267 lines
40 KiB
XML
267 lines
40 KiB
XML
<document id="7AC9477343FC583113185AED7F6E3626" ID-DOI="10.5070/P9401057774" ID-ISSN="0031-0298" ID-Zenodo-Dep="10913295" ID-ZooBank="11600574-2B0E-4C13-BC08-A3A5EF9EE562" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="operationResults" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1712167020321" checkinUser="felipe" docAuthor="Groves, Lindsey & Squires, Richard L." docDate="2023" docId="921FD94CFFF1FF8BFC49FF78FD0BFE68" docLanguage="en" docName="PaleoBios.40.10.1-52.pdf" docOrigin="PaleoBios 40 (10)" docSource="http://dx.doi.org/10.5070/p9401057774" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.3:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="3" docTitle="Luponovula maniobraensis" docType="treatment" docVersion="8" lastPageNumber="33" masterDocId="6E26A134FFD3FFA8FFA2FFBBFFCAFFE5" masterDocTitle="Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover" masterLastPageNumber="52" masterPageNumber="1" pageNumber="33" updateTime="1727793805776" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="4641276AAB0322AA97B9D90BBCD11E63">Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover</mods:title>
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<mods:namePart id="172B3992D452F508FD8916423F1FA5E7">Squires, Richard L.</mods:namePart>
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<mods:title id="A4220A9910419BAE073BB5278A0DB340">PaleoBios</mods:title>
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<mods:date id="6F97A2105E63129CD0B7E3E3847ABA00">2023</mods:date>
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<treatment id="921FD94CFFF1FF8BFC49FF78FD0BFE68" LSID="urn:lsid:plazi:treatment:921FD94CFFF1FF8BFC49FF78FD0BFE68" httpUri="http://treatment.plazi.org/id/921FD94CFFF1FF8BFC49FF78FD0BFE68" lastPageId="35" lastPageNumber="33" pageId="34" pageNumber="33">
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<subSubSection id="52AC3BD1FFF1FF8AFC49FF78FA5EFEE5" pageId="34" pageNumber="33" type="nomenclature">
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<paragraph id="1A09685AFFF1FF8AFC49FF78FA8FFF38" blockId="34.[926,1428,195,291]" box="[1003,1349,195,221]" pageId="34" pageNumber="33">
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<heading id="4141DF36FFF1FF8AFC49FF78FA8FFF38" allCaps="true" box="[1003,1349,195,221]" centered="true" fontSize="10" level="2" pageId="34" pageNumber="33" reason="2">
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<taxonomicName id="DDB613D9FFF1FF8AFC49FF78FA8FFF38" authority="(SQUIRES AND ADVOCATE, 1986)" box="[1003,1349,195,221]" class="Gastropoda" family="Cypraeidae" genus="Luponovula" kingdom="Animalia" order="Littorinimorpha" pageId="34" pageNumber="33" phylum="Mollusca" rank="species" species="maniobraensis">
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<emphasis id="28C2B448FFF1FF8AFC49FF78FA8FFF38" box="[1003,1349,195,221]" italics="true" pageId="34" pageNumber="33">LUPONOVULA MANIOBRAENSIS</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="1A09685AFFF1FF8AFC3CFF5DFA5EFEE5" blockId="34.[926,1428,195,291]" box="[926,1428,230,257]" pageId="34" pageNumber="33">(SQUIRES AND ADVOCATE, 1986) N. COMB.</paragraph>
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</subSubSection>
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<subSubSection id="52AC3BD1FFF1FF8AFBF6FEB2FB16FEC6" box="[1108,1244,265,291]" pageId="34" pageNumber="33" type="description">
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<paragraph id="1A09685AFFF1FF8AFBF6FEB2FB16FEC6" blockId="34.[926,1428,195,291]" box="[1108,1244,265,291]" pageId="34" pageNumber="33">
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<figureCitation id="828D74DFFFF1FF8AFBF6FEB2FB16FEC6" box="[1108,1244,265,291]" captionStart="Figure 9" captionStartId="31.[112,199,200,226]" captionTargetId="figure-12@30.[141,1485,192,1920]" captionTargetPageId="30" captionText="Figure 9A–B. Grovesia castacensis (Stewart, 1926 [1927]), Tejon Formation (Eocene, “Tejon Stage”), Tehachapi Mountains, Kern County, southern California. Dorsal (A) and left (columellar) (B) views, 3.4x, length 15.5 mm, width 9.1 mm, height 6.9 mm, of hypotype LACMIP 41206.2, LACMIP Type 14941, base of Metralla Sandstone Member, Live Oak Canyon, LACMIP Locality 41206 [ex CSUN Locality 1206]. C–L. Grovesia mathewsonii (Gabb, 1869), Muir Sandstone (Eocene, “Domengine Stage” to “Tejon Stage”), near Martinez, Contra Costa County, northern California. Basal (C), dorsal (D), right (labral) views, 1x, length 10.7 mm, width 7.5 mm, height 4.7 mm, of holotype ANSP 4217. F–G. Basal (F) [as published] and dorsal (G) views, 2.9x, length 17.9 mm, width 11.6 mm, of holotype CASG 245.02 [ex CAS 816] of Cypraea kerniana Anderson and Hanna, 1925, Tejon Formation (Eocene, “Tejon Stage”) Grapevine Canyon, Kern County, southern California. H–J. Basal (H), dorsal (I), left (columellar), (J) left (columellar) views, 2.7x, length 14.6 mm, width 10.8 mm, height 7.8 mm, of hypotype LACMIP 7155.9, LACMIP type 14930, Tejon Formation, Grapevine Canyon, LACMIP Locality 7155, Kern County, southern California. K-L. Basal (K), dorsal (L) views, 3.9x, length 14 mm, width 10.3 mm, of hypotype UCMP 30903, UCMP Locality 5085, Ardath Shale (Eocene, “Domengine Stage”), San Diego County, southern California. M–P. Luponovula maniobraensis (Squires and Advocate, 1986), (M–O). Maniobra Formation (Eocene, “Capay Stage”), Orocopia Mountains, Riverside County, southern California. Basal (M), dorsal (N), and right (labral) (O) views, 1.3x, length 39.6 mm, width 22.8 mm, height 21.2 mm, of holotype LACMIP 10566.4, LACMIP Type 10566 [ex UCLA 48431], LACMIP Locality 23779. (P). Basal view, Llajas Formation (Eocene, “Capay Stage”), north side Simi Valley, Ventura County, southern California, 2.1x, length 26.6 mm, width 15.8 mm, height 10.9 mm, of hypotype LACMIP 40491.1, LACMIP Type 14945 (LACMIP Locality 40491 [ex CSUN Locality 491])." figureDoi="http://doi.org/10.5281/zenodo.13869606" httpUri="https://zenodo.org/record/13869606/files/figure.png" pageId="34" pageNumber="33">FIGS. 9M–P</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="52AC3BD1FFF1FF8AFCC2FEEBFB5BFD42" pageId="34" pageNumber="33" type="reference_group">
