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<document id="23EB871B7A7749ECFF7F436323122A7D" ID-CLB-Dataset="28457" ID-DOI="http://dx.doi.org/10.3897/nl.42.28505" ID-GBIF-Dataset="144fe477-72f6-4128-8a2f-b41b143486cc" ID-Pensoft-Pub="2367-5365-1-17" ID-ZooBank="D9A5B42189804807B3E40AF0AA205C02" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="2367-5365-1-17" ModsDocOrigin="Nota Lepidopterologica 42 (1)" ModsDocTitle="Holcophora Staudinger, 1871, a senior synonym of Aponoea Walsingham, 1905, syn. n., (Lepidoptera, Gelechioidea, Gelechiidae): with Holcophorainderskella (Caradja, 1920), comb. n., transferred from Blastobasis Zeller, 1855 (Blastobasidae)" checkinTime="1552155030140" checkinUser="pensoft" docAuthor="Adamski, David &amp; Sattler, Klaus" docDate="2019" docId="8FCA9615BAE55C5E2C5BFF8F2D559E55" docLanguage="en" docName="NotaLepidopt 42(1): 17-25" docOrigin="Nota Lepidopterologica 42 (1)" docSource="http://dx.doi.org/10.3897/nl.42.28505" docTitle="Holcophora statices Staudinger 1871" docType="treatment" docVersion="6" lastPageNumber="24" masterDocId="FFB7DC2EFF96AE1AFFAD2705A30A1676" masterDocTitle="Holcophora Staudinger, 1871, a senior synonym of Aponoea Walsingham, 1905, syn. n., (Lepidoptera, Gelechioidea, Gelechiidae): with Holcophorainderskella (Caradja, 1920), comb. n., transferred from Blastobasis Zeller, 1855 (Blastobasidae)" masterLastPageNumber="25" masterPageNumber="17" pageNumber="23" updateTime="1701383041549" updateUser="ExternalLinkService">
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<mods:title id="B0C5CF3368E414004587B542A46F56DA">Holcophora Staudinger, 1871, a senior synonym of Aponoea Walsingham, 1905, syn. n., (Lepidoptera, Gelechioidea, Gelechiidae): with Holcophorainderskella (Caradja, 1920), comb. n., transferred from Blastobasis Zeller, 1855 (Blastobasidae)</mods:title>
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<subSubSection id="257E015410FACC1039F726656AFF47B9" pageId="6" pageNumber="23" type="nomenclature">
<paragraph id="439DCF2513B676B9345F8627A42A7AAA" pageId="6" pageNumber="23">
<taxonomicName id="3B8951366C94E4B7F1C9864F137D7450" ID-CoL="6M9WQ" authority="Staudinger, 1871" authorityName="Staudinger" authorityYear="1871" class="Insecta" family="Gelechiidae" genus="Holcophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holcophora statices" order="Lepidoptera" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="statices">Holcophora statices Staudinger, 1871</taxonomicName>
</paragraph>
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<subSubSection id="C2839D22943DE11B8124E837C784DEE4" pageId="6" pageNumber="23" type="reference_group">
<paragraph id="654167E0F1E5DD1ECB26DFD5F95A5DAF" pageId="6" pageNumber="23">
<taxonomicName id="7F61DC44491A0B24DD759218734204E9" class="Insecta" family="Gelechiidae" genus="Holcophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holcophora statices" order="Lepidoptera" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="statices">Holcophora statices</taxonomicName>
Staudinger, 1871: 313. Lectotype ♂, RUSSIAN FEDERATION: S Russia, Volgograd, Krasnoarmeysk (
<normalizedToken id="4F7D4D463C896BBA13AE0FE5015CD550" originalValue="Sarepta">'Sarepta'</normalizedToken>
) (
<taxonomicName id="F8211F0F093D38BAB5E1B88C3C653ABF" family="Gelechiidae" genus="Christoph" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Christoph" order="Lepidoptera" pageId="6" pageNumber="23" rank="genus">Christoph</taxonomicName>
), PRESENT DESIGNATION. (ZMHU, Berlin).
</paragraph>
</subSubSection>
<subSubSection id="20097F811638940BBAABBAFB2130EC74" pageId="6" pageNumber="23" type="host">
<paragraph id="1E5D4324162821B1B5EBE8353CDA0BA2" pageId="6" pageNumber="23">Hostplant(s).</paragraph>
<paragraph id="B1A3E9857C3E61BD40A0886D4FAA0C35" pageId="6" pageNumber="23">
Unknown.
<taxonomicName id="6843D29D0098DE06A80B908BDABD852A" class="Magnoliopsida" family="Plumbaginaceae" genus="Limonium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Limonium" order="Caryophyllales" pageId="6" pageNumber="23" phylum="Tracheophyta" rank="genus">Limonium</taxonomicName>
sp. ('
<taxonomicName id="70803C0281E928C1ED3FAB2BA5A94794" class="Magnoliopsida" family="Plumbaginaceae" genus="Statice" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Statice" order="Caryophyllales" pageId="6" pageNumber="23" phylum="Tracheophyta" rank="genus">Statice</taxonomicName>
') sp. suspected (Staudinger 1871: 313) -
<taxonomicName id="B09FA3A09D741F79B4B74754659C85AC" family="Plumbaginaceae" lsidName="" pageId="6" pageNumber="23" rank="family">Plumbaginaceae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A1DC635AB99D7B1DC4A6CA10735BEE71" pageId="6" pageNumber="23" type="distribution">
<paragraph id="DFB03321350DC1011C80CE8C826FAFF7" pageId="6" pageNumber="23">Distribution.</paragraph>
<paragraph id="C638CA52E090603ACC92391B96657C9A" pageId="6" pageNumber="23">Palaearctic: Europe (France, Hungary, Romania, Ukraine, Russia), Kazakhstan, China.</paragraph>
</subSubSection>
<subSubSection id="0A252D779451595A8A99E8FCAED784D4" pageId="6" pageNumber="23" type="remarks">
<paragraph id="469CF999E5F81C4AECB642CE67F1D324" pageId="6" pageNumber="23">Remarks.</paragraph>
<paragraph id="57503BBD5B585A3041931E2441CD3BA5" pageId="6" pageNumber="23">
<taxonomicName id="A5EB5DD1F1BB4FFCF257830DBAE7A6DC" class="Insecta" family="Gelechiidae" genus="Holcophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Holcophora statices" order="Lepidoptera" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="statices">Holcophora statices</taxonomicName>
was described by Staudinger from 15 specimens, both sexes, he had received from H.
<taxonomicName id="423D5509CCC799234510D21D8FA410FA" genus="Christoph" lsidName="Christoph" pageId="6" pageNumber="23" rank="genus">Christoph</taxonomicName>
under the manuscript name
<taxonomicName id="2ADE73D394AE7D7B4F3D394632B17BB2" class="Insecta" family="Plutellidae" genus="Ypsolophus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ypsolophus statices" order="Lepidoptera" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="statices">Ypsolophus statices</taxonomicName>
. As there was only one female amongst the 15 specimens Staudinger concluded that
<taxonomicName id="716EF3917F00BCEF496263E0D4D7C2F5" genus="Christoph" lsidName="Christoph" pageId="6" pageNumber="23" rank="genus">Christoph</taxonomicName>
may not have bred the species but collected the adults from
<taxonomicName id="73649E0429DD475610B02E9FD1FA518F" class="Magnoliopsida" family="Plumbaginaceae" genus="Limonium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Limonium" order="Caryophyllales" pageId="6" pageNumber="23" phylum="Tracheophyta" rank="genus">Limonium</taxonomicName>
('
<taxonomicName id="4B0E1FB0C1F3C6EB08BC6C8732278C4B" class="Magnoliopsida" family="Plumbaginaceae" genus="Statice" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Statice" order="Caryophyllales" pageId="6" pageNumber="23" phylum="Tracheophyta" rank="genus">Statice</taxonomicName>
')
<taxonomicName id="6C15C8E50D1886E4B7F6C5C95F7273B4" class="Magnoliopsida" family="Plumbaginaceae" genus="Statice" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Statice" order="Caryophyllales" pageId="6" pageNumber="23" phylum="Tracheophyta" rank="genus">Statice</taxonomicName>
flowers.
</paragraph>
</subSubSection>
<subSubSection id="30ED16D8AF2CCBFBA8A58B6BF1723A76" lastPageId="7" lastPageNumber="24" pageId="6" pageNumber="23" type="head structures">
<paragraph id="2B9F14C475D2A0CC6BE60CBE7AE6A773" pageId="6" pageNumber="23">Head structures.</paragraph>
<paragraph id="CAF0324FC1D553B5824BA137C239A3E7" lastPageId="7" lastPageNumber="24" pageId="6" pageNumber="23">
Various modifications of the head structure, culminating in distinct frontal processes are widespread in
<taxonomicName id="E8952C09F568022D3651F6C0FC74E49C" lsidName="" order="Lepidoptera" pageId="6" pageNumber="23" rank="order">Lepidoptera</taxonomicName>
and have independently evolved in many families. If present they are equally developed in both sexes. Their function, although plausibly interpreted in a few instances as that of a cocoon cutter, is largely obscure. Sand grains found on the head around such processes suggest that in some species they may assist the adult in drilling through soil on eclosion from the pupa. In
<taxonomicName id="A218189766D0BF3298189B13B43834FC" lsidName="Gelechioidea" pageId="6" pageNumber="23" rank="superfamily" superfamily="Gelechioidea">Gelechioidea</taxonomicName>
frontal processes are known in
<taxonomicName id="7608E791F7B6E299D37A93157E0EAF78" family="Autostichidae" lsidName="" pageId="6" pageNumber="23" rank="family">Autostichidae</taxonomicName>
(
<taxonomicName id="6EE6895D21F80FD33A0865F7911414E4" lsidName="" pageId="6" pageNumber="23" rank="subfamily" subfamily="Symmocinae">Symmocinae</taxonomicName>
),
<taxonomicName id="026EC2CB39DB1CCD247BB52BDB2B1908" family="Cosmopterigidae" lsidName="" pageId="6" pageNumber="23" rank="family">Cosmopterigidae</taxonomicName>
and
<taxonomicName id="E534FB43A14958B8DF92E5C9A16CF8AE" family="Gelechiidae" lsidName="" pageId="6" pageNumber="23" rank="family">Gelechiidae</taxonomicName>
. In the last family they occur in several genera unrelated to each other, such as
<taxonomicName id="B33B67EC200AFFD60DEAC6FFE5A4DCA9" class="Insecta" family="Gelechiidae" genus="Athrips" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Athrips" order="Lepidoptera" pageId="6" pageNumber="23" phylum="Arthropoda" rank="genus">Athrips</taxonomicName>
Billberg, 1820 (1 of 70+ spp., Palaearctic),
<taxonomicName id="ED9013D8E0FB5C42FED9B57410CE89DD" class="Insecta" family="Gelechiidae" genus="Catatinagma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatinagma" order="Lepidoptera" pageId="6" pageNumber="23" phylum="Arthropoda" rank="genus">Catatinagma</taxonomicName>
Rebel, 1903 (1
<pageBreakToken id="CCE5452D944C13BA7A48BBB9628DDE52" pageId="7" pageNumber="24" start="start">of</pageBreakToken>
5 spp., Palaearctic),
<taxonomicName id="704FC22200F01BCF3680988056695497" class="Insecta" family="Gelechiidae" genus="Caulastrocecis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caulastrocecis" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Caulastrocecis</taxonomicName>
<normalizedToken id="CD17F94665D3EAA09F28C87DA6D483EC" originalValue="Chrétien">Chretien</normalizedToken>
, 1931 (2 of 7 spp., Palaearctic),
<taxonomicName id="F9E824EE4F1035320887B937C66896AC" class="Insecta" family="Gelechiidae" genus="Cerofrontia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerofrontia" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Cerofrontia</taxonomicName>
Janse, 1951 (1 of 1 sp., Afrotropical),
<taxonomicName id="393AB78CF34444F48D59A21CF3DF9ECF" class="Insecta" family="Gelechiidae" genus="Leistogenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leistogenes" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Leistogenes</taxonomicName>
Meyrick, 1927 (1 of 1 sp., Neotropical),
<taxonomicName id="F6630143E312F6D460649958D6CFC24B" class="Insecta" family="Gelechiidae" genus="Ornativalva" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ornativalva" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Ornativalva</taxonomicName>
<normalizedToken id="50E327764230D7F1239D8830492E8F28" originalValue="Gozmány">Gozmany</normalizedToken>
, 1955 (about 15 of 60+ spp., Palaearctic),
<taxonomicName id="78B8728D2536A524DADF07B881645318" class="Insecta" family="Gelechiidae" genus="Prolita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prolita" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Prolita</taxonomicName>
Leraut, 1993 (8 Nearctic, of 22 spp., Holarctic),
<taxonomicName id="1CE2F9BC6F5A4E1193203899FAEF3463" class="Insecta" family="Gelechiidae" genus="Scrobipalpa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scrobipalpa" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Scrobipalpa</taxonomicName>
Janse, 1951 (2 Palaearctic, of 300+ spp. worldwide) and others. Examples were illustrated by
<bibRefCitation id="FFF634A928D6F5EB90FB56CB1FE57C15" author="Hodges, RW" journalOrPublisher="Proceedings of the United States National Museum" pageId="7" pageNumber="24" pagination="1 - 66" title="Revision of Nearctic Gelechiidae, I. The Lita group (Lepidoptera: Gelechioidea)." url="https://biodiversitylibrary.org/page/7760379" volume="119" year="1966">Hodges (1966</bibRefCitation>
, pl. 8,
<taxonomicName id="10F4EECE7AF86A4FB0434BF10862808E" class="Insecta" family="Gelechiidae" genus="Prolita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prolita" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Prolita</taxonomicName>
) and
<bibRefCitation id="98AAFA5D9CDABBE5E01C91AA4BA37492" author="Sattler, K" journalOrPublisher="Bulletin of the British Museum (Natural History) (Entomology)" pageId="8" pageNumber="25" pagination="85 - 152" title="A taxonomic revision of the genus Ornativalva Gozmany, 1955 (Lepidoptera: Gelechiidae)." url="https://biodiversitylibrary.org/page/41004900" volume="34" year="1976">Sattler (1976</bibRefCitation>
, pls 6-12,
<taxonomicName id="4EF11FBBD2D8BB18E150FC476A5FB855" class="Insecta" family="Gelechiidae" genus="Ornativalva" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ornativalva" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Ornativalva</taxonomicName>
). Such specializations of the head structure can provide useful characters for species recognition but they are not important at the generic level as in several genera (
<taxonomicName id="E10ECE26A8E86A5D380370C798383A4D" class="Insecta" family="Gelechiidae" genus="Athrips" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Athrips" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Athrips</taxonomicName>
,
<taxonomicName id="E7B8DB2F81D6FC05387951D9651ADB97" class="Insecta" family="Gelechiidae" genus="Ornativalva" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ornativalva" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Ornativalva</taxonomicName>
,
<taxonomicName id="27AAB24035313C6CF232505EA5C2EDEF" class="Insecta" family="Gelechiidae" genus="Prolita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prolita" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Prolita</taxonomicName>
,
<taxonomicName id="E1AC7944A58F3CCFF94BEB29ACECD00C" class="Insecta" family="Gelechiidae" genus="Scrobipalpa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scrobipalpa" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Scrobipalpa</taxonomicName>
) there exist species with processes next to such without. In
<taxonomicName id="10A58D4F8647B0978DCF16EB4E49EA29" family="Gelechiidae" lsidName="" pageId="7" pageNumber="24" rank="family">Gelechiidae</taxonomicName>
there seems to be a concentration of species with frontal processes in arid parts of the Palaearctic and Nearctic regions.
</paragraph>
<paragraph id="BCBE9C4D8628B79F7815231CE67AB790" pageId="7" pageNumber="24">
Frontal modifications are only obvious once the head is freed of scales, and even a prominent frontal process can otherwise remain hidden in the scale cover. On the denuded head the transfrontal sulcus, the border between frons and vertex, normally extends in an almost straight line between the antennal sockets and is clearly visible as a narrow band free of scale bases. If a discrete frontal process is developed it usually arises beneath the transfrontal sulcus which it tends to push dorsad. Various teeth and protuberances, when present, originate from enlarged scale bases, and the central process may be surrounded by such enlarged scale sockets which can be arranged into a distinctive arc. An additional arc of teeth sometimes arises above the transfrontal sulcus. The whole range of such developments is demonstrated in the genus
<taxonomicName id="5A600C66EBC9BB66B56010AE9AEEF755" class="Insecta" family="Gelechiidae" genus="Ornativalva" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ornativalva" order="Lepidoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">Ornativalva</taxonomicName>
<normalizedToken id="B628B69FE400D95ACC1611410F9E70AF" originalValue="Gozmány">Gozmany</normalizedToken>
, 1955 (
<bibRefCitation id="769F28C357BB600DA0339475FEA83D14" author="Sattler, K" journalOrPublisher="Bulletin of the British Museum (Natural History) (Entomology)" pageId="8" pageNumber="25" pagination="85 - 152" title="A taxonomic revision of the genus Ornativalva Gozmany, 1955 (Lepidoptera: Gelechiidae)." url="https://biodiversitylibrary.org/page/41004900" volume="34" year="1976">Sattler 1976</bibRefCitation>
: 92, pls 5-12), where the arrangement in
<taxonomicName id="74B1DBF04B6AD5E135DFE90F6D734CF1" lsidName="O. cornifrons" pageId="7" pageNumber="24" rank="species" species="cornifrons">O. cornifrons</taxonomicName>
Sattler, 1976, is not unlike that in the unrelated
<taxonomicName id="8D7D0C0E454ACE6A4FCD18F5D984494D" lsidName="H. inderskella" pageId="7" pageNumber="24" rank="species" species="inderskella">H. inderskella</taxonomicName>
.
</paragraph>
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