163 lines
19 KiB
XML
163 lines
19 KiB
XML
<document id="CA85CF93C666208CBFCE2B1A25DA2207" ID-DOI="10.5281/zenodo.13652021" ID-ISSN="1732-2421" ID-Zenodo-Dep="13652021" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1723141930295" checkinUser="felipe" docAuthor="Ernst, Andrej & Minwegen, Elke" docDate="2006" docId="92125B5E2400FF92FFD067A473896B15" docLanguage="en" docName="ActaPalaeontolPol.51.3.569-588.pdf" docOrigin="Acta Palaeontologica Polonica 51 (3)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Coscinium hermidensis Ernst & Minwegen, 2006, sp. nov." docType="treatment" docVersion="3" lastPageNumber="571" masterDocId="6E2B23262402FF90FFB6610777056E4B" masterDocTitle="Late Carboniferous bryozoans from La Hermida, Spain" masterLastPageNumber="588" masterPageNumber="569" pageNumber="571" updateTime="1725393044933" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="6E897EB6C5A06AE5028E1AB6E8846A92">
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<mods:title id="D55ACEF23893B507361F088ABB8A14E9">Late Carboniferous bryozoans from La Hermida, Spain</mods:title>
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<mods:namePart id="D4F4A4035F2FD9D1340A53861438A4F9">Ernst, Andrej</mods:namePart>
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<mods:namePart id="AD2BBB98290048DAE76F9F24222D4A6A">Minwegen, Elke</mods:namePart>
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<mods:typeOfResource id="5F10D810D851B0DFB1449670D1529E7E">text</mods:typeOfResource>
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<mods:title id="F7DA2C379B75D10804117677AE69AD9A">Acta Palaeontologica Polonica</mods:title>
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</mods:titleInfo>
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<mods:part id="B790B43ED3856EEE426FE4432ACF75D5">
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<mods:date id="3080F85BC86B49D21216720273ED4F79">2006</mods:date>
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<mods:number id="B537AD18AF5F94AB363436C50F1BFB31">51</mods:number>
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<mods:identifier id="3DFF88591C2CDDF11A9589D1CC9E4463" type="DOI">10.5281/zenodo.13652021</mods:identifier>
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<mods:identifier id="FF15218A770AF99639E641A20AC43259" type="ISSN">1732-2421</mods:identifier>
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<mods:identifier id="2151BDF4F257365F607E73341CA9ACAC" type="Zenodo-Dep">13652021</mods:identifier>
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<treatment id="92125B5E2400FF92FFD067A473896B15" ID-DOI="http://doi.org/10.5281/zenodo.13652025" ID-Zenodo-Dep="13652025" LSID="urn:lsid:plazi:treatment:92125B5E2400FF92FFD067A473896B15" httpUri="http://treatment.plazi.org/id/92125B5E2400FF92FFD067A473896B15" lastPageNumber="571" pageId="2" pageNumber="571">
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<subSubSection id="52A1B9C32400FF92FFD067A476FB68F5" box="[102,510,1698,1727]" pageId="2" pageNumber="571" type="nomenclature">
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<paragraph id="1A04EA482400FF92FFD067A476FB68F5" blockId="2.[102,510,1698,1759]" box="[102,510,1698,1727]" pageId="2" pageNumber="571">
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<heading id="414C5D242400FF92FFD067A476FB68F5" box="[102,510,1698,1727]" fontSize="12" level="2" pageId="2" pageNumber="571" reason="9">
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<taxonomicName id="DDBB91CB2400FF92FFD067A4769768F4" ID-CoL="86FZK" authorityName="Ernst & Minwegen" authorityYear="2006" box="[102,402,1699,1727]" class="Stenolaemata" family="Hexagonellidae" genus="Coscinium" kingdom="Animalia" order="Cystoporida" pageId="2" pageNumber="571" phylum="Bryozoa" rank="species" species="hermidensis" status="sp. nov.">
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<emphasis id="28CF365A2400FF92FFD067A4769768F4" box="[102,402,1699,1727]" italics="true" pageId="2" pageNumber="571">Coscinium hermidensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="33FC8B212400FF92FE2F67A576FB68F5" box="[409,510,1698,1726]" pageId="2" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="52A1B9C32400FF92FFD067CD73896B15" pageId="2" pageNumber="571" type="description">
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<paragraph id="1A04EA482400FF92FFD067CD76216894" blockId="2.[102,510,1698,1759]" box="[102,292,1738,1759]" pageId="2" pageNumber="571">
