146 lines
15 KiB
XML
146 lines
15 KiB
XML
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<mods:title id="05AB5F7543A5C2A4B07B17B64C81242B">A review of the Heteroclinus heptaeolus complex (Pisces: Blennioidei: Clinidae), with three new species and discussion of use of proportions in taxonomic studies</mods:title>
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<mods:namePart id="727C756419310B2D86C6621F4E3B6F08">Hoese, Douglass F.</mods:namePart>
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<mods:affiliation id="246C946A79142ED1254ACBA99F7CF18D">Australian Museum Research Institute, Australian Museum, 1 William Street, Sydney NSW 2010, Australia</mods:affiliation>
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<mods:namePart id="BD8B48B427F5E7AE3E7F142C33E8F509">Hay, Amanda</mods:namePart>
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<mods:affiliation id="566AA620C4ECA15CFF252C49ED93DE1D">Australian Museum Research Institute, Australian Museum, 1 William Street, Sydney NSW 2010, Australia</mods:affiliation>
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<mods:namePart id="8785BA8F788A79E1C0489ADD7E3A76A8">Dibattista, Joseph D.</mods:namePart>
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<mods:affiliation id="7502C0CEEAD8050F52EC1014454DEA46">Australian Museum Research Institute, Australian Museum, 1 William Street, Sydney NSW 2010, Australia</mods:affiliation>
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<mods:title id="B52CA7F44D9AB3157F0BD141F2D6193A">Zootaxa</mods:title>
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<mods:date id="D146B65636A6E5465370AE30FFC7AFA3">2024</mods:date>
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<mods:number id="789B950FDE2D743B1C7D6E92127AEC43">2024-04-02</mods:number>
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<treatment id="933987E1FFCAFFCE80A3999FFA2DA0A8" ID-DOI="http://doi.org/10.5281/zenodo.10909612" ID-Zenodo-Dep="10909612" LSID="urn:lsid:plazi:treatment:933987E1FFCAFFCE80A3999FFA2DA0A8" httpUri="http://treatment.plazi.org/id/933987E1FFCAFFCE80A3999FFA2DA0A8" lastPageNumber="338" pageId="37" pageNumber="338">
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<subSubSection id="538A657CFFCAFFCE80A3999FFC8FA7EF" box="[151,831,726,753]" pageId="37" pageNumber="338" type="nomenclature">
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<paragraph id="1B2F36F7FFCAFFCE80A3999FFC8FA7EF" blockId="37.[151,831,726,753]" box="[151,831,726,753]" pageId="37" pageNumber="338">
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<heading id="4067819BFFCAFFCE80A3999FFC8FA7EF" bold="true" box="[151,831,726,753]" fontSize="11" level="1" pageId="37" pageNumber="338" reason="1">
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<emphasis id="29E4EAE5FFCAFFCE80A3999FFC8FA7EF" bold="true" box="[151,831,726,753]" pageId="37" pageNumber="338">
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Key to Species of the
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<taxonomicName id="DC904D74FFCAFFCE8196999EFD77A7EF" ID-CoL="3L8KQ" baseAuthorityName="Ogilby" baseAuthorityYear="1885" box="[418,711,726,753]" family="Clinidae" genus="Heteroclinus" kingdom="Animalia" order="Perciformes" pageId="37" pageNumber="338" phylum="Chordata" rank="species" species="heptaeolus">
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<emphasis id="29E4EAE5FFCAFFCE8196999EFD77A7EF" bold="true" box="[418,711,726,753]" italics="true" pageId="37" pageNumber="338">Heteroclinus heptaeolus</emphasis>
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complex.
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</emphasis>
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<subSubSection id="538A657CFFCAFFCE80A39854FA2DA0A8" pageId="37" pageNumber="338" type="key">
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<paragraph id="1B2F36F7FFCAFFCE80A39854FA2DA675" blockId="37.[151,1437,796,1463]" pageId="37" pageNumber="338">
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<keyLead id="A06195C2FFCAFFCE80A39854FA2DA675" pageId="37" pageNumber="338">
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1a. Orbital tentacle absent; last dorsal ray connected by membrane to caudal peduncle about halfway from base of last ray to base of caudal fin, well away from caudal fin; body with stripes or horizontally elongate oval spots in live; pectoral rays 13................................................................................
