231,256]" pageId="10" pageNumber="203">Venter</collectorName>
4614
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FAC4FF3BF383B766" box="[1373,1435,230,256]" pageId="10" pageNumber="203">PRE</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FF11FED7F345B742" ID-GBIF-Occurrence="4041537316" box="[136,1373,266,293]" collectingDate="1993-03-15" collectionCode="PRE, PRU" collectorName="Van Wyk" lastPageId="10" lastPageNumber="203" location="Ndumo" municipality="Ndumu" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
Corridor between
<collectingMunicipality id="FC30ACC7ED1AFF90FECEFED7F7B3B742" box="[343,427,266,292]" pageId="10" pageNumber="203">Ndumu</collectingMunicipality>
[
<location id="19346066ED1AFF90FE20FED7F40BB742" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FE20FED7F40BB742" box="[441,531,266,292]" municipality="Ndumu" name="Ndumo" pageId="10" pageNumber="203">Ndumo</location>
] and
<location id="19346066ED1AFF90FDCBFED7F4B9B742" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FDCBFED7F4B9B742" box="[594,673,266,292]" municipality="Ndumu" name="Tembe" pageId="10" pageNumber="203">Tembe</location>
, (2732AB),
<date id="6855107DED1AFF90FCB7FED7F5C7B742" box="[814,991,266,293]" pageId="10" pageNumber="203" value="1993-03-15">
<collectingDate id="7811E995ED1AFF90FCB7FED7F5C7B742" box="[814,991,266,293]" pageId="10" pageNumber="203" value="1993-03-15">15 March 1993</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FC75FED6F2B7B742" box="[1004,1199,266,292]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FC75FED6F254B742" box="[1004,1100,266,292]" pageId="10" pageNumber="203">Van Wyk</collectorName>
BSA748
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FB26FED7F2E3B742" box="[1215,1275,266,292]" pageId="10" pageNumber="203">PRE</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FA95FED7F354B742" box="[1292,1356,266,292]" pageId="10" pageNumber="203">PRU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FAFCFED7F5C3B72E" ID-GBIF-Occurrence="4041537324" collectingDate="1999-05-16" collectionCode="AA" collectorName="Schmidt" lastPageId="10" lastPageNumber="203" location="Jobe's Kraal" pageId="10" pageNumber="202" specimenCount="1" stateProvince="Jobe's Kraal" typeStatus="paratype">
<collectingRegion id="DE2FF85FED1AFF90FAFCFED7F6D1B72E" pageId="10" pageNumber="203">Jobe’s Kraal</collectingRegion>
, (2732
<collectionCode id="7AFAAE78ED1AFF90FE8EFEF3F75AB72E" box="[279,322,302,328]" pageId="10" pageNumber="203">AA</collectionCode>
),
<date id="6855107DED1AFF90FECCFEF3F7F5B72E" box="[341,493,302,329]" pageId="10" pageNumber="203" value="1999-05-16">
<collectingDate id="7811E995ED1AFF90FECCFEF3F7F5B72E" box="[341,493,302,329]" pageId="10" pageNumber="203" value="1999-05-16">16 May 1999</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FE61FEF3F489B72E" box="[504,657,302,328]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FE61FEF3F44AB72E" box="[504,594,302,328]" pageId="10" pageNumber="203">Schmidt</collectorName>
2406
</emphasis>
(Ernst Schmidt Herbarium!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FC7BFEF3F51EB70A" ID-GBIF-Occurrence="4041537326" collectingDate="1981-11-09" collectionCode="NU" collectorName="Cunningham" lastPageId="10" lastPageNumber="203" location="Sihangwane Nature Conservation Camp" municipality="Sihangwane Nature Conservation Camp" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FC7BFEF3F3BFB72E" box="[994,1447,302,329]" pageId="10" pageNumber="203">Sihangwane Nature Conservation Camp</collectingMunicipality>
, (2732AB),
<date id="6855107DED1AFF90FE93FE8FF7C4B70A" box="[266,476,338,364]" pageId="10" pageNumber="203" value="1981-11-09">
<collectingDate id="7811E995ED1AFF90FE93FE8FF7C4B70A" box="[266,476,338,364]" pageId="10" pageNumber="203" value="1981-11-09">9 November 1981</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FE72FE8FF4B7B70A" box="[491,687,338,364]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FE72FE8FF464B70A" box="[491,636,338,364]" pageId="10" pageNumber="203">Cunningham</collectorName>
518
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FD5BFE8FF4ECB70A" box="[706,756,338,364]" pageId="10" pageNumber="203">NU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FC96FE8FF72CB7F6" ID-GBIF-Occurrence="4041537329" collectingDate="1995-04-23" collectionCode="PRU" collectorName="Van Wyk" lastPageId="10" lastPageNumber="203" location="Tembe Elephant Park" municipality="Tembe Elephant Park" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FC96FE8FF210B70A" box="[783,1032,338,364]" pageId="10" pageNumber="203">Tembe Elephant Park</collectingMunicipality>
, (2732AB),
<date id="6855107DED1AFF90FB01FE8FF324B70B" box="[1176,1340,338,365]" pageId="10" pageNumber="203" value="1995-04-23">
<collectingDate id="7811E995ED1AFF90FB01FE8FF324B70B" box="[1176,1340,338,365]" pageId="10" pageNumber="203" value="1995-04-23">23 April 1995</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FAD3FE8EF6D6B7F6" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FAD3FE8EF3B4B70A" box="[1354,1452,338,364]" pageId="10" pageNumber="203">Van Wyk</collectorName>
12786
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FF7BFEABF73BB7F6" box="[226,291,374,400]" pageId="10" pageNumber="203">PRU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FEA7FEABF6CCB7D2" ID-GBIF-Occurrence="4041537308" collectingDate="1995-10-23" collectionCode="NH" collectorName="Kruger &amp; Symmonds" lastPageId="10" lastPageNumber="203" location="Park" municipality="Tembe Elephant Reserve" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FEA7FEABF446B7F7" box="[318,606,374,401]" pageId="10" pageNumber="203">Tembe Elephant Reserve</collectingMunicipality>
[
<location id="19346066ED1AFF90FDE9FEABF4B3B7F6" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FDE9FEABF4B3B7F6" box="[624,683,374,400]" municipality="Tembe Elephant Reserve" name="Park" pageId="10" pageNumber="203">Park</location>
], (2732AB),
<date id="6855107DED1AFF90FCDFFEABF215B7F7" box="[838,1037,374,401]" pageId="10" pageNumber="203" value="1995-10-23">
<collectingDate id="7811E995ED1AFF90FCDFFEABF215B7F7" box="[838,1037,374,401]" pageId="10" pageNumber="203" value="1995-10-23">23 October 1995</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FB85FEABF3B3B7F6" box="[1052,1451,374,401]" italics="true" pageId="10" pageNumber="203">
<location id="19346066ED1AFF90FB85FEABF26CB7F6" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FB85FEABF26CB7F6" box="[1052,1140,374,400]" municipality="Tembe Elephant Reserve" name="Nichols" pageId="10" pageNumber="203">Nichols</location>
,
<collectorName id="B11E536BED1AFF90FB1DFEAAF361B7F6" box="[1156,1401,374,401]" pageId="10" pageNumber="203">Kruger &amp; Symmonds</collectorName>
s.n.
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FF08FE47F6DBB7D2" box="[145,195,410,436]" pageId="10" pageNumber="203">NH</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FF43FE47F217B7D2" ID-GBIF-Occurrence="4041537334" box="[218,1039,410,436]" collectingDate="1996-02-13" collectionCode="PRU" collectorName="Prozesky" lastPageId="10" lastPageNumber="203" location="Tembe Elephant Park" municipality="Tembe Elephant Park" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FF43FE47F7D4B7D2" box="[218,460,410,436]" pageId="10" pageNumber="203">Tembe Elephant Park</collectingMunicipality>
, (2732AB),
<date id="6855107DED1AFF90FDCCFE47F506B7D2" box="[597,798,410,436]" pageId="10" pageNumber="203" value="1996-02-13">
<collectingDate id="7811E995ED1AFF90FDCCFE47F506B7D2" box="[597,798,410,436]" pageId="10" pageNumber="203" value="1996-02-13">13 February 1996</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FCB1FE47F5B6B7D2" box="[808,942,410,436]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FCB1FE47F594B7D2" box="[808,908,410,436]" pageId="10" pageNumber="203">Prozesky</collectorName>
50
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FC24FE47F5E6B7D2" box="[957,1022,410,436]" pageId="10" pageNumber="203">PRU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FB8DFE47F427B7BE" ID-GBIF-Occurrence="4041537333" collectingDate="1997-10-10" collectionCode="PRU" collectorName="Matthews" lastPageId="10" lastPageNumber="203" location="Tembe Elephant Park" municipality="Tembe Elephant Park" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FB8DFE47F31EB7D2" box="[1044,1286,410,436]" pageId="10" pageNumber="203">Tembe Elephant Park</collectingMunicipality>
, (2732AB),
<date id="6855107DED1AFF90FA16FE47F73DB7BE" pageId="10" pageNumber="203" value="1997-10-10">
<collectingDate id="7811E995ED1AFF90FA16FE47F73DB7BE" pageId="10" pageNumber="203" value="1997-10-10">10 October 1997</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FEA8FE63F7C5B7BE" box="[305,477,446,472]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FEA8FE63F786B7BE" box="[305,414,446,472]" pageId="10" pageNumber="203">Matthews</collectorName>
1304
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FE74FE63F436B7BE" box="[493,558,446,472]" pageId="10" pageNumber="203">PRU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FDDFFE63F7C9B79A" ID-GBIF-Occurrence="4041537317" collectingDate="1999-01-17" collectorName="Schmidt" lastPageId="10" lastPageNumber="203" location="Tembe Elephant Reserve" municipality="Tembe Elephant Reserve" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FDDFFE63F547B7BF" box="[582,863,446,473]" pageId="10" pageNumber="203">Tembe Elephant Reserve</collectingMunicipality>
[Park], (2732AB?),
<date id="6855107DED1AFF90FBD1FE63F31EB7BE" box="[1096,1286,446,473]" pageId="10" pageNumber="203" value="1999-01-17">
<collectingDate id="7811E995ED1AFF90FBD1FE63F31EB7BE" box="[1096,1286,446,473]" pageId="10" pageNumber="203" value="1999-01-17">17 January 1999</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FA88FE63F3B3B7BE" box="[1297,1451,446,472]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FA88FE63F373B7BE" box="[1297,1387,446,472]" pageId="10" pageNumber="203">Schmidt</collectorName>
2185
</emphasis>
(Ernst Schmidt Herbarium!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FE43FE3FF2F3B79A" ID-GBIF-Occurrence="4041537330" box="[474,1259,482,508]" collectingDate="2011-03-30" collectionCode="PRU" collectorName="Wright" lastPageId="10" lastPageNumber="203" location="Tembe Elephant Park" municipality="Tembe Elephant Park" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FE43FE3FF4CAB79A" box="[474,722,482,508]" pageId="10" pageNumber="203">Tembe Elephant Park</collectingMunicipality>
, (2732AB),
<date id="6855107DED1AFF90FCFAFE3FF20DB79A" box="[867,1045,482,508]" pageId="10" pageNumber="203" value="2011-03-30">
<collectingDate id="7811E995ED1AFF90FCFAFE3FF20DB79A" box="[867,1045,482,508]" pageId="10" pageNumber="203" value="2011-03-30">30 March 2011</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FBBAFE3FF29EB79A" box="[1059,1158,482,508]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FBBAFE3FF277B79A" box="[1059,1135,482,508]" pageId="10" pageNumber="203">Wright</collectorName>
1
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FB00FE3FF2C2B79A" box="[1177,1242,482,508]" pageId="10" pageNumber="203">PRU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FB6BFE3FF24CB446" ID-GBIF-Occurrence="4041537321" collectingDate="1958-01-15" collectionCode="NH, PRE" collectorName="Tinley" lastPageId="10" lastPageNumber="203" location="Lake Sibayi" municipality="Amanzimnyama Pan" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FB6BFE3FF6AAB447" pageId="10" pageNumber="203">Amanzimnyama Pan</collectingMunicipality>