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<figureCitation id="8280F6CD2400FF92FFD067CD77C96894" box="[102,204,1738,1759]" captionStart="Fig" captionStartId="4.[102,135,1809,1829]" captionText="Fig. 3. Cystoporid bryozoans Fistulipora, Coscinium, and Cystodictya from the La Hermida locality, Spain, Late Carboniferous (Moscovian). A–C. Fistulipora petaloida Schulga−Nesterenko, 1955. A. SMF 1737, longitudinal section displaying styles in outer granular skeleton and skeletal vesicles. B. SMF 1759, tangential section of the colony surface: close view of a single aperture (B1) and view of several apertures (B2). C. SMF 1737, longitudinal section of autozooecial chamber showing diaphragm and vesicular skeleton. D–F. Coscinium hermidensis sp. nov. D. Holotype SMF 1729, tangential section of the colony surface displaying apertures (D1, arrow—lunarium) and fused branches (D2). E. SMF 1736, longitudinal section displaying autozooecial chambers with diaphragms and vesicular skeleton. F. Paratype SMF 1775, cross section of the branch displaying mesotheca with rods (F1), autozooecial chambers and vesicular skeleton (F2). G, H. Cystodyctia pustulosa sp. nov. G. Holotype SMF 1776, tangential section of the colony displaying apertures. H. Paratype SMF 1743, cross section of the branch." pageId="2" pageNumber="571">Fig. 3D–F</figureCitation>
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;
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<tableCitation id="5739DFF32400FF92FF6367CD76256894" box="[213,288,1738,1759]" captionStart="Table2" captionStartId="2.[812,867,232,253]" captionText="Table2.Measurementsof Cosciniumhermidensis sp.nov.N,number of measurements; X, mean; SD, standard deviation; CV, coefficient of variation; MIN, minimum value; MAX, maximum value." pageId="2" pageNumber="571">Table 2</tableCitation>
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.
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</paragraph>
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<paragraph id="1A04EA482400FF92FFD067F6752F694E" blockId="2.[102,775,1776,1953]" box="[102,554,1776,1798]" pageId="2" pageNumber="571">
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<emphasis id="28CF365A2400FF92FFD067F67641694D" box="[102,324,1777,1798]" italics="true" pageId="2" pageNumber="571">Derivation of the name</emphasis>
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: After the
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<typeStatus id="C50054EA2400FF92FE1A67F776D0694E" box="[428,469,1776,1797]" pageId="2" pageNumber="571">type</typeStatus>
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locality.
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</paragraph>
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<paragraph id="1A04EA482400FF92FFD0661577DE6908" blockId="2.[102,775,1776,1953]" pageId="2" pageNumber="571">
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<emphasis id="28CF365A2400FF92FFD0661577EC696C" box="[102,233,1810,1831]" italics="true" pageId="2" pageNumber="571">Type material</emphasis>
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:
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<materialsCitation id="AAD3E0152400FF92FF45661577DE6908" collectionCode="SMF" pageId="2" pageNumber="571" specimenCode="SMF 1729, SMF 1775, SMF 1736" specimenCount="2" typeStatus="holotype">
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<typeStatus id="C50054EA2400FF92FF4566157649696C" box="[243,332,1810,1831]" pageId="2" pageNumber="571" type="holotype">Holotype</typeStatus>
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(
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<specimenCode id="4A1D42332400FF92FEE1661576B8696C" box="[343,445,1810,1831]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="571">SMF 1729</specimenCode>
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) and
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<specimenCount id="0CBD21C12400FF92FE5866157574696C" box="[494,625,1810,1831]" count="2" pageId="2" pageNumber="571" type="generic" typeStatus="paratype">two paratypes</specimenCount>
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(
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<specimenCode id="4A1D42332400FF92FDCD661575E5696C" box="[635,736,1810,1831]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="571">SMF 1775</specimenCode>
