266 lines
22 KiB
XML
266 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/BDJ.3.e4596" ID-PMC="PMC4698461" ID-Pensoft-Pub="1314-2828--4596" ID-PubMed="26752964" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1314-2828--e4596" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="Nine new species of Itaplectops (Diptera: Tachinidae) reared from caterpillars in Area de Conservación Guanacaste, northwestern Costa Rica, with a key to Itaplectops species" checkinTime="1585357573225" checkinUser="pensoft" docAuthor="Fleming, AJ, Wood, D. Monty, Smith, M. Alex, Janzen, Daniel H. & Hallwachs, Winnie" docDate="2015" docId="91566459FB069A4B671EC03E13A90833" docLanguage="en" docName="BiodivDatJour 3: e4596" docOrigin="Biodiversity Data Journal 3" docSource="http://dx.doi.org/10.3897/BDJ.3.e4596" docTitle="Itaplectops argentifrons Fleming & Wood 2014, sp. n." docType="treatment" docVersion="3" lastPageNumber="4596" masterDocId="A4229B60FFCCFFBA4464FFC3AE4D9158" masterDocTitle="Nine new species of Itaplectops (Diptera: Tachinidae) reared from caterpillars in Area de Conservacion Guanacaste, northwestern Costa Rica, with a key to Itaplectops species" masterLastPageNumber="4596" masterPageNumber="4596" pageNumber="4596" updateTime="1668122940553" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Nine new species of Itaplectops (Diptera: Tachinidae) reared from caterpillars in Area de Conservacion Guanacaste, northwestern Costa Rica, with a key to Itaplectops species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fleming, AJ</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wood, D. Monty</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Smith, M. Alex</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Janzen, Daniel H.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hallwachs, Winnie</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Biodiversity Data Journal</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>4596</mods:start>
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<mods:end>4596</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/BDJ.3.e4596</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.3.e4596</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2828--4596</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:91566459FB069A4B671EC03E13A90833" httpUri="http://treatment.plazi.org/id/91566459FB069A4B671EC03E13A90833" lastPageNumber="4596" pageId="0" pageNumber="4596">
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<subSubSection pageId="0" pageNumber="4596" type="nomenclature">
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<paragraph pageId="0" pageNumber="4596">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:B8B22BAF-46B8-4544-B7E5-9061A72A1445" authority="Fleming & Wood, 2014" authorityName="Fleming & Wood" authorityYear="2014" class="Insecta" family="Tachinidae" genus="Itaplectops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Itaplectops argentifrons" order="Diptera" pageId="0" pageNumber="4596" phylum="Arthropoda" rank="species" species="argentifrons">Itaplectops argentifrons Fleming & Wood, 2014</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="4596">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="4596" type="materials_examined">
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<paragraph pageId="0" pageNumber="4596">Materials</paragraph>
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<paragraph pageId="0" pageNumber="4596">
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<materialsCitation collectingDate="2011-08-25" collectingMethod="reared from caterpillar of Eucleamesoamericana (Limacodidae)" collectionCode="CNC, Insects" collectorName="D. H. Janzen & W. Hallwachs, Minor Carmona" country="Costa Rica" latitude="10.989" location="Sector Rincon Rain Forest" longitude="-85.426" pageId="0" pageNumber="4596" specimenCode="11 - SRNP- 32160" specimenCount="1" typeStatus="Holotype">
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Type status:
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<typeStatus pageId="0" pageNumber="4596">Holotype</typeStatus>
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. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
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<specimenCode pageId="0" pageNumber="4596">11-SRNP-32160</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="4596">D.H. Janzen & W. Hallwachs, Minor Carmona</collectorName>
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; individualID: 11-SRNP-32160; individualCount:
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<specimenCount pageId="0" pageNumber="4596">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="4596">M</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="4596">adult</specimenType>
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; preparations: pinned; Taxon: scientificName: Itaplectopsargentifrons; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Itaplectops; specificEpithet: argentifrons; scientificNameAuthorship: Fleming & Wood; Location: continent: Central America; country:
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<collectingCountry pageId="0" pageNumber="4596">Costa Rica</collectingCountry>
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; countryCode: CR; stateProvince: Alajuela; county: Area de Conservacion Guanacaste; locality:
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<location LSID="urn:lsid:plazi:treatment:91566459FB069A4B671EC03E13A90833:26C722127D4D062947DF145657B99DA0" country="Costa Rica" latitude="10.989" longitude="-85.426" name="Sector Rincon Rain Forest" pageId="0" pageNumber="4596">Sector Rincon Rain Forest</location>