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<taxonomicName id="DC904D74FFCAFFCE8447981EFAD0A675" authority="Hoese & Hay & Dibattista, 2024" authorityName="Hoese & Hay & Dibattista" authorityYear="2024" box="[1139,1376,854,875]" family="Clinidae" genus="Heteroclinus" kingdom="Animalia" order="Perciformes" pageId="37" pageNumber="338" phylum="Chordata" rank="species" species="longicauda" status="sp. nov.">
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<emphasis id="29E4EAE5FFCAFFCE8447981EFAD0A675" box="[1139,1376,854,875]" italics="true" pageId="37" pageNumber="338">Heteroclinus longicauda</emphasis>
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</taxonomicName>
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<emphasis id="29E4EAE5FFCAFFCE8552981EFA2DA675" bold="true" box="[1382,1437,854,875]" pageId="37" pageNumber="338">
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<taxonomicNameLabel id="32D7579EFFCAFFCE8552981EFA2DA675" box="[1382,1437,854,875]" pageId="37" pageNumber="338" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="1B2F36F7FFCAFFCE80A3983CFA2CA6B8" blockId="37.[151,1437,796,1463]" pageId="37" pageNumber="338">
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<keyLead id="A06195C2FFCAFFCE80A3983CFA2CA6B8" pageId="37" pageNumber="338">1b Orbital tentacle present above middle of eye; last dorsal ray connected by membrane to caudal peduncle just before base of caudal fin; body with irregular vertical bars, stripes or spots.................................................... 2</keyLead>
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</paragraph>
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<keyStep id="A0642E52FFCAFFCE80A398E6FA2CA105" pageId="37" pageNumber="338">
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<paragraph id="1B2F36F7FFCAFFCE80A398E6FA2CA6FF" blockId="37.[151,1437,796,1463]" pageId="37" pageNumber="338">
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<keyLead id="A06195C2FFCAFFCE80A398E6FA2CA6FF" pageId="37" pageNumber="338">2a. First dorsal fin elevated, second spine distinctly longer than anterior spines of second dorsal fin; second dorsal spine 6.9–11.5% SL; orbital tentacle usually greater than half the eye diameter in adults; pectoral fin-rays 12–13....................... 3</keyLead>
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</paragraph>
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<paragraph id="1B2F36F7FFCAFFCE80A398A1FA2CA105" blockId="37.[151,1437,796,1463]" pageId="37" pageNumber="338">
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<keyLead id="A06195C2FFCAFFCE80A398A1FA2CA105" pageId="37" pageNumber="338">2b. First dorsal fin low, second spine slightly longer than anterior spines of second dorsal fin; second dorsal spine 10.5–16.7% SL; orbital tentacle usually less than half eye diameter in adults; pectoral rays 12...................................... 4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="A0642E52FFCAFFCE80A39F6CFA2CA1D5" pageId="37" pageNumber="338">
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<paragraph id="1B2F36F7FFCAFFCE80A39F6CFA2DA16D" blockId="37.[151,1437,796,1463]" pageId="37" pageNumber="338">
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<keyLead id="A06195C2FFCAFFCE80A39F6CFA2DA16D" pageId="37" pageNumber="338">
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3a. Pectoral rays 13; second dorsal fin with 24–25 spines; anal fin II, 18–19, second dorsal fin not incised; in life head with a thin white or silver bar below eye, followed posteriorly by broad dark bar and a thin white to silver stripe from anteroventral margin of eye to middle of jaws; body with dark spots or vertical bands on midside and dorsally only..........
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<taxonomicName id="DC904D74FFCAFFCE84DF9F17FAD0A16D" authority="Hoese & Hay & Dibattista, 2024" authorityName="Hoese & Hay & Dibattista" authorityYear="2024" box="[1259,1376,1118,1140]" family="Clinidae" genus="Heteroclinus" kingdom="Animalia" order="Perciformes" pageId="37" pageNumber="338" phylum="Chordata" rank="species" species="colemani" status="n. sp.">
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<emphasis id="29E4EAE5FFCAFFCE84DF9F17FAD0A16D" box="[1259,1376,1118,1140]" italics="true" pageId="37" pageNumber="338">H. colemani</emphasis>
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</taxonomicName>
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<emphasis id="29E4EAE5FFCAFFCE85529F16FA2DA16D" bold="true" box="[1382,1437,1118,1139]" pageId="37" pageNumber="338">
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<taxonomicNameLabel id="32D7579EFFCAFFCE85529F16FA2DA16D" box="[1382,1437,1118,1139]" pageId="37" pageNumber="338" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="1B2F36F7FFCAFFCE80A39F34FA2CA1D5" blockId="37.[151,1437,796,1463]" pageId="37" pageNumber="338">
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<keyLead id="A06195C2FFCAFFCE80A39F34FA2CA1D5" pageId="37" pageNumber="338">
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3b. Pectoral rays 12; second dorsal fin with 25–28 spines, rarely 25; anal fin II, 20–22; second dorsal fin slightly incised; in life head with a broad bar from eye to near posterior end of jaw, sometimes bordered with white to silver thin bars and no bar from eye to middle of jaws; body with irregular vertical dark bands extending between dorsal and anal fines..........