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<paragraph id="1A09685AFFF1FF8AFCC2FEEBFB9AFE55" blockId="34.[830,1520,334,1947]" pageId="34" pageNumber="33">
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<taxonomicName id="DDB613D9FFF1FF8AFCC2FEEBFC4BFE68" ID-CoL="C5KVV" authority="Squires and Advocate, 1986" authorityName="Squires and Advocate" authorityYear="1986" class="Gastropoda" family="Eocypraeidae" genus="Eocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="34" pageNumber="33" phylum="Mollusca" rank="species" species="maniobraensis">
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<emphasis id="28C2B448FFF1FF8AFCC2FEEBFC2DFE89" box="[864,999,336,364]" italics="true" pageId="34" pageNumber="33">Eocypraea</emphasis>
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?
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<emphasis id="28C2B448FFF1FF8AFBA2FEF5FB77FE8F" box="[1024,1213,334,362]" italics="true" pageId="34" pageNumber="33">maniobraensis</emphasis>
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<bibRefCitation id="7E2715ABFFF1FF8AFB6AFEEBFC4BFE68" author="Squires, R. L. & D. M. Advocate" pageId="34" pageNumber="33" pagination="851 - 864" refId="ref43291" refString="Squires, R. L. and D. M. Advocate 1986. New early Eocene mollusks from the Orocopia Mountains, southern California. Journal of Paleontology 60 (4): 851 - 864, figs, 1 - 3." type="journal article" year="1986">Squires and Advocate, 1986</bibRefCitation>
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</taxonomicName>
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. pp. 856, 857, figs. 2.5, 2.6.
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<bibRefCitation id="7E2715ABFFF1FF8AFB42FEC8FA47FE68" author="Squires, R. L." box="[1248,1421,371,397]" pageId="34" pageNumber="33" pagination="217 - 226" refId="ref42749" refString="Squires, R. L. 1991. Molluscan paleontology of the lower Eocene Maniobra Formation, Orocopia Mountains, southern California. In P. L. Abbott and J. A. May (eds.). Eocene Geologic History San Diego Region. Pacific Section, Society of Economic Paleontologists and Mineralogists. 68: 217 - 226, pls. 1 - 2." type="journal article" year="1991">Squires, 1991</bibRefCitation>
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. p. 219, 220, 225; pl. 1, fig. 23.
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</paragraph>
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<paragraph id="1A09685AFFF1FF8AFCC2FE01FA3AFDD8" blockId="34.[830,1520,334,1947]" pageId="34" pageNumber="33">
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<taxonomicName id="DDB613D9FFF1FF8AFCC2FE01FBAEFE12" ID-CoL="C5KVV" authority="Squires and Advocate. Groves, 1992" authorityName="Squires and Advocate. Groves" authorityYear="1992" class="Gastropoda" family="Eocypraeidae" genus="Eocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="34" pageNumber="33" phylum="Mollusca" rank="species" species="maniobraensis" subGenus="Eocypraea">
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<emphasis id="28C2B448FFF1FF8AFCC2FE01FC2EFE33" box="[864,996,442,470]" italics="true" pageId="34" pageNumber="33">Eocypraea</emphasis>
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(
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<emphasis id="28C2B448FFF1FF8AFC58FE01FBB4FE33" box="[1018,1150,442,470]" italics="true" pageId="34" pageNumber="33">Eocypraea</emphasis>
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)
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<emphasis id="28C2B448FFF1FF8AFB36FE03FA86FE31" box="[1172,1356,440,468]" italics="true" pageId="34" pageNumber="33">maniobraensis</emphasis>
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Squires and Advocate.
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<bibRefCitation id="7E2715ABFFF1FF8AFC62FE66FBAEFE12" author="Groves, L. T." box="[960,1124,477,503]" pageId="34" pageNumber="33" pagination="101 - 107" refId="ref37814" refString="Groves, L. T. 1992. California cowries (Cypraeacea): Past and present, with notes on recent tropical eastern Pacific species. The Festivus 24 (9): 101 - 107, figs. 1 - 3." type="journal article" year="1992">Groves, 1992</bibRefCitation>
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</taxonomicName>
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. p. 106 [not 1987 as in table 1].
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<bibRefCitation id="7E2715ABFFF1FF8AFCE2FDBBFC28FDFF" author="Groves, L. T." box="[832,994,512,538]" pageId="34" pageNumber="33" pagination="11 - 14" refId="ref37858" refString="Groves, L. T. 1993. Fossil and recent species of eastern Pacific Cypraeacea (Pediculariidae, Cypraeidae, and Ovulidae). The Western Society of Malacologists Annual Report 25: 11 - 14." type="book chapter" year="1993">Groves, 1993</bibRefCitation>
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. p. 11 [not 1983 as in text].