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; verbatimLocality: Puente Rio Negro; verbatimElevation:
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<elevation pageId="0" pageNumber="4596">675</elevation>
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; verbatimLatitude: 10.989; verbatimLongitude: -85.426; verbatimCoordinateSystem: Decimal; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="4596" value="10.989">10.989</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="4596" value="-85.426">-85.426</geoCoordinate>
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; Identification: identifiedBy:
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<determinerName pageId="0" pageNumber="4596">AJ Fleming</determinerName>
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; dateIdentified: 2014; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="4596">reared from caterpillar of Eucleamesoamericana (Limacodidae)</collectingMethod>
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; verbatimEventDate:
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<collectingDate pageId="0" pageNumber="4596" value="2011-08-25">25-Aug-2011</collectingDate>
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; Record Level: language: en; institutionCode:
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<collectionCode pageId="0" pageNumber="4596">CNC</collectionCode>
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; collectionCode:
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<collectionCode pageId="0" pageNumber="4596">Insects</collectionCode>
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; basisOfRecord: Pinned Specimen
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</materialsCitation>
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<materialsCitation collectingDate="2006-10-09" collectingMethod="reared from caterpillar of Eucleamesoamericana (Limacodidae)" collectionCode="CNC, Insects" collectorName="D. H. Janzen & W. Hallwachs, Freddy Quesada" country="Costa Rica" latitude="10.904" location="Sector Pitilla" longitude="-85.303" pageId="0" pageNumber="4596" specimenCode="06 - SRNP- 43634" specimenCount="1" typeStatus="Paratype">
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Type status:
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<typeStatus pageId="0" pageNumber="4596">Paratype</typeStatus>
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. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
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<specimenCode pageId="0" pageNumber="4596">06-SRNP-43634</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="4596">D.H. Janzen & W. Hallwachs, Freddy Quesada</collectorName>
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; individualID: 06-SRNP-43634; individualCount:
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<specimenCount pageId="0" pageNumber="4596">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="4596">M</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="4596">adult</specimenType>
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; preparations: pinned; Taxon: scientificName: Itaplectopsargentifrons; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Itaplectops; specificEpithet: argentifrons; scientificNameAuthorship: Fleming & Wood; Location: continent: Central America; country:
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<collectingCountry pageId="0" pageNumber="4596">Costa Rica</collectingCountry>
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; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservacion Guanacaste; locality:
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<location LSID="urn:lsid:plazi:treatment:91566459FB069A4B671EC03E13A90833:331DC4A7D2D0CA66596A2D74D346F046" country="Costa Rica" latitude="10.904" longitude="-85.303" name="Sector Pitilla" pageId="0" pageNumber="4596">Sector Pitilla</location>
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; verbatimLocality:
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<normalizedToken originalValue="Estación">Estacion</normalizedToken>
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Pitilla; verbatimElevation:
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<elevation pageId="0" pageNumber="4596">340</elevation>
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; verbatimLatitude: 10.904; verbatimLongitude: -85.303; verbatimCoordinateSystem: Decimal; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="4596" value="10.904">10.904</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="4596" value="-85.303">-85.303</geoCoordinate>
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; Identification: identifiedBy:
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<determinerName pageId="0" pageNumber="4596">AJ Fleming</determinerName>
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; dateIdentified: 2014; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="4596">reared from caterpillar of Eucleamesoamericana (Limacodidae)</collectingMethod>
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; verbatimEventDate:
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<collectingDate pageId="0" pageNumber="4596" value="2006-10-09">09-Oct-2006</collectingDate>
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; Record Level: language: en; institutionCode:
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<collectionCode pageId="0" pageNumber="4596">CNC</collectionCode>