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<taxonomicName id="DC904D74FFCAFFCE850E9FFFFA2CA1D5" ID-CoL="3L8L4" baseAuthorityName="Lucas" baseAuthorityYear="1891" box="[1338,1436,1206,1228]" family="Clinidae" genus="Heteroclinus" kingdom="Animalia" order="Perciformes" pageId="37" pageNumber="338" phylum="Chordata" rank="species" species="wilsoni">
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<emphasis id="29E4EAE5FFCAFFCE850E9FFFFA2CA1D5" box="[1338,1436,1206,1228]" italics="true" pageId="37" pageNumber="338">H. wilsoni</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="A0642E52FFCAFFCE80A39F9CFA2DA0A8" pageId="37" pageNumber="338">
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<paragraph id="1B2F36F7FFCAFFCE80A39F9CFA2CA05F" blockId="37.[151,1437,796,1463]" pageId="37" pageNumber="338">
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<keyLead id="A06195C2FFCAFFCE80A39F9CFA2CA05F" pageId="37" pageNumber="338">
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4a. In life body with irregular vertical dark bands; round dark spots on back below dorsal fin narrower than spaces between spots; broad light areas on second dorsal and anal fins containing 2 or 3 spines and irregular dark pigment; disappearing in preserved material; second dorsal spines 25–28, rarely 25 (less than 5% of specimens counted); anal fin II, 18–21, rarely 18 (in less than 5% of individuals counted); total gill rakers on first arch 6–10, rarely 10................................
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<taxonomicName id="DC904D74FFCAFFCE85239E64FA2CA05F" ID-CoL="3L8KQ" baseAuthorityName="Ogilby" baseAuthorityYear="1885" box="[1303,1436,1324,1345]" family="Clinidae" genus="Heteroclinus" kingdom="Animalia" order="Perciformes" pageId="37" pageNumber="338" phylum="Chordata" rank="species" species="heptaeolus">
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<emphasis id="29E4EAE5FFCAFFCE85239E64FA2CA05F" box="[1303,1436,1324,1345]" italics="true" pageId="37" pageNumber="338">H. heptaeolus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="1B2F36F7FFCAFFCE80A39E01FA2DA0A8" blockId="37.[151,1437,796,1463]" pageId="37" pageNumber="338">
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<keyLead id="A06195C2FFCAFFCE80A39E01FA2DA0A8" pageId="37" pageNumber="338">
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4b. In life body with stripes or horizontally elongated spots; round dark spots on back broader than spaces between spots; second dorsal and anal fins with distinct clear windows between spines, sometimes with window containing one spine, windows typically remaining in preserved material; second dorsal spines 23–25; anal fin II,15–18; total gill rakers on first arch 8–12, usually 10 or 11.........................................................................
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<taxonomicName id="DC904D74FFCAFFCE84C39EEAFAD0A0A8" authority="Hoese & Hay & Dibattista, 2024" authorityName="Hoese & Hay & Dibattista" authorityYear="2024" box="[1271,1376,1441,1463]" family="Clinidae" genus="Heteroclinus" kingdom="Animalia" order="Perciformes" pageId="37" pageNumber="338" phylum="Chordata" rank="species" species="whitleyi" status="n. sp.">
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<emphasis id="29E4EAE5FFCAFFCE84C39EEAFAD0A0A8" box="[1271,1376,1441,1463]" italics="true" pageId="37" pageNumber="338">H. whitleyi</emphasis>
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</taxonomicName>
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<emphasis id="29E4EAE5FFCAFFCE85529EE9FA2DA0A8" bold="true" box="[1382,1437,1441,1462]" pageId="37" pageNumber="338">
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<taxonomicNameLabel id="32D7579EFFCAFFCE85529EE9FA2DA0A8" box="[1382,1437,1441,1462]" pageId="37" pageNumber="338" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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</subSubSection>
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</treatment>
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