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and
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<specimenCode id="4A1D42332400FF92FFD0662977D56908" box="[102,208,1838,1859]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="571">SMF 1736</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="1A04EA482400FF92FFD0664876A9692F" blockId="2.[102,775,1776,1953]" box="[102,428,1871,1892]" pageId="2" pageNumber="571">
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<materialsCitation id="AAD3E0152400FF92FFD0664876AD692F" box="[102,424,1871,1892]" collectorName="La Hermida" country="Spain" location="Spain" pageId="2" pageNumber="571" specimenCount="1" typeStatus="holotype">
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<emphasis id="28CF365A2400FF92FFD0664877E7692F" box="[102,226,1871,1892]" italics="true" pageId="2" pageNumber="571">
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<typeStatus id="C50054EA2400FF92FFD066487790692F" box="[102,149,1871,1892]" pageId="2" pageNumber="571">Type</typeStatus>
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locality
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</emphasis>
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:
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<collectorName id="B74E8F9E2400FF92FF5966487661692F" box="[239,356,1871,1892]" pageId="2" pageNumber="571">La Hermida</collectorName>
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,
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<collectingCountry id="62ACAAD82400FF92FED8664876AD692F" box="[366,424,1871,1892]" name="Spain" pageId="2" pageNumber="571">Spain</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="1A04EA482400FF92FFD0667677A469EA" blockId="2.[102,775,1776,1953]" pageId="2" pageNumber="571">
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<emphasis id="28CF365A2400FF92FFD0667677E469CD" box="[102,225,1905,1926]" italics="true" pageId="2" pageNumber="571">
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<typeStatus id="C50054EA2400FF92FFD06676779069CD" box="[102,149,1905,1926]" pageId="2" pageNumber="571">Type</typeStatus>
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horizon
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</emphasis>
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: Picos de Europa Formation, Moscovian, Upper Carboniferous.
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</paragraph>
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<paragraph id="1A04EA482400FF92FFD066B576A269A1" blockId="2.[102,774,1969,2026]" pageId="2" pageNumber="571">
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<emphasis id="28CF365A2400FF92FFD066B576096981" box="[102,268,1970,1994]" italics="true" pageId="2" pageNumber="571">Other material</emphasis>
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.—SMF 1725, 1746, 1752, 1761, 1771, 1796, and 21 additional fragments.
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</paragraph>
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<paragraph id="1A04EA482400FF92FC9A631E73CB6CBA" blockId="2.[812,1485,536,753]" pageId="2" pageNumber="571">
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<emphasis id="28CF365A2400FF92FC9A631E749B6C7B" box="[812,926,537,560]" italics="true" pageId="2" pageNumber="571">Diagnosis</emphasis>
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.—
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<taxonomicName id="DDBB91CB2400FF92FC77631F73C96C7B" authorityName="Ernst & Minwegen" authorityYear="2006" box="[961,1228,536,560]" class="Stenolaemata" family="Hexagonellidae" genus="Coscinium" kingdom="Animalia" order="Cystoporida" pageId="2" pageNumber="571" phylum="Bryozoa" rank="species" species="hermidensis" status="sp. nov.">
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<emphasis id="28CF365A2400FF92FC77631F73C96C7B" box="[961,1228,536,560]" italics="true" pageId="2" pageNumber="571">Coscinium hermidensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="33FC8B212400FF92FB6E631E72336C7B" box="[1240,1334,537,560]" pageId="2" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
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differs from the most similar species
