282 lines
37 KiB
XML
282 lines
37 KiB
XML
<document id="723399DED1D07DA7330AA66C63585D51" ID-DOI="10.11646/zootaxa.5424.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821580" ID-ZooBank="B70EEB3B-A24B-4756-BB0C-CA4406B921D6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710501405233" checkinUser="plazi" docAuthor="Shavrin, Alexey V. & Medina, Milton Norman D." docDate="2024" docId="9117327D93644403B9FBF9D9CC8BDC9E" docLanguage="en" docName="zootaxa.5424.4.7.pdf" docOrigin="Zootaxa 5424 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.4.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orphnebius (Deroleptus) poncei Shavrin & Medina 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="488" masterDocId="6D2E4A0593674406B96CFFE1C82FDF57" masterDocTitle="Two new species of the genus Orphnebius Motschulsky, 1858 from Mindanao, Philippines (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="489" masterPageNumber="483" pageNumber="486" updateTime="1710502021849" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="DD1971D998E65441F87A1E8E7F594E32" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C6AD24498E6927ECCEA5ADF8FC0AC669">
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<mods:title id="4E8CB989BD7F17591DC0E61D06EBAD69">Two new species of the genus Orphnebius Motschulsky, 1858 from Mindanao, Philippines (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:name id="0762BB1C5A1F5777D3829A63CF174171" type="personal">
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<mods:roleTerm id="F609DCC79D4F2A05CD8A57E8681E7FB4">Author</mods:roleTerm>
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<mods:namePart id="88D1A9B7D2273C1CCDB9FAF55C9647ED">Shavrin, Alexey V.</mods:namePart>
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<mods:affiliation id="264F0DE5101F9AFF4BC1DDC5E150FED9">Institute of Life Sciences and Technologies, Daugavpils University, Vienibas 13, Daugavpils, LV- 5401, Latvia</mods:affiliation>
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<mods:roleTerm id="043FDF1CBE63F21085511012FB9F2400">Author</mods:roleTerm>
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<mods:namePart id="5281A133BE0A718F5DEB7601D8CC74DD">Medina, Milton Norman D.</mods:namePart>
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<mods:affiliation id="C71D98B61B69143BCF71FC8BD6C377D9">Tropical Genomics Laboratory-URESCOM, and Faculty of Agriculture and Life Sciences, Davao Oriental State University, Mati City, 8200, Philippines & National Museum of Natural History, Ermita, 1000 Philippines</mods:affiliation>
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<mods:typeOfResource id="0112B3985CEFFB4272AE547334695408">text</mods:typeOfResource>
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<mods:title id="FA78E759DDBFFC9680F11941BC2CC03D">Zootaxa</mods:title>
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<mods:part id="B0DBA04F1F8E133D53D9DB5154260EC4">
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<mods:date id="984829A74DD6554FF8937D9EE08966DD">2024</mods:date>
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<mods:number id="3B380C86EDD3F1D9803623452A35D14D">2024-03-15</mods:number>
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<mods:number id="D474B437F18CDB348C2D5AD136494574">5424</mods:number>
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<mods:url id="2E90F0DDED478409E425D04247C1DA25">http://dx.doi.org/10.11646/zootaxa.5424.4.7</mods:url>
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<mods:classification id="DB63ABBBF4CAB5FFDA355C349B63A7AE">journal article</mods:classification>
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<mods:identifier id="8816F3C5F9CECA503952DD5ABDEF7F4D" type="DOI">10.11646/zootaxa.5424.4.7</mods:identifier>
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<treatment id="9117327D93644403B9FBF9D9CC8BDC9E" ID-DOI="http://doi.org/10.5281/zenodo.10821571" ID-Zenodo-Dep="10821571" LSID="urn:lsid:plazi:treatment:9117327D93644403B9FBF9D9CC8BDC9E" httpUri="http://treatment.plazi.org/id/9117327D93644403B9FBF9D9CC8BDC9E" lastPageId="5" lastPageNumber="488" pageId="3" pageNumber="486">
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<subSubSection id="51A4D0E093644405B9FBF9D9CA49D904" box="[151,614,1592,1619]" pageId="3" pageNumber="486" type="nomenclature">
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<paragraph id="1901836B93644405B9FBF9D9CA49D904" blockId="3.[151,614,1592,1655]" box="[151,614,1592,1619]" pageId="3" pageNumber="486">
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<heading id="4249340793644405B9FBF9D9CA49D904" bold="true" box="[151,614,1592,1619]" fontSize="11" level="1" pageId="3" pageNumber="486" reason="1">
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<emphasis id="2BCA5F7993644405B9FBF9D9CA49D904" bold="true" box="[151,614,1592,1619]" pageId="3" pageNumber="486">
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<taxonomicName id="DEBEF8E893644405B9FBF9D9CA32D904" authority="Shavrin & Medina, 2024" authorityName="Shavrin & Medina" authorityYear="2024" box="[151,541,1592,1619]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="486" phylum="Arthropoda" rank="species" species="poncei" status="sp. nov." subGenus="Deroleptus">
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<emphasis id="2BCA5F7993644405B9FBF9D9C909D904" bold="true" box="[151,294,1592,1619]" italics="true" pageId="3" pageNumber="486">Orphnebius</emphasis>
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(