area,
<location id="19346066ED1AFF90FF63FDDBF79BB446" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FF63FDDBF79BB446" box="[250,387,518,544]" municipality="Amanzimnyama Pan" name="Lake Sibayi" pageId="10" pageNumber="203">Lake Sibayi</location>
[
<location id="19346066ED1AFF90FE0BFDDBF7FEB446" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FE0BFDDBF7FEB446" box="[402,486,518,544]" municipality="Amanzimnyama Pan" name="Sibaya" pageId="10" pageNumber="203">Sibaya</location>
], (2732BC),
<date id="6855107DED1AFF90FDE2FDDBF523B447" box="[635,827,518,545]" pageId="10" pageNumber="203" value="1958-01-15">
<collectingDate id="7811E995ED1AFF90FDE2FDDBF523B447" box="[635,827,518,545]" pageId="10" pageNumber="203" value="1958-01-15">15 January 1958</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FCD1FDDBF5A9B446" box="[840,945,518,544]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FCD1FDDBF594B446" box="[840,908,518,544]" pageId="10" pageNumber="203">Tinley</collectorName>
59
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FC5AFDDBF5ECB446" box="[963,1012,518,544]" pageId="10" pageNumber="203">NH</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FB9FFDDBF25BB446" box="[1030,1091,518,544]" pageId="10" pageNumber="203">PRE</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FBC5FDDAF4C6B422" ID-GBIF-Occurrence="4041537303" collectingDate="1967-11-28" collectionCode="NH" collectorName="Nicholson" lastPageId="10" lastPageNumber="203" location="Sibaya" municipality="Lake Sibayi" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
E. side
<collectingMunicipality id="FC30ACC7ED1AFF90FB2DFDDBF325B446" box="[1204,1341,518,544]" pageId="10" pageNumber="203">Lake Sibayi</collectingMunicipality>
[
<location id="19346066ED1AFF90FAD5FDDBF3B8B446" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FAD5FDDBF3B8B446" box="[1356,1440,518,544]" municipality="Lake Sibayi" name="Sibaya" pageId="10" pageNumber="203">Sibaya</location>
], (2732BC),
<date id="6855107DED1AFF90FE9CFDF7F7C7B422" box="[261,479,554,580]" pageId="10" pageNumber="203" value="1967-11-28">
<collectingDate id="7811E995ED1AFF90FE9CFDF7F7C7B422" box="[261,479,554,580]" pageId="10" pageNumber="203" value="1967-11-28">28 November 1967</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FE70FDF7F493B422" box="[489,651,554,580]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FE70FDF7F443B422" box="[489,603,554,580]" pageId="10" pageNumber="203">Nicholson</collectorName>
621
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FD02FDF7F4D5B422" box="[667,717,554,580]" pageId="10" pageNumber="203">NH</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FD7DFDF7F756B40E" ID-GBIF-Occurrence="4041537315" collectingDate="1967-11-28" collectionCode="K, NH, PRE" collectorName="Strey &amp; Moll" lastPageId="10" lastPageNumber="203" location="Sibaya Lake" municipality="Sibaya Lake" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FD7DFDF7F56AB422" box="[740,882,554,580]" pageId="10" pageNumber="203">Sibaya Lake</collectingMunicipality>
, (2732BC),
<date id="6855107DED1AFF90FC60FDF7F2CBB422" box="[1017,1235,554,580]" pageId="10" pageNumber="203" value="1967-11-28">
<collectingDate id="7811E995ED1AFF90FC60FDF7F2CBB422" box="[1017,1235,554,580]" pageId="10" pageNumber="203" value="1967-11-28">28 November 1967</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FB44FDF6F3B3B422" box="[1245,1451,554,581]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FB44FDF6F375B422" box="[1245,1389,554,581]" pageId="10" pageNumber="203">Strey &amp; Moll</collectorName>
3977
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FF0BFD93F6B5B40E" box="[146,173,590,616]" pageId="10" pageNumber="203">K</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FF26FD93F6F7B40E" box="[191,239,590,616]" pageId="10" pageNumber="203">NH</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FE99FD93F725B40E" box="[256,317,590,616]" pageId="10" pageNumber="203">PRE</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FECFFD92F3B4B40E" ID-GBIF-Occurrence="4041537302" box="[342,1452,590,617]" collectingDate="1971-10-26" collectionCode="K, NH, PRE" collectorName="Moll &amp; Nel" lastPageId="10" lastPageNumber="203" location="E. Shores Lake Sibayi" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<location id="19346066ED1AFF90FECFFD92F44CB40E" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FECFFD92F44CB40E" box="[342,596,590,617]" name="E. Shores Lake Sibayi" pageId="10" pageNumber="203">E. Shores Lake Sibayi</location>
[
<location id="19346066ED1AFF90FDFAFD93F4AFB40E" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FDFAFD93F4AFB40E" box="[611,695,590,616]" name="Sibaya" pageId="10" pageNumber="203">Sibaya</location>
], (2732BC),
<date id="6855107DED1AFF90FCD2FD93F214B40E" box="[843,1036,590,616]" pageId="10" pageNumber="203" value="1971-10-26">
<collectingDate id="7811E995ED1AFF90FCD2FD93F214B40E" box="[843,1036,590,616]" pageId="10" pageNumber="203" value="1971-10-26">26 October 1971</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FB83FD93F2C5B40E" box="[1050,1245,590,617]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FB83FD93F285B40E" box="[1050,1181,590,617]" pageId="10" pageNumber="203">Moll &amp; Nel</collectorName>
5618
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FB69FD93F313B40E" box="[1264,1291,590,616]" pageId="10" pageNumber="203">K</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FA85FD93F354B40E" box="[1308,1356,590,616]" pageId="10" pageNumber="203">NH</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FAC7FD93F383B40E" box="[1374,1435,590,616]" pageId="10" pageNumber="203">PRE</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FF11FDAFF5C4B4EA" ID-GBIF-Occurrence="4041537312" box="[136,988,626,653]" collectingDate="1987-01-20" collectionCode="NU" collectorName="Cunningham" lastPageId="10" lastPageNumber="203" location="Lake Sibayi" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<location id="19346066ED1AFF90FF11FDAFF715B4EA" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FF11FDAFF715B4EA" box="[136,269,626,652]" name="Lake Sibayi" pageId="10" pageNumber="203">Lake Sibayi</location>
[
<location id="19346066ED1AFF90FE80FDAFF775B4EA" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FE80FDAFF775B4EA" box="[281,365,626,652]" name="Sibaya" pageId="10" pageNumber="203">Sibaya</location>
], (2732BC),
<date id="6855107DED1AFF90FE63FDAFF4ABB4EA" box="[506,691,626,653]" pageId="10" pageNumber="203" value="1987-01-20">
<collectingDate id="7811E995ED1AFF90FE63FDAFF4ABB4EA" box="[506,691,626,653]" pageId="10" pageNumber="203" value="1987-01-20">20 January 1987</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FD25FDAFF592B4EA" box="[700,906,626,652]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FD25FDAFF555B4EA" box="[700,845,626,652]" pageId="10" pageNumber="203">Cunningham</collectorName>
2200
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FC00FDAFF5D3B4EA" box="[921,971,626,652]" pageId="10" pageNumber="203">NU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FC78FDAFF42EB4D6" ID-GBIF-Occurrence="4041537320" collectingDate="1971-06-28" collectionCode="NH, NU, PRE, UDW" collectorName="Ward" lastPageId="10" lastPageNumber="203" location="False Bay Park" pageId="10" pageNumber="202" specimenCount="1" stateProvince="False Bay Park" typeStatus="paratype">
<collectingRegion id="DE2FF85FED1AFF90FC78FDAFF292B4EA" box="[993,1162,626,653]" pageId="10" pageNumber="203">False Bay Park</collectingRegion>
, (2732CD),
<date id="6855107DED1AFF90FA8BFDAFF3BEB4EA" box="[1298,1446,626,653]" pageId="10" pageNumber="203" value="1971-06-28">
<collectingDate id="7811E995ED1AFF90FA8BFDAFF3BEB4EA" box="[1298,1446,626,653]" pageId="10" pageNumber="203" value="1971-06-28">28 June 1971</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FF11FD4BF6E7B4D6" box="[136,255,662,688]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FF11FD4BF6DBB4D6" box="[136,195,662,688]" pageId="10" pageNumber="203">Ward</collectorName>
7114
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FE96FD4BF758B4D6" box="[271,320,662,688]" pageId="10" pageNumber="203">NH</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FED6FD4BF767B4D6" box="[335,383,662,688]" pageId="10" pageNumber="203">NU</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FE17FD4BF7D2B4D6" box="[398,458,662,688]" pageId="10" pageNumber="203">PRE</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FE41FD4BF43DB4D6" box="[472,549,662,688]" pageId="10" pageNumber="203">UDW</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FDA5FD4BF3B4B4D6" ID-GBIF-Occurrence="4041537306" box="[572,1452,662,689]" collectingDate="1944-12" collectionCode="PRE" collectorName="Gerstner" lastPageId="10" lastPageNumber="203" location="False Bay" municipality="False Bay" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FDA5FD4BF4B4B4D7" box="[572,684,662,689]" pageId="10" pageNumber="203">False Bay</collectingMunicipality>
, birds sanctuary, (2732DC),
<date id="6855107DED1AFF90FC75FD4BF2B9B4D6" box="[1004,1185,662,688]" pageId="10" pageNumber="203" value="1944-12">
<collectingDate id="7811E995ED1AFF90FC75FD4BF2B9B4D6" box="[1004,1185,662,688]" pageId="10" pageNumber="203" value="1944-12">December 1944</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FB32FD4AF356B4D6" box="[1195,1358,662,689]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FB32FD4AF317B4D7" box="[1195,1295,663,689]" pageId="10" pageNumber="203">Gerstner</collectorName>
5222
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FAC4FD4BF383B4D6" box="[1373,1435,662,688]" pageId="10" pageNumber="203">PRE</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FF11FD67F591B4B2" ID-GBIF-Occurrence="4041537322" box="[136,905,698,725]" collectingDate="1985-06-16" collectionCode="NH" collectorName="Gordon" lastPageId="10" lastPageNumber="203" location="Sodwana State Forest" municipality="Sodwana State Forest" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FF11FD67F766B4B3" box="[136,382,698,725]" pageId="10" pageNumber="203">Sodwana State Forest</collectingMunicipality>
, (2732DC),
<date id="6855107DED1AFF90FD93FD67F4BBB4B3" box="[522,675,698,725]" pageId="10" pageNumber="203" value="1985-06-16">
<collectingDate id="7811E995ED1AFF90FD93FD67F4BBB4B3" box="[522,675,698,725]" pageId="10" pageNumber="203" value="1985-06-16">16 June 1985</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FD37FD67F52EB4B2" box="[686,822,698,724]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FD37FD67F51CB4B2" box="[686,772,698,724]" pageId="10" pageNumber="203">Gordon</collectorName>
203
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FCDFFD67F560B4B2" box="[838,888,698,724]" pageId="10" pageNumber="203">NH</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FC08FD67F504B49E" ID-GBIF-Occurrence="4041537304" collectingDate="1985-12-22" collectionCode="NH, PRE" collectorName="Gordon" lastPageId="10" lastPageNumber="203" location="Air Force" municipality="Sodwana State Forest" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FC08FD67F29FB4B3" box="[913,1159,698,725]" pageId="10" pageNumber="203">Sodwana State Forest</collectingMunicipality>
, south of
<location id="19346066ED1AFF90FB6DFD67F379B4B3" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FB6DFD67F379B4B3" box="[1268,1377,698,725]" municipality="Sodwana State Forest" name="Air Force" pageId="10" pageNumber="203">Air Force</location>