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; collectionCode:
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<collectionCode pageId="0" pageNumber="4596">Insects</collectionCode>
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; basisOfRecord: Pinned Specimen
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</materialsCitation>
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<materialsCitation collectingDate="2006-10-09" collectingMethod="reared from caterpillar of Eucleamesoamericana (Limacodidae)" collectionCode="CNC, Insects" collectorName="D. H. Janzen & W. Hallwachs, Freddy Quesada" country="Costa Rica" latitude="10.904" location="Sector Pitilla" longitude="-85.303" pageId="0" pageNumber="4596" specimenCode="DHJPAR 0016106" specimenCount="1" typeStatus="Paratype">
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Type status:
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<typeStatus pageId="0" pageNumber="4596">Paratype</typeStatus>
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. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
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<specimenCode pageId="0" pageNumber="4596">DHJPAR0016106</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="4596">D.H. Janzen & W. Hallwachs, Freddy Quesada</collectorName>
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; individualID: DHJPAR0016106; individualCount:
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<specimenCount pageId="0" pageNumber="4596">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="4596">M</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="4596">adult</specimenType>
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; preparations: pinned; otherCatalogNumbers: ASTAP135-06, 06-SRNP-43634; Taxon: scientificName: Itaplectopsargentifrons; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Itaplectops; specificEpithet: argentifrons; scientificNameAuthorship: Fleming & Wood; Location: continent: Central America; country:
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<collectingCountry pageId="0" pageNumber="4596">Costa Rica</collectingCountry>
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; countryCode: CR; stateProvince: Guanacaste; county: Area de Conservacion Guanacaste; locality:
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<location LSID="urn:lsid:plazi:treatment:91566459FB069A4B671EC03E13A90833:332D70AA0292049AD8281226DDA2BA52" country="Costa Rica" latitude="10.904" longitude="-85.303" name="Sector Pitilla" pageId="0" pageNumber="4596">Sector Pitilla</location>
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; verbatimLocality:
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<normalizedToken originalValue="Estación">Estacion</normalizedToken>
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Pitilla; verbatimElevation:
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<elevation pageId="0" pageNumber="4596">340</elevation>
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; verbatimLatitude: 10.904; verbatimLongitude: -85.303; verbatimCoordinateSystem: Decimal; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="4596" value="10.904">10.904</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="4596" value="-85.303">-85.303</geoCoordinate>
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; Identification: identifiedBy:
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<determinerName pageId="0" pageNumber="4596">AJ Fleming</determinerName>
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; dateIdentified: 2014; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="4596">reared from caterpillar of Eucleamesoamericana (Limacodidae)</collectingMethod>
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; verbatimEventDate:
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<collectingDate pageId="0" pageNumber="4596" value="2006-10-09">09-Oct-2006</collectingDate>
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; Record Level: language: en; institutionCode:
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<collectionCode pageId="0" pageNumber="4596">CNC</collectionCode>
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; collectionCode:
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<collectionCode pageId="0" pageNumber="4596">Insects</collectionCode>
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; basisOfRecord: Pinned Specimen
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</materialsCitation>
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<materialsCitation collectingDate="2011-08-25" collectingMethod="reared from caterpillar of Eucleamesoamericana (Limacodidae)" collectionCode="CNC, Insects" collectorName="D. H. Janzen & W. Hallwachs, Minor Carmona" country="Costa Rica" latitude="10.989" location="Sector Rincon Rain Forest" longitude="-85.426" pageId="0" pageNumber="4596" specimenCode="DHJPAR 0044962" specimenCount="1" typeStatus="Paratype">
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Type status:
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<typeStatus pageId="0" pageNumber="4596">Paratype</typeStatus>
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. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
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<specimenCode pageId="0" pageNumber="4596">DHJPAR0044962</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="4596">D.H. Janzen & W. Hallwachs, Minor Carmona</collectorName>
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; individualID: DHJPAR0044962; individualCount:
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<specimenCount pageId="0" pageNumber="4596">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="4596">F</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="4596">adult</specimenType>
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; preparations: pinned; otherCatalogNumbers: ACGAZ186-11, 11-SRNP-32160; Taxon: scientificName: Itaplectopsargentifrons; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Itaplectops; specificEpithet: argentifrons; scientificNameAuthorship: Fleming & Wood; Location: continent: Central America; country:
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<collectingCountry pageId="0" pageNumber="4596">Costa Rica</collectingCountry>
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; countryCode: CR; stateProvince: Alajuela; county: Area de Conservacion Guanacaste; locality:
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<location LSID="urn:lsid:plazi:treatment:91566459FB069A4B671EC03E13A90833:1D2BE906769C45D7D5A603268905D9E7" country="Costa Rica" latitude="10.989" longitude="-85.426" name="Sector Rincon Rain Forest" pageId="0" pageNumber="4596">Sector Rincon Rain Forest</location>
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; verbatimLocality: Puente Rio Negro; verbatimElevation:
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<elevation pageId="0" pageNumber="4596">675</elevation>
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; verbatimLatitude: 10.989; verbatimLongitude: -85.426; verbatimCoordinateSystem: Decimal; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="4596" value="10.989">10.989</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="4596" value="-85.426">-85.426</geoCoordinate>
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; Identification: identifiedBy:
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<determinerName pageId="0" pageNumber="4596">AJ Fleming</determinerName>
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; dateIdentified: 2014; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="4596">reared from caterpillar of Eucleamesoamericana (Limacodidae)</collectingMethod>
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; verbatimEventDate:
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<collectingDate pageId="0" pageNumber="4596" value="2011-08-25">25-Aug-2011</collectingDate>
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; Record Level: language: en; institutionCode:
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<collectionCode pageId="0" pageNumber="4596">CNC</collectionCode>
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; collectionCode:
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<collectionCode pageId="0" pageNumber="4596">Insects</collectionCode>
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; basisOfRecord: Pinned Specimen
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="4596" type="description">
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<paragraph pageId="0" pageNumber="4596">Description</paragraph>
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<paragraph pageId="0" pageNumber="4596">Males and females</paragraph>
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<paragraph pageId="0" pageNumber="4596">Length: male 4-5mm; female 5mm.</paragraph>
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<paragraph pageId="0" pageNumber="4596">Head (Fig. 5c): proclinate orbital bristles present in both males and females; first flagellomere entirely dark or brownish orange over at least 1/2 of its surface; arista dark brown over 1/2 of its length, with gradual taper; first flagellomere reaching facial margin; ocellar bristles reduced, almost hair-like, no longer than length of pedicel, arising behind anterior ocellus; ocellar triangle covered in small proclinate hairs; frontal vitta approximately 2x as wide as fronto-orbital plate; facial ridge bearing 3-4 stout, decumbent bristles; fronto-orbital plate and parafacial entirely silver; parafacial bare; fronto-orbital plate of male with fine hairs confined to a row lateral to frontal bristles, these not extending past lowest frontal bristle; absent in female.</paragraph>
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<paragraph pageId="0" pageNumber="4596">Thorax (Fig. 5a): three postsutural supra-alar bristles, anteriormost greatly reduced to an almost hair-like structure; katepisternum with 2 bristles, anteriormost reduced in size and arising slightly behind suture; apical scutellar bristles long, up to 3/4 length of subapical scutellars; subapical scutellar bristles parallel or convergent (often crossed); scutellum with 1-2 pairs of widely separated discal bristles.</paragraph>
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<paragraph pageId="0" pageNumber="4596">Wings (Fig. 5a): smoky yellow.</paragraph>
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<paragraph pageId="0" pageNumber="4596">Legs (Fig. 5b): appearing dark overall, at least 1/2 of femur yellow, tibia yellow, and tarsi yellow (although these may appear dark due to hirsuteness); dorso-ventral margin of hind tarsi with yellow tufts of bristles apically.</paragraph>
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<paragraph pageId="0" pageNumber="4596">Abdomen (Fig. 5a, b): T1+2 with mid-dorsal depression extending along 2/3 of its length, but not reaching tergal margin; median marginal bristles present on T4 and T5 but absent on T1+2 and T3; discal bristles absent from all tergites; silver tomentosity on margins of abdominal segments T3 and T4 only visible under certain angles, and not extending beyond 1/3 of tergal surface.</paragraph>
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<paragraph pageId="0" pageNumber="4596">
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Male terminalia (Fig. 5d, e): both cerci tightly juxtaposed basally and diverging at their tips; haired up to tapering point, and bare until the tip; cercus, in lateral view, with a downward bend at 1/3 its length then curving back upwards apically,forming a very slight hook at its tip; surstylus 9/10 the length of the cercus, outwardly convex at its center so as to appear outwardly bowed with a slight inward bend apically; surstylus vaguely
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<normalizedToken originalValue="“S”">"S"</normalizedToken>
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shaped, with a downwardly hooked tip when viewed laterally; densely bristled along its entire length; phallus 2x as long as cercus, with a downward bend.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="4596" type="diagnosis">
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<paragraph pageId="0" pageNumber="4596">Diagnosis</paragraph>
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<paragraph pageId="0" pageNumber="4596">
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<taxonomicName class="Insecta" family="Tachinidae" genus="Itaplectops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Itaplectops argentifrons" order="Diptera" pageId="0" pageNumber="4596" phylum="Arthropoda" rank="species" species="argentifrons">Itaplectops argentifrons</taxonomicName>
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can be distinguished by the following combination of traits: proclinate orbital bristles present in males; fronto-orbital plate with silver sheen; first flagellomere brown/black over at least 1/2 of surface; three postsutural supra-alar bristles, anteriormost greatly reduced; median marginal bristles absent on T1+2 and T3 but present on T4 and T5; discal bristles absent from all tergites; silver tomentosity present on margins of abdominal segments T3 and T4. It can be distinguished from its most similar congener,
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<taxonomicName class="Insecta" family="Tachinidae" genus="Itaplectops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Itaplectops aurifrons" order="Diptera" pageId="0" pageNumber="4596" phylum="Arthropoda" rank="species" species="aurifrons">Itaplectops aurifrons</taxonomicName>
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, following couplet 7 in the key to
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<taxonomicName class="Insecta" family="Tachinidae" genus="Itaplectops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Itaplectops" order="Diptera" pageId="0" pageNumber="4596" phylum="Arthropoda" rank="genus">Itaplectops</taxonomicName>
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(below).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="4596" type="etymology">
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<paragraph pageId="0" pageNumber="4596">Etymology</paragraph>
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<paragraph pageId="0" pageNumber="4596">
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From the Latin adjective, argentea, meaning
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<normalizedToken originalValue="“silver-bearing”">"silver-bearing"</normalizedToken>
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, and the Latin noun frons, meaning "forehead" (a common term in insect anatomy), in reference to the solid silver tomentosity of the fronto-orbital plate and parafacial.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="4596" type="distribution">
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<paragraph pageId="0" pageNumber="4596">Distribution</paragraph>
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<paragraph pageId="0" pageNumber="4596">Costa Rica, ACG, Prov. Alajuela and Guanacaste, rain forest and dry forest.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="4596" type="ecology">
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<paragraph pageId="0" pageNumber="4596">Ecology</paragraph>
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<subSection pageId="0" pageNumber="4596" type="hosts">
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<paragraph pageId="0" pageNumber="4596">Hosts</paragraph>
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<paragraph pageId="0" pageNumber="4596">
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Reared from caterpillars of the
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<taxonomicName family="Limacodidae" lsidName="" pageId="0" pageNumber="4596" rank="family">Limacodidae</taxonomicName>
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<taxonomicName class="Insecta" family="Limacodidae" genus="Euclea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euclea mesoamericana" order="Lepidoptera" pageId="0" pageNumber="4596" phylum="Arthropoda" rank="species" species="mesoamericana">Euclea mesoamericana</taxonomicName>
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Corrales & Epstein, 2004.
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</paragraph>
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</subSection>
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</subSubSection>
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</treatment>
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</document> |