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<bibRefCitation id="7E2715ABFFF1FF8AFA8FFDBBFA05FDFF" author="Groves, L. T." box="[1325,1487,512,538]" pageId="34" pageNumber="33" pagination="7 - 10" refId="ref38049" refString="Groves, L. T. 1997. Fossil and recent species of eastern Pacific Cypraeacea (Cypraeidae and Eocypraeinae [Ovulidae]): An update. The Western Society of Malacology Annual Report 29: 7 - 10." type="book chapter" year="1997">Groves, 1997</bibRefCitation>
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. p. 8 [as
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<emphasis id="28C2B448FFF1FF8AFCDDFD9AFC5CFDD8" box="[895,918,545,573]" italics="true" pageId="34" pageNumber="33">E.</emphasis>
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<taxonomicName id="DDB613D9FFF1FF8AFC38FD98FBB5FDD8" ID-CoL="C5KVV" authorityName="Squires and Advocate. Groves" authorityYear="1992" box="[922,1151,545,573]" class="Gastropoda" family="Eocypraeidae" genus="Eocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="34" pageNumber="33" phylum="Mollusca" rank="species" species="maniobraensis" subGenus="Eocypraea">
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(
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<emphasis id="28C2B448FFF1FF8AFC07FD9AFC7CFDD8" box="[933,950,545,573]" italics="true" pageId="34" pageNumber="33">E</emphasis>
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||
.)
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<emphasis id="28C2B448FFF1FF8AFC6EFD9AFBB5FDD8" box="[972,1151,545,573]" italics="true" pageId="34" pageNumber="33">maniobraensis</emphasis>
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</taxonomicName>
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]. Groves, 2011. p. 46 (table 1).
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</paragraph>
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<paragraph id="1A09685AFFF1FF8AFCC2FDFDFB5BFD42" blockId="34.[830,1520,334,1947]" pageId="34" pageNumber="33">
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||
<taxonomicName id="DDB613D9FFF1FF8AFCC2FDFDFC4BFD66" ID-CoL="C5KVV" authority="Squires and Advocate. Fehse, 2001" authorityName="Squires and Advocate. Fehse" authorityYear="2001" class="Gastropoda" family="Eocypraeidae" genus="Eocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="34" pageNumber="33" phylum="Mollusca" rank="species" species="maniobraensis">
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<emphasis id="28C2B448FFF1FF8AFCC2FDFDFB59FD84" box="[864,1171,581,610]" italics="true" pageId="34" pageNumber="33">Eocypraea maniobraensis</emphasis>
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Squires and Advocate.
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<bibRefCitation id="7E2715ABFFF1FF8AFA01FDFDFC4BFD66" author="Fehse, D." pageId="34" pageNumber="33" pagination="1 - 51" refId="ref37011" refString="Fehse, D. 2001. Beitrage zur Kenntnis der Ovuliade (Mollusca: Cypraeoidea). 8. Einleitung zur familie sowie Katalog, Taxonomy und Bibliographie und Bemerkungen zu verwandten Gruppen. Acta Conchyliorum 5: 1 - 51, figs. 1 - 3." type="journal article" year="2001">Fehse, 2001</bibRefCitation>
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||
</taxonomicName>
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||
. p. 20.
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<bibRefCitation id="7E2715ABFFF1FF8AFC79FDD2FAD1FD66" author="Groves, L. T. & R. L. Squires" box="[987,1307,617,643]" pageId="34" pageNumber="33" pagination="1 - 360" refId="ref38553" refString="Groves, L. T. and R. L. Squires. 2021. Checklist of California Paleogene-Neogene marine Mollusca since Keen and Bentson (1944). PaleoBios 38: 1 - 360, figs. 1 - 3." type="journal article" year="2021">Groves and Squires, 2021</bibRefCitation>
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||
. p. 231 (as
|
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<taxonomicName id="DDB613D9FFF1FF8AFA0FFDD1FC4DFD42" ID-CoL="7P3H3" authorityName="Cossmann" authorityYear="1903" class="Gastropoda" family="Eocypraeidae" genus="Eocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="34" pageNumber="33" phylum="Mollusca" rank="genus">
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<emphasis id="28C2B448FFF1FF8AFA0FFDD1FC4DFD42" italics="true" pageId="34" pageNumber="33">Eocypraea</emphasis>
|
||
</taxonomicName>
|
||
?).
|
||
<bibRefCitation id="7E2715ABFFF1FF8AFC09FD37FB8BFD42" author="Fehse, D." box="[939,1089,652,679]" pageId="34" pageNumber="33" pagination="1 - 12" refId="ref37236" refString="Fehse, D. 2021. Zur systemstischen stellung der Eocy- praeidae F. A. Schilder 1924. Part 2. (Mollusca: Gas- tropoda: Cypraeoidea). Privately published online, Berlin, Germany 46 p., pls. 1 - 12." type="book chapter" year="2021">Fehse, 2021</bibRefCitation>
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. p. 42.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="52AC3BD1FFF1FF8AFCC2FD0BFAA9FA6F" pageId="34" pageNumber="33" type="materials_examined">
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||
<paragraph id="1A09685AFFF1FF8AFCC2FD0BFB56FC9F" blockId="34.[830,1520,334,1947]" pageId="34" pageNumber="33">
|
||
<materialsCitation id="AADE6207FFF1FF8AFCC2FD0BFB56FC9F" collectionCode="LACMIP, UCLA, CSUN" county="Riverside County" location="Maniobra Formation" municipality="Locality" pageId="34" pageNumber="33" specimenCode="LACMIP10566.4, UCLA 48431" specimenCount="1" stateProvince="California" typeStatus="holotype">
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||
<emphasis id="28C2B448FFF1FF8AFCC2FD0BFB15FD2C" bold="true" box="[864,1247,687,714]" pageId="34" pageNumber="33">
|
||
<typeStatus id="C50DD6F8FFF1FF8AFCC2FD0BFC13FD2F" box="[864,985,688,714]" pageId="34" pageNumber="33" type="holotype">Holotype</typeStatus>
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||
and Type Locality—
|
||
</emphasis>
|
||
<specimenCode id="4A10C021FFF1FF8AFB7DFD0BFA66FD2F" box="[1247,1452,687,714]" collectionCode="LACMIP" pageId="34" pageNumber="33">LACMIP10566.4</specimenCode>
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||
, LAC- MIP Type 10566 (
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||
<emphasis id="28C2B448FFF1FF8AFB80FD6AFBF6FD08" box="[1058,1084,721,749]" italics="true" pageId="34" pageNumber="33">ex</emphasis>
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||
<specimenCode id="4A10C021FFF1FF8AFBE1FD68FB29FD08" box="[1091,1251,723,749]" collectionCode="UCLA" country="USA" httpUri="http://grbio.org/cool/irpn-5wg7" name="University of California at Los Angeles" pageId="34" pageNumber="33">UCLA 48431</specimenCode>
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||
) (
|
||
<figureCitation id="828D74DFFFF1FF8AFB5EFD68FA47FD08" box="[1276,1421,723,749]" captionStart="Figure 9" captionStartId="31.[112,199,200,226]" captionTargetId="figure-12@30.[141,1485,192,1920]" captionTargetPageId="30" captionText="Figure 9A–B. Grovesia castacensis (Stewart, 1926 [1927]), Tejon Formation (Eocene, “Tejon Stage”), Tehachapi Mountains, Kern County, southern California. Dorsal (A) and left (columellar) (B) views, 3.4x, length 15.5 mm, width 9.1 mm, height 6.9 mm, of hypotype LACMIP 41206.2, LACMIP Type 14941, base of Metralla Sandstone Member, Live Oak Canyon, LACMIP Locality 41206 [ex CSUN Locality 1206]. C–L. Grovesia mathewsonii (Gabb, 1869), Muir Sandstone (Eocene, “Domengine Stage” to “Tejon Stage”), near Martinez, Contra Costa County, northern California. Basal (C), dorsal (D), right (labral) views, 1x, length 10.7 mm, width 7.5 mm, height 4.7 mm, of holotype ANSP 4217. F–G. Basal (F) [as published] and dorsal (G) views, 2.9x, length 17.9 mm, width 11.6 mm, of holotype CASG 245.02 [ex CAS 816] of Cypraea kerniana Anderson and Hanna, 1925, Tejon Formation (Eocene, “Tejon Stage”) Grapevine Canyon, Kern County, southern California. H–J. Basal (H), dorsal (I), left (columellar), (J) left (columellar) views, 2.7x, length 14.6 mm, width 10.8 mm, height 7.8 mm, of hypotype LACMIP 7155.9, LACMIP type 14930, Tejon Formation, Grapevine Canyon, LACMIP Locality 7155, Kern County, southern California. K-L. Basal (K), dorsal (L) views, 3.9x, length 14 mm, width 10.3 mm, of hypotype UCMP 30903, UCMP Locality 5085, Ardath Shale (Eocene, “Domengine Stage”), San Diego County, southern California. M–P. Luponovula maniobraensis (Squires and Advocate, 1986), (M–O). Maniobra Formation (Eocene, “Capay Stage”), Orocopia Mountains, Riverside County, southern California. Basal (M), dorsal (N), and right (labral) (O) views, 1.3x, length 39.6 mm, width 22.8 mm, height 21.2 mm, of holotype LACMIP 10566.4, LACMIP Type 10566 [ex UCLA 48431], LACMIP Locality 23779. (P). Basal view, Llajas Formation (Eocene, “Capay Stage”), north side Simi Valley, Ventura County, southern California, 2.1x, length 26.6 mm, width 15.8 mm, height 10.9 mm, of hypotype LACMIP 40491.1, LACMIP Type 14945 (LACMIP Locality 40491 [ex CSUN Locality 491])." figureDoi="http://doi.org/10.5281/zenodo.13869606" httpUri="https://zenodo.org/record/13869606/files/figure.png" pageId="34" pageNumber="33">Figs. 9M–O</figureCitation>
|
||
), length
|
||
<quantity id="DD4EC5BFFFF1FF8AFCE2FD4DFC7AFCF4" box="[832,944,758,785]" metricMagnitude="-2" metricUnit="m" metricValue="3.96" pageId="34" pageNumber="33" unit="mm" value="39.6">39.6 mm</quantity>
|
||
, width
|
||
<quantity id="DD4EC5BFFFF1FF8AFBAEFD4DFBB6FCF4" box="[1036,1148,758,785]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="34" pageNumber="33" unit="mm" value="22.8">22.8 mm</quantity>
|
||
, height 21.1.