camp, (2732DC),
<date id="6855107DED1AFF90FE90FD03F7FDB49F" box="[265,485,734,761]" pageId="10" pageNumber="203" value="1985-12-22">
<collectingDate id="7811E995ED1AFF90FE90FD03F7FDB49F" box="[265,485,734,761]" pageId="10" pageNumber="203" value="1985-12-22">22 December 1985</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FE6BFD03F463B49E" box="[498,635,734,760]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FE6BFD03F450B49E" box="[498,584,734,760]" pageId="10" pageNumber="203">Gordon</collectorName>
259
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FD15FD03F4A5B49E" box="[652,701,734,760]" pageId="10" pageNumber="203">NH</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FD57FD03F513B49E" box="[718,779,734,760]" pageId="10" pageNumber="203">PRE</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FCBCFD03F472B57A" ID-GBIF-Occurrence="4041537309" collectingDate="2014-04-30" collectionCode="NH" collectorName="Boon" lastPageId="10" lastPageNumber="203" location="Mphophomeni Trail" pageId="10" pageNumber="202" specimenCount="1" stateProvince="False Bay Park" typeStatus="paratype">
<collectingRegion id="DE2FF85FED1AFF90FCBCFD03F5CCB49E" box="[805,980,734,761]" pageId="10" pageNumber="203">False Bay Park</collectingRegion>
, near the end of the
<location id="19346066ED1AFF90FB5BFD03F3BFB49E" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FB5BFD03F3BFB49E" box="[1218,1447,734,760]" name="Mphophomeni Trail" pageId="10" pageNumber="203" stateProvince="False Bay Park">Mphophomeni Trail</location>
, (2732DC),
<date id="6855107DED1AFF90FE90FCDFF7B2B57A" box="[265,426,770,796]" pageId="10" pageNumber="203" value="2014-04-30">
<collectingDate id="7811E995ED1AFF90FE90FCDFF7B2B57A" box="[265,426,770,796]" pageId="10" pageNumber="203" value="2014-04-30">30 April 2014</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FE2FFCDEF40DB57A" box="[438,533,770,796]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FE2FFCDEF7E9B57A" box="[438,497,771,796]" pageId="10" pageNumber="203">Boon</collectorName>
63
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FDBEFCDFF441B57A" box="[551,601,770,796]" pageId="10" pageNumber="203">NH</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FDEBFCDFF3B3B57A" ID-GBIF-Occurrence="4041537311" box="[626,1451,770,797]" collectingDate="1956-01-07" collectionCode="NU" collectorName="Ward" lastPageId="10" lastPageNumber="203" location="Hluhluwe Game Reserve" pageId="10" pageNumber="202" specimenCount="1" stateProvince="Hluhluwe Game Reserve" typeStatus="paratype">
<collectingRegion id="DE2FF85FED1AFF90FDEBFCDFF589B57B" box="[626,913,770,797]" pageId="10" pageNumber="203">Hluhluwe Game Reserve</collectingRegion>
, (2832AA),
<date id="6855107DED1AFF90FB86FCDEF2C8B57B" box="[1055,1232,770,797]" pageId="10" pageNumber="203" value="1956-01-07">
<collectingDate id="7811E995ED1AFF90FB86FCDEF2C8B57B" box="[1055,1232,770,797]" pageId="10" pageNumber="203" value="1956-01-07">7 January 1956</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FB45FCDFF34FB57A" box="[1244,1367,770,796]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FB45FCDFF30FB57A" box="[1244,1303,770,796]" pageId="10" pageNumber="203">Ward</collectorName>
2947
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FAF1FCDFF382B57A" box="[1384,1434,770,796]" pageId="10" pageNumber="203">NU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FF11FCFBF233B526" ID-GBIF-Occurrence="4041537328" box="[136,1067,806,833]" collectingDate="1959-11-28" collectionCode="PRE, NU" collectorName="Ward" lastPageId="10" lastPageNumber="203" location="Hluhluwe Game Reserve" pageId="10" pageNumber="202" specimenCount="1" stateProvince="Hluhluwe Game Reserve" typeStatus="paratype">
<collectingRegion id="DE2FF85FED1AFF90FF11FCFBF7BCB527" box="[136,420,806,833]" pageId="10" pageNumber="203">Hluhluwe Game Reserve</collectingRegion>
, (2832AA),
<date id="6855107DED1AFF90FDB6FCFBF511B527" box="[559,777,806,833]" pageId="10" pageNumber="203" value="1959-11-28">
<collectingDate id="7811E995ED1AFF90FDB6FCFBF511B527" box="[559,777,806,833]" pageId="10" pageNumber="203" value="1959-11-28">28 November 1959</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FC8DFCFBF595B526" box="[788,909,806,832]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FC8DFCFBF557B526" box="[788,847,806,832]" pageId="10" pageNumber="203">Ward</collectorName>
3327
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FC05FCFBF5C0B526" box="[924,984,806,832]" pageId="10" pageNumber="203">PRE</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FC71FCFBF201B526" box="[1000,1049,806,832]" pageId="10" pageNumber="203">NU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FBA8FCFBF5A7B502" ID-GBIF-Occurrence="4041537301" collectingDate="1971-03-02" collectionCode="K, NH, PRE" collectorName="Moll" lastPageId="10" lastPageNumber="203" location="Monsia" pageId="10" pageNumber="202" specimenCount="1" stateProvince="Hluhluwe Game Reserve" typeStatus="paratype">
<collectingRegion id="DE2FF85FED1AFF90FBA8FCFBF355B527" box="[1073,1357,806,833]" pageId="10" pageNumber="203">Hluhluwe Game Reserve</collectingRegion>
,
<location id="19346066ED1AFF90FACEFCFBF3B3B526" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FACEFCFBF3B3B526" box="[1367,1451,806,832]" name="Monsia" pageId="10" pageNumber="203" stateProvince="Hluhluwe Game Reserve">Monsia</location>
[
<location id="19346066ED1AFF90FF16FC97F6EDB502" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FF16FC97F6EDB502" box="[143,245,842,868]" name="Mansiya" pageId="10" pageNumber="203" stateProvince="Hluhluwe Game Reserve">Mansiya</location>
]
<location id="19346066ED1AFF90FE9AFC97F75BB502" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FE9AFC97F75BB502" box="[259,323,842,868]" name="River" pageId="10" pageNumber="203" stateProvince="Hluhluwe Game Reserve">River</location>
, (2832AA),
<date id="6855107DED1AFF90FE48FC97F469B502" box="[465,625,842,868]" pageId="10" pageNumber="203" value="1971-03-02">
<collectingDate id="7811E995ED1AFF90FE48FC97F469B502" box="[465,625,842,868]" pageId="10" pageNumber="203" value="1971-03-02">2 March 1971</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FDE7FC97F4EBB502" box="[638,755,842,868]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FDE7FC97F4ABB502" box="[638,691,842,868]" pageId="10" pageNumber="203">Moll</collectorName>
5291
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FC9CFC97F538B502" box="[773,800,842,868]" pageId="10" pageNumber="203">K</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FCA8FC97F579B502" box="[817,865,842,868]" pageId="10" pageNumber="203">NH</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FCE8FC97F5B6B502" box="[881,942,842,868]" pageId="10" pageNumber="203">PRE</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FC5EFC97F5DDB5EE" ID-GBIF-Occurrence="4041537305" collectingDate="1994-02-22" collectionCode="PCE, PRU" collectorName="Abbott" lastPageId="10" lastPageNumber="203" location="Reserve" municipality="Bonamanzi Game Ranch" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FC5EFC97F2F9B502" box="[967,1249,842,868]" pageId="10" pageNumber="203">Bonamanzi Game Ranch</collectingMunicipality>
[
<location id="19346066ED1AFF90FB76FC96F349B503" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FB76FC96F349B503" box="[1263,1361,843,869]" municipality="Bonamanzi Game Ranch" name="Reserve" pageId="10" pageNumber="203">Reserve</location>
],
<locationDeviation id="0DE62F9FED1AFF90FAFDFC97F6AEB5EE" bearing="180.0" distance="10000.0" pageId="10" pageNumber="203">
<quantity id="DB139B58ED1AFF90FAFDFC97F3B4B502" box="[1380,1452,842,868]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="203" unit="km" value="10.0">10 km</quantity>
S of
</locationDeviation>
<location id="19346066ED1AFF90FF24FCB3F736B5EE" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FF24FCB3F736B5EE" box="[189,302,878,904]" municipality="Bonamanzi Game Ranch" name="Hluhluwe" pageId="10" pageNumber="203">Hluhluwe</location>
, (2832AA),
<date id="6855107DED1AFF90FE23FCB3F49CB5EE" box="[442,644,878,904]" pageId="10" pageNumber="203" value="1994-02-22">
<collectingDate id="7811E995ED1AFF90FE23FCB3F49CB5EE" box="[442,644,878,904]" pageId="10" pageNumber="203" value="1994-02-22">22 February 1994</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FD16FCB3F501B5EE" box="[655,793,878,904]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FD16FCB3F4C2B5EE" box="[655,730,878,904]" pageId="10" pageNumber="203">Abbott</collectorName>
6317
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FCB1FCB3F57CB5EE" box="[808,868,878,904]" collectionName="PCE" pageId="10" pageNumber="203">PCE</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FCEAFCB3F5ABB5EE" box="[883,947,878,904]" pageId="10" pageNumber="203">PRU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FC52FCB3F278B5CA" ID-GBIF-Occurrence="4041537327" collectingDate="1994-11-05" collectionCode="PCE, PRU" collectorName="Abbott" lastPageId="10" lastPageNumber="203" location="Reserve" municipality="Bonamanzi Game Ranch" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FC52FCB3F2FBB5EE" box="[971,1251,878,904]" pageId="10" pageNumber="203">Bonamanzi Game Ranch</collectingMunicipality>
[
<location id="19346066ED1AFF90FB69FCB2F34AB5EF" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FB69FCB2F34AB5EF" box="[1264,1362,879,905]" municipality="Bonamanzi Game Ranch" name="Reserve" pageId="10" pageNumber="203">Reserve</location>
],
<locationDeviation id="0DE62F9FED1AFF90FAFCFCB3F6C0B5CA" bearing="112.5" distance="10000.0" pageId="10" pageNumber="203">
<quantity id="DB139B58ED1AFF90FAFCFCB3F3B4B5EE" box="[1381,1452,878,904]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="203" unit="km" value="10.0">10 km</quantity>
ESE of
</locationDeviation>
<location id="19346066ED1AFF90FF46FC4FF748B5CA" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FF46FC4FF748B5CA" box="[223,336,914,940]" municipality="Bonamanzi Game Ranch" name="Hluhluwe" pageId="10" pageNumber="203">Hluhluwe</location>
, (2832AA) [2832AB],
<date id="6855107DED1AFF90FDCDFC4FF538B5CA" box="[596,800,914,941]" pageId="10" pageNumber="203" value="1994-11-05">
<collectingDate id="7811E995ED1AFF90FDCDFC4FF538B5CA" box="[596,800,914,941]" pageId="10" pageNumber="203" value="1994-11-05">5 November 1994</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FCB2FC4FF5ACB5CA" box="[811,948,914,940]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FCB2FC4FF56EB5CA" box="[811,886,914,940]" pageId="10" pageNumber="203">Abbott</collectorName>
6498
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FC5AFC4FF5E7B5CA" box="[963,1023,914,940]" collectionName="PCE" pageId="10" pageNumber="203">PCE</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FB96FC4FF257B5CA" box="[1039,1103,914,940]" pageId="10" pageNumber="203">PRU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FBFFFC4FF6C1B592" ID-GBIF-Occurrence="4041537337" collectingDate="1996-03-05" collectionCode="PCE, PRU" collectorName="Abbott" elevation="150" lastPageId="10" lastPageNumber="203" location="Bonamanzi Game" municipality="Hluhluwe" pageId="10" pageNumber="202" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="FC30ACC7ED1AFF90FBFFFC4FF2CFB5CA" box="[1126,1239,914,940]" pageId="10" pageNumber="203">Hluhluwe</collectingMunicipality>
,
<location id="19346066ED1AFF90FB7BFC4FF3B3B5CA" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FB7BFC4FF3B3B5CA" box="[1250,1451,914,940]" municipality="Hluhluwe" name="Bonamanzi Game" pageId="10" pageNumber="203">Bonamanzi Game</location>
<location id="19346066ED1AFF90FF11FC6BF6A3B5B6" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FF11FC6BF6A3B5B6" box="[136,187,950,976]" municipality="Hluhluwe" name="Park" pageId="10" pageNumber="203">Park</location>
[
<location id="19346066ED1AFF90FF52FC6AF735B5B7" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FF52FC6AF735B5B7" box="[203,301,951,977]" municipality="Hluhluwe" name="Reserve" pageId="10" pageNumber="203">Reserve</location>
], on left of road
<quantity id="DB139B58ED1AFF90FE76FC6BF420B5B7" box="[495,568,950,977]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="203" unit="m" value="150.0">
<elevation id="97C6D18EED1AFF90FE76FC6BF420B5B7" box="[495,568,950,977]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="203" unit="m" value="150.0">150 m</elevation>
</quantity>
beyond Game Lodge turnoff, (2832AA),
<date id="6855107DED1AFF90FB80FC6BF2A5B5B6" box="[1049,1213,950,977]" pageId="10" pageNumber="203" value="1996-03-05">
<collectingDate id="7811E995ED1AFF90FB80FC6BF2A5B5B6" box="[1049,1213,950,977]" pageId="10" pageNumber="203" value="1996-03-05">5 March 1996</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FB53FC6BF34DB5B6" box="[1226,1365,950,976]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FB53FC6BF30DB5B6" box="[1226,1301,950,976]" pageId="10" pageNumber="203">Abbott</collectorName>
6930
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FAFEFC6BF3BBB5B6" box="[1383,1443,950,976]" collectionName="PCE" pageId="10" pageNumber="203">PCE</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FF11FC07F6D0B592" box="[136,200,986,1012]" pageId="10" pageNumber="203">PRU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FF7AFC07F335B592" ID-GBIF-Occurrence="4041537318" box="[227,1325,986,1013]" collectingDate="1996-03-05" collectionCode="PRU" collectorName="Abbott" lastPageId="10" lastPageNumber="203" location="Reserve" municipality="Bonamanzi Game Ranch" pageId="10" pageNumber="202" specimenCount="1" stateProvince="Hluhluwe" typeStatus="paratype">
<collectingRegion id="DE2FF85FED1AFF90FF7AFC07F74CB592" box="[227,340,986,1012]" pageId="10" pageNumber="203">Hluhluwe</collectingRegion>
,
<collectingMunicipality id="FC30ACC7ED1AFF90FEFBFC07F467B592" box="[354,639,986,1012]" pageId="10" pageNumber="203">Bonamanzi Game Ranch</collectingMunicipality>
[
<location id="19346066ED1AFF90FD16FC06F4E9B593" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FD16FC06F4E9B593" box="[655,753,987,1013]" municipality="Bonamanzi Game Ranch" name="Reserve" pageId="10" pageNumber="203" stateProvince="Hluhluwe">Reserve</location>
], (2832AA),
<date id="6855107DED1AFF90FC10FC07F237B592" box="[905,1071,986,1013]" pageId="10" pageNumber="203" value="1996-03-05">
<collectingDate id="7811E995ED1AFF90FC10FC07F237B592" box="[905,1071,986,1013]" pageId="10" pageNumber="203" value="1996-03-05">5 March 1996</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FBA5FC07F2D0B592" box="[1084,1224,986,1012]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FBA5FC07F29FB592" box="[1084,1159,986,1012]" pageId="10" pageNumber="203">Abbott</collectorName>
6931
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FB42FC07F304B592" box="[1243,1308,986,1012]" pageId="10" pageNumber="203">PRU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FAAFFC07F241B27E" ID-GBIF-Occurrence="4041537332" collectingDate="1996-03-05" collectionCode="PRU" collectorName="Abbott" lastPageId="10" lastPageNumber="203" location="Reserve" municipality="Bonamanzi Game Ranch" pageId="10" pageNumber="202" specimenCount="1" stateProvince="Hluhluwe" typeStatus="paratype">
<collectingRegion id="DE2FF85FED1AFF90FAAFFC07F3BFB592" box="[1334,1447,986,1012]" pageId="10" pageNumber="203">Hluhluwe</collectingRegion>
,
<collectingMunicipality id="FC30ACC7ED1AFF90FF11FC23F7BFB27E" box="[136,423,1022,1048]" pageId="10" pageNumber="203">Bonamanzi Game Ranch</collectingMunicipality>
[
<location id="19346066ED1AFF90FE2EFC22F401B27F" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FE2EFC22F401B27F" box="[439,537,1023,1049]" municipality="Bonamanzi Game Ranch" name="Reserve" pageId="10" pageNumber="203" stateProvince="Hluhluwe">Reserve</location>
], (2832AA),
<date id="6855107DED1AFF90FD2AFC23F541B27E" box="[691,857,1022,1049]" pageId="10" pageNumber="203" value="1996-03-05">
<collectingDate id="7811E995ED1AFF90FD2AFC23F541B27E" box="[691,857,1022,1049]" pageId="10" pageNumber="203" value="1996-03-05">5 March 1996</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FCFEFC23F5ECB27E" box="[871,1012,1022,1048]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FCFEFC23F5AAB27E" box="[871,946,1022,1048]" pageId="10" pageNumber="203">Abbott</collectorName>
6932
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FB9EFC23F250B27E" box="[1031,1096,1022,1048]" pageId="10" pageNumber="203">PRU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FBFAFC23F212B25A" ID-GBIF-Occurrence="4041537331" collectingDate="1963-01-16" collectionCode="NH, NU" collectorName="Dutton" lastPageId="10" lastPageNumber="203" location="Dlozi" pageId="10" pageNumber="202" specimenCount="1" stateProvince="Hluhluwe" typeStatus="paratype">
<collectingRegion id="DE2FF85FED1AFF90FBFAFC23F2CCB27E" box="[1123,1236,1022,1048]" pageId="10" pageNumber="203">Hluhluwe</collectingRegion>
,
<location id="19346066ED1AFF90FB7AFC23F339B27E" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FB7AFC23F339B27E" box="[1251,1313,1022,1048]" name="Dlozi" pageId="10" pageNumber="203" stateProvince="Hluhluwe">Dlozi</location>
(or
<location id="19346066ED1AFF90FACFFC23F3BCB27E" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FACFFC23F3BCB27E" box="[1366,1444,1022,1048]" name="Dhlozi" pageId="10" pageNumber="203" stateProvince="Hluhluwe">Dhlozi</location>
)
<location id="19346066ED1AFF90FF11FBFFF6E5B25A" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FF11FBFFF6E5B25A" box="[136,253,1058,1084]" name="Plantation" pageId="10" pageNumber="203" stateProvince="Hluhluwe">Plantation</location>
,
<location id="19346066ED1AFF90FE9EFBFFF7A2B25A" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FE9EFBFFF7A2B25A" box="[263,442,1058,1084]" name="Hlabisa District" pageId="10" pageNumber="203" stateProvince="Hluhluwe">Hlabisa District</location>
, (2832AB),
<date id="6855107DED1AFF90FDDAFBFFF4E6B25A" box="[579,766,1058,1085]" pageId="10" pageNumber="203" value="1963-01-16">
<collectingDate id="7811E995ED1AFF90FDDAFBFFF4E6B25A" box="[579,766,1058,1085]" pageId="10" pageNumber="203" value="1963-01-16">16 January 1963</collectingDate>
</date>
,
<emphasis id="2E9FEAAFED1AFF90FC91FBFEF560B25A" box="[776,888,1058,1084]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FC91FBFEF54EB25A" box="[776,854,1059,1084]" pageId="10" pageNumber="203">Dutton</collectorName>
88
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FC1EFBFFF5A0B25A" box="[903,952,1058,1084]" pageId="10" pageNumber="203">NH</collectionCode>
!,
<collectionCode id="7AFAAE78ED1AFF90FC5EFBFFF5E0B25A" box="[967,1016,1058,1084]" pageId="10" pageNumber="203">NU</collectionCode>
!);
</materialsCitation>
<materialsCitation id="AC833CE0ED1AFF90FB89FBFFF33DB206" ID-GBIF-Occurrence="4041537336" collectingDate="1997-02-15" collectionCode="NH" collectorName="Ward" lastPageId="10" lastPageNumber="203" location="St. Lucia" municipality="Lake Bhangazi" pageId="10" pageNumber="202" specimenCount="1" stateProvince="North Coast" typeStatus="paratype">
<collectingRegion id="DE2FF85FED1AFF90FB89FBFFF283B25A" box="[1040,1179,1058,1084]" pageId="10" pageNumber="203">North Coast</collectingRegion>
,
<location id="19346066ED1AFF90FB3CFBFFF312B25A" LSID="urn:lsid:plazi:treatment:944287ABED12FF97FF11FE69F3B3B21A:19346066ED1AFF90FB3CFBFFF312B25A" box="[1189,1290,1058,1084]" municipality="Lake Bhangazi" name="St. Lucia" pageId="10" pageNumber="203" stateProvince="North Coast">St. Lucia</location>
system (sensu lato),
<collectingMunicipality id="FC30ACC7ED1AFF90FF50FB9BF76AB206" box="[201,370,1094,1120]" pageId="10" pageNumber="203">Lake Bhangazi</collectingMunicipality>
(South) area, eastern side of lake,
<date id="6855107DED1AFF90FD6FFB9BF5D8B206" box="[758,960,1094,1121]" pageId="10" pageNumber="203" value="1997-02-15">
<collectingDate id="7811E995ED1AFF90FD6FFB9BF5D8B206" box="[758,960,1094,1121]" pageId="10" pageNumber="203" value="1997-02-15">15 February 1997</collectingDate>
</date>
, (2832BA),
<emphasis id="2E9FEAAFED1AFF90FBD2FB9BF2CAB206" box="[1099,1234,1094,1120]" italics="true" pageId="10" pageNumber="203">
<collectorName id="B11E536BED1AFF90FBD2FB9BF29EB206" box="[1099,1158,1094,1120]" pageId="10" pageNumber="203">Ward</collectorName>
13915
</emphasis>
(
<collectionCode id="7AFAAE78ED1AFF90FB7BFB9BF30CB206" box="[1250,1300,1094,1120]" pageId="10" pageNumber="203">NH</collectionCode>
!).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED1AFF90FF24FBB7F250B08E" pageId="10" pageNumber="203" type="discussion">
<paragraph id="1C5436BDED1AFF90FF24FBB7F33EB3AF" blockId="10.[136,1452,158,2093]" pageId="10" pageNumber="203">
<emphasis id="2E9FEAAFED1AFF90FF24FBB7F4D7B2E2" bold="true" box="[189,719,1130,1157]" pageId="10" pageNumber="203">
Notes and key to the species of
<taxonomicName id="DBEB4D3EED1AFF90FDADFBB7F4DEB2E3" authorityName="Herzog" authorityYear="1903" baseAuthorityName="Weberbauer" baseAuthorityYear="1895" box="[564,710,1130,1157]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="genus">
<emphasis id="2E9FEAAFED1AFF90FDADFBB7F4DEB2E3" bold="true" box="[564,710,1130,1157]" italics="true" pageId="10" pageNumber="203">Phyllogeiton</emphasis>
</taxonomicName>
:
</emphasis>
—
<taxonomicName id="DBEB4D3EED1AFF90FD72FBB7F21BB2E2" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[747,1027,1130,1156]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED1AFF90FD72FBB7F21BB2E2" box="[747,1027,1130,1156]" italics="true" pageId="10" pageNumber="203">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
is morphologically most similar to
<taxonomicName id="DBEB4D3EED1AFF90FA0EFBB6F6FCB2CE" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1AFF90FA0EFBB6F6FCB2CE" italics="true" pageId="10" pageNumber="203">P. discolor</emphasis>
</taxonomicName>
(
<figureCitation id="84D02A38ED1AFF90FF6AFB53F720B2CE" box="[243,312,1166,1192]" captionStart="FIGURE 6" captionStartId="11.[136,229,2011,2033]" captionTargetBox="[177,1410,317,1986]" captionTargetId="figure-81@11.[177,1410,317,1986]" captionTargetPageId="11" captionText="FIGURE 6. Phyllogeiton discolor. A. Mature bark pattern. B. Flowers. C. Fruit; note leaves with upper surface of blade glossy and dark green. D. Leaves viewed from below, showing a greyish bloom. Photographs: R.G.C. Boon (A, C) &amp; A.E. van Wyk (B, D)." figureDoi="http://doi.org/10.5281/zenodo.7690767" httpUri="https://zenodo.org/record/7690767/files/figure.png" pageId="10" pageNumber="203">Fig. 6</figureCitation>
), but the species’ known ranges are allopatric and
<taxonomicName id="DBEB4D3EED1AFF90FCE9FB52F214B2CE" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[880,1036,1166,1192]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED1AFF90FCE9FB52F214B2CE" box="[880,1036,1166,1192]" italics="true" pageId="10" pageNumber="203">P. trachybasis</emphasis>
</taxonomicName>
is for now considered to be endemic to the Maputaland Centre of Endemism, as well as to KwaZulu-Natal.
<taxonomicName id="DBEB4D3EED1AFF90FC2BFB6FF2BEB2AA" box="[946,1190,1202,1228]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1AFF90FC2BFB6FF2BEB2AA" box="[946,1190,1202,1228]" italics="true" pageId="10" pageNumber="203">Phyllogeiton discolor</emphasis>
</taxonomicName>
has not been collected from KwaZulu-Natal nor has it been recorded from
<collectingCountry id="64FC762DED1AFF90FD7DFB0BF55EB296" box="[740,838,1238,1264]" name="Swaziland" pageId="10" pageNumber="203">Eswatini</collectingCountry>
(
<bibRefCitation id="787A4B4CED1AFF90FCCEFB0BF27DB297" author="Loffler, L. &amp; Loffler, P." box="[855,1125,1238,1265]" pageId="10" pageNumber="203" refId="ref10117" refString="Loffler, L. &amp; Loffler, P. (2005) Swaziland tree atlas - including shrubs and climbers, 2 nd ed. South African Botanical Diversity Network Report No. 35, SABONET, Pretoria, 196 pp." type="book" year="2005">Loffler &amp; Loffler 2005</bibRefCitation>
, L. Loffler, pers. comm.). A specimen at Herb. SDNH (
<emphasis id="2E9FEAAFED1AFF90FE5CFB26F465B372" box="[453,637,1274,1301]" italics="true" pageId="10" pageNumber="203">Compton 28939</emphasis>
) tentatively identified as
<taxonomicName id="DBEB4D3EED1AFF90FC30FB27F290B372" box="[937,1160,1274,1300]" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1AFF90FC30FB27F290B372" box="[937,1160,1274,1300]" italics="true" pageId="10" pageNumber="203">Berchemia discolor</emphasis>
</taxonomicName>
is of
<taxonomicName id="DBEB4D3EED1AFF90FB4BFB26F35AB372" authorityName="Grubov" authorityYear="1949" baseAuthorityName="Sonder" baseAuthorityYear="1860" box="[1234,1346,1274,1300]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="zeyheri">
<emphasis id="2E9FEAAFED1AFF90FB4BFB26F35AB372" box="[1234,1346,1274,1300]" italics="true" pageId="10" pageNumber="203">P. zeyheri</emphasis>
</taxonomicName>
(
<figureCitation id="84D02A38ED1AFF90FACCFB27F385B372" box="[1365,1437,1274,1300]" captionStart="FIGURE 7" captionStartId="12.[136,229,1953,1975]" captionTargetBox="[151,1435,190,1929]" captionTargetId="figure-19@12.[151,1435,190,1929]" captionTargetPageId="12" captionText="FIGURE 7. Phyllogeiton zeyheri. A. Mature bark pattern. B. Flowers; note leaves with upper surface of blade dull and with a greyish bloom. C. Fruit. D. Leaves viewed from below, showing a greyish bloom similar to that of the upper surface. Photographs: R.G.C. Boon (A) &amp; A.E. van Wyk (B–D)." figureDoi="http://doi.org/10.5281/zenodo.7690769" httpUri="https://zenodo.org/record/7690769/files/figure.png" pageId="10" pageNumber="203">Fig. 7</figureCitation>
). There is one correctly labelled
<taxonomicName id="DBEB4D3EED1AFF90FE70FAC2F479B35E" box="[489,609,1310,1336]" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1AFF90FE70FAC2F479B35E" box="[489,609,1310,1336]" italics="true" pageId="10" pageNumber="203">P. discolor</emphasis>
</taxonomicName>
specimen at LMA from the southern part of
<collectingCountry id="64FC762DED1AFF90FBC6FAC3F2EBB35E" box="[1119,1267,1310,1336]" name="Mozambique" pageId="10" pageNumber="203">Mozambique</collectingCountry>
(covered by the provinces of
<collectingRegion id="DE2FF85FED1AFF90FE82FA9EF76DB33B" box="[283,373,1347,1373]" country="Mozambique" name="Maputo" pageId="10" pageNumber="203">Maputo</collectingRegion>
,
<collectingRegion id="DE2FF85FED1AFF90FE19FA9FF7A2B33A" box="[384,442,1346,1372]" country="Mozambique" name="Gaza" pageId="10" pageNumber="203">Gaza</collectingRegion>
and
<collectingRegion id="DE2FF85FED1AFF90FE68FA9FF46BB33A" box="[497,627,1346,1372]" country="Mozambique" name="Inhambane" pageId="10" pageNumber="203">Inhambane</collectingRegion>
). This specimen was collected at Massingir in
<collectingRegion id="DE2FF85FED1AFF90FBE7FA9FF2A0B33A" box="[1150,1208,1346,1372]" country="Mozambique" name="Gaza" pageId="10" pageNumber="203">Gaza</collectingRegion>
(
<emphasis id="2E9FEAAFED1AFF90FB50FA9EF383B33A" box="[1225,1435,1346,1373]" italics="true" pageId="10" pageNumber="203">Lousã &amp; Rosa 301</emphasis>
), which is about
<quantity id="DB139B58ED1AFF90FEAAFABBF790B3E6" box="[307,392,1382,1409]" metricMagnitude="5" metricUnit="m" metricValue="3.25" pageId="10" pageNumber="203" unit="km" value="325.0">325 km</quantity>
north of the nearest known
<taxonomicName id="DBEB4D3EED1AFF90FD5AFABAF578B3E6" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[707,864,1382,1408]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED1AFF90FD5AFABAF578B3E6" box="[707,864,1382,1408]" italics="true" pageId="10" pageNumber="203">P. trachybasis</emphasis>
</taxonomicName>
plants in KwaZulu-Natal. Specimens at Herb. LMA labelled
<taxonomicName id="DBEB4D3EED1AFF90FF70FA56F77DB3C2" box="[233,357,1418,1444]" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1AFF90FF70FA56F77DB3C2" box="[233,357,1418,1444]" italics="true" pageId="10" pageNumber="203">B. discolor</emphasis>
</taxonomicName>
from Catuane, which is in
<collectingCountry id="64FC762DED1AFF90FD04FA57F529B3C2" box="[669,817,1418,1444]" name="Mozambique" pageId="10" pageNumber="203">Mozambique</collectingCountry>
just north of Ndumo Game Reserve in KwaZulu-Natal, and Namaacha (west of
<collectingRegion id="DE2FF85FED1AFF90FE0EFA72F7F6B3AF" box="[407,494,1455,1481]" country="Mozambique" name="Maputo" pageId="10" pageNumber="203">Maputo</collectingRegion>
near the border with
<collectingCountry id="64FC762DED1AFF90FD47FA73F55AB3AE" box="[734,834,1454,1480]" name="Swaziland" pageId="10" pageNumber="203">Eswatini</collectingCountry>
) are misidentified collections of
<taxonomicName id="DBEB4D3EED1AFF90FB2AFA72F307B3AE" authorityName="Grubov" authorityYear="1949" baseAuthorityName="Sonder" baseAuthorityYear="1860" box="[1203,1311,1454,1480]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="zeyheri">
<emphasis id="2E9FEAAFED1AFF90FB2AFA72F307B3AE" box="[1203,1311,1454,1480]" italics="true" pageId="10" pageNumber="203">P. zeyheri</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1C5436BDED1AFF90FF24FA0FF250B08E" blockId="10.[136,1452,158,2093]" pageId="10" pageNumber="203">
The habitat of the new species and
<taxonomicName id="DBEB4D3EED1AFF90FDD5FA0EF4DCB38A" box="[588,708,1490,1516]" class="Magnoliopsida" family="Rhamnaceae" genus="Berchemia" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1AFF90FDD5FA0EF4DCB38A" box="[588,708,1490,1516]" italics="true" pageId="10" pageNumber="203">P. discolor</emphasis>
</taxonomicName>
also differs.
<taxonomicName id="DBEB4D3EED1AFF90FCCEFA0FF277B38A" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[855,1135,1490,1516]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED1AFF90FCCEFA0FF277B38A" box="[855,1135,1490,1516]" italics="true" pageId="10" pageNumber="203">Phyllogeiton trachybasis</emphasis>
</taxonomicName>
prefers sandy soils of Early to Late Pleistocene coastal dunes (
<bibRefCitation id="787A4B4CED1AFF90FD8DFA2BF528B076" author="Van Wyk, A. E. &amp; Smith, G. F." box="[532,816,1526,1552]" pageId="10" pageNumber="203" refId="ref11475" refString="Van Wyk, A. E. &amp; Smith, G. F. (2001) Regions of floristic endemism in southern Africa: a review with emphasis on succulents. Umdaus Press, Hatfield, Pretoria, 199 pp." type="book" year="2001">Van Wyk &amp; Smith 2001</bibRefCitation>
), although it has also been collected from Scarp Forest and once from riverine forest.