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<emphasis id="2BCA5F7993644405B854F9D9C993D904" bold="true" box="[312,444,1592,1619]" italics="true" pageId="3" pageNumber="486">Deroleptus</emphasis>
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)
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<emphasis id="2BCA5F7993644405B8A1F9D8CA32D904" bold="true" box="[461,541,1593,1619]" italics="true" pageId="3" pageNumber="486">poncei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="30F9E20293644405BB48F9D8CA49D904" box="[548,614,1593,1619]" pageId="3" pageNumber="486" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="51A4D0E093644405B9FBF9BCC936D920" box="[151,281,1629,1655]" pageId="3" pageNumber="486" type="description">
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<paragraph id="1901836B93644405B9FBF9BCC936D920" blockId="3.[151,614,1592,1655]" box="[151,281,1629,1655]" pageId="3" pageNumber="486">
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(
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<figureCitation id="81859FEE93644405B9F3F9BCC93ED920" box="[159,273,1629,1655]" captionStart-0="FIGURE 9" captionStart-1="FIGURES 10–17" captionStartId-0="3.[151,250,1270,1294]" captionStartId-1="4.[151,264,1327,1351]" captionTargetBox-0="[196,1379,181,1238]" captionTargetBox-1="[162,1424,194,1291]" captionTargetId-0="figure-19@3.[196,1392,181,1246]" captionTargetId-1="figure-21@4.[151,1435,181,1303]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 9. Distribution of Orphnebius in Mindanao: O. barsevskisi (circle), O. bulbosus (square), O. poncei (triangle)." captionText-1="FIGURES 10–17. Habitus of Orphnebius (s.str.) poncei (holotype): 10—habitus, 11—male abdominal tergite VII, 12—male abdominal sternite VII, 13—male abdominal tergite VIII, 14—male abdominal sternite VIII, 15—aedeagus, ventral view, 16— aedeagus, lateral view, 17—paramere. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs. 2–8)." figureDoi-0="http://doi.org/10.5281/zenodo.10821584" figureDoi-1="http://doi.org/10.5281/zenodo.10821586" httpUri-0="https://zenodo.org/record/10821584/files/figure.png" httpUri-1="https://zenodo.org/record/10821586/files/figure.png" pageId="3" pageNumber="486">Figs 9–17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="51A4D0E093644405B9FBF944CA75D87C" pageId="3" pageNumber="486" type="materials_examined">
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<paragraph id="1901836B93644405B9FBF944CA75D87C" blockId="3.[151,1437,1701,1979]" pageId="3" pageNumber="486">
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<emphasis id="2BCA5F7993644405B9FBF944C969D9E8" bold="true" box="[151,326,1701,1727]" pageId="3" pageNumber="486">Type material.</emphasis>
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<typeStatus id="C6053DC993644405B823F944C998D9E8" box="[335,439,1701,1727]" pageId="3" pageNumber="486" type="holotype">Holotype</typeStatus>
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<specimenCount id="0FB848E293644405B8ACF944C9FAD9E8" box="[448,469,1701,1727]" count="1" pageId="3" pageNumber="486" type="male">♂</specimenCount>
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(
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<figureCitation id="81859FEE93644405B889F944CA15D9E8" box="[485,570,1701,1727]" captionStart="FIGURES 10–17" captionStartId="4.[151,264,1327,1351]" captionTargetBox="[162,1424,194,1291]" captionTargetId="figure-21@4.[151,1435,181,1303]" captionTargetPageId="4" captionText="FIGURES 10–17. Habitus of Orphnebius (s.str.) poncei (holotype): 10—habitus, 11—male abdominal tergite VII, 12—male abdominal sternite VII, 13—male abdominal tergite VIII, 14—male abdominal sternite VIII, 15—aedeagus, ventral view, 16— aedeagus, lateral view, 17—paramere. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs. 2–8)." figureDoi="http://doi.org/10.5281/zenodo.10821586" httpUri="https://zenodo.org/record/10821586/files/figure.png" pageId="3" pageNumber="486">Fig. 10</figureCitation>
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; dissected): ‘
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<collectingCountry id="61A9C3FB93644405BBA0F944CB7DD9E8" box="[716,850,1701,1727]" name="Philippines" pageId="3" pageNumber="486">Philippines</collectingCountry>
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, Mindanao,
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<collectingRegion id="DB7A4D8993644405BAB3F944CC07D9E8" box="[991,1064,1701,1727]" country="Philippines" name="Davao" pageId="3" pageNumber="486">Davao</collectingRegion>
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| Oriental, Tibanga River Pool, | Cawa-Cawa
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<collectionCode id="7FAF1BAE93644405B84AF928C913D9B4" box="[294,316,1737,1763]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="486">R</collectionCode>
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.
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<geoCoordinate id="7C8AE5AC93644405B82BF928C9E9D9B3" box="[327,454,1737,1764]" degrees="6.744185" direction="north" orientation="latitude" pageId="3" pageNumber="486" precision="1" value="6.744185">6.744185N</geoCoordinate>
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, |
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<geoCoordinate id="7C8AE5AC93644405B8B1F928CA58D9B4" box="[477,631,1737,1763]" degrees="126.179798" direction="east" orientation="longitude" pageId="3" pageNumber="486" precision="1" value="126.179794">126.179798E</geoCoordinate>
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.
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<date id="6D00A5AB93644405BBEDF928CB2FD9B3" box="[641,768,1737,1764]" pageId="3" pageNumber="486" value="2023-11-05">11.05.2023</date>
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, | Shavrin A.
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<collectionCode id="7FAF1BAE93644405BAE1F928CB8CD9B4" box="[909,931,1737,1763]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="486" type="Museum">V</collectionCode>
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. leg.’ <printed>, ‘Sifting of wet leaf litter and deb- | ris on slope near the river’ <printed>, ‘
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<typeStatus id="C6053DC993644405BBFCF90CCB06D850" box="[656,809,1773,1799]" pageId="3" pageNumber="486" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="DEBEF8E893644405BA52F90CCC78D850" authorityName="Bernhauer" authorityYear="1915" box="[830,1111,1773,1799]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="486" phylum="Arthropoda" rank="subGenus" subGenus="Deroleptus">
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<emphasis id="2BCA5F7993644405BA52F90CCBEDD850" box="[830,962,1773,1799]" italics="true" pageId="3" pageNumber="486">Orphnebius</emphasis>
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(
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<emphasis id="2BCA5F7993644405BABFF90CCC61D850" box="[979,1102,1773,1799]" italics="true" pageId="3" pageNumber="486">Deroleptus</emphasis>
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)
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</taxonomicName>
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<taxonomicName id="DEBEF8E893644405BD01F90CCCC6D850" authority="Shavrin & Medina, 2024" authorityName="Shavrin & Medina" authorityYear="2024" box="[1133,1257,1773,1799]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="486" phylum="Arthropoda" rank="species" species="barsevskisi" status="sp. nov.">
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<emphasis id="2BCA5F7993644405BD01F90CCCC6D850" box="[1133,1257,1773,1799]" italics="true" pageId="3" pageNumber="486">barsevskisi</emphasis>
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</taxonomicName>
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<emphasis id="2BCA5F7993644405BD9DF90CCD1ED850" bold="true" box="[1265,1329,1773,1799]" pageId="3" pageNumber="486">
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<taxonomicNameLabel id="30F9E20293644405BD9DF90CCD1ED850" box="[1265,1329,1773,1799]" pageId="3" pageNumber="486" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.
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<collectionCode id="7FAF1BAE93644405B9DDF8F0C8E8D87C" box="[177,199,1809,1835]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="486" type="Museum">V</collectionCode>
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. des. 2024’ <red, printed> (
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<collectionCode id="7FAF1BAE93644405B896F8F0CA61D87C" box="[506,590,1809,1835]" pageId="3" pageNumber="486">DUBC</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="51A4D0E093644403B9ABF8D4CA4EDC6E" lastPageId="5" lastPageNumber="488" pageId="3" pageNumber="486" type="description">
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<paragraph id="1901836B93644405B9ABF8D4CBC5D824" blockId="3.[151,1437,1701,1979]" pageId="3" pageNumber="486">
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<emphasis id="2BCA5F7993644405B9ABF8D4C976D818" bold="true" box="[199,345,1845,1871]" pageId="3" pageNumber="486">Description.</emphasis>
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Measurements: HW: 0.40; HL: 0.23; OL: 0.15; TL: 0.05; AL: 0.47; PL: 0.18; PW: 0.32; ESL: 0.17; EW: 0.42; MTbL: 0.37; MTrL: 0.35; AW: 0.42; AedL: 0.27; BL: 4.30.
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</paragraph>
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<paragraph id="1901836B93644405B9ABF89CCD35D8EC" blockId="3.[151,1437,1701,1979]" pageId="3" pageNumber="486">Body reddish-brown, with distinctly darker head and latero-posterior portions of elytra;mouthparts, antennomeres and legs yellowish (basal portions of tibiae slightly darker, apical antennomere and tarsi slightly paler).</paragraph>
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<caption id="4DC1D3E393634402B9FBFACECC24DAC7" ID-DOI="http://doi.org/10.5281/zenodo.10821586" ID-Zenodo-Dep="10821586" httpUri="https://zenodo.org/record/10821586/files/figure.png" pageId="4" pageNumber="487" startId="4.[151,264,1327,1351]" targetBox="[162,1424,194,1291]" targetPageId="4" targetType="figure">
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<paragraph id="1901836B93634402B9FBFACECC24DAC7" blockId="4.[151,1436,1327,1424]" pageId="4" pageNumber="487">
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<emphasis id="2BCA5F7993634402B9FBFACEC97ADA10" bold="true" box="[151,341,1327,1351]" pageId="4" pageNumber="487">FIGURES 10–17.</emphasis>
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Habitus of
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<taxonomicName id="DEBEF8E893634402B8BCFACECAF5DA10" authorityName="Shavrin & Medina" authorityYear="2024" box="[464,730,1327,1352]" class="Insecta" family="Staphylinidae" genus="Orphnebius" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="poncei">
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<emphasis id="2BCA5F7993634402B8BCFACECA67DA10" box="[464,584,1327,1351]" italics="true" pageId="4" pageNumber="487">Orphnebius</emphasis>
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(s.str.)