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<taxonomicName id="DDBB91CB2400FF92FB8B633F727D6C1A" authority="Keyserling, 1846" authorityName="Keyserling" authorityYear="1846" box="[1085,1400,568,593]" class="Stenolaemata" family="Hexagonellidae" genus="Coscinium" kingdom="Animalia" order="Cystoporida" pageId="2" pageNumber="571" phylum="Bryozoa" rank="species" species="cyclops">
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<emphasis id="28CF365A2400FF92FB8B633F73B46C1B" box="[1085,1201,568,592]" italics="true" pageId="2" pageNumber="571">C. cyclops</emphasis>
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<bibRefCitation id="7E2A97B92400FF92FB0E633F727D6C1A" author="Keyserling, A." box="[1208,1400,568,593]" pageId="2" pageNumber="571" refId="ref12228" refString="Keyserling, A. 1846. Wissenschaftliche Beobachtungen auf einer Reise in das Petschora-Land im Jahre 1843. 465 pp. Geognostische Bobachtungen, St. Petersburg." type="book" year="1846">
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Keyserling,
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<quantity id="DD4347AD2400FF92FAF6633F727D6C1A" box="[1344,1400,568,593]" metricMagnitude="1" metricUnit="m" metricValue="4.68884" pageId="2" pageNumber="571" unit="in" value="1846.0">1846</quantity>
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</bibRefCitation>
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</taxonomicName>
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in having smaller apertures (0.14–0.19
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<emphasis id="28CF365A2400FF92FB1E635E73EA6C3B" box="[1192,1263,601,624]" italics="true" pageId="2" pageNumber="571">versus</emphasis>
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||
<quantity id="DD4347AD2400FF92FB4F635F72656C3B" box="[1273,1376,600,625]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="571" unit="mm" value="0.25">0.25 mm</quantity>
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in
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<taxonomicName id="DDBB91CB2400FF92FA3E635F74626CDB" authorityName="Keyserling" authorityYear="1846" class="Stenolaemata" family="Hexagonellidae" genus="Coscinium" kingdom="Animalia" order="Cystoporida" pageId="2" pageNumber="571" phylum="Bryozoa" rank="species" species="cyclops">
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<emphasis id="28CF365A2400FF92FA3E635F74626CDB" italics="true" pageId="2" pageNumber="571">C. cyclops</emphasis>
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</taxonomicName>
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) and in more closely spaced apertures (12–13
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<emphasis id="28CF365A2400FF92FA33637E72C96CDB" box="[1413,1484,633,656]" italics="true" pageId="2" pageNumber="571">versus</emphasis>
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average 8 per
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||
<quantity id="DD4347AD2400FF92FC7C639F730F6CFB" box="[970,1034,664,689]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="571" unit="mm" value="5.0">5 mm</quantity>
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longitudinally). The new species differs from
|
||
<taxonomicName id="DDBB91CB2400FF92FCDF63BF73D96C9A" authority="Stuckenberg, 1895" authorityName="Stuckenberg" authorityYear="1895" box="[873,1244,696,721]" class="Stenolaemata" family="Hexagonellidae" genus="Coscinium" kingdom="Animalia" order="Cystoporida" pageId="2" pageNumber="571" phylum="Bryozoa" rank="species" species="keyserlingi">
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<emphasis id="28CF365A2400FF92FCDF63BF73016C9B" box="[873,1028,696,720]" italics="true" pageId="2" pageNumber="571">C. keyserlingi</emphasis>
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||
<bibRefCitation id="7E2A97B92400FF92FBBC63BF73D96C9A" author="Stuckenberg, A." box="[1034,1244,696,721]" pageId="2" pageNumber="571" pagination="22 - 24" refId="ref13549" refString="Stuckenberg, A. 1895. Korallen und Bryozoen der Steinkohlen-Ablagerungen des Urals und des Timan [in Russian]. Trudy Geologiceskogo Komiteta 10: 22 - 24, 120 - 127, 137 - 177." type="journal article" year="1895">
|
||
Stuckenberg,
|
||
<quantity id="DD4347AD2400FF92FB1263BF73D96C9A" box="[1188,1244,696,721]" metricMagnitude="1" metricUnit="m" metricValue="4.8133" pageId="2" pageNumber="571" unit="in" value="1895.0">1895</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
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||
in having smaller apertures and poorly developed lunaria.