<taxonomicName id="DBEB4D3EED1AFF90FE7BF9C7F4CDB052" box="[482,725,1562,1588]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1AFF90FE7BF9C7F4CDB052" box="[482,725,1562,1588]" italics="true" pageId="10" pageNumber="203">Phyllogeiton discolor</emphasis>
</taxonomicName>
also occurs in bushveld (savanna), but is usually found on river banks or termitaria (Coates
<bibRefCitation id="787A4B4CED1AFF90FE57F9E3F46EB03E" author="Palgrave, M." box="[462,630,1598,1624]" pageId="10" pageNumber="203" refId="ref9035" refString="Palgrave, M. (2002) Keith Coates Palgrave trees of southern Africa, 3 rd ed. Struik, Cape Town, 1212 pp." type="book" year="2002">Palgrave 2002</bibRefCitation>
,
<bibRefCitation id="787A4B4CED1AFF90FD1DF9E3F531B03E" author="Schmidt, E." box="[644,809,1598,1624]" pageId="10" pageNumber="203" pagination="544 - 552" refId="ref10878" refString="Schmidt, E. (2018) Rhamnaceae. In: Burrows, J. E., Burrows, S. M., Lotter, M. C. &amp; Schmidt, E. (Eds.) Trees and shrubs Mozambique. Publishing Print Matters, Noordhoek, Cape Town, pp. 544 - 552." type="book chapter" year="2018">Schmidt 2018</bibRefCitation>
). The two species differ in a number of morphological features including bark, leaf shape, sepal and petal position at anthesis, and fruit size, shape and colour. In addition,
<taxonomicName id="DBEB4D3EED1AFF90FA0EF9BEF711B0C6" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED1AFF90FA0EF9BEF711B0C6" italics="true" pageId="10" pageNumber="203">P. trachybasis</emphasis>
</taxonomicName>
has ± concolorous leaves without any bloom and the fruit are often faintly longitudinally ribbed, whereas
<taxonomicName id="DBEB4D3EED1AFF90FF11F976F719B0A2" box="[136,257,1706,1732]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1AFF90FF11F976F719B0A2" box="[136,257,1706,1732]" italics="true" pageId="10" pageNumber="203">P. discolor</emphasis>
</taxonomicName>
has leaves that are distinctly paler due to the presence of a greyish bloom and the fruit surface is plane. Further distinctions, also from
<taxonomicName id="DBEB4D3EED1AFF90FE7BF912F457B08E" authorityName="Grubov" authorityYear="1949" baseAuthorityName="Sonder" baseAuthorityYear="1860" box="[482,591,1742,1768]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="zeyheri">
<emphasis id="2E9FEAAFED1AFF90FE7BF912F457B08E" box="[482,591,1742,1768]" italics="true" pageId="10" pageNumber="203">P. zeyheri</emphasis>
</taxonomicName>
, are supplied in the identification key below.
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED1AFF97FF24F92FF6F9B742" lastPageId="13" lastPageNumber="206" pageId="10" pageNumber="203" type="reference_group">
<paragraph id="1C5436BDED1AFF91FF24F92FF769B742" blockId="10.[136,1452,158,2093]" lastBlockId="11.[136,1451,158,292]" lastPageId="11" lastPageNumber="204" pageId="10" pageNumber="203">
<taxonomicName id="DBEB4D3EED1AFF90FF24F92FF7ABB16A" ID-CoL="77FR2" authority="Klotzsch 1862: 110" box="[189,435,1778,1804]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1AFF90FF24F92FF7ABB16A" box="[189,435,1778,1804]" italics="true" pageId="10" pageNumber="203">Phyllogeiton discolor</emphasis>
</taxonomicName>
(
<figureCitation id="84D02A38ED1AFF90FE5FF92FF415B16A" box="[454,525,1778,1804]" captionStart="FIGURE 6" captionStartId="11.[136,229,2011,2033]" captionTargetBox="[177,1410,317,1986]" captionTargetId="figure-81@11.[177,1410,317,1986]" captionTargetPageId="11" captionText="FIGURE 6. Phyllogeiton discolor. A. Mature bark pattern. B. Flowers. C. Fruit; note leaves with upper surface of blade glossy and dark green. D. Leaves viewed from below, showing a greyish bloom. Photographs: R.G.C. Boon (A, C) &amp; A.E. van Wyk (B, D)." figureDoi="http://doi.org/10.5281/zenodo.7690767" httpUri="https://zenodo.org/record/7690767/files/figure.png" pageId="10" pageNumber="203">Fig. 6</figureCitation>
) and
<taxonomicName id="DBEB4D3EED1AFF90FDCAF92EF4DBB16A" ID-CoL="77G2Q" authorityName="Grubov" authorityYear="1949" baseAuthorityName="Sonder" baseAuthorityYear="1860" box="[595,707,1778,1804]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="zeyheri">
<emphasis id="2E9FEAAFED1AFF90FDCAF92EF4DBB16A" box="[595,707,1778,1804]" italics="true" pageId="10" pageNumber="203">P. zeyheri</emphasis>
</taxonomicName>
(
<figureCitation id="84D02A38ED1AFF90FD4CF92FF504B16A" box="[725,796,1778,1804]" captionStart="FIGURE 7" captionStartId="12.[136,229,1953,1975]" captionTargetBox="[151,1435,190,1929]" captionTargetId="figure-19@12.[151,1435,190,1929]" captionTargetPageId="12" captionText="FIGURE 7. Phyllogeiton zeyheri. A. Mature bark pattern. B. Flowers; note leaves with upper surface of blade dull and with a greyish bloom. C. Fruit. D. Leaves viewed from below, showing a greyish bloom similar to that of the upper surface. Photographs: R.G.C. Boon (A) &amp; A.E. van Wyk (B–D)." figureDoi="http://doi.org/10.5281/zenodo.7690769" httpUri="https://zenodo.org/record/7690769/files/figure.png" pageId="10" pageNumber="203">Fig. 7</figureCitation>
) differ markedly in the colour and fluorescence of their hardwood (
<bibRefCitation id="787A4B4CED1AFF90FE9FF8CBF798B156" author="Dyer, S. T." box="[262,384,1814,1840]" pageId="10" pageNumber="203" pagination="75 - 87" refId="ref9309" refString="Dyer, S. T. (1988) Wood fluorescence of indigenous South African trees. IAWA Bulletin n. s. 9: 75 - 87." type="book chapter" year="1988">Dyer 1988</bibRefCitation>
,
<bibRefCitation id="787A4B4CED1AFF90FE13F8CAF459B156" author="Dyer, S. &amp; James, B. &amp; James, D." box="[394,577,1814,1841]" pageId="10" pageNumber="203" refId="ref9273" refString="Dyer, S., James, B. &amp; James, D. (2016) Guide to the properties and uses of southern African wood. Briza Publications, Pretoria, 336 pp." type="book" year="2016">
Dyer
<emphasis id="2E9FEAAFED1AFF90FE51F8CAF418B156" box="[456,512,1814,1840]" italics="true" pageId="10" pageNumber="203">et al.</emphasis>
2016
</bibRefCitation>
), a distinction reflected in their English and Afrikaans common names. Known as “red ivory”/
<emphasis id="2E9FEAAFED1AFF90FEB7F8E6F7B8B132" box="[302,416,1851,1876]" italics="true" pageId="10" pageNumber="203">rooi-ivoor</emphasis>
, the heartwood of
<taxonomicName id="DBEB4D3EED1AFF90FDECF8E6F4FAB132" ID-CoL="77G2Q" authorityName="Grubov" authorityYear="1949" baseAuthorityName="Sonder" baseAuthorityYear="1860" box="[629,738,1850,1876]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="zeyheri">
<emphasis id="2E9FEAAFED1AFF90FDECF8E6F4FAB132" box="[629,738,1850,1876]" italics="true" pageId="10" pageNumber="203">P. zeyheri</emphasis>
</taxonomicName>
ranges in colour from pale pink to various shades of bright red and is much sought-after for making furniture and curios (
<bibRefCitation id="787A4B4CED1AFF90FC87F883F5C6B11E" author="Dyer, S. &amp; James, B. &amp; James, D." box="[798,990,1886,1912]" pageId="10" pageNumber="203" refId="ref9273" refString="Dyer, S., James, B. &amp; James, D. (2016) Guide to the properties and uses of southern African wood. Briza Publications, Pretoria, 336 pp." type="book" year="2016">
Dyer
<emphasis id="2E9FEAAFED1AFF90FCF9F882F581B11E" box="[864,921,1886,1912]" italics="true" pageId="10" pageNumber="203">et al.</emphasis>
2016
</bibRefCitation>
). On the other hand, the heartwood of
<taxonomicName id="DBEB4D3EED1AFF90FA0EF882F6FCB1FA" ID-CoL="77FR2" authority="Klotzsch 1862: 110" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1AFF90FA0EF882F6FCB1FA" italics="true" pageId="10" pageNumber="203">P. discolor</emphasis>
</taxonomicName>
(“brown ivory”/
<emphasis id="2E9FEAAFED1AFF90FE39F85FF400B1FA" box="[416,536,1922,1948]" italics="true" pageId="10" pageNumber="203">bruinivoor</emphasis>
) is yellowish brown to deep reddish brown. Heartwood of both species fluoresce when viewed under longwave ultraviolet light (ca.
<quantity id="DB139B58ED1AFF90FD23F87BF50AB1A6" box="[698,786,1958,1985]" metricMagnitude="-7" metricUnit="m" metricValue="3.65" pageId="10" pageNumber="203" unit="nm" value="365.0">365 nm</quantity>
), a distinction that has been successfully used to confirm the identification of commercial consignments of these two woods (
<bibRefCitation id="787A4B4CED1AFF90FCCEF817F5CCB182" author="Dyer, S. T." box="[855,980,1994,2020]" pageId="10" pageNumber="203" pagination="75 - 87" refId="ref9309" refString="Dyer, S. T. (1988) Wood fluorescence of indigenous South African trees. IAWA Bulletin n. s. 9: 75 - 87." type="book chapter" year="1988">Dyer 1988</bibRefCitation>
).
<taxonomicName id="DBEB4D3EED1AFF90FC71F817F2D7B182" ID-CoL="77G2Q" authorityName="Grubov" authorityYear="1949" baseAuthorityName="Sonder" baseAuthorityYear="1860" box="[1000,1231,1994,2020]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="zeyheri">
<emphasis id="2E9FEAAFED1AFF90FC71F817F2D7B182" box="[1000,1231,1994,2020]" italics="true" pageId="10" pageNumber="203">Phyllogeiton zeyheri</emphasis>
</taxonomicName>
has a characteristic orange fluorescence, while
<taxonomicName id="DBEB4D3EED1AFF90FE23F832F428BE6E" ID-CoL="77FR2" authority="Klotzsch 1862: 110" box="[442,560,2030,2056]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1AFF90FE23F832F428BE6E" box="[442,560,2030,2056]" italics="true" pageId="10" pageNumber="203">P. discolor</emphasis>
</taxonomicName>
fluoresces with a green colour. Because of its rarity, the colour and fluorescense of the mature heartwood of
<taxonomicName id="DBEB4D3EED1AFF90FE23F7CEF44EBE4A" ID-CoL="B8QZ7" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[442,598,2066,2092]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="203" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED1AFF90FE23F7CEF44EBE4A" box="[442,598,2066,2092]" italics="true" pageId="10" pageNumber="203">P. trachybasis</emphasis>
</taxonomicName>
still needs to be established with certainty. Heartwood in a transverse section of a long-dead branch (ca.