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<emphasis id="2BCA5F7993634402BBFAFAD1CAF5DA10" box="[662,730,1328,1351]" italics="true" pageId="4" pageNumber="487">poncei</emphasis>
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</taxonomicName>
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(holotype): 10—habitus, 11—male abdominal tergite VII, 12—male abdominal sternite VII, 13—male abdominal tergite VIII, 14—male abdominal sternite VIII, 15—aedeagus, ventral view, 16— aedeagus, lateral view, 17—paramere. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs. 2–8).
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</paragraph>
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</caption>
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<paragraph id="1901836B93634402B9ABFA03C9F4D9AF" blockId="4.[151,1437,1506,2036]" pageId="4" pageNumber="487">Head strongly transverse, 1.7 times as broad as long, with rounded postocular portions and posterior margin widely truncate in middle.Punctation indistinct, fine and very sparse; middle and posterior portions without punctures. Microsculpture absent except of fine, indistinct and transverse meshes in mediolateral portions. Clypeus distinctly sclerotized. Eyes large and convex. Temples about three times shorter than longitudinal length of eyes. Antenna almost reaching basal margin of pronotum when reclined; antennomeres 4–10 slightly transverse; antennomere 4 distinctly broader than 3, 5–6 slightly shorter than 4, 7–10 slightrly longer and indistinctly broader than 6, apical antennomere moderately long, slightly shorter than three preceding segments, from widest middle portion gradually narrowed toward acute apex.</paragraph>
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<paragraph id="1901836B93634402B9ABF8E3CD2FD817" blockId="4.[151,1437,1506,2036]" pageId="4" pageNumber="487">Pronotum distinctly transverse, 1.7 times as broad as long, broader at level of anterior angles. Disc with indistinct, very fine and sparse punctation; middle portion without punctures. Microsculpture absent.</paragraph>
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<paragraph id="1901836B93634402B9ABF8ABCCFBD8DF" blockId="4.[151,1437,1506,2036]" pageId="4" pageNumber="487">Elytra indistinctly shorter than pronotum, transverse, about two and a half times as broad as long. Punctation fine and very sparse, indistinct along suture. Microsculpture absent. Hind wings fully developed.</paragraph>
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<paragraph id="1901836B93634402B9ABF873CBF4D8FB" blockId="4.[151,1437,1506,2036]" box="[199,987,1938,1964]" pageId="4" pageNumber="487">Legs very long and slender; metatarsus slightly shorter than metatibia.</paragraph>
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<paragraph id="1901836B93634402B9ABF857CC0FD8A3" blockId="4.[151,1437,1506,2036]" pageId="4" pageNumber="487">Abdomen as broad as elytra, with very fine punctation and without microsculpture; abdominal tergite IV distinctly longer than III, V slightly longer than IV, VI indistinctly longer than V.</paragraph>
|
||
<paragraph id="1901836B93624403B9ABFF76CAB0DE4A" blockId="5.[151,1437,151,969]" pageId="5" pageNumber="488">
|
||
Male. Abdominal tergite VII (
|
||
<figureCitation id="81859FEE93624403BB4CFF76CA5DDFE6" box="[544,626,151,177]" captionStart="FIGURES 10–17" captionStartId="4.[151,264,1327,1351]" captionTargetBox="[162,1424,194,1291]" captionTargetId="figure-21@4.[151,1435,181,1303]" captionTargetPageId="4" captionText="FIGURES 10–17. Habitus of Orphnebius (s.str.) poncei (holotype): 10—habitus, 11—male abdominal tergite VII, 12—male abdominal sternite VII, 13—male abdominal tergite VIII, 14—male abdominal sternite VIII, 15—aedeagus, ventral view, 16— aedeagus, lateral view, 17—paramere. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs. 2–8)." figureDoi="http://doi.org/10.5281/zenodo.10821586" httpUri="https://zenodo.org/record/10821586/files/figure.png" pageId="5" pageNumber="488">Fig. 11</figureCitation>
|
||
) and tergite VIII
|
||
<figureCitation id="81859FEE93624403BA2CFF76CBBDDFE6" box="[832,914,151,177]" captionStart="FIGURES 10–17" captionStartId="4.[151,264,1327,1351]" captionTargetBox="[162,1424,194,1291]" captionTargetId="figure-21@4.[151,1435,181,1303]" captionTargetPageId="4" captionText="FIGURES 10–17. Habitus of Orphnebius (s.str.) poncei (holotype): 10—habitus, 11—male abdominal tergite VII, 12—male abdominal sternite VII, 13—male abdominal tergite VIII, 14—male abdominal sternite VIII, 15—aedeagus, ventral view, 16— aedeagus, lateral view, 17—paramere. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs. 2–8)." figureDoi="http://doi.org/10.5281/zenodo.10821586" httpUri="https://zenodo.org/record/10821586/files/figure.png" pageId="5" pageNumber="488">Fig. 13</figureCitation>
|
||
) with distinctly serrate latero-apical margins. Abdominal sternite VII with dense and large punctation in middle (
|
||
<figureCitation id="81859FEE93624403BAC9FF5ACBD4DF82" box="[933,1019,187,213]" captionStart="FIGURES 10–17" captionStartId="4.[151,264,1327,1351]" captionTargetBox="[162,1424,194,1291]" captionTargetId="figure-21@4.[151,1435,181,1303]" captionTargetPageId="4" captionText="FIGURES 10–17. Habitus of Orphnebius (s.str.) poncei (holotype): 10—habitus, 11—male abdominal tergite VII, 12—male abdominal sternite VII, 13—male abdominal tergite VIII, 14—male abdominal sternite VIII, 15—aedeagus, ventral view, 16— aedeagus, lateral view, 17—paramere. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs. 2–8)." figureDoi="http://doi.org/10.5281/zenodo.10821586" httpUri="https://zenodo.org/record/10821586/files/figure.png" pageId="5" pageNumber="488">Fig. 12</figureCitation>
|
||
). Abdominal sternite VIII as in
|
||
<figureCitation id="81859FEE93624403BC1CFF5AC899DFAE" captionStart="FIGURES 10–17" captionStartId="4.[151,264,1327,1351]" captionTargetBox="[162,1424,194,1291]" captionTargetId="figure-21@4.[151,1435,181,1303]" captionTargetPageId="4" captionText="FIGURES 10–17. Habitus of Orphnebius (s.str.) poncei (holotype): 10—habitus, 11—male abdominal tergite VII, 12—male abdominal sternite VII, 13—male abdominal tergite VIII, 14—male abdominal sternite VIII, 15—aedeagus, ventral view, 16— aedeagus, lateral view, 17—paramere. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs. 2–8)." figureDoi="http://doi.org/10.5281/zenodo.10821586" httpUri="https://zenodo.org/record/10821586/files/figure.png" pageId="5" pageNumber="488">Fig. 14</figureCitation>
|
||
. Aedeagus elongate, from widest basal portion gradually narrowed toward apex (
|
||
<figureCitation id="81859FEE93624403BD20FF3ECC8FDFAE" box="[1100,1184,222,249]" captionStart="FIGURES 10–17" captionStartId="4.[151,264,1327,1351]" captionTargetBox="[162,1424,194,1291]" captionTargetId="figure-21@4.[151,1435,181,1303]" captionTargetPageId="4" captionText="FIGURES 10–17. Habitus of Orphnebius (s.str.) poncei (holotype): 10—habitus, 11—male abdominal tergite VII, 12—male abdominal sternite VII, 13—male abdominal tergite VIII, 14—male abdominal sternite VIII, 15—aedeagus, ventral view, 16— aedeagus, lateral view, 17—paramere. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs. 2–8)." figureDoi="http://doi.org/10.5281/zenodo.10821586" httpUri="https://zenodo.org/record/10821586/files/figure.png" pageId="5" pageNumber="488">Fig. 15</figureCitation>
|
||
). Lateral aspect of the aedeagus as in
|
||
<figureCitation id="81859FEE93624403B82CFEE2C9BDDE4A" box="[320,402,259,285]" captionStart="FIGURES 10–17" captionStartId="4.[151,264,1327,1351]" captionTargetBox="[162,1424,194,1291]" captionTargetId="figure-21@4.[151,1435,181,1303]" captionTargetPageId="4" captionText="FIGURES 10–17. Habitus of Orphnebius (s.str.) poncei (holotype): 10—habitus, 11—male abdominal tergite VII, 12—male abdominal sternite VII, 13—male abdominal tergite VIII, 14—male abdominal sternite VIII, 15—aedeagus, ventral view, 16— aedeagus, lateral view, 17—paramere. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs. 2–8)." figureDoi="http://doi.org/10.5281/zenodo.10821586" httpUri="https://zenodo.org/record/10821586/files/figure.png" pageId="5" pageNumber="488">Fig. 16</figureCitation>
|
||
. Paramere as in
|
||
<figureCitation id="81859FEE93624403BB25FEE2CAB4DE4A" box="[585,667,259,285]" captionStart="FIGURES 10–17" captionStartId="4.[151,264,1327,1351]" captionTargetBox="[162,1424,194,1291]" captionTargetId="figure-21@4.[151,1435,181,1303]" captionTargetPageId="4" captionText="FIGURES 10–17. Habitus of Orphnebius (s.str.) poncei (holotype): 10—habitus, 11—male abdominal tergite VII, 12—male abdominal sternite VII, 13—male abdominal tergite VIII, 14—male abdominal sternite VIII, 15—aedeagus, ventral view, 16— aedeagus, lateral view, 17—paramere. Scale bars: 1.0 mm (Fig. 1), 0.1 mm (Figs. 2–8)." figureDoi="http://doi.org/10.5281/zenodo.10821586" httpUri="https://zenodo.org/record/10821586/files/figure.png" pageId="5" pageNumber="488">Fig. 17</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1901836B93624403B9ABFEC6C9BEDE16" blockId="5.[151,1437,151,969]" box="[199,401,295,321]" pageId="5" pageNumber="488">Female unknown.</paragraph>
|
||
<paragraph id="1901836B93624403B9ABFEABCA4EDC6E" blockId="5.[151,1437,151,969]" pageId="5" pageNumber="488">
|
||
<emphasis id="2BCA5F7993624403B9ABFEABC99CDE32" bold="true" box="[199,435,330,357]" pageId="5" pageNumber="488">Comparative notes.</emphasis>
|
||
Based on the general shape of the head with wide posterior portions and the shape of the aedeagus,
|
||
<emphasis id="2BCA5F7993624403B855FE8EC962DEDE" box="[313,333,367,393]" italics="true" pageId="5" pageNumber="488">O</emphasis>
|
||
.
|
||
<taxonomicName id="DEBEF8E893624403B831FE8EC9F0DEDF" authority="Shavrin & Medina, 2024" authorityName="Shavrin & Medina" authorityYear="2024" box="[349,479,367,393]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="poncei" status="sp. nov." subGenus="Deroleptus">
|
||
(
|
||
<emphasis id="2BCA5F7993624403B80BFE8EC954DEDF" box="[359,379,367,392]" italics="true" pageId="5" pageNumber="488">D</emphasis>
|
||
.)
|
||
<emphasis id="2BCA5F7993624403B8F8FE8EC9F0DEDF" box="[404,479,367,392]" italics="true" pageId="5" pageNumber="488">poncei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2BCA5F7993624403B884FE8ECA06DEDE" bold="true" box="[488,553,367,393]" pageId="5" pageNumber="488">
|
||
<taxonomicNameLabel id="30F9E20293624403B884FE8ECA06DEDE" box="[488,553,367,393]" pageId="5" pageNumber="488" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<emphasis id="2BCA5F7993624403BBAFFE8ECAF8DEDE" box="[707,727,367,393]" italics="true" pageId="5" pageNumber="488">O</emphasis>
|
||
.
|
||
<taxonomicName id="DEBEF8E893624403BB8BFE8ECC10DEDE" authority="Assing, 2021" authorityName="Assing" authorityYear="2021" box="[743,1087,367,393]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="tuberculifer" subGenus="Deroleptus">
|
||
(
|
||
<emphasis id="2BCA5F7993624403BB9DFE8ECB2ADEDF" box="[753,773,367,392]" italics="true" pageId="5" pageNumber="488">D</emphasis>
|
||
.)
|
||
<emphasis id="2BCA5F7993624403BA72FE8ECB8EDEDE" box="[798,929,367,393]" italics="true" pageId="5" pageNumber="488">tuberculifer</emphasis>
|
||
<bibRefCitation id="7D2FFE9A93624403BAC5FE8ECC10DEDE" author="Assing, V." box="[937,1087,367,393]" pageId="5" pageNumber="488" pagination="1 - 193" refId="ref3488" refString="Assing, V. (2021) On the Lomechusini of the Palaearctic and Oriental regions XXVIII. New genera, species, and records, with a focus of the fauna of Laos and Borneo (Coleoptera: Staphylinidae: Aleocharinae). Acta Musei Moraviae, Scientiae biologicae, 106 (1), 1 - 193." type="journal article" year="2021">Assing, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="2BCA5F7993624403BD15FE8ECCA2DEDE" box="[1145,1165,367,393]" italics="true" pageId="5" pageNumber="488">O</emphasis>
|
||
.
|
||
<taxonomicName id="DEBEF8E893624403BDF1FE8EC8FDDEFA" authority="Assing, 2021" authorityName="Assing" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="tuberosus" subGenus="Deroleptus">
|
||
(
|
||
<emphasis id="2BCA5F7993624403BDCBFE8ECC94DEDF" box="[1191,1211,367,392]" italics="true" pageId="5" pageNumber="488">D</emphasis>
|
||
.)
|
||
<emphasis id="2BCA5F7993624403BDB8FE8ECD6FDEDE" box="[1236,1344,367,393]" italics="true" pageId="5" pageNumber="488">tuberosus</emphasis>
|
||
<bibRefCitation id="7D2FFE9A93624403BC2BFE8EC8FDDEFA" author="Assing, V." pageId="5" pageNumber="488" pagination="1 - 193" refId="ref3488" refString="Assing, V. (2021) On the Lomechusini of the Palaearctic and Oriental regions XXVIII. New genera, species, and records, with a focus of the fauna of Laos and Borneo (Coleoptera: Staphylinidae: Aleocharinae). Acta Musei Moraviae, Scientiae biologicae, 106 (1), 1 - 193." type="journal article" year="2021">Assing, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
, both recently described from
|
||
<collectingCountry id="61A9C3FB93624403BB5DFE72CAB5DEFA" box="[561,666,403,429]" name="Malaysia" pageId="5" pageNumber="488">Malaysia</collectingCountry>
|
||
,
|
||
<collectingCountry id="61A9C3FB93624403BBC4FE72CB6BDEFA" box="[680,836,403,429]" name="Malaysia" pageId="5" pageNumber="488">North Borneo</collectingCountry>
|
||
(
|
||
<bibRefCitation id="7D2FFE9A93624403BA39FE72CBC5DEFA" author="Assing, V." box="[853,1002,403,429]" pageId="5" pageNumber="488" pagination="1 - 193" refId="ref3488" refString="Assing, V. (2021) On the Lomechusini of the Palaearctic and Oriental regions XXVIII. New genera, species, and records, with a focus of the fauna of Laos and Borneo (Coleoptera: Staphylinidae: Aleocharinae). Acta Musei Moraviae, Scientiae biologicae, 106 (1), 1 - 193." type="journal article" year="2021">Assing 2021</bibRefCitation>
|
||
). It can be distinguished from
|
||
<emphasis id="2BCA5F7993624403BC26FE72CD71DEFA" box="[1354,1374,403,429]" italics="true" pageId="5" pageNumber="488">O</emphasis>
|
||
.
|
||
<taxonomicName id="DEBEF8E893624403BC02FE72C935DE86" authorityName="Assing" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="tuberculifer" subGenus="Deroleptus">
|
||
(
|
||
<emphasis id="2BCA5F7993624403BC14FE72CDA3DEFB" box="[1400,1420,403,428]" italics="true" pageId="5" pageNumber="488">D</emphasis>
|
||
.)
|
||
<emphasis id="2BCA5F7993624403B9FBFE56C935DE86" box="[151,282,439,465]" italics="true" pageId="5" pageNumber="488">tuberculifer</emphasis>
|
||
</taxonomicName>
|
||
by the broader apical portion of the pronotum, and from
|
||
<emphasis id="2BCA5F7993624403BAC6FE56CB91DE86" box="[938,958,439,465]" italics="true" pageId="5" pageNumber="488">O</emphasis>
|
||
.
|
||
<taxonomicName id="DEBEF8E893624403BAA1FE56CC5FDE86" authorityName="Assing" authorityYear="2021" box="[973,1136,439,465]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="tuberosus" subGenus="Deroleptus">
|
||
(
|
||
<emphasis id="2BCA5F7993624403BABBFE56CBC4DE87" box="[983,1003,439,464]" italics="true" pageId="5" pageNumber="488">D</emphasis>
|
||
.)
|
||
<emphasis id="2BCA5F7993624403BD68FE56CC5FDE86" box="[1028,1136,439,465]" italics="true" pageId="5" pageNumber="488">tuberosus</emphasis>
|
||
</taxonomicName>
|
||
by the distinctly narrower median lobe of the aedeagus. From both species it differs by the paler coloration of the larger body, distinctly longer eyes, the absence of tubercles on the dorsal surface of the abdominal tergite VIII, narrower median lobe of the aedeagus and different structure of the internal sac. Based on the general shape of the forebody and the aedeagus, the presence of widely rounded posterior portions of head and the absence of tubercles on the dorsal surface of all apical abdominal tergites,
|
||
<emphasis id="2BCA5F7993624403B88BFD8AC9D4DDD2" box="[487,507,619,645]" italics="true" pageId="5" pageNumber="488">O</emphasis>
|
||
.
|
||
<taxonomicName id="DEBEF8E893624403BB67FD8ACAA3DDD3" authority="Shavrin & Medina, 2024" authorityName="Shavrin & Medina" authorityYear="2024" box="[523,652,619,645]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="poncei" status="sp. nov." subGenus="Deroleptus">
|
||
(
|
||
<emphasis id="2BCA5F7993624403BB78FD8ACA07DDD3" box="[532,552,619,644]" italics="true" pageId="5" pageNumber="488">D</emphasis>
|
||
.)
|
||
<emphasis id="2BCA5F7993624403BB2DFD8ACAA3DDD3" box="[577,652,619,644]" italics="true" pageId="5" pageNumber="488">poncei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2BCA5F7993624403BBF8FD8ACAFADDD2" bold="true" box="[660,725,619,645]" pageId="5" pageNumber="488">
|
||
<taxonomicNameLabel id="30F9E20293624403BBF8FD8ACAFADDD2" box="[660,725,619,645]" pageId="5" pageNumber="488" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is also similar to
|
||
<emphasis id="2BCA5F7993624403BACFFD8ACB98DDD2" box="[931,951,619,645]" italics="true" pageId="5" pageNumber="488">O</emphasis>
|
||
.
|
||
<taxonomicName id="DEBEF8E893624403BAAAFD8ACC56DDD2" authority="Shavrin & Medina, 2024" authorityName="Shavrin & Medina" authorityYear="2024" box="[966,1145,619,645]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="barsevskisi" status="sp. nov." subGenus="Deroleptus">
|
||
(
|
||
<emphasis id="2BCA5F7993624403BABCFD8ACBCBDDD3" box="[976,996,619,644]" italics="true" pageId="5" pageNumber="488">D</emphasis>
|
||
.)
|
||
<emphasis id="2BCA5F7993624403BA91FD8ACC56DDD2" box="[1021,1145,619,645]" italics="true" pageId="5" pageNumber="488">barsevskisi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2BCA5F7993624403BDEDFD8ACCEDDDD2" bold="true" box="[1153,1218,619,645]" pageId="5" pageNumber="488">
|
||
<taxonomicNameLabel id="30F9E20293624403BDEDFD8ACCEDDDD2" box="[1153,1218,619,645]" pageId="5" pageNumber="488" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, from which it can be distinguished by the narrower body, less broader posterior portions of head, absence of microsculpture on the darker elytra, slightly narrower median lobe and details of the structure of the internal sac. Besides that, it can be distinguished from
|
||
<emphasis id="2BCA5F7993624403B818FD36C9A7DDA6" box="[372,392,727,753]" italics="true" pageId="5" pageNumber="488">O</emphasis>
|
||
.
|
||
<taxonomicName id="DEBEF8E893624403B8FBFD36CAFFDDA6" authority="Assing, 2021" authorityName="Assing" authorityYear="2021" box="[407,720,727,753]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="bulbosus" subGenus="Deroleptus">
|
||
(
|
||
<emphasis id="2BCA5F7993624403B8CDFD36C99ADDA7" box="[417,437,727,752]" italics="true" pageId="5" pageNumber="488">D</emphasis>
|
||
.)
|
||
<emphasis id="2BCA5F7993624403B8A2FD36CA1DDDA6" box="[462,562,727,753]" italics="true" pageId="5" pageNumber="488">bulbosus</emphasis>
|
||
<bibRefCitation id="7D2FFE9A93624403BB54FD36CAFFDDA6" author="Assing, V." box="[568,720,727,753]" pageId="5" pageNumber="488" pagination="1 - 193" refId="ref3488" refString="Assing, V. (2021) On the Lomechusini of the Palaearctic and Oriental regions XXVIII. New genera, species, and records, with a focus of the fauna of Laos and Borneo (Coleoptera: Staphylinidae: Aleocharinae). Acta Musei Moraviae, Scientiae biologicae, 106 (1), 1 - 193." type="journal article" year="2021">Assing, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
, recently described from Mindanao (
|
||
<bibRefCitation id="7D2FFE9A93624403BD18FD36CD2BDDA6" author="Assing, V." box="[1140,1284,727,753]" pageId="5" pageNumber="488" pagination="1 - 193" refId="ref3488" refString="Assing, V. (2021) On the Lomechusini of the Palaearctic and Oriental regions XXVIII. New genera, species, and records, with a focus of the fauna of Laos and Borneo (Coleoptera: Staphylinidae: Aleocharinae). Acta Musei Moraviae, Scientiae biologicae, 106 (1), 1 - 193." type="journal article" year="2021">Assing 2021</bibRefCitation>
|
||
) by the paler coloration, distinctly longer and broader postocular portions of the head, broader apical portion of the pronotum, and distinctly shorter preapical antennomere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="51A4D0E093624403B9ABFCA3CCDFDC0A" box="[199,1264,834,861]" pageId="5" pageNumber="488" type="distribution">
|
||
<paragraph id="1901836B93624403B9ABFCA3CCDFDC0A" blockId="5.[151,1437,151,969]" box="[199,1264,834,861]" pageId="5" pageNumber="488">
|
||
<emphasis id="2BCA5F7993624403B9ABFCA3C94EDC0B" bold="true" box="[199,353,834,860]" pageId="5" pageNumber="488">Distribution.</emphasis>
|
||
The new species is known only from the
|
||
<typeStatus id="C6053DC993624403BA58FCA2CB4BDC0A" box="[820,868,835,861]" pageId="5" pageNumber="488">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="DB7A4D8993624403BA8FFCA2CCA0DC0A" box="[995,1167,835,861]" country="Philippines" name="Davao Oriental" pageId="5" pageNumber="488">Davao Oriental</collectingRegion>
|
||
(
|
||
<figureCitation id="81859FEE93624403BDF2FCA2CCCDDC0A" box="[1182,1250,835,861]" captionStart="FIGURE 9" captionStartId="3.[151,250,1270,1294]" captionTargetBox="[196,1379,181,1238]" captionTargetId="figure-19@3.[196,1392,181,1246]" captionTargetPageId="3" captionText="FIGURE 9. Distribution of Orphnebius in Mindanao: O. barsevskisi (circle), O. bulbosus (square), O. poncei (triangle)." figureDoi="http://doi.org/10.5281/zenodo.10821584" httpUri="https://zenodo.org/record/10821584/files/figure.png" pageId="5" pageNumber="488">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="51A4D0E093624403B9ABFC86CC8BDC9E" pageId="5" pageNumber="488" type="etymology">
|
||
<paragraph id="1901836B93624403B9ABFC86CAB4DCF2" blockId="5.[151,1437,151,969]" pageId="5" pageNumber="488">
|
||
<emphasis id="2BCA5F7993624403B9ABFC86C961DCD6" bold="true" box="[199,334,871,897]" pageId="5" pageNumber="488">Etymology.</emphasis>
|
||
The new species is named after our colleague Roy G. Ponce, the president of
|
||
<collectingRegion id="DB7A4D8993624403BDDEFC86CDB3DCD6" box="[1202,1436,871,897]" country="Philippines" name="Davao Oriental" pageId="5" pageNumber="488">Davao Oriental State</collectingRegion>
|
||
College of Science and Technology (DOrSU).
|
||
</paragraph>
|
||
<paragraph id="1901836B93624403B9ABFC4FCC8BDC9E" blockId="5.[151,1437,151,969]" box="[199,1188,942,969]" pageId="5" pageNumber="488">
|
||
<emphasis id="2BCA5F7993624403B9ABFC4FC964DC9F" bold="true" box="[199,331,942,968]" pageId="5" pageNumber="488">Bionomics.</emphasis>
|
||
The specimen was sifted from wet litter and debris on a slope near the river.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |