treatments-xml/data/94/3E/87/943E87C0FFEDFFB3FECB6CC3FD28F639.xml

<document id="1D78AE7259BE341D1FEB769400874E00" ID-DOI="10.5281/zenodo.5372196" ID-ISSN="1638-9395" ID-Zenodo-Dep="5372196" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630271865429" checkinUser="marcus" docAuthor="Dumitrica, Paulian &amp; Zügel, Peter" docDate="2003" docId="943E87C0FFEDFFB3FECB6CC3FD28F639" docLanguage="en" docName="Geodiversitas.2003.25.1.5-72.pdf" docOrigin="Geodiversitas 25 (1)" docStyle="DocumentStyle:EA833BFD04EE0E3F39962C07E351CEA5.3:Geodiversitas.2000-2008.journal_article.type1" docStyleId="EA833BFD04EE0E3F39962C07E351CEA5" docStyleName="Geodiversitas.2000-2008.journal_article.type1" docStyleVersion="3" docTitle="Tetrarchiplagia ramosa Dumitrica &amp; Zügel 2003, n. sp." docType="treatment" docVersion="4" lastPageNumber="18" masterDocId="6807FFB8FFE6FFBEFFCA6929FFB2F469" masterDocTitle="Lower Tithonian mono- and dicyrtid Nassellaria (Radiolaria) from the Solnhofen area (southern Germany)" masterLastPageNumber="72" masterPageNumber="5" pageNumber="16" updateTime="1699207405888" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="60BFAE1A9F42CFF43AEA13019B4E6DF1">Lower Tithonian mono- and dicyrtid Nassellaria (Radiolaria) from the Solnhofen area (southern Germany)</mods:title>
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<mods:namePart id="EC0BA948A351D487526B39F788BCD9D9">Dumitrica, Paulian</mods:namePart>
<mods:affiliation id="22B7A96D5DA6331AA701D97A0A316EBC">Institut de Géologie et Paléontologie, BFSH 2, UNIL, CH- 1015 Lausanne (Switzerland) Paulian. Dumitrica @ igp. unil. ch</mods:affiliation>
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<mods:namePart id="BCA3CEF63B029448334DD9760950029A">Zügel, Peter</mods:namePart>
<mods:affiliation id="7D7F1D0332676D76B5714CF42798FF84">Geologisch-Paläontologisches Institut, Johann Wolfgang Goethe-Universität, Senckenberganlage 32 - 34, D- 60054 Frankfurt am Main (Germany) zuegel @ em. uni-frankfurt. de.</mods:affiliation>
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<mods:title id="25CFF824995E3BAA898A4D0A6171327A">Geodiversitas</mods:title>
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<mods:date id="43C01B7EF5DA39AD4CAC44C26D2B3CDB">2003</mods:date>
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<treatment id="943E87C0FFEDFFB3FECB6CC3FD28F639" LSID="urn:lsid:plazi:treatment:943E87C0FFEDFFB3FECB6CC3FD28F639" httpUri="http://treatment.plazi.org/id/943E87C0FFEDFFB3FECB6CC3FD28F639" lastPageId="13" lastPageNumber="18" pageId="11" pageNumber="16">
<subSubSection id="548D655DFFEDFFB5FECB6CC3FE69F24A" pageId="11" pageNumber="16" type="nomenclature">
<paragraph id="1C2836D6FFEDFFB5FECB6CC3FE69F24A" blockId="11.[151,684,1513,1669]" pageId="11" pageNumber="16">
<heading id="476081BAFFEDFFB5FECB6CC3FE69F24A" centered="true" fontSize="11" level="2" pageId="11" pageNumber="16" reason="2">
<taxonomicName id="DB974D55FFEDFFB5FECB6CC3FE4DF26A" authority="Dumitrica &amp; Zügel, 2003" authorityName="Dumitrica &amp; Zügel" authorityYear="2003" box="[257,511,1513,1540]" class="Polycystina" family="Plagiacanthidae" genus="Tetrarchiplagia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="11" pageNumber="16" phylum="Radiozoa" rank="species" species="ramosa" status="sp. nov.">
<emphasis id="2EE3EAC4FFEDFFB5FECB6CC3FE4DF26A" bold="true" box="[257,511,1513,1540]" italics="true" pageId="11" pageNumber="16">Tetrarchiplagia ramosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35D057BFFFEDFFB5FDCD6CC0FDF0F26A" box="[519,578,1513,1539]" pageId="11" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="84AC2A53FFEDFFB5FEA56F20FE1FF24A" box="[367,429,1545,1571]" captionStart="FIG" captionStartId="12.[151,162,1543,1560]" captionTargetBox="[194,1199,180,1510]" captionTargetId="figure-19@12.[194,1200,180,1510]" captionTargetPageId="12" captionText="FIG. 9. — A-I, Tetrarchiplagia ramosa n. sp.; A, holotype, right lateral view; B-I, paratypes; B, C, left lateral and apical views of the same specimen; D, E, left lateral view and detail of apical spine of the same specimen; F, right lateral view of a specimen showing a smaller angle between D and L; G, oblique ventral view; H, dorsal view of a small specimen with strongly curved D and L; I, left lateral view, detail of the central part showing the slender ventral spine and Ll broken off; J, Tetrarchiplagia tithoniana n. sp., paratype, apical view. Abbreviations: D, dorsal spine; L, primary lateral spine; Ll, left primary lateral spine. Scale bars: A-D, F-H, J, 100 µm; E, I, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.5372214" httpUri="https://zenodo.org/record/5372214/files/figure.png" pageId="11" pageNumber="16">Fig. 9</figureCitation>
A-I)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="548D655DFFEDFFB5FF5D6F1BFD28F20D" pageId="11" pageNumber="16" type="materials_examined">
<paragraph id="1C2836D6FFEDFFB5FF5D6F1BFD28F20D" blockId="11.[151,684,1513,1669]" pageId="11" pageNumber="16">
<materialsCitation id="ACFF3C8BFFEDFFB5FF5D6F1BFD28F20D" collectorName="Photo No. &amp; Lausanne, No." location="Musee de Geologie" pageId="11" pageNumber="16" specimenCount="1" typeStatus="holotype">
<typeStatus id="C32C8874FFEDFFB5FF5D6F1BFEB6F221" box="[151,260,1586,1609]" pageId="11" pageNumber="16" type="holotype">HOLOTYPE</typeStatus>
. — 
<collectorName id="B1625300FFEDFFB5FEFC6F1AFE16F220" box="[310,420,1586,1611]" pageId="11" pageNumber="16">Photo No.</collectorName>
58432; stub Mue 22/21; 
<location id="1948600DFFEDFFB5FF5D6F65FEE7F20D" LSID="urn:lsid:plazi:treatment:943E87C0FFEDFFB3FECB6CC3FD28F639:1948600DFFEDFFB5FF5D6F65FEE7F20D" box="[151,341,1612,1636]" name="Musee de Geologie" pageId="11" pageNumber="16">Musée de Géologie</location>
, 
<collectorName id="B1625300FFEDFFB5FE956F62FE5DF20B" box="[351,495,1611,1634]" pageId="11" pageNumber="16">Lausanne, No.</collectorName>
74376 (
<figureCitation id="84AC2A53FFEDFFB5FD886F65FD3DF20D" box="[578,655,1612,1636]" captionStart="FIG" captionStartId="12.[151,162,1543,1560]" captionTargetBox="[194,1199,180,1510]" captionTargetId="figure-19@12.[194,1200,180,1510]" captionTargetPageId="12" captionText="FIG. 9. — A-I, Tetrarchiplagia ramosa n. sp.; A, holotype, right lateral view; B-I, paratypes; B, C, left lateral and apical views of the same specimen; D, E, left lateral view and detail of apical spine of the same specimen; F, right lateral view of a specimen showing a smaller angle between D and L; G, oblique ventral view; H, dorsal view of a small specimen with strongly curved D and L; I, left lateral view, detail of the central part showing the slender ventral spine and Ll broken off; J, Tetrarchiplagia tithoniana n. sp., paratype, apical view. Abbreviations: D, dorsal spine; L, primary lateral spine; Ll, left primary lateral spine. Scale bars: A-D, F-H, J, 100 µm; E, I, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.5372214" httpUri="https://zenodo.org/record/5372214/files/figure.png" pageId="11" pageNumber="16">Fig. 9A</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="548D655DFFEDFFB5FF5D6F45FD20F2EC" box="[151,658,1644,1669]" pageId="11" pageNumber="16" type="etymology">
<paragraph id="1C2836D6FFEDFFB5FF5D6F45FD20F2EC" blockId="11.[151,684,1513,1669]" box="[151,658,1644,1669]" pageId="11" pageNumber="16">
ETYMOLOGY. — From the Latin 
<emphasis id="2EE3EAC4FFEDFFB5FE126F45FD90F2EA" box="[472,546,1644,1667]" italics="true" pageId="11" pageNumber="16">ramosus</emphasis>
: branched.
</paragraph>
</subSubSection>
<subSubSection id="548D655DFFEDFFB5FD006985FCC1F5EC" pageId="11" pageNumber="16" type="materials_examined">
<paragraph id="1C2836D6FFEDFFB5FD006985FC78F4B7" blockId="11.[714,1247,172,389]" pageId="11" pageNumber="16">
MATERIAL EXAMINED. — 
<specimenCount id="0A91FD5FFFEDFFB5FC036985FBFCF4AD" box="[969,1102,172,196]" pageId="11" pageNumber="16" type="generic">16 specimens</specimenCount>
from the 
<typeStatus id="C32C8874FFEDFFB5FB7F6985FB6DF4AD" box="[1205,1247,172,196]" pageId="11" pageNumber="16">type</typeStatus>
horizon (sample Mue 22).
</paragraph>
<paragraph id="1C2836D6FFEDFFB5FD0069C4FCC1F5EC" blockId="11.[714,1247,172,389]" pageId="11" pageNumber="16">
DIMENSIONS (
<collectionCode id="7A86AE13FFEDFFB5FC9D69DAFCDEF56A" box="[855,876,243,259]" pageId="11" pageNumber="16">IN</collectionCode>
µM). — Length of apical spine 140- 235 (av. 195), of dorsal spine 190-245 (av. 225), of primary lateral spines 160-245 (av. 210), of ventral spine 20-75 (av. 50), of median bar 4-6; distance of first verticil from 
<collectionCode id="7A86AE13FFEDFFB5FCB0687AFC12F503" box="[890,928,339,362]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="11" pageNumber="16">MB</collectionCode>
on D and L 30-50 (av. 40), on A 55-80 (av. 65).
</paragraph>
</subSubSection>
<subSubSection id="548D655DFFEDFFB5FD006883FB6DF02C" pageId="11" pageNumber="16" type="description">
<paragraph id="1C2836D6FFEDFFB5FD006883FCE9F5AB" blockId="11.[714,1248,426,1093]" box="[714,859,426,452]" pageId="11" pageNumber="16">DESCRIPTION</paragraph>
<paragraph id="1C2836D6FFEDFFB5FD0068E2FB6DF02C" blockId="11.[714,1248,426,1093]" pageId="11" pageNumber="16">Spicule with apical, dorsal, ventral, and both primary lateral spines originating in a very short median bar. A, D, and L stout, long, subequal, with a number of verticils oriented at about right angles to the spines, and generally decreasing in length to the distal part of the spines. Ventral spine small, thin, directed obliquely upward, distally slightly curved downwards when longer. Apical spine straight with three verticils of which the first has generally five branches, the second three, rarely more, and the third two or three. Branches of first and second verticils with one or two verticils of spinules. Dorsal and primary lateral spines equal, longer than A, and curved downward. They bear four verticils of which the first has generally three branches, the second two or three, and the third and fourth two branches. Branches of first verticil, rarely of the second, bear spinules. Except V all spines increase slowly in diametre to the last verticil, then taper distally.</paragraph>
</subSubSection>
<subSubSection id="548D655DFFEDFFB3FD006D43FD28F639" lastPageId="13" lastPageNumber="18" pageId="11" pageNumber="16" type="discussion">
<paragraph id="1C2836D6FFEDFFB5FD006D43FC9CF0EB" blockId="11.[714,1248,1130,1669]" box="[714,814,1130,1156]" pageId="11" pageNumber="16">REMARKS</paragraph>
<paragraph id="1C2836D6FFEDFFB5FD006DA2FB7EF2EC" blockId="11.[714,1248,1130,1669]" pageId="11" pageNumber="16">
<taxonomicName id="DB974D55FFEDFFB5FD006DA2FC0EF0CA" authority="Dumitrica &amp; Zügel, 2003" authorityName="Dumitrica &amp; Zügel" authorityYear="2003" box="[714,956,1162,1189]" class="Polycystina" family="Plagiacanthidae" genus="Tetrarchiplagia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="11" pageNumber="16" phylum="Radiozoa" rank="species" species="ramosa" status="sp. nov.">
<emphasis id="2EE3EAC4FFEDFFB5FD006DA2FC0EF0CA" box="[714,956,1162,1189]" italics="true" pageId="11" pageNumber="16">Tetrarchiplagia ramosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35D057BFFFEDFFB5FC0E6DA3FBB3F0CD" box="[964,1025,1162,1188]" pageId="11" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
resembles somehow the Triassic species 
<taxonomicName id="DB974D55FFEDFFB5FC7F6D83FCB7F08C" authority="Dumitrica, 1982" authorityName="Dumitrica" authorityYear="1982" class="Polycystina" family="Plagiacanthidae" genus="Tetrarchiplagia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="11" pageNumber="16" phylum="Radiozoa" rank="species" species="verticillata">
<emphasis id="2EE3EAC4FFEDFFB5FC7F6D83FC75F0AA" box="[949,967,1194,1219]" italics="true" pageId="11" pageNumber="16">T</emphasis>
. 
<emphasis id="2EE3EAC4FFEDFFB5FC136D82FBE1F0AC" box="[985,1107,1195,1221]" italics="true" pageId="11" pageNumber="16">verticillata</emphasis>
Dumitrica, 1982
</taxonomicName>
and 
<taxonomicName id="DB974D55FFEDFFB5FC896DE3FB08F08C" authority="Dumitrica, 1982" authorityName="Dumitrica" authorityYear="1982" box="[835,1210,1226,1253]" class="Polycystina" family="Plagiacanthidae" genus="Tetrarchiplagia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="11" pageNumber="16" phylum="Radiozoa" rank="species" species="abietinoides">
<emphasis id="2EE3EAC4FFEDFFB5FC896DE3FCE7F08A" box="[835,853,1226,1251]" italics="true" pageId="11" pageNumber="16">T</emphasis>
. 
<emphasis id="2EE3EAC4FFEDFFB5FCAC6DE2FC58F08C" box="[870,1002,1227,1253]" italics="true" pageId="11" pageNumber="16">abietinoides</emphasis>
Dumitrica, 1982
</taxonomicName>
by having verticils of three or four branches, but differs from both by the number of verticils, size of branches, presence of spinules on some branches, etc. Together with the following species, this is the first Jurassic species of the family 
<taxonomicName id="DB974D55FFEDFFB5FD006CA2FCC1F1CC" authorityName="Hertwig" authorityYear="1879" box="[714,883,1419,1445]" class="Polycystina" family="Plagiacanthidae" kingdom="Chromista" order="Nassellaria" pageId="11" pageNumber="16" phylum="Radiozoa" rank="family">Plagiacanthidae</taxonomicName>
so far described bridging the preexisting long gap between the Middle Triassic and the Paleogene. It is noteworthy, however, that they are not the only species of the family occurring in the Jurassic. Work in progress of the senior author proves that, although much sparser than in the Middle Triassic, at least three more species occur in the Lower and Middle Jurassic.
</paragraph>
<caption id="48E8665EFFEAFFB2FF5D6F2EFF45F2EA" ID-DOI="http://doi.org/10.5281/zenodo.5372214" ID-Zenodo-Dep="5372214" httpUri="https://zenodo.org/record/5372214/files/figure.png" pageId="12" pageNumber="17" startId="12.[151,162,1543,1560]" targetBox="[194,1199,180,1510]" targetPageId="12">
<paragraph id="1C2836D6FFEAFFB2FF5D6F2EFF45F2EA" blockId="12.[151,1249,1543,1667]" pageId="12" pageNumber="17">
FIG. 9. — 
<emphasis id="2EE3EAC4FFEAFFB2FF206F2EFEB6F271" bold="true" box="[234,260,1543,1560]" pageId="12" pageNumber="17">A -I</emphasis>
, 
<taxonomicName id="DB974D55FFEAFFB2FEC56F2EFE7FF271" authority="Dumitrica &amp; Zügel, 2003" authorityName="Dumitrica &amp; Zügel" authorityYear="2003" box="[271,461,1543,1560]" class="Polycystina" family="Plagiacanthidae" genus="Tetrarchiplagia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="12" pageNumber="17" phylum="Radiozoa" rank="species" species="ramosa" status="sp. nov.">
<emphasis id="2EE3EAC4FFEAFFB2FEC56F2EFE7FF271" box="[271,461,1543,1560]" italics="true" pageId="12" pageNumber="17">Tetrarchiplagia ramosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35D057BFFFEAFFB2FE196F2EFDB4F271" box="[467,518,1543,1560]" pageId="12" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
; 
<emphasis id="2EE3EAC4FFEAFFB2FDC46F2EFDA9F271" bold="true" box="[526,539,1543,1560]" pageId="12" pageNumber="17">A</emphasis>
, holotype, right lateral view; 
<emphasis id="2EE3EAC4FFEAFFB2FCC46F2EFC9BF271" bold="true" box="[782,809,1543,1560]" pageId="12" pageNumber="17">B -I</emphasis>
, paratypes; 
<emphasis id="2EE3EAC4FFEAFFB2FC586F2EFC2DF271" bold="true" box="[914,927,1543,1560]" pageId="12" pageNumber="17">B</emphasis>
, 
<emphasis id="2EE3EAC4FFEAFFB2FC606F2EFC0AF271" bold="true" box="[938,952,1543,1560]" pageId="12" pageNumber="17">C</emphasis>
, left lateral and apical views of the same specimen; 
<emphasis id="2EE3EAC4FFEAFFB2FEEF6F34FE81F247" bold="true" box="[293,307,1565,1582]" pageId="12" pageNumber="17">D</emphasis>
, 
<emphasis id="2EE3EAC4FFEAFFB2FEF76F34FEFBF247" bold="true" box="[317,329,1565,1582]" pageId="12" pageNumber="17">E</emphasis>
, left lateral view and detail of apical spine of the same specimen; 
<emphasis id="2EE3EAC4FFEAFFB2FCA76F34FCCAF247" bold="true" box="[877,888,1565,1582]" pageId="12" pageNumber="17">F</emphasis>
, right lateral view of a specimen showing a smaller angle between D and L; 
<emphasis id="2EE3EAC4FFEAFFB2FE6B6F1BFE1DF22A" bold="true" box="[417,431,1586,1603]" pageId="12" pageNumber="17">G</emphasis>
, oblique ventral view; 
<emphasis id="2EE3EAC4FFEAFFB2FDA26F1BFDC4F22A" bold="true" box="[616,630,1586,1603]" pageId="12" pageNumber="17">H</emphasis>
, dorsal view of a small specimen with strongly curved D and L; 
<emphasis id="2EE3EAC4FFEAFFB2FB4F6F1BFB39F22A" bold="true" box="[1157,1163,1586,1603]" pageId="12" pageNumber="17">I</emphasis>
, left lateral view, detail of the central part showing the slender ventral spine and Ll broken off; 
<emphasis id="2EE3EAC4FFEAFFB2FCA26F6EFCC0F231" bold="true" box="[872,882,1607,1624]" pageId="12" pageNumber="17">J</emphasis>
, 
<taxonomicName id="DB974D55FFEAFFB2FCB46F6EFBFDF231" authority="Dumitrica &amp; Zügel, 2003" authorityName="Dumitrica &amp; Zügel" authorityYear="2003" box="[894,1103,1607,1624]" class="Polycystina" family="Plagiacanthidae" genus="Tetrarchiplagia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="12" pageNumber="17" phylum="Radiozoa" rank="species" species="tithoniana" status="sp. nov.">
<emphasis id="2EE3EAC4FFEAFFB2FCB46F6EFBFDF231" box="[894,1103,1607,1624]" italics="true" pageId="12" pageNumber="17">Tetrarchiplagia tithoniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35D057BFFFEAFFB2FB9C6F6EFB3BF231" box="[1110,1161,1607,1624]" pageId="12" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
, paratype, apical view. Abbreviations: 
<emphasis id="2EE3EAC4FFEAFFB2FEB46F74FE3EF207" bold="true" box="[382,396,1629,1646]" pageId="12" pageNumber="17">D</emphasis>
, dorsal spine; 
<emphasis id="2EE3EAC4FFEAFFB2FDC36F74FDA6F207" bold="true" box="[521,532,1629,1646]" pageId="12" pageNumber="17">L</emphasis>
, primary lateral spine; 
<emphasis id="2EE3EAC4FFEAFFB2FD1D6F74FD55F207" bold="true" box="[727,743,1629,1646]" pageId="12" pageNumber="17">Ll</emphasis>
, left primary lateral spine. Scale bars: A-D, F-H, J, 100 µm; E, I, 30 µm.
</paragraph>
</caption>
<paragraph id="1C2836D6FFEBFFB3FD416B15FD28F639" blockId="13.[651,666,572,592]" box="[651,666,572,592]" pageId="13" pageNumber="18">D</paragraph>
</subSubSection>
</treatment>
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