<quantity id="DB139B58ED1BFF91FE5BFF43F43AB6DF" box="[450,546,158,185]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="204" unit="mm" value="130.0">130 mm</quantity>
in diameter) of this species (
<figureCitation id="84D02A38ED1BFF91FCEEFF43F5D6B6DE" box="[887,974,158,184]" captionStart="FIGURE 2" captionStartId="5.[136,229,1955,1977]" captionTargetBox="[164,1423,190,1931]" captionTargetId="figure-18@5.[164,1423,190,1931]" captionTargetPageId="5" captionText="FIGURE 2. Phyllogeiton trachybasis, bark morphology illustrating variation (A–D) and colour of the wood (E). A. Bark on trunk of old, mature tree. B. Bark on trunk of relatively young tree (sunny habitat). C. Bark on trunk of relatively young tree (shady habitat). D. Exceedingly rough and flaking bark on trunk of mature tree. E. Finely sanded, untreated wood in transverse section; from dead branch ± 130 mm in diameter. Photographs: R.G.C. Boon (A, D), F. du Randt (B, C), &amp; D. Bishop (E)." figureDoi="http://doi.org/10.5281/zenodo.7690761" httpUri="https://zenodo.org/record/7690761/files/figure.png" pageId="11" pageNumber="204">Fig. 2E</figureCitation>
) is yellowish brown with a reddish tinge towards the sapwood, thus bearing a close resemblance to the heartwood of
<taxonomicName id="DBEB4D3EED1BFF91FC71FF1EF278B6BA" ID-CoL="77FR2" authority="Klotzsch 1862: 110" box="[1000,1120,194,220]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="11" pageNumber="204" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1BFF91FC71FF1EF278B6BA" box="[1000,1120,194,220]" italics="true" pageId="11" pageNumber="204">P. discolor</emphasis>
</taxonomicName>
. The narrow sapwood in this particular branch was bleached to a greyish colour by exposure to the elements and was also extensively damaged by wood boring insects.
</paragraph>
<caption id="48946635ED1BFF91FF11F806F330BE72" ID-DOI="http://doi.org/10.5281/zenodo.7690767" ID-Zenodo-Dep="7690767" httpUri="https://zenodo.org/record/7690767/files/figure.png" pageId="11" pageNumber="204" startId="11.[136,229,2011,2033]" targetBox="[177,1410,317,1986]" targetPageId="11">
<paragraph id="1C5436BDED1BFF91FF11F806F330BE72" blockId="11.[136,1452,2011,2069]" pageId="11" pageNumber="204">
<emphasis id="2E9FEAAFED1BFF91FF11F806F6E6B196" bold="true" box="[136,254,2011,2033]" pageId="11" pageNumber="204">FIGURE 6.</emphasis>
<taxonomicName id="DBEB4D3EED1BFF91FE9DF806F7CBB196" ID-CoL="77FR2" box="[260,467,2011,2032]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="11" pageNumber="204" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1BFF91FE9DF806F7CBB196" box="[260,467,2011,2032]" italics="true" pageId="11" pageNumber="204">Phyllogeiton discolor</emphasis>
</taxonomicName>
.
<emphasis id="2E9FEAAFED1BFF91FE46F806F7EFB196" bold="true" box="[479,503,2011,2032]" pageId="11" pageNumber="204">A.</emphasis>
Mature bark pattern.
<emphasis id="2E9FEAAFED1BFF91FD5EF806F4C6B196" bold="true" box="[711,734,2011,2032]" pageId="11" pageNumber="204">B.</emphasis>
Flowers.
<emphasis id="2E9FEAAFED1BFF91FCA4F806F54CB197" bold="true" box="[829,852,2011,2033]" pageId="11" pageNumber="204">C.</emphasis>
Fruit; note leaves with upper surface of blade glossy and dark green.
<emphasis id="2E9FEAAFED1BFF91FF50F822F6F9BE72" bold="true" box="[201,225,2047,2068]" pageId="11" pageNumber="204">D.</emphasis>
Leaves viewed from below, showing a greyish bloom. Photographs: R.G.C. Boon (A, C) &amp; A.E. van Wyk (B, D).
</paragraph>
</caption>
<caption id="48946635ED1CFF96FF11F87CF780B198" ID-DOI="http://doi.org/10.5281/zenodo.7690769" ID-Zenodo-Dep="7690769" httpUri="https://zenodo.org/record/7690769/files/figure.png" pageId="12" pageNumber="205" startId="12.[136,229,1953,1975]" targetBox="[151,1435,190,1929]" targetPageId="12">
<paragraph id="1C5436BDED1CFF96FF11F87CF780B198" blockId="12.[136,1452,1953,2047]" pageId="12" pageNumber="205">
<emphasis id="2E9FEAAFED1CFF96FF11F87CF6E7B1D0" bold="true" box="[136,255,1953,1975]" pageId="12" pageNumber="205">FIGURE 7.</emphasis>
<taxonomicName id="DBEB4D3EED1CFF96FE9FF87CF7D5B1D0" ID-CoL="77G2Q" authorityName="Grubov" authorityYear="1949" baseAuthorityName="Sonder" baseAuthorityYear="1860" box="[262,461,1953,1974]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="12" pageNumber="205" phylum="Tracheophyta" rank="species" species="zeyheri">
<emphasis id="2E9FEAAFED1CFF96FE9FF87CF7D5B1D0" box="[262,461,1953,1974]" italics="true" pageId="12" pageNumber="205">Phyllogeiton zeyheri</emphasis>
</taxonomicName>
.
<emphasis id="2E9FEAAFED1CFF96FE42F87CF7EBB1D0" bold="true" box="[475,499,1953,1974]" pageId="12" pageNumber="205">A.</emphasis>
Mature bark pattern.
<emphasis id="2E9FEAAFED1CFF96FD50F87CF4C7B1D0" bold="true" box="[713,735,1953,1974]" pageId="12" pageNumber="205">B.</emphasis>
Flowers; note leaves with upper surface of blade dull and with a greyish bloom.
<emphasis id="2E9FEAAFED1CFF96FF4BF818F6F1B1BD" bold="true" box="[210,233,1989,2011]" pageId="12" pageNumber="205">C.</emphasis>
Fruit.
<emphasis id="2E9FEAAFED1CFF96FEB3F818F75AB1BC" bold="true" box="[298,322,1989,2010]" pageId="12" pageNumber="205">D.</emphasis>
Leaves viewed from below, showing a greyish bloom similar to that of the upper surface. Photographs: R.G.C. Boon (A) &amp; A.E. van Wyk (B–D).
</paragraph>
</caption>
<paragraph id="1C5436BDED1DFF97FF24FF42F6F9B742" blockId="13.[136,1452,158,580]" pageId="13" pageNumber="206">
Two herbarium collections of
<taxonomicName id="DBEB4D3EED1DFF97FD97FF42F4B3B6DE" ID-CoL="B8QZ7" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[526,683,158,184]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="206" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED1DFF97FD97FF42F4B3B6DE" box="[526,683,158,184]" italics="true" pageId="13" pageNumber="206">P. trachybasis</emphasis>
</taxonomicName>
refer to ripe purple fruit (
<emphasis id="2E9FEAAFED1DFF97FC55FF43F258B6DE" box="[972,1088,158,184]" italics="true" pageId="13" pageNumber="206">
Moll
<quantity id="DB139B58ED1DFF97FB91FF43F258B6DE" box="[1032,1088,158,184]" metricMagnitude="2" metricUnit="m" metricValue="1.343914" pageId="13" pageNumber="206" unit="in" value="5291.0">5291</quantity>
</emphasis>
in Herbs K, NH, PRE) and pink fruit (
<emphasis id="2E9FEAAFED1DFF97FF52FF1FF74DB6BA" box="[203,341,194,220]" italics="true" pageId="13" pageNumber="206">
Gordon
<quantity id="DB139B58ED1DFF97FEB2FF1FF74DB6BA" box="[299,341,194,220]" metricMagnitude="0" metricUnit="m" metricValue="6.5786" pageId="13" pageNumber="206" unit="in" value="259.0">259</quantity>
</emphasis>
in Herbs NH, PRE). These observations are curious because in all other collections where fruit colour is mentioned it is recorded as yellow, which matches known images and our own field experience with the species.
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED1DFF97FF24FEF3F41DB422" pageId="13" pageNumber="206" type="description">
<paragraph id="1C5436BDED1DFF97FF24FEF3F41DB422" blockId="13.[136,1452,158,580]" pageId="13" pageNumber="206">
The conservation outlook for
<taxonomicName id="DBEB4D3EED1DFF97FD8CFEF2F4ACB72E" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[533,692,302,328]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="206" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED1DFF97FD8CFEF2F4ACB72E" box="[533,692,302,328]" italics="true" pageId="13" pageNumber="206">P. trachybasis</emphasis>
</taxonomicName>
seems poor. There are few known trees, the overall population is fragmented, most sub-populations consist of few individuals, a number of trees are dead or in poor health, and there is an apparent lack of recruitment. Given these population demographics and the excessive and destructive harvesting of bark, traditional biocultural uses of the species are clearly unsustainable. Conservation actions should include ensuring that trees in protected areas are not harvested, supplementing populations with cultivated specimens, and testing whether plants will recruit where browsing mammals are excluded. There is also a need to assess the current situation with trees in the dune forest between Lake Sibayi and
<collectingRegion id="DE2FF85FED1DFF97FCCCFDDBF597B446" box="[853,911,518,544]" country="South Africa" name="Western Cape" pageId="13" pageNumber="206">Cape</collectingRegion>
Vidal and the population at the Dlozi Peninsula in the iSimangaliso Wetland Park.
</paragraph>
</subSubSection>
<subSubSection id="54F16536ED1DFF97FF11FDAFF3B3B21A" pageId="13" pageNumber="206" type="key">
<paragraph id="1C5436BDED1DFF97FF11FDAFF4AEB4EB" blockId="13.[136,694,626,653]" box="[136,694,626,653]" pageId="13" pageNumber="206">
<emphasis id="2E9FEAAFED1DFF97FF11FDAFF4AEB4EB" bold="true" box="[136,694,626,653]" pageId="13" pageNumber="206">
Identification key to the species of
<taxonomicName id="DBEB4D3EED1DFF97FDBDFDAFF4AEB4EB" ID-CoL="7NCM2" authorityName="Herzog" authorityYear="1903" baseAuthorityName="Weberbauer" baseAuthorityYear="1895" box="[548,694,626,653]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="206" phylum="Tracheophyta" rank="genus">
<emphasis id="2E9FEAAFED1DFF97FDBDFDAFF4AEB4EB" bold="true" box="[548,694,626,653]" italics="true" pageId="13" pageNumber="206">Phyllogeiton</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<key id="687A618CED1DFF97FF11FD6AF3B3B21A" pageId="13" pageNumber="206">
<keyStep id="A71F2E18ED1DFF97FF11FD6AF3B3B53A" pageId="13" pageNumber="206">
<paragraph id="1C5436BDED1DFF97FF11FD6AF3B2B563" blockId="13.[136,1452,694,1149]" pageId="13" pageNumber="206">
<keyLead id="A71A9588ED1DFF97FF11FD6AF3B2B563" pageId="13" pageNumber="206">
1. Leaf blade usually to 40 ×
<quantity id="DB139B58ED1DFF97FE4CFD6BF404B4AA" box="[469,540,694,716]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="206" unit="mm" value="25.0">25 mm</quantity>
; flowers with ovary not immersed in disc; fruit ripening red (very often yellowish in immature fruit), to ca. 13 ×
<quantity id="DB139B58ED1DFF97FEE5FD0EF7AEB48F" box="[380,438,723,745]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="206" unit="mm" value="5.0">5 mm</quantity>
; receptacle at point of fruit stalk’s attachment at base of fruit markedly convex (see
<figureCitation id="84D02A38ED1DFF97FB42FD0EF330B48F" box="[1243,1320,723,745]" captionStart="FIGURE 4" captionStartId="7.[136,229,1953,1975]" captionTargetBox="[136,1450,190,1932]" captionTargetId="figure-18@7.[151,1436,190,1928]" captionTargetPageId="7" captionText="FIGURE 4. Phyllogeiton trachybasis (A–E), with fruit of P. discolor (F) and P. zeyheri (G). A. Fruiting branchlet (fruit almost mature). B. Flower. C. Petal. D. Stamen. E–G. Comparison of the fruit (bottom) of the three species of Phyllogeiton, with corresponding enlarged persistent receptacle and fruit stalk (top). Scale bar = 10 mm (A), 1 mm (B–D), or 4 mm (E–G). Drawings based on photographs and liquid-preserved fresh material deposited in Herb. PRU. Artist: Daleen Roodt." figureDoi="http://doi.org/10.5281/zenodo.7702415" httpUri="https://zenodo.org/record/7702415/files/figure.png" pageId="13" pageNumber="206">Figs 4G</figureCitation>
&amp;
<figureCitation id="84D02A38ED1DFF97FADEFD09F37CB48F" box="[1351,1380,724,745]" captionStart="FIGURE 7" captionStartId="12.[136,229,1953,1975]" captionTargetBox="[151,1435,190,1929]" captionTargetId="figure-19@12.[151,1435,190,1929]" captionTargetPageId="12" captionText="FIGURE 7. Phyllogeiton zeyheri. A. Mature bark pattern. B. Flowers; note leaves with upper surface of blade dull and with a greyish bloom. C. Fruit. D. Leaves viewed from below, showing a greyish bloom similar to that of the upper surface. Photographs: R.G.C. Boon (A) &amp; A.E. van Wyk (B–D)." figureDoi="http://doi.org/10.5281/zenodo.7690769" httpUri="https://zenodo.org/record/7690769/files/figure.png" pageId="13" pageNumber="206">7C</figureCitation>
) .......... ..............................................................................................................................................................................................
<taxonomicName id="DBEB4D3EED1DFF97FAD7FD2CF3B2B563" ID-CoL="77G2Q" authorityName="Grubov" authorityYear="1949" baseAuthorityName="Sonder" baseAuthorityYear="1860" box="[1358,1450,752,774]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="206" phylum="Tracheophyta" rank="species" species="zeyheri">
<emphasis id="2E9FEAAFED1DFF97FAD7FD2CF3B2B563" box="[1358,1450,752,774]" italics="true" pageId="13" pageNumber="206">P. zeyheri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1C5436BDED1DFF97FF11FCD3F3B3B53A" blockId="13.[136,1452,694,1149]" pageId="13" pageNumber="206">
<keyLead id="A71A9588ED1DFF97FF11FCD3F3B3B53A" pageId="13" pageNumber="206">
- Leaf blade usually to 100 ×
<quantity id="DB139B58ED1DFF97FE73FCD0F429B545" box="[490,561,781,803]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="206" unit="mm" value="40.0">40 mm</quantity>
; flowers with ovary immersed in disc; fruit ripening pale yellow, yellow, orange-yellow or brownish yellow,&gt;13 ×
<quantity id="DB139B58ED1DFF97FE27FCF7F7E1B559" box="[446,505,810,832]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="206" unit="mm" value="5.0">5 mm</quantity>
; receptacle at point of fruit stalk’s attachment at base of fruit plane or markedly concave (see
<figureCitation id="84D02A38ED1DFF97FA1BFCF7F702B53A" captionStart="FIGURE 4" captionStartId="7.[136,229,1953,1975]" captionTargetBox="[136,1450,190,1932]" captionTargetId="figure-18@7.[151,1436,190,1928]" captionTargetPageId="7" captionText="FIGURE 4. Phyllogeiton trachybasis (A–E), with fruit of P. discolor (F) and P. zeyheri (G). A. Fruiting branchlet (fruit almost mature). B. Flower. C. Petal. D. Stamen. E–G. Comparison of the fruit (bottom) of the three species of Phyllogeiton, with corresponding enlarged persistent receptacle and fruit stalk (top). Scale bar = 10 mm (A), 1 mm (B–D), or 4 mm (E–G). Drawings based on photographs and liquid-preserved fresh material deposited in Herb. PRU. Artist: Daleen Roodt." figureDoi="http://doi.org/10.5281/zenodo.7702415" httpUri="https://zenodo.org/record/7702415/files/figure.png" pageId="13" pageNumber="206">Figs 4E &amp; F</figureCitation>
) ..............................................................................................................................................................................................2.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A71F2E18ED1DFF97FF11FCB9F3B3B21A" pageId="13" pageNumber="206">
<paragraph id="1C5436BDED1DFF97FF11FCB9F3B3B58A" blockId="13.[136,1452,694,1149]" pageId="13" pageNumber="206">
<keyLead id="A71A9588ED1DFF97FF11FCB9F3B3B58A" pageId="13" pageNumber="206">
2. Bark towards base of trunk on mature trees very rough, flaking in large patches, almost black (sometimes with pale crustose lichen); leaves fresh green and without a bloom, ± concolorous, apex acuminate; flowers with petals spreading to ascending at anthesis; fruit usually ellipsoid, very often slightly ribbed longitudinally, rich orange-yellow, to 25 ×
<quantity id="DB139B58ED1DFF97FB0DFC40F2C1B5D5" box="[1172,1241,925,947]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="206" unit="mm" value="19.0">19 mm</quantity>
; receptacle at point of fruit stalk’s attachment at base of fruit plane (see
<figureCitation id="84D02A38ED1DFF97FD3FFC67F4E8B5A9" box="[678,752,954,976]" captionStart="FIGURE 4" captionStartId="7.[136,229,1953,1975]" captionTargetBox="[136,1450,190,1932]" captionTargetId="figure-18@7.[151,1436,190,1928]" captionTargetPageId="7" captionText="FIGURE 4. Phyllogeiton trachybasis (A–E), with fruit of P. discolor (F) and P. zeyheri (G). A. Fruiting branchlet (fruit almost mature). B. Flower. C. Petal. D. Stamen. E–G. Comparison of the fruit (bottom) of the three species of Phyllogeiton, with corresponding enlarged persistent receptacle and fruit stalk (top). Scale bar = 10 mm (A), 1 mm (B–D), or 4 mm (E–G). Drawings based on photographs and liquid-preserved fresh material deposited in Herb. PRU. Artist: Daleen Roodt." figureDoi="http://doi.org/10.5281/zenodo.7702415" httpUri="https://zenodo.org/record/7702415/files/figure.png" pageId="13" pageNumber="206">Fig. 4E</figureCitation>
); currently known with certainty only from
<collectingRegion id="DE2FF85FED1DFF97FB09FC67F331B5B6" box="[1168,1321,954,976]" country="South Africa" name="KwaZulu-Natal" pageId="13" pageNumber="206">KwaZulu-Natal</collectingRegion>
,
<collectingCountry id="64FC762DED1DFF97FAABFC67F3B3B5B6" box="[1330,1451,954,976]" name="South Africa" pageId="13" pageNumber="206">South Africa</collectingCountry>
(Maputaland Centre of Endemism) ...............................................................................................................................
<taxonomicName id="DBEB4D3EED1DFF97FABCFC0AF3B3B58A" ID-CoL="B8QZ7" authorityName="R. G. C. Boon &amp; A. E. van Wyk" authorityYear="2023" box="[1317,1451,983,1004]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="206" phylum="Tracheophyta" rank="species" species="trachybasis">
<emphasis id="2E9FEAAFED1DFF97FABCFC0AF3B3B58A" box="[1317,1451,983,1004]" italics="true" pageId="13" pageNumber="206">P. trachybasis</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="1C5436BDED1DFF97FF11FC29F3B3B21A" blockId="13.[136,1452,694,1149]" pageId="13" pageNumber="206">
<keyLead id="A71A9588ED1DFF97FF11FC29F3B3B21A" pageId="13" pageNumber="206">
- Bark towards base of trunk on mature trees rough, grey-brown, if flaking then in small patches; leaves dark green, distinctly paler with a greyish bloom below, apex acute to obtuse; flowers with petals reflexed at anthesis; fruit usually ovoid, surface plane, pale yellow to yellow, often turning brownish yellow when overripe, to 20 ×
<quantity id="DB139B58ED1DFF97FC0BFBF0F5C0B225" box="[914,984,1069,1091]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="206" unit="mm" value="11.0">11 mm</quantity>
; receptacle at point of fruit stalk’s attachment at base of fruit markedly concave (see
<figureCitation id="84D02A38ED1DFF97FDB0FB97F468B239" box="[553,624,1098,1120]" captionStart="FIGURE 4" captionStartId="7.[136,229,1953,1975]" captionTargetBox="[136,1450,190,1932]" captionTargetId="figure-18@7.[151,1436,190,1928]" captionTargetPageId="7" captionText="FIGURE 4. Phyllogeiton trachybasis (A–E), with fruit of P. discolor (F) and P. zeyheri (G). A. Fruiting branchlet (fruit almost mature). B. Flower. C. Petal. D. Stamen. E–G. Comparison of the fruit (bottom) of the three species of Phyllogeiton, with corresponding enlarged persistent receptacle and fruit stalk (top). Scale bar = 10 mm (A), 1 mm (B–D), or 4 mm (E–G). Drawings based on photographs and liquid-preserved fresh material deposited in Herb. PRU. Artist: Daleen Roodt." figureDoi="http://doi.org/10.5281/zenodo.7702415" httpUri="https://zenodo.org/record/7702415/files/figure.png" pageId="13" pageNumber="206">Fig. 4F</figureCitation>
); widespread, from southern through central, eastern and northeastern Africa to Arabia, also in
<collectingCountry id="64FC762DED1DFF97FE81FBBAF796B21B" box="[280,398,1127,1149]" name="Madagascar" pageId="13" pageNumber="206">Madagascar</collectingCountry>
, but not yet recorded in
<collectingRegion id="DE2FF85FED1DFF97FDEFFBBAF517B21B" box="[630,783,1127,1149]" country="South Africa" name="KwaZulu-Natal" pageId="13" pageNumber="206">KwaZulu-Natal</collectingRegion>
,
<collectingCountry id="64FC762DED1DFF97FC80FBBAF58BB21B" box="[793,915,1127,1149]" name="South Africa" pageId="13" pageNumber="206">South Africa</collectingCountry>
........................................................................
<taxonomicName id="DBEB4D3EED1DFF97FADDFBBAF3B3B21A" ID-CoL="77FR2" box="[1348,1451,1127,1148]" class="Magnoliopsida" family="Rhamnaceae" genus="Phyllogeiton" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="206" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="2E9FEAAFED1DFF97FADDFBBAF3B3B21A" box="[1348,1451,1127,1148]" italics="true" pageId="13" pageNumber="206">P. discolor</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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