|
||
</paragraph>
|
||
<paragraph id="1A04EA482400FF92FC9A620173896B15" blockId="2.[812,1486,773,1374]" pageId="2" pageNumber="571">
|
||
<emphasis id="28CF365A2400FF92FC9A620174B56D56" box="[812,944,774,797]" italics="true" pageId="2" pageNumber="571">Description</emphasis>
|
||
.—Bifoliate frondescent colony. Branches lens−shaped in cross−section, 2.63–4.00 mm wide and
|
||
<quantity id="DD4347AD2400FF92FAEA6222745D6D16" metricMagnitude="-4" metricUnit="m" metricValue="8.65" metricValueMax="10.5" metricValueMin="6.8" pageId="2" pageNumber="571" unit="mm" value="0.865" valueMax="1.05" valueMin="0.68">0.68–1.05 mm</quantity>
|
||
thick, occasionally anastomosing to give oval fenestrules,
|
||
<quantity id="DD4347AD2400FF92FCCC626274D66D36" box="[890,979,869,894]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="571" unit="mm" value="0.9">0.9 mm</quantity>
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||
wide and 2.0 mm long. Mesotheca straight, three−layered, consisting of dark medial layer and two outer pale layers,
|
||
<quantity id="DD4347AD2400FF92FC0062A273596DF6" box="[950,1116,933,958]" metricMagnitude="-5" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="571" unit="mm" value="0.045" valueMax="0.05" valueMin="0.04">0.04–0.05 mm</quantity>
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||
thick, containing abundant hyaline rods. Rods
|
||
<quantity id="DD4347AD2400FF92FC5562C2734C6D96" box="[995,1097,965,990]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="571" unit="mm" value="0.01">0.01 mm</quantity>
|
||
in diameter, spaced densely (
|
||
<figureCitation id="8280F6CD2400FF92FA1662C2744F6DB5" captionStart="Fig" captionStartId="4.[102,135,1809,1829]" captionText="Fig. 3. Cystoporid bryozoans Fistulipora, Coscinium, and Cystodictya from the La Hermida locality, Spain, Late Carboniferous (Moscovian). A–C. Fistulipora petaloida Schulga−Nesterenko, 1955. A. SMF 1737, longitudinal section displaying styles in outer granular skeleton and skeletal vesicles. B. SMF 1759, tangential section of the colony surface: close view of a single aperture (B1) and view of several apertures (B2). C. SMF 1737, longitudinal section of autozooecial chamber showing diaphragm and vesicular skeleton. D–F. Coscinium hermidensis sp. nov. D. Holotype SMF 1729, tangential section of the colony surface displaying apertures (D1, arrow—lunarium) and fused branches (D2). E. SMF 1736, longitudinal section displaying autozooecial chambers with diaphragms and vesicular skeleton. F. Paratype SMF 1775, cross section of the branch displaying mesotheca with rods (F1), autozooecial chambers and vesicular skeleton (F2). G, H. Cystodyctia pustulosa sp. nov. G. Holotype SMF 1776, tangential section of the colony displaying apertures. H. Paratype SMF 1743, cross section of the branch." pageId="2" pageNumber="571">Fig. 3F</figureCitation>
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<figureCitation id="8280F6CD2400FF92FCFC62F674566A4A" box="[842,851,1009,1025]" captionStart="Fig" captionStartId="1.[102,135,1099,1119]" captionTargetBox="[103,1483,227,1070]" captionTargetId="figure-260@1.[104,1479,229,1071]" captionTargetPageId="1" captionText="Fig. 1. Geographical position of the at locality La Hermida, Spain." figureDoi="http://doi.org/10.5281/zenodo.13652023" httpUri="https://zenodo.org/record/13652023/files/figure.png" pageId="2" pageNumber="571">
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<subScript id="863FE80D2400FF92FCFC62F674566A4A" attach="left" box="[842,851,1009,1025]" fontSize="7" pageId="2" pageNumber="571">1</subScript>
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</figureCitation>
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). Indistinct longitudinal crests present on mesotheca. Autozooecia short, budding parallel to the mesotheca for a distance of about three zooecial diameters, semicircular in basal cross section, completely separated by vesicular skeleton, arranged in 8–10 rows on branches. Apertures oval, spaced 9–10 longitudinally and 12–13 diagonally in
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<quantity id="DD4347AD2400FF92FA3C658272C86AD6" box="[1418,1485,1157,1182]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="571" unit="mm" value="5.0">5 mm</quantity>
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distance. Lunaria weakly developed. Autozooecia surrounded by a thick layer of granular skeleton. Autozooecial diaphragms rare to common, thin, planar. Skeletal vesicles relatively large, with rounded roofs, covered at the colony surface by a thick layer of dense calcitic material, arranged in 2–3 rows between autozooecia.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |