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<document id="5882D979A4DB2C991DC4D866CA479D41" ID-CLB-Dataset="35343" ID-DOI="http://dx.doi.org/10.3897/zookeys.329.4961" ID-GBIF-Dataset="1e4ab867-cf67-4105-9918-ebb52be7f061" ID-PMC="PMC3800826" ID-Pensoft-Pub="1313-2970-329-33" ID-PubMed="24146552" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-329-33" ModsDocOrigin="ZooKeys 329" ModsDocTitle="A revision of the Larainae (Coleoptera, Elmidae) of Venezuela, with description of nine new species" checkinTime="1451246973851" checkinUser="pensoft" docAuthor="Maier, Crystal A." docDate="2013" docId="90A529DB66E4EA2F6F3D7ABD45E6618A" docLanguage="en" docName="ZooKeys 329: 33-91" docOrigin="ZooKeys 329" docSource="http://dx.doi.org/10.3897/zookeys.329.4961" docTitle="Pharceonus volcanus Spangler &amp; Santiago-Fragoso 1992" docType="treatment" docVersion="5" lastPageNumber="82" masterDocId="307FFFAA5B6E686D5B2FFF86DF568114" masterDocTitle="A revision of the Larainae (Coleoptera, Elmidae) of Venezuela, with description of nine new species" masterLastPageNumber="91" masterPageNumber="33" pageNumber="80" updateTime="1732672090439" updateUser="ExternalLinkService">
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<mods:title id="41AEA48D77BA51A4FFF13ADEA8FB5D83">A revision of the Larainae (Coleoptera, Elmidae) of Venezuela, with description of nine new species</mods:title>
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<mods:namePart id="180B2ECE2C67E9477E2B648824B868A9">Maier, Crystal A.</mods:namePart>
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<mods:identifier id="98AC864A02E51D48F6A5AF8CF59E0385" type="DOI">http://dx.doi.org/10.3897/zookeys.329.4961</mods:identifier>
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<treatment id="90A529DB66E4EA2F6F3D7ABD45E6618A" ID-GBIF-Taxon="152047481" LSID="urn:lsid:plazi:treatment:90A529DB66E4EA2F6F3D7ABD45E6618A" httpUri="http://treatment.plazi.org/id/90A529DB66E4EA2F6F3D7ABD45E6618A" lastPageId="49" lastPageNumber="82" pageId="47" pageNumber="80" scope_family="Elmidae" scope_order="Coleoptera" scope_subFamily="Larainae">
<subSubSection id="56174FBF5533D54BBF3B77EBA7284BFF" pageId="47" pageNumber="80" type="nomenclature">
<paragraph id="E73E3B504791D557633F9F50FA0F7D2A" pageId="47" pageNumber="80">
<taxonomicName id="38765269B72465CB30EACEBEB525A74B" ID-CoL="6V8QW" LSID="http://species-id.net/wiki/Pharceonus_volcanus" authority="Spangler &amp; Santiago-Fragoso, 1992" authorityName="Spangler &amp; Santiago-Fragoso" authorityYear="1992" class="Insecta" family="Elmidae" genus="Pharceonus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pharceonus volcanus" order="Coleoptera" pageId="47" pageNumber="80" phylum="Arthropoda" rank="species" species="volcanus">Pharceonus volcanus Spangler &amp; Santiago-Fragoso, 1992</taxonomicName>
Figs 5, 85, 86, 87
</paragraph>
</subSubSection>
<subSubSection id="54B8BBD635B2776302915B0834A51307" pageId="47" pageNumber="80" type="">
<paragraph id="C92C0448E571069A157B241C6E76A9BE" pageId="47" pageNumber="80">
See
<bibRefCitation id="6FD36E03338879685E04A838852A3CFE" author="Spangler, PJ" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="56" pageNumber="89" title="The Aquatic Beetle Subfamily Larainae (Coleoptera: Elmidae) in Mexico, Central America, and the West Indies." volume="528" year="1992">Spangler and Santiago-Fragoso 1992</bibRefCitation>
for complete species description and genitalia illustrations.
</paragraph>
</subSubSection>
<subSubSection id="21830F38F1913FB55BE36D555C51E629" pageId="47" pageNumber="80" type="material examined">
<paragraph id="E3247406F4EC2D72F6B73DB73FA04E21" pageId="47" pageNumber="80">Material examined.</paragraph>
<paragraph id="9523204AA723D2EEE9B1218B793AF006" pageId="47" pageNumber="80">
PANAMA: Chiriqui: &quot;PANAMA, Chiriqui; Volcan (26km W); 1380 m, small brook; at culvert, 3 June1983&quot; &quot;Collectors; P.J. Spangler; R.A. Faitoute; W.E. Steiner&quot; (1 Holotype USNM). VENEZUELA: Zulia State: &quot;VENEZUELA: Zulia; SW of Machiques; 31 XII 05; 435m;
<geoCoordinate id="4E6989A4AF707D9EA590C18B99547A35" direction="north" orientation="latitude" precision="925" value="10.05">10°03'N</geoCoordinate>
,
<geoCoordinate id="CADDAC034D5BC174DD2E8087DA282737" direction="west" orientation="longitude" precision="925" value="-72.71667">72°43'W</geoCoordinate>
; trib[utary] of Rio Negro&quot; &quot;A.E.Z. Short; P.J. Torres; collectors&quot; (5 specimens SEMC). &quot;VENEZUELA: Zulia State;
<geoCoordinate id="40378B56F04CE5E230E68EF46C5EDABF" direction="north" orientation="latitude" precision="1" value="10.050966">10°03.058'N</geoCoordinate>
,
<geoCoordinate id="BA4A92D5441616D09AECFD0F58F36585" direction="west" orientation="longitude" precision="1" value="-72.71623">72°42.974'W</geoCoordinate>
; 435m; Perija National Park; Toromo; 31.xii.2005; leg. A.E.Z. Short; Small stream &amp; seep; AS-06-001&quot; (3 specimens SEMC).
</paragraph>
</subSubSection>
<subSubSection id="DA9AAA68D055C73E2C0FEB6A45C4ABA7" lastPageId="48" lastPageNumber="81" pageId="47" pageNumber="80" type="diagnosis">
<paragraph id="C101A0A390B1E98DA7433C2BD7016D47" pageId="47" pageNumber="80">Diagnosis.</paragraph>
<paragraph id="FAD5FBF8DF5D5F32173E97A4A6852624" lastPageId="48" lastPageNumber="81" pageId="47" pageNumber="80">
This species can be distinguished from all other species of
<taxonomicName id="6F8DE1976AF3F3BBCB514BE347E53692" class="Insecta" family="Elmidae" genus="Pharceonus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pharceonus" order="Coleoptera" pageId="48" pageNumber="81" phylum="Arthropoda" rank="genus">
<pageBreakToken id="66063274EBCC137FD5D4A56A91AC3473" pageId="48" pageNumber="81" start="start">Pharceonus</pageBreakToken>
</taxonomicName>
by its small size (3.1-3.5 mm), dark brown color, genitalia (aedeagus + phallobase) of medium width (4.3
<normalizedToken id="3A8372E35E747E46DF855F78064FD094" originalValue="×">x</normalizedToken>
as long as wide), and pronotum with only a shallow transverse impression across apical third (Fig. 87).
</paragraph>
</subSubSection>
<subSubSection id="AD932FB92806D08F8573BF697B3ACE94" pageId="48" pageNumber="81" type="intraspecific variation">
<paragraph id="74C6660F4C8647BD0EB421557A1D7E9F" pageId="48" pageNumber="81">Intraspecific variation.</paragraph>
<paragraph id="A2EF3CBABFE592B5DAA926CB23DFD69A" pageId="48" pageNumber="81">This species varies slightly in color, from dark brown to medium brown, and size, from 3.1-3.5 mm.</paragraph>
</subSubSection>
<subSubSection id="C94DC58CD457B42B2FDE6ECF19437704" pageId="48" pageNumber="81" type="distribution">
<paragraph id="00ECAAA9FD862B1D2A76CF557FF7FA7B" pageId="48" pageNumber="81">Distribution and habitat.</paragraph>
<paragraph id="4A44571D6F5C1A29387C8A2B8FBE6402" pageId="48" pageNumber="81">This species is found on roots in small to medium forested streams, and on roots in seeps in the Sierra de Perija in Venezuela (Fig. 5). This appears to be the southern extent of its range. Outside of Venezuela, this species is known from as far north as Costa Rica. Specimens have been collected in leaf packs and in the benthos in shallow streams.</paragraph>
</subSubSection>
<subSubSection id="4382B74F4AC62FF68F6AC029F809DCD1" pageId="49" pageNumber="82" type="associated species">
<paragraph id="4A892E3ABD6BBDB44FE75FC22309CEBF" pageId="49" pageNumber="82">
<pageBreakToken id="5FA89C89B952A8A94CAE0BDE9B30A83A" pageId="49" pageNumber="82" start="start">Associated</pageBreakToken>
species.
</paragraph>
<paragraph id="5DFE967B298023DEA9B6B80F1829D6DB" pageId="49" pageNumber="82">
This species was collected at the same sites as following taxa:
<taxonomicName id="5E9B25A1C88E5ED1C888F716F843C589" class="Insecta" family="Hydrophilidae" genus="Oocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oocyclus" order="Coleoptera" pageId="49" pageNumber="82" phylum="Arthropoda" rank="genus">Oocyclus</taxonomicName>
spp.,
<taxonomicName id="5B4EB372FE26964936C638C12B51EB2E" class="Insecta" family="Hydrophilidae" genus="Anacaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anacaena" order="Coleoptera" pageId="49" pageNumber="82" phylum="Arthropoda" rank="genus">Anacaena</taxonomicName>
spp.,
<taxonomicName id="6629F9AA6884343593ED7418D3FD3707" class="Insecta" family="Hydrophilidae" genus="Enochrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enochrus" order="Coleoptera" pageId="49" pageNumber="82" phylum="Arthropoda" rank="genus">Enochrus</taxonomicName>
spp.,
<taxonomicName id="0ABB1587F6DFEBA3A343FADE2491677F" class="Insecta" family="Hydrophilidae" genus="Notionotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notionotus" order="Coleoptera" pageId="49" pageNumber="82" phylum="Arthropoda" rank="genus">Notionotus</taxonomicName>
spp., and
<taxonomicName id="ADF91AD6B9CCFDC5FF03736B3C2E18A9" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus" order="Coleoptera" pageId="49" pageNumber="82" phylum="Arthropoda" rank="genus">Chasmogenus</taxonomicName>
spp. (
<taxonomicName id="01E5C3D1CE91151921B335D7097F9741" class="Insecta" family="Hydrophilidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="49" pageNumber="82" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
), and
<taxonomicName id="E1D91AD50A746390BAB3897377DCC54C" class="Insecta" family="Hydraenidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="49" pageNumber="82" phylum="Arthropoda" rank="family">Hydraenidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>