|
||
<collectionCode id="7CA7F09FFFF1FF8AFA83FD4DFA4FFCF5" box="[1313,1413,758,784]" pageId="34" pageNumber="33">LACMIP</collectionCode>
|
||
<collectingMunicipality id="FA6DF220FFF1FF8AFA2FFD4DFA3AFCF5" box="[1421,1520,758,784]" pageId="34" pageNumber="33">Locality</collectingMunicipality>
|
||
23779 [
|
||
<emphasis id="28C2B448FFF1FF8AFC06FCA3FC74FCD1" box="[932,958,792,820]" italics="true" pageId="34" pageNumber="33">ex</emphasis>
|
||
<collectionCode id="7CA7F09FFFF1FF8AFC65FCA2FBC7FCD6" box="[967,1037,793,819]" country="USA" httpUri="http://grbio.org/cool/irpn-5wg7" name="University of California at Los Angeles" pageId="34" pageNumber="33">UCLA</collectionCode>
|
||
Locality 3779;
|
||
<emphasis id="28C2B448FFF1FF8AFB70FCA3FB26FCD1" box="[1234,1260,792,820]" italics="true" pageId="34" pageNumber="33">ex</emphasis>
|
||
<collectionCode id="7CA7F09FFFF1FF8AFB57FCA2FAF1FCD6" box="[1269,1339,793,819]" country="USA" httpUri="http://grbio.org/cool/sgeh-wrgd" name="California State University, Northridge" pageId="34" pageNumber="33" type="University or college">CSUN</collectionCode>
|
||
Locality 662],
|
||
<location id="1F693E81FFF1FF8AFCE2FC87FB84FCB3" LSID="urn:lsid:plazi:treatment:921FD94CFFF1FF8BFC49FF78FD0BFE68:1F693E81FFF1FF8AFCE2FC87FB84FCB3" box="[832,1102,828,854]" county="Riverside County" municipality="Locality" name="Maniobra Formation" pageId="34" pageNumber="33" stateProvince="California">Maniobra Formation</location>
|
||
,
|
||
<location id="1F693E81FFF1FF8AFBFCFC87FAA2FCB3" LSID="urn:lsid:plazi:treatment:921FD94CFFF1FF8BFC49FF78FD0BFE68:1F693E81FFF1FF8AFBFCFC87FAA2FCB3" box="[1118,1384,828,854]" county="Riverside County" municipality="Locality" name="Orocopia Mountains" pageId="34" pageNumber="33" stateProvince="California">Orocopia Mountains</location>
|
||
,
|
||
<collectingCounty id="F36810D6FFF1FF8AFAD5FC87FC53FC9F" pageId="34" pageNumber="33">Riverside County</collectingCounty>
|
||
, southern
|
||
<collectingRegion id="D872A6B8FFF1FF8AFBBEFCDBFB53FC9F" box="[1052,1177,864,890]" country="United States of America" name="California" pageId="34" pageNumber="33">California</collectingRegion>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="1A09685AFFF1FF8AFCC2FC38FA3AFB52" blockId="34.[830,1520,334,1947]" pageId="34" pageNumber="33">
|
||
<emphasis id="28C2B448FFF1FF8AFCC2FC38FA4BFC79" bold="true" box="[864,1409,898,925]" pageId="34" pageNumber="33">Referred Specimen—Hypotype (herein)</emphasis>
|
||
<materialsCitation id="AADE6207FFF1FF8AFA29FC38FC62FC25" collectionCode="LACMIP" pageId="34" pageNumber="33" specimenCode="LACMIP 40491.1" specimenCount="1" typeStatus="holotype">
|
||
<specimenCode id="4A10C021FFF1FF8AFA29FC38FC62FC25" collectionCode="LACMIP" pageId="34" pageNumber="33">LACMIP 40491.1</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="AADE6207FFF1FF8AFC10FC1DFAD0FC25" box="[946,1306,934,961]" collectionCode="LACMIP" pageId="34" pageNumber="33" specimenCount="1" typeStatus="holotype">
|
||
<collectionCode id="7CA7F09FFFF1FF8AFC10FC1DFBDCFC25" box="[946,1046,934,960]" pageId="34" pageNumber="33">LACMIP</collectionCode>
|
||
<typeStatus id="C50DD6F8FFF1FF8AFBB8FC1CFB92FC24" box="[1050,1112,935,961]" pageId="34" pageNumber="33">Type</typeStatus>
|
||
14945 (
|
||
<figureCitation id="828D74DFFFF1FF8AFB19FC1DFADBFC25" box="[1211,1297,934,960]" captionStart="Figure 9" captionStartId="31.[112,199,200,226]" captionTargetId="figure-12@30.[141,1485,192,1920]" captionTargetPageId="30" captionText="Figure 9A–B. Grovesia castacensis (Stewart, 1926 [1927]), Tejon Formation (Eocene, “Tejon Stage”), Tehachapi Mountains, Kern County, southern California. Dorsal (A) and left (columellar) (B) views, 3.4x, length 15.5 mm, width 9.1 mm, height 6.9 mm, of hypotype LACMIP 41206.2, LACMIP Type 14941, base of Metralla Sandstone Member, Live Oak Canyon, LACMIP Locality 41206 [ex CSUN Locality 1206]. C–L. Grovesia mathewsonii (Gabb, 1869), Muir Sandstone (Eocene, “Domengine Stage” to “Tejon Stage”), near Martinez, Contra Costa County, northern California. Basal (C), dorsal (D), right (labral) views, 1x, length 10.7 mm, width 7.5 mm, height 4.7 mm, of holotype ANSP 4217. F–G. Basal (F) [as published] and dorsal (G) views, 2.9x, length 17.9 mm, width 11.6 mm, of holotype CASG 245.02 [ex CAS 816] of Cypraea kerniana Anderson and Hanna, 1925, Tejon Formation (Eocene, “Tejon Stage”) Grapevine Canyon, Kern County, southern California. H–J. Basal (H), dorsal (I), left (columellar), (J) left (columellar) views, 2.7x, length 14.6 mm, width 10.8 mm, height 7.8 mm, of hypotype LACMIP 7155.9, LACMIP type 14930, Tejon Formation, Grapevine Canyon, LACMIP Locality 7155, Kern County, southern California. K-L. Basal (K), dorsal (L) views, 3.9x, length 14 mm, width 10.3 mm, of hypotype UCMP 30903, UCMP Locality 5085, Ardath Shale (Eocene, “Domengine Stage”), San Diego County, southern California. M–P. Luponovula maniobraensis (Squires and Advocate, 1986), (M–O). Maniobra Formation (Eocene, “Capay Stage”), Orocopia Mountains, Riverside County, southern California. Basal (M), dorsal (N), and right (labral) (O) views, 1.3x, length 39.6 mm, width 22.8 mm, height 21.2 mm, of holotype LACMIP 10566.4, LACMIP Type 10566 [ex UCLA 48431], LACMIP Locality 23779. (P). Basal view, Llajas Formation (Eocene, “Capay Stage”), north side Simi Valley, Ventura County, southern California, 2.1x, length 26.6 mm, width 15.8 mm, height 10.9 mm, of hypotype LACMIP 40491.1, LACMIP Type 14945 (LACMIP Locality 40491 [ex CSUN Locality 491])." figureDoi="http://doi.org/10.5281/zenodo.13869606" httpUri="https://zenodo.org/record/13869606/files/figure.png" pageId="34" pageNumber="33">Fig. 9P</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="7CA7F09FFFF1FF8AFA87FC1DFA43FC25" box="[1317,1417,934,960]" pageId="34" pageNumber="33">LACMIP</collectionCode>
|
||
<materialsCitation id="AADE6207FFF1FF8AFA2FFC1DFA20FB52" collectionCode="CSUN" county="Ventura County" location="Las Llajas Canyon" municipality="Llajas Formation" pageId="34" pageNumber="33" specimenCount="1" stateProvince="California" typeStatus="holotype">
|
||
Locality 40491 [
|
||
<emphasis id="28C2B448FFF1FF8AFC03FC73FC71FC01" box="[929,955,968,996]" italics="true" pageId="34" pageNumber="33">ex</emphasis>
|
||
<collectionCode id="7CA7F09FFFF1FF8AFC63FC72FBCDFC06" box="[961,1031,969,995]" country="USA" httpUri="http://grbio.org/cool/sgeh-wrgd" name="California State University, Northridge" pageId="34" pageNumber="33" type="University or college">CSUN</collectionCode>
|
||
Locality 491],
|
||
<collectingMunicipality id="FA6DF220FFF1FF8AFB1EFC72FA45FC06" box="[1212,1423,969,995]" pageId="34" pageNumber="33">Llajas Formation</collectingMunicipality>
|
||
, is from a faulted area low in the stratigraphic section of this for- mation (see geologic map in
|
||
<bibRefCitation id="7E2715ABFFF1FF8AFB3FFBABFA8AFBCF" author="Squires, R. L." box="[1181,1344,1040,1066]" pageId="34" pageNumber="33" pagination="1 - 76" refId="ref42615" refString="Squires, R. L. 1984. Megapaleontology of the Eocene Lla- jas Formation, Simi Valley, California. Contributions in Science, Natural History Museum of Los Angeles County 350: 1 - 76, figs. 1 - 18." type="journal article" year="1984">Squires 1984</bibRefCitation>
|
||
: fig. 16) in
|
||
<location id="1F693E81FFF1FF8AFA67FBABFC26FBA8" LSID="urn:lsid:plazi:treatment:921FD94CFFF1FF8BFC49FF78FD0BFE68:1F693E81FFF1FF8AFA67FBABFC26FBA8" county="Ventura County" municipality="Llajas Formation" name="Las Llajas Canyon" pageId="34" pageNumber="33" stateProvince="California">Las Llajas Canyon</location>
|
||
, north side of
|
||
<location id="1F693E81FFF1FF8AFB38FB88FAE8FBA8" LSID="urn:lsid:plazi:treatment:921FD94CFFF1FF8BFC49FF78FD0BFE68:1F693E81FFF1FF8AFB38FB88FAE8FBA8" box="[1178,1314,1075,1101]" county="Ventura County" municipality="Llajas Formation" name="Simi Valley" pageId="34" pageNumber="33" stateProvince="California">Simi Valley</location>
|
||
,
|
||
<collectingCounty id="F36810D6FFF1FF8AFA89FB88FA26FBA8" box="[1323,1516,1075,1101]" pageId="34" pageNumber="33">Ventura County</collectingCounty>
|
||
, southern
|
||
<collectingRegion id="D872A6B8FFF1FF8AFC1BFBEDFBF9FB95" box="[953,1075,1110,1136]" country="United States of America" name="California" pageId="34" pageNumber="33">California</collectingRegion>
|
||
[new information].
|
||
<location id="1F693E81FFF1FF8AFA8CFBEDFA7FFB95" LSID="urn:lsid:plazi:treatment:921FD94CFFF1FF8BFC49FF78FD0BFE68:1F693E81FFF1FF8AFA8CFBEDFA7FFB95" box="[1326,1461,1110,1136]" county="Ventura County" municipality="Llajas Formation" name="This Llajas" pageId="34" pageNumber="33" stateProvince="California">This Llajas</location>
|
||
For- mation specimen has been crushed but retains enough morphological features to identify it as
|
||
<taxonomicName id="DDB613D9FFF1FF8AFABCFB20FA20FB52" authorityName="Squires and Advocate. Fehse" authorityYear="2001" box="[1310,1514,1179,1207]" class="Gastropoda" family="Eocypraeidae" genus="Eocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="34" pageNumber="33" phylum="Mollusca" rank="species" species="maniobraensis">
|
||
<emphasis id="28C2B448FFF1FF8AFABCFB20FA20FB52" box="[1310,1514,1179,1207]" italics="true" pageId="34" pageNumber="33">L. maniobraensis</emphasis>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1A09685AFFF1FF8AFCC2FB04FAA9FA6F" blockId="34.[830,1520,334,1947]" pageId="34" pageNumber="33">
|
||
<emphasis id="28C2B448FFF1FF8AFCC2FB04FBDDFB3C" bold="true" box="[864,1047,1215,1241]" pageId="34" pageNumber="33">Occurrence—</emphasis>
|
||
Lower Eocene (“Capay Stage”), Llajas Formation, southwestern Santa Susana Mountains, Los Angeles County, southern
|
||
<collectingRegion id="D872A6B8FFF1FF8AFB35FABDFADCFAC5" box="[1175,1302,1286,1312]" country="United States of America" name="California" pageId="34" pageNumber="33">California</collectingRegion>
|
||
, Ventura County (new information) to Maniobra Formation, Orocopia Mountains, Riverside County, southern
|
||
<collectingRegion id="D872A6B8FFF1FF8AFAC8FAF7FA25FA83" box="[1386,1519,1356,1382]" country="United States of America" name="California" pageId="34" pageNumber="33">California</collectingRegion>
|
||
(
|
||
<bibRefCitation id="7E2715ABFFF1FF8AFCE9FACBFB55FA6F" author="Squires, R. L. & D. M. Advocate" box="[843,1183,1392,1418]" pageId="34" pageNumber="33" pagination="851 - 864" refId="ref43291" refString="Squires, R. L. and D. M. Advocate 1986. New early Eocene mollusks from the Orocopia Mountains, southern California. Journal of Paleontology 60 (4): 851 - 864, figs, 1 - 3." type="journal article" year="1986">Squires and Advocate 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E2715ABFFF1FF8AFB0FFACBFA98FA6F" author="Squires, R. L." box="[1197,1362,1392,1418]" pageId="34" pageNumber="33" pagination="217 - 226" refId="ref42749" refString="Squires, R. L. 1991. Molluscan paleontology of the lower Eocene Maniobra Formation, Orocopia Mountains, southern California. In P. L. Abbott and J. A. May (eds.). Eocene Geologic History San Diego Region. Pacific Section, Society of Economic Paleontologists and Mineralogists. 68: 217 - 226, pls. 1 - 2." type="journal article" year="1991">Squires 1991</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52AC3BD1FFF1FF8AFCC2FA28FA12FA35" pageId="34" pageNumber="33" type="etymology">
|
||
<paragraph id="1A09685AFFF1FF8AFCC2FA28FA12FA35" blockId="34.[830,1520,334,1947]" pageId="34" pageNumber="33">
|
||
<emphasis id="28C2B448FFF1FF8AFCC2FA28FBC1FA48" bold="true" box="[864,1035,1427,1453]" pageId="34" pageNumber="33">Etymology—</emphasis>
|
||
Originally named for the Eocene Manio- bra Formation, Riverside County, southern
|
||
<collectingRegion id="D872A6B8FFF1FF8AFAF5FA0DFA1EFA35" box="[1367,1492,1462,1488]" country="United States of America" name="California" pageId="34" pageNumber="33">California</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52AC3BD1FFF1FF8AFCC2FA62FA3AF923" pageId="34" pageNumber="33" type="description">
|
||
<paragraph id="1A09685AFFF1FF8AFCC2FA62FA3AF923" blockId="34.[830,1520,334,1947]" pageId="34" pageNumber="33">
|
||
<emphasis id="28C2B448FFF1FF8AFCC2FA62FBD2FA16" bold="true" box="[864,1048,1497,1523]" pageId="34" pageNumber="33">Description—</emphasis>
|
||
Shell size medium; very inflated; ovatepyriform. Shell widest medially. Base inflated. Spire in- volute. Aperture narrow and near right margin of shell; aperture elongate anteriorly and curved posteriorly. Aperture narrow and prominently curved. Outer lip nar- row, with numerous teeth that become stronger toward anterior of aperture. Inner lip with numerous small teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52AC3BD1FFF1FF8BFCC2F96BFD0BFE68" lastPageId="35" lastPageNumber="34" pageId="34" pageNumber="33" type="discussion">
|
||
<paragraph id="1A09685AFFF1FF8BFCC2F96BFD0BFE68" blockId="34.[830,1520,334,1947]" lastBlockId="35.[112,800,193,397]" lastPageId="35" lastPageNumber="34" pageId="34" pageNumber="33">
|
||
<emphasis id="28C2B448FFF1FF8AFCC2F96BFC33F90F" bold="true" box="[864,1017,1744,1770]" pageId="34" pageNumber="33">Remarks—</emphasis>
|
||
This is an uncommon species. The an- teriormost part of the
|
||
<typeStatus id="C50DD6F8FFF1FF8AFBC0F948FB1BF8E8" box="[1122,1233,1779,1805]" pageId="34" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
is missing. The dorsal view of this species is shown for the first time in
|
||
<figureCitation id="828D74DFFFF1FF8AFA3DF8ADFCA1F8B6" captionStart="Figure 4" captionStartId="9.[112,199,195,221]" captionTargetId="figure-13@8.[123,1510,192,1968]" captionTargetPageId="8" captionText="Figure 4 (previous page). A–C. Nucleolaria cowlitzensis Groves, 1994, Cowlitz Formation (middle Eocene, upper “Tejon Stage”), Vader area, south-central Lewis County, Washington. Basal (A), dorsal (B), and left (columellar) side (C) views, 1.9x, length 27.2 mm, width 17.3 mm, height 11.3 mm, of holotype UCMP 39837, UCMP Locality D-8040. D–H. Propustularia kemperae (Nelson, 1925), Santa Susana Formation, (Paleocene, Selandian Stage), north side Simi Hills, Ventura County, southern California. Basal (D), dorsal (E), and right (labral) side (F), and views, 2.2x, length 21 mm, width 17.3, height 12 mm, of holotype CASG 391.01 [ex CASG 987], Locality CASG 391. Dorsal (G) view, 2.6x, length 18.2 mm, width 14.9, height 11.2 mm, of hypotype LACMIP 22307.6, LACMIP Type 14918 (LACMIP Locality 22307). Basal (H) view, 2.2x, length 21 mm, width 17.3, height 12 mm, of paratype UCMP 30541, UCMP Locality 3764. I–N. Subepona goedertorum (Groves and Squires, 1995), Crescent Forma- tion (Eocene, “Capay Stage”), Black Hills, Thurston County, Washington. Basal (I), dorsal (J), left (columellar) lateral (K), and right (labral) (L) views, 2.8x, length 19.4 mm, width 12.3, height 10.8 mm, of holotype LACMIP 16655.23, LACMIP Type 12375, LACMIP Locality 16655 [=LACMIP Locality 41563; ex CSUN Locality 1563]. M–N, Juncal Formation, Canton Canyon, Whitaker Peak area, Ventura County, southern California. Basal (M) and dorsal (N) views, both 2.4x, length 19.4 mm, width 13.9, height 9.4 mm, of hypotype LACMIP 40807.1, LACMIP Type 14935 (LACMIP Locality 40807 [= LACMIP Locality 16191, ex CSUN Locality 807]). O–Q, Subepona leahae n. sp., Llajas Formation (Eocene, “Domengine Stage”), north side Simi Valley, Ventura County, southern California. Basal (O), dorsal (P), and left (labral) side (Q) views, 1.6x, length 29.5 mm, width 13.9, height 15.1 mm, of holotype LACMIP 40374.79, LACMIP Type 14936, (LACMIP Locality 40374)." figureDoi="http://doi.org/10.5281/zenodo.13869592" httpUri="https://zenodo.org/record/13869592/files/figure.png" pageId="34" pageNumber="33">
|
||
Figure
|
||
<specimenCount id="0CB0A3D3FFF1FF8AFCE2F882FCA5F8B6" box="[832,879,1849,1875]" count="4" pageId="34" pageNumber="33" type="generic">4M.</specimenCount>
|
||
</figureCitation>
|
||
<taxonomicName id="DDB613D9FFF1FF8AFCD5F883FB08F8B1" baseAuthorityName="Squires and Advocate" baseAuthorityYear="1986" box="[887,1218,1848,1876]" class="Gastropoda" family="Eocypraeidae" genus="Luponovula" kingdom="Animalia" order="Littorinimorpha" pageId="34" pageNumber="33" phylum="Mollusca" rank="species" species="maniobraensis">
|
||
<emphasis id="28C2B448FFF1FF8AFCD5F883FB08F8B1" box="[887,1218,1848,1876]" italics="true" pageId="34" pageNumber="33">Luponovula maniobraensis</emphasis>
|
||
</taxonomicName>
|
||
most closely resembles
|
||
<taxonomicName id="DDB613D9FFF1FF8AFCE2F8E0FC05F89C" box="[832,975,1883,1913]" class="Gastropoda" family="Eocypraeidae" genus="Luponovula" kingdom="Animalia" order="Littorinimorpha" pageId="34" pageNumber="33" phylum="Mollusca" rank="species" species="oligovata">
|
||
<emphasis id="28C2B448FFF1FF8AFCE2F8E0FC05F89C" box="[832,975,1883,1913]" italics="true" pageId="34" pageNumber="33">L. oligovata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E2715ABFFF1FF8AFC40F8E6FBBAF892" author="Sacco, F." box="[994,1136,1885,1911]" pageId="34" pageNumber="33" refId="ref41565" refString="Sacco, F. 1894. I molluschi dei terreni terziarii del Piemonte e della Liguria. Partie 15: (Cypraeidae, ed Amphiperasidae). Torino, Carlo Clausen, 74 pp., 3 pls." type="book" year="1894">Sacco 1894</bibRefCitation>
|
||
: pl. 3, fig. 25;
|
||
<bibRefCitation id="7E2715ABFFF1FF8AFABDF8E6FA22F892" author="Cossmann, A. E. M." box="[1311,1512,1885,1911]" pageId="34" pageNumber="33" refId="ref35846" refString="Cossmann, A. E. M. 1903. Essais de paleoconchologie comparee. Vol. 5. Privately published. Paris. 215 pp., 16 figs., 9 pls." type="book" year="1903">Cossmann 1903</bibRefCitation>
|
||
: pl. 7, fig. 2) of late early Oligocene age (Latorfian) from the
|
||
<collectingRegion id="D872A6B8FFF0FF8BFF3FFF78FF3CFF38" box="[157,246,195,221]" country="Italy" name="Liguria" pageId="35" pageNumber="34">Liguria</collectingRegion>
|
||
and
|
||
<collectingRegion id="D872A6B8FFF0FF8BFE8CFF78FE6DFF38" box="[302,423,195,221]" country="Italy" name="Piemonte" pageId="35" pageNumber="34">Piedmont</collectingRegion>
|
||
regions of
|
||
<collectingCountry id="62A128CAFFF0FF8BFD89FF78FDA9FF38" box="[555,611,195,221]" name="Italy" pageId="35" pageNumber="34">Italy</collectingCountry>
|
||
and
|
||
<taxonomicName id="DDB613D9FFF0FF8BFD38FF7AFCEAFF38" baseAuthorityName="de Gregorio" baseAuthorityYear="1880" box="[666,800,193,221]" class="Gastropoda" family="Eocypraeidae" genus="Luponovula" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="34" phylum="Mollusca" rank="species" species="normalis">
|
||
<emphasis id="28C2B448FFF0FF8BFD38FF7AFCEAFF38" box="[666,800,193,221]" italics="true" pageId="35" pageNumber="34">L. normalis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E2715ABFFF0FF8BFFDBFF5DFE90FEE5" author="De Gregorio, A." box="[121,346,230,256]" pageId="35" pageNumber="34" pagination="1 - 9" refId="ref36098" refString="De Gregorio, A. 1880. Fauna di S. Giovanni Ilarione (Pa- risiano). Parte 1: Cefalopodi e Gasteropodi. Palermo. XXVIII + 1 - 106, pls. 1 - 9." type="book chapter" year="1880">de Gregorio 1880</bibRefCitation>
|
||
: pl. 6, fig. 14) of middle Eocene age (Lutetian) from San Giovanni Illarione,
|
||
<collectingRegion id="D872A6B8FFF0FF8BFDFDFEB1FD7DFEC1" box="[607,695,266,292]" country="Italy" name="Veneto" pageId="35" pageNumber="34">Veneto</collectingRegion>
|
||
,
|
||
<collectingCountry id="62A128CAFFF0FF8BFD61FEB2FD31FEC6" box="[707,763,265,291]" name="Italy" pageId="35" pageNumber="34">Italy</collectingCountry>
|
||
. In comparison,
|
||
<taxonomicName id="DDB613D9FFF0FF8BFEAFFE90FD9BFEA2" baseAuthorityName="Squires and Advocate" baseAuthorityYear="1986" box="[269,593,299,327]" class="Gastropoda" family="Eocypraeidae" genus="Luponovula" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="34" phylum="Mollusca" rank="species" species="maniobraensis">
|
||
<emphasis id="28C2B448FFF0FF8BFEAFFE90FD9BFEA2" box="[269,593,299,327]" italics="true" pageId="35" pageNumber="34">Luponovula maniobraensis</emphasis>
|
||
</taxonomicName>
|
||
differs by having fewer teeth on its columellar lip, as well as having fewer and shorter teeth on the interior of its outer lip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |