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<document id="90A9B19AAEA07B0774FC413434E8350E" ID-CLB-Dataset="302074" ID-DOI="10.3897/phytokeys.246.129476" ID-GBIF-Dataset="b381880a-e0bb-4b02-ad0f-2f90a0b6c995" ID-publisher-id="129476" URI-arpha="7631F16E-4CD8-5A76-9F28-45E5F398CE20" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1725298343909" checkinUser="pensoft" docAuthor="Meerts, Pierre J. &amp; Paton, Alan J." docDate="2024" docId="94284EB1C58150AF96A8792D3DB21BC9" docLanguage="en" docName="PhytoKeys 246: 71-178" docOrigin="PhytoKeys 246" docSource="https://phytokeys.pensoft.net/article/129476/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Coleus shirensis Gurke" docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="178" masterDocId="7631F16E4CD85A769F2845E5F398CE20" masterDocTitle="The genus Coleus (Lamiaceae) in Central Africa (Democratic Republic of the Congo, Rwanda, Burundi), with the description of 15 new species" masterLastPageNumber="178" masterPageNumber="71" pageNumber="71" updateTime="1725322260938" updateUser="ExternalLinkService">
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<mods:title id="A121C91D78AF27F0C88D9AD4DD8CE7DD">The genus Coleus (Lamiaceae) in Central Africa (Democratic Republic of the Congo, Rwanda, Burundi), with the description of 15 new species</mods:title>
</mods:titleInfo>
<mods:name id="BFADAFA688FFD874B8744B6A4CE01B09" type="personal">
<mods:role id="7AD0B38271C35E5AD0BD1860B8CEB531">
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<mods:namePart id="82A1973E2534E090500C201789A811D3">Meerts, Pierre J.</mods:namePart>
<mods:nameIdentifier id="7DA5C55FCA37EAF48D57197976C69131" type="ORCID">0000-0003-4215-027X</mods:nameIdentifier>
<mods:affiliation id="3D4B087470A65A8AA34B639DA5F81F04">Meise Botanic Garden, Nieuwelaan 38, BE- 1860 Meise, Belgium &amp; Université Libre de Bruxelles, Av. F. D. Roosevelt 50 CP 244, BE- 1050 Brussels, Belgium &amp; Fédération Wallonie-Bruxelles, Service général de lEnseignement supérieur et de la Recherche scientifique, Brussels, Belgium</mods:affiliation>
</mods:name>
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<mods:role id="6B53369106BA6DE9575919195DE263DF">
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</mods:role>
<mods:namePart id="A8AA3EA73D2143CAA6D120D388E26537">Paton, Alan J.</mods:namePart>
<mods:nameIdentifier id="24FCAACCA454365CC614184B82AD5132" type="ORCID">0000-0002-6052-6675</mods:nameIdentifier>
<mods:affiliation id="78F417A9D1FA7E910AA50DE3BA6428A3">The Herbarium, Royal Botanic Gardens, Kew, Richmond TW 9 3 AB, UK</mods:affiliation>
</mods:name>
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<mods:titleInfo id="C053D14C49D8981E4812B49845B0F024">
<mods:title id="719E3F9184B2BD205A907438825DF8AB">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="784D08A9A8D600F50883F1BB28B55F5A">
<mods:date id="1B3A792052F1C77961FB8F20E1CCF0A4">2024</mods:date>
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<mods:number id="FB4CB6213DFA9719E9D06992E774C5F4">2024-09-02</mods:number>
</mods:detail>
<mods:detail id="0A6141F1A35C12B8B2BA4EBBB23540F0" type="volume">
<mods:number id="9484FBEFC2CC13E9F6B170E7843555FB">246</mods:number>
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</mods:extent>
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<mods:classification id="FBE743C53FEE594263C843FA6E0FA906">journal article</mods:classification>
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<mods:identifier id="E51B45CD704BF6702C513C9783897FBE" type="DOI">10.3897/phytokeys.246.129476</mods:identifier>
<mods:identifier id="1DD59D9A03AE7FA4D4CEC840EBF612FF" type="GBIF-Dataset">b381880a-e0bb-4b02-ad0f-2f90a0b6c995</mods:identifier>
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<treatment id="94284EB1C58150AF96A8792D3DB21BC9" ID-DOI="http://doi.org/10.5281/zenodo.13629900" ID-GBIF-Taxon="240061412" ID-Zenodo-Dep="13629900" ID-arpha="94284EB1-C581-50AF-96A8-792D3DB21BC9" LSID="urn:lsid:plazi:treatment:94284EB1C58150AF96A8792D3DB21BC9" httpUri="http://treatment.plazi.org/id/94284EB1C58150AF96A8792D3DB21BC9">
<subSubSection id="685E8E7FC9C1BC32E1EFD1AECC5B6501" type="nomenclature">
<paragraph id="61ED8DACB1096DC09EC2DEAE7C3B0DF4">
<treatmentCitation id="07EA01789649338F2F911DC459C3C0C4" author="Gurke" journal="Bot. Jahrb. Syst." page="216" score="144.29617767960897" volume="19" year="1894">
<taxonomicName id="3DA3F73998BDBCCD910A271C861C3038" ID-CoL="5ZLNR" authority="Gurke" authorityName="Gurke" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="shirensis">
<emphasis id="B66657058E77B84D2C426DDC3921C230" italics="true">Coleus shirensis</emphasis>
Gürke
</taxonomicName>
, Bot. Jahrb. Syst. 19: 216. 1894
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A965F85A5BABE5D6494842662761902E" type="reference_group">
<paragraph id="D97252EA006EC4748ABA35BC29C7D5A0">
<treatmentCitationGroup id="36B5E89D278A1E7DCFA9006712473227">
<treatmentCitation id="B3A2E8AA2A5416D4B317877E4067F218" author="A. J. Paton" journal="SABONET Rep. Ser." page="189" score="0.30547891401646055" volume="31" year="2005">
<taxonomicName id="2DAB4C481FE3B85D82051FC7FB0E6055" ID-CoL="4K4LV" authority="(Gurke) A. J. Paton" authorityName="A. J. Paton" baseAuthorityName="Gurke" class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="shirensis">
<emphasis id="3696B5AA699D191BB74167B45E07A55A" italics="true">Plectranthus shirensis</emphasis>
(Gürke) A. J. Paton
</taxonomicName>
, SABONET Rep. Ser. 31: 189. 2005
</treatmentCitation>
. Type:
<collectingCountry id="05CCB5F380680F236E28F75FA23305B7" name="Malawi">Malawi</collectingCountry>
, 1891,
<emphasis id="5D25BE66E4C38B42054F6AD6BB6F60E2" italics="true">J. Buchanan 376</emphasis>
(
<typeStatus id="1DD8626F2AC94AB105BF6C563B07BA80" type="lectotype">lectotype</typeStatus>
<collectionCode id="18EB3268F897943D896877D02A5AAA48" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
[
<specimenCode id="3A28BAD2A2CA50CD46E057AF49504875">
<collectionCode id="A3B7BD0FF48DBB6B68F8DF88419FA7A9" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
000070492
</specimenCode>
];
<typeStatus id="68061B67384F07D62BE07F2BBD616AAA" type="isolectotype">isolectotype</typeStatus>
<collectionCode id="BF27B2B6D85F6B1387CC91336E8FD1EC" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" type="Herbarium">E</collectionCode>
[
<collectionCode id="1854BB64525591AFE4500BE1D61CBBAC" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" type="Herbarium">E</collectionCode>
00193509], designated by
<bibRefCitation id="0F68C4B8DCC8848354EAC5F615656760" DOI="10.2307/4109002" author="Mathew B" firstAuthor="Mathew" refId="B16" refString="Mathew B (1976) Notes on some Solenostemon and Plectranthus (Labiatae). Kew Bulletin 31: 174. https://doi.org/10.2307/4109002" year="1976">Mathew [1976]</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3CIM" type="description">
<paragraph id="3A98ADC8CA5DD8665454665E14E2E345">
<heading id="0C58FA1D20B9AB3BF60037450C2B9030" reason="title">Description.</heading>
</paragraph>
<paragraph id="15BE4587B902C255C5E5B505A8BC68B6">
<bibRefCitation id="F0538E19E828FD27F368FB87D589EE1D" author="Paton AJ &amp; Bramley G &amp; Ryding O &amp; Polhill RM &amp; Harvey YB &amp; Iwarsson M &amp; Willis F &amp; Phillipson PB &amp; Balkwill K &amp; Lukhoba CW &amp; Otieno D &amp; Harley RM" editor="Beentje HJ &amp; Ghazanfar SA &amp; Polhill RM" etAl="et al." firstAuthor="Paton" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" page="336" pagination="1-430" refId="B20" refString="Paton AJ, Bramley G, Ryding O, Polhill RM, Harvey YB, Iwarsson M, Willis F, Phillipson PB, Balkwill K, Lukhoba CW, Otieno D, Harley RM (2009) Lamiaceae (Labiatae). In: Beentje HJ, Ghazanfar SA, Polhill RM (Eds) Flora of Tropical East Africa. Royal Botanic Gardens, Kew, Richmond, 1430." title="Lamiaceae (Labiatae)." volumeTitle="Flora of Tropical East Africa." year="2009">Paton et al. (2009: 336)</bibRefCitation>
,
<bibRefCitation id="0650C08655367FC11CF38CF6BBDB85C6" author="Paton AJ &amp; Bramley G &amp; Ryding O &amp; Polhill RM &amp; Harvey YB &amp; Iwarsson M &amp; Willis F &amp; Phillipson PB &amp; Balkwill K &amp; Lukhoba CW &amp; Otieno D &amp; Harley RM" editor="Timberlake J" etAl="et al." firstAuthor="Paton" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" page="272" pagination="1-346" refId="B21" refString="Paton AJ, Bramley G, Ryding O, Polhill RM, Harvey YB, Iwarsson M, Willis F, Phillipson PB, Balkwill K, Lukhoba CW, Otieno D, Harley RM (2013) Lamiaceae. In: Timberlake J (Ed.) Flora Zambesiaca 8 (8). Royal Botanic Gardens, Kew, Richmond, 1346." title="Lamiaceae." volumeTitle="Flora Zambesiaca 8 (8)." year="2013">Paton et al. (2013: 272)</bibRefCitation>
, as
<taxonomicName id="AAB300FF255A18F1B61CC02D8194FE0B" authority="(Gurke) A. J. Paton." authorityName="A. J. Paton." baseAuthorityName="Gurke" class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="shirensis">
<emphasis id="022FE500589FA612C8DEDAD930B51458" italics="true">Plectranthus shirensis</emphasis>
(Gürke) A. J. Paton.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVDIM" type="distribution">
<paragraph id="AC0C4FED959BCAA4AEEFC245ABCE7B32">
<heading id="F31EF70FE340ED8134B1765B1D4BF4DF" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="C57DDB7E58549E18D01339052AC5708E">
<collectionCode id="632CBCD4DE2853C77B618F53FA5CAC00">SW</collectionCode>
&amp; S
<collectingCountry id="6D5FD9BA97C43EACEA3D3018A431DEEF" name="Tanzania">Tanzania</collectingCountry>
to S Tropical Africa.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1DIM" type="habitat">
<paragraph id="5E551C2386DACE294DF28B61BD465A90">
<heading id="91EEE56B39A5ABB68F55EF2F0B00ADAB" reason="title">Habitat and еcology.</heading>
</paragraph>
<paragraph id="55B5A6775993BAB8304FBD98A513D91D">
Riparian forest, swamp savannah, forest margins;
<quantity id="CDCF4545040294DC339884F387394576" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" unit="m" value="1400.0" valueMax="1800.0" valueMin="1000.0">10001800 m</quantity>
elev.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6DIM" type="Additional specimens">
<paragraph id="CDEC55B68F923656C81663FEB8083185">
<heading id="DBA789E0C254928434FA6C3DD5623E3A" reason="title">Additional specimens.</heading>
</paragraph>
<paragraph id="FC47761D29EDBF3077A7BFFB3159A5A6">
<collectingCountry id="56C8915BEA0BDEB18C680047C7DDD91B">
DR.
<collectingRegion id="EB9BA88B10DE742E3C10FAF5105BE9CE" country="Republic of the Congo" name="Bas-Congo">Congo</collectingRegion>
</collectingCountry>
,
<emphasis id="AFF1AD5F8F41972AF0D6ACAC39DF5F18" bold="true" italics="true">
<collectingRegion id="671EE6F64A4284929F71E245315DE296" country="Republic of the Congo" name="Bas-Congo">Bas-Congo</collectingRegion>
</emphasis>
, Kinkosi,
<date id="11FF94188F722F645E933D9B7E5B3C4C" value="1959-02-26">26 Feb 1959</date>
,
<emphasis id="AC392B5E5FBD53BE716C441E59F494FA" italics="true">L. Pauwels 1600</emphasis>
(
<collectingRegion id="76503F123BCE88F64C09FDDDC9826C61" country="Serbia" name="Bor">BR</collectingRegion>
);
<emphasis id="6FDB69C62C8559F1B5FAE075414E54E0" bold="true" italics="true">
<collectingRegion id="F07948964BEE8FC4C98C403E7DC92771" country="Republic of the Congo" name="Kasai-Oriental">Kasaï</collectingRegion>
</emphasis>
, Entre Kwango-Lufuna,
<date id="2F800DBA6D39570824E93F3E6B3514EB" value="1944-08-04">4 Aug 1944</date>
,
<emphasis id="F845AFC21AAF762A1FAF72C352462374" italics="true">
<collectionCode id="BE686B8DCBF161F834E54D3483E1B9AB" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Germain 2643
</emphasis>
(
<collectingRegion id="BEEB47A6B1239F260C58F90106735E37" country="Serbia" name="Bor">BR</collectingRegion>
);
<collectingRegion id="CDB16305962AE27476290A6B5CFD572D" country="Republic of the Congo" name="Bas-Congo">Kwango</collectingRegion>
,
<date id="468B6883E91F106A4F5627F0C2E95E5C" value="1955-07-21">21 Jul 1955</date>
,
<emphasis id="95AD146DD1B860977688C0AFC658A454" italics="true">
<collectionCode id="3AB6333EDEE3927017B0EE854E46084D" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Devred 2290
</emphasis>
(
<collectingRegion id="322560AFCC9B639954A63D28F4C38A2F" country="Serbia" name="Bor">BR</collectingRegion>
);
<emphasis id="43C44BED7C6E832E9ECCD33A94ABA565" bold="true" italics="true">Bas-Katanga</emphasis>
, Gandajika,
<date id="012E91190BCF9492A2B6947D7129D1B6" value="1951-06-20">20 Jun 1951</date>
,
<emphasis id="CADACB9016FEF19F819A14ED21B76CBC" italics="true">Chalon 326</emphasis>
(
<collectingRegion id="DB8BB7430A21A4961E83E6B10D260DC4" country="Serbia" name="Bor">BR</collectingRegion>
); Kindele,
<date id="6553E37BEB8F132832915F82A9CBBAF2" value="1951-09">Sep 1951</date>
,
<emphasis id="75F794A24954335A5C61DA4CBCB4FE35" italics="true">
<collectionCode id="AF9D49D927FD5B10FD1D39FB872B2602" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Desenfans 2013
</emphasis>
(
<collectingRegion id="6C71BC8707270FC0F5FC2439D248F334" country="Serbia" name="Bor">BR</collectingRegion>
,
<collectionCode id="632B6CE78DA82D3EA42D127F9AE4FEE3" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
);
<quantity id="B3676F8EB46F1171DCEF84066926AFF0" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
<collectionCode id="1FBD2EEF4CD1F2CF3F347EF767FF4947">SW</collectionCode>
Gandajika,
<date id="3BBB287813EB5FE1C5FBC6904EFA62F4" value="1945-08-09">9 Aug 1945</date>
,
<emphasis id="49FA67FF08217DD2336530C0FD151515" italics="true">F. Luxen 534</emphasis>
(
<collectingRegion id="3184398F269A070B8B010EA3A555C161" country="Serbia" name="Bor">BR</collectingRegion>
); Lovoi, Kamina,
<date id="A9CBD9CE2B57AC5919910843CDCA932C" value="1932-03">
<collectingRegion id="5F5D667B62ED4DF9D2D446F1D28FA60B" country="South Sudan" name="Lakes">Mar</collectingRegion>
1932
</date>
,
<emphasis id="0970571F7E6BDAF33EA3958F1E934EE7" italics="true">P. Quarré 2929</emphasis>
(
<collectingRegion id="EE9F241981EF12B1476B13E35AB40D3B" country="Serbia" name="Bor">BR</collectingRegion>
);
<emphasis id="0629CA9AD3492E1118AF708BF586F680" bold="true" italics="true">
<collectingRegion id="CF0467E6E6671A326B26899279414949">Haut-Katanga</collectingRegion>
</emphasis>
, Mulumbi, riv. Dona,
<date id="BA4BBB1FEA19913341301A52CBD3C95B" value="1953-08-30">30 Aug 1953</date>
,
<emphasis id="6C6AC36D4A09C9FEF603638E083A4AE7" italics="true">
<collectionCode id="5A8542C41C9A04541E3D25B9FCFFA799" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Desenfans 3731
</emphasis>
(
<collectionCode id="322B2ED870D94FDF3411CBE5679D3A40" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
);
<quantity id="49B20C9B247E7AB3241B0E2008D520D9" metricMagnitude="4" metricUnit="m" metricValue="1.4" unit="km" value="14.0">14 km</quantity>
N de Mitwaba,
<date id="5ECCC46B0EB3F2413219A86EA62FAF08" value="1960-01-16">16 Jan 1960</date>
,
<emphasis id="973B9A320CAD8DC92C884DEA1E4186FF" italics="true">P. Duvigneaud 5077 Co</emphasis>
(
<collectionCode id="5BF5F0BF0C99E1F66109D629A0193498" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
); Kundelungu,
<quantity id="C3D6E0DCF8DB821AACB8F5A1A046B187" metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
NNW poste Katshupa,
<date id="327FC5B9FAA3F53A9090EE329A00E413" value="1966-07-29">29 Jul 1966</date>
,
<emphasis id="2149725F35AAEDE0D5D36FA76C45C1CC" italics="true">F. Malaisse 4213 a</emphasis>
(
<collectingRegion id="22C26787774AA25AEFD8D72627653BF9" country="Serbia" name="Bor">BR</collectingRegion>
;
<collectionCode id="E6B88E99D0C531C75882350840FA1E1F" country="Democratic Republic of the Congo" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14451" lsid="urn:lsid:biocol.org:col:14451" name="Université Nationale du Zaïre" type="Herbarium">LSHI</collectionCode>
); Kisanga,
<date id="F3693A79AF4678A79F0F21C0AE805920" value="1933-08">Aug 1933</date>
,
<emphasis id="AC51455B06A574518B53A02D04F650C2" italics="true">P. Quarré 3466</emphasis>
(
<collectingRegion id="A56E086C7221500D5F62DD2BC812E21A" country="Serbia" name="Bor">BR</collectingRegion>
); Upemba, Lusinga, route de Mitwaba,
<date id="2F86D41231B91C01113167D38EE8928F" value="1948-09-14">14 Sep 1948</date>
,
<emphasis id="498AB88B9A246333AF40051521C2A6FD" italics="true">
<collectionCode id="B9A78752FD2F01FD283B08C3500BCA68" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" type="Herbarium">W</collectionCode>
. Robyns 3590
</emphasis>
(
<collectingRegion id="1B201946A8785A206A04FD19A15F1085" country="Serbia" name="Bor">BR</collectingRegion>
); Keyberg,
<date id="01AF298DAC2194417E4654B6F9091E03" value="1947-06-24">24 Jun 1947</date>
,
<emphasis id="A6DDECD025C2F23B54DB5AE110DCC4CE" italics="true">A. Schmitz 732</emphasis>
(
<collectingRegion id="FA81C6F21D3A112064ABDA1B6EEF1BC2" country="Serbia" name="Bor">BR</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2FIM" type="notes">
<paragraph id="0981D7CD0A63C94D64E7FAB220B3AD41">
<heading id="556F2D8F90AE724F758264480C1650C3" reason="title">Note.</heading>
</paragraph>
<paragraph id="BF47E12AE76B96AD4EF871AFAFCC80D7">
At the northern limit of the distribution range, some specimens are unusual in lacking long hairs on pedicel and calyx (
<emphasis id="95F19D243DF3DE2C8C7C6B005A4BD7E3" italics="true">Herman 2308</emphasis>
,
<emphasis id="F067730CAD63B9E0D946602898D041D2" italics="true">
<collectionCode id="F9A0F6902AFE00A810B20BFA90D0AC0B" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
. Schaijes 1885, P. Quarré 2929
</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>