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<document id="3547C83F32EDA4587B38C2702B9D9D39" ID-CLB-Dataset="33832" ID-DOI="http://dx.doi.org/10.3897/zookeys.415.6406" ID-GBIF-Dataset="aefb1cac-93e1-4a01-8735-f8ba09948390" ID-PMC="PMC4089821" ID-Pensoft-Pub="1313-2970-415-81" ID-PubMed="25009425" ID-ZooBank="372DF48DD1634742AABF5D7E4913050C" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-415-81" ModsDocOrigin="ZooKeys 415" ModsDocTitle="A cladistically based reinterpretation of the taxonomy of two Afrotropical tenebrionid genera Ectateus Koch, 1956 and Selinus Mulsant &amp; Rey, 1853 (Coleoptera, Tenebrionidae, Platynotina)" checkinTime="1451245791699" checkinUser="pensoft" docAuthor="Kaminski, Marcin Jan" docDate="2014" docId="8F7242045FDBA385A7269FC6EFEE6E5B" docLanguage="en" docName="ZooKeys 415: 81-132" docOrigin="ZooKeys 415" docSource="http://dx.doi.org/10.3897/zookeys.415.6406" docTitle="Ectateus calcaripes Gebien 1904, comb. n." docType="treatment" docVersion="6" lastPageNumber="90" masterDocId="FF91FFC18F381D1915336F07FFCA965D" masterDocTitle="A cladistically based reinterpretation of the taxonomy of two Afrotropical tenebrionid genera Ectateus Koch, 1956 and Selinus Mulsant &amp; Rey, 1853 (Coleoptera, Tenebrionidae, Platynotina)" masterLastPageNumber="132" masterPageNumber="81" pageNumber="88" updateTime="1732876459651" updateUser="ExternalLinkService">
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<mods:title id="A41909499319F34B7A274F6B5C3CADB4">A cladistically based reinterpretation of the taxonomy of two Afrotropical tenebrionid genera Ectateus Koch, 1956 and Selinus Mulsant &amp; Rey, 1853 (Coleoptera, Tenebrionidae, Platynotina)</mods:title>
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<treatment id="8F7242045FDBA385A7269FC6EFEE6E5B" ID-GBIF-Taxon="152053554" LSID="urn:lsid:plazi:treatment:8F7242045FDBA385A7269FC6EFEE6E5B" httpUri="http://treatment.plazi.org/id/8F7242045FDBA385A7269FC6EFEE6E5B" lastPageId="9" lastPageNumber="90" pageId="7" pageNumber="88" scope_family="Tenebrionidae" scope_order="Coleoptera">
<subSubSection id="F9D3609F7939972193509783EBDF2FCF" pageId="7" pageNumber="88" type="nomenclature">
<paragraph id="047D4749A51816ACB260DBA01422E268" pageId="7" pageNumber="88">
<taxonomicName id="24DA4DEA051362E5ACA65A84F1FA8928" LSID="http://species-id.net/wiki/Ectateus_calcaripes" authority="Gebien, 1904" authorityName="Gebien" authorityYear="1904" genus="Ectateus" lsidName="Ectateus calcaripes" pageId="7" pageNumber="88" rank="species" species="calcaripes">Ectateus calcaripes (Gebien, 1904)</taxonomicName>
<taxonomicNameLabel id="2425F67ABB3D511B5276854908128EAC" pageId="7" pageNumber="88">comb. n.</taxonomicNameLabel>
Figs 3, 20, 27, 31, 41, 45
</paragraph>
</subSubSection>
<subSubSection id="B36C49887038DD1C6E8FC661FC99546A" pageId="7" pageNumber="88" type="reference_group">
<paragraph id="4D4DD24A2EC776F18A064B063019A4B7" pageId="7" pageNumber="88">
<taxonomicName id="A5A852F2804999D3A87F5280CAEBC660" genus="Selinus" lsidName="Selinus calcaripes" pageId="7" pageNumber="88" rank="species" species="calcaripes">Selinus calcaripes</taxonomicName>
Gebien, 1904: 3. -
<bibRefCitation id="4265B9F74E77E3C4F58328D52ACF282E" author="Gebien, H" journalOrPublisher="Berlin" pageId="40" pageNumber="121" pagination="167 - 354" title="W. Junk, S. Schenkling, Coleopterorum Catalogus" year="1910">Gebien 1910</bibRefCitation>
: 277,
<bibRefCitation id="D63B3162005DDE9A720368608446F6D1" author="Gebien, H" journalOrPublisher="Mitteilungen der Muncher Entomologischen Gesellschaft" pageId="41" pageNumber="122" title="Katalog der Tenebrioniden. Teil II" volume="28" year="1938">1938</bibRefCitation>
: 297;
<bibRefCitation id="1E2FD1E4114C5612470E05B5C8584E6D" author="Koch, C" journalOrPublisher="Bruxelles" pageId="42" pageNumber="123" title="Exploration du Parc National de l'Upemba. II. Tenebrionidae (Coleoptera, Polyphaga), Opatrinae, First part: Platynotini, Litoborini and Loensini" year="1956">Koch 1956</bibRefCitation>
: 238;
<bibRefCitation id="56C296CE86C2A80372BC62DCD3D814A8" author="Kulzer, H" journalOrPublisher="Entomologischen Arbeiten aus dem Museum G. Frey" pageId="42" pageNumber="123" pagination="375 - 434" title="Verzeichnis des Typenmaterials der Tenebrionidensammlung des Museums G. Frey." volume="14" year="1963">Kulzer 1963</bibRefCitation>
: 425,
<bibRefCitation id="9DD56FD1EA3EBB9DC44DE6E32D15C85E" author="Iwan, D" journalOrPublisher="Genus" pageId="41" pageNumber="122" pagination="219 - 323" title="Catalogue of the World Platynotini (Coleoptera: Tenebrionidae)." volume="13" year="2002 b">Iwan 2002b</bibRefCitation>
: 265.
</paragraph>
</subSubSection>
<subSubSection id="D6458D9AAC158CC05C77D6BCD502D3DA" pageId="7" pageNumber="88" type="notes">
<paragraph id="E233463B142B786626F26C76F0D22EAE" pageId="7" pageNumber="88">Notes.</paragraph>
<paragraph id="49062E6F25B0B720022F69C951035C1C" pageId="7" pageNumber="88">
The types of
<taxonomicName id="9E2044AF1AC2B2A88A46A2F41AEA130D" genus="Selinus" lsidName="Selinus calcaripes" pageId="7" pageNumber="88" rank="species" species="calcaripes">Selinus calcaripes</taxonomicName>
seems to be lost. According to the information provided by
<bibRefCitation id="3D474B185D102388BC1C3EEF19A01CA3" author="Iwan, D" journalOrPublisher="Genus" pageId="41" pageNumber="122" pagination="219 - 323" title="Catalogue of the World Platynotini (Coleoptera: Tenebrionidae)." volume="13" year="2002 b">Iwan (2002b)</bibRefCitation>
they should be deposited in Naturhistorisches Museum collection (Basel, Switzerland). Unfortunately, the curators do not confirm this statement. Additionally, during the preparation of my recent scientific project - Phylogeny, biogeography and generic classification of the
<taxonomicName id="31BEAD2A206A23D0E5B4A9F37B76365F" genus="Ectateus" lsidName="Ectateus" pageId="7" pageNumber="88" rank="genus">Ectateus</taxonomicName>
generic group (
<taxonomicName id="5D178E590443A3EB98BE775200D004D6" genus="Coleoptera" lsidName="Coleoptera" pageId="7" pageNumber="88" rank="genus">Coleoptera</taxonomicName>
:
<taxonomicName id="32B9DE70612EEE789451AFAAE1766400" class="Insecta" family="Tenebrionidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
:
<taxonomicName id="9D1890CC274C2848DAE24867ED63A211" genus="Platynotina" lsidName="Platynotina" pageId="7" pageNumber="88" rank="genus">Platynotina</taxonomicName>
) - I have studied diverse entomological material concerning the subtribe
<taxonomicName id="E19E536772A9F9606967C196ADF971F6" genus="Platynotina" lsidName="Platynotina" pageId="7" pageNumber="88" rank="genus">Platynotina</taxonomicName>
from several collections across the World and I did not menaged to locate these specimens.
</paragraph>
<paragraph id="984E62B93A94CC2044B127138CB2A256" pageId="7" pageNumber="88">
Based only on the original species descriptions
<bibRefCitation id="0C1A576EC0EA4F6AD1FEF2084448E631" author="Koch, C" journalOrPublisher="Bruxelles" pageId="42" pageNumber="123" title="Exploration du Parc National de l'Upemba. II. Tenebrionidae (Coleoptera, Polyphaga), Opatrinae, First part: Platynotini, Litoborini and Loensini" year="1956">Koch (1956)</bibRefCitation>
proposed to consider
<taxonomicName id="98E60AF0A64F28F3AFB887C021433C0F" genus="Selinus" lsidName="Selinus calcaripes" pageId="7" pageNumber="88" rank="species" species="calcaripes">Selinus calcaripes</taxonomicName>
as a synonym of
<taxonomicName id="DC49EFBEC8640493836A52CAD710C0C1" genus="Ectateus" lsidName="Ectateus curtulus" pageId="7" pageNumber="88" rank="species" species="curtulus">Ectateus curtulus</taxonomicName>
(Fairmaire, 1893). Unfortunately, the morphology of the holotype of
<taxonomicName id="CDFE7AA54EE96DB971F0B37AEE75FAC0" genus="Ectateus" lsidName="Ectateus curtulus" pageId="7" pageNumber="88" rank="species" species="curtulus">Ectateus curtulus</taxonomicName>
(damaged female - Fig. 47) do not correspond to
<bibRefCitation id="150279BC8CD61FEB36CF44B2128DEB12" author="Gebien, H" journalOrPublisher="Arkiv Fur Zoologi" pageId="40" pageNumber="121" pagination="1 - 31" title="Beitraege zur Kenntnis der Insektenfauna von Kamerun. No 28. Verzeichnis der von Professor Dr. Yngve Sjoestedt in Kamerun gesammelten Tenebrioniden." volume="2" year="1904">
<normalizedToken id="9F0E5762360E63D94E473AAA81AD11BF" originalValue="Gebiens">Gebien's</normalizedToken>
(1904)
</bibRefCitation>
description of
<taxonomicName id="2E7476479A6DBB5B0FCB3208C12B70CB" genus="Selinus" lsidName="Selinus calcaripes" pageId="7" pageNumber="88" rank="species" species="calcaripes">Selinus calcaripes</taxonomicName>
and
<bibRefCitation id="3B1C7E58CD340A93F1AE9C3D23B4CC23" author="Koch, C" journalOrPublisher="Bruxelles" pageId="42" pageNumber="123" title="Exploration du Parc National de l'Upemba. II. Tenebrionidae (Coleoptera, Polyphaga), Opatrinae, First part: Platynotini, Litoborini and Loensini" year="1956">
<normalizedToken id="76E519833F188268D45C2F4377EED0BE" originalValue="Kochs">Koch's</normalizedToken>
(1956)
</bibRefCitation>
interpretation of
<taxonomicName id="88AEF35C72F2462FBB3E490615744E1F" genus="Ectateus" lsidName="Ectateus curtulus" pageId="7" pageNumber="88" rank="species" species="curtulus">Ectateus curtulus</taxonomicName>
. Both publications refer rather to a morphological form that is very closely to
<taxonomicName id="2BBB3A66749103B35B711B2D9C5B81C6" genus="Ectateus" lsidName="Ectateus modestus" pageId="7" pageNumber="88" rank="species" species="modestus">Ectateus modestus</taxonomicName>
and differers from it mainly by the structure of male protibiae (Figs 31-32). A consistent to the above mentioned descriptions morph was found in the studied material. It was included in the cladistic analysis as
<taxonomicName id="3F606B5CC603BC6F1FCA100059385BFA" genus="Selinus" lsidName="Selinus calcaripes" pageId="7" pageNumber="88" rank="species" species="calcaripes">Selinus calcaripes</taxonomicName>
.
</paragraph>
<paragraph id="9F9A5BF5C80B19779FF2BF1CB4633ED4" pageId="7" pageNumber="88">
The results of a cladistic analysis confirmed the aforementioned assumption that
<taxonomicName id="335DA388010A71B6FB839988CBDEDDA0" genus="Ectateus" lsidName="Ectateus curtulus" pageId="7" pageNumber="88" rank="species" species="curtulus">Ectateus curtulus</taxonomicName>
and
<taxonomicName id="1B5474AFD1B327085F05651F67E64E68" genus="Selinus" lsidName="Selinus calcaripes" pageId="7" pageNumber="88" rank="species" species="calcaripes">Selinus calcaripes</taxonomicName>
represent two distinct morphological forms (Fig. 25). They can be easily distinguished by the structure of head (char. 3, 5), pronotum (char. 9) and elytra (elytral intervals with conspicuous punctures in
<taxonomicName id="5A5C69C50ABBCF790520B1587AFBD615" genus="Ectateus" lsidName="Ectateus curtulus" pageId="7" pageNumber="88" rank="species" species="curtulus">Ectateus curtulus</taxonomicName>
). Additionally, the results shows that
<taxonomicName id="CCAEA12E8878578529044258F25C3C1F" genus="Selinus" lsidName="Selinus calcaripes" pageId="7" pageNumber="88" rank="species" species="calcaripes">Selinus calcaripes</taxonomicName>
is very closely related to
<taxonomicName id="82C3C67A397A8A809F01AF62DBF7B2D8" genus="Ectateus" lsidName="Ectateus modestus" pageId="7" pageNumber="88" rank="species" species="modestus">Ectateus modestus</taxonomicName>
- which is consistent with
<bibRefCitation id="D4E3E5583510ADFF0858B19226C9A149" author="Gebien, H" journalOrPublisher="Arkiv Fur Zoologi" pageId="40" pageNumber="121" pagination="1 - 31" title="Beitraege zur Kenntnis der Insektenfauna von Kamerun. No 28. Verzeichnis der von Professor Dr. Yngve Sjoestedt in Kamerun gesammelten Tenebrioniden." volume="2" year="1904">
<normalizedToken id="100D22802A1EA3FD6A6350105B9A8B8F" originalValue="Gebiens">Gebien's</normalizedToken>
(1904)
</bibRefCitation>
description and
<bibRefCitation id="C4722362C6C91D62D73ADFF282BADB9C" author="Koch, C" journalOrPublisher="Bruxelles" pageId="42" pageNumber="123" title="Exploration du Parc National de l'Upemba. II. Tenebrionidae (Coleoptera, Polyphaga), Opatrinae, First part: Platynotini, Litoborini and Loensini" year="1956">
<normalizedToken id="FA6BFE782BA3F646F5D1BCB909F1B5C6" originalValue="Kochs">Koch's</normalizedToken>
(1956)
</bibRefCitation>
interpretation.
</paragraph>
<paragraph id="7FEBBDB793E3126F9802D5DF24E31728" pageId="7" pageNumber="88">
Taking into consideration the difficulties associated with
<taxonomicName id="1E8EE76D3FB207FD349C00C57554A702" genus="Selinus" lsidName="Selinus calcaripes" pageId="7" pageNumber="88" rank="species" species="calcaripes">Selinus calcaripes</taxonomicName>
I propose to designate a neotype to clarify the taxonomic status of this species. Additionally, on the basis of the results of a cladistic analysis I propose to treat this taxon as a independent species - not as a synonym of
<taxonomicName id="E8C765D633F663B72727922EEB41CE2C" genus="Ectateus" lsidName="Ectateus curtulus" pageId="7" pageNumber="88" rank="species" species="curtulus">Ectateus curtulus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C2450798722D53F1D0C253239CAB3453" lastPageId="8" lastPageNumber="89" pageId="7" pageNumber="88" type="studied material">
<paragraph id="8AC9539A92D20E94699955E5467F4A27" pageId="7" pageNumber="88">Studied material.</paragraph>
<paragraph id="8AA20CC65790DE1A3F18C2F7D21FAA3C" lastPageId="8" lastPageNumber="89" pageId="7" pageNumber="88">
Neotype designated here, male (MNHN): &quot;Rep. Centrafric., La Maboke, 2.IX.1967, L. Matile rec.&quot;. Other material: male (MNHN):
<normalizedToken id="55D521D472898902817E5F883B9B8D0E" originalValue="“Musée">&quot;Musee</normalizedToken>
du Congo, Haut-Uele: Yebo moto, VII-1926, L. Burgeon&quot;, male (MRAC):
<normalizedToken id="9C1E15314684C6BB94BCB2A4010E9670" originalValue="“Musée">&quot;Musee</normalizedToken>
du Congo, Haut-Uele: Yebo moto, V-1926, L. Burgeon&quot;, male (MRAC):
<normalizedToken id="2F42D67797A119F2B5BA182CDA286099" originalValue="“Musée">&quot;Musee</normalizedToken>
du Congo, Bambesa, 15-X-1933, J. V. Leroy&quot;, male and 3 females (MRAC): &quot;Coll. Mus. Tervuren, Oubanghi-Chari: Bangui I/III.1968, ex. coll. Breuning&quot;, male and female (RBINS): &quot;R.I.Sc.Nat. Belg.,I.G. 16.364&quot;, 4 females (MNHN): &quot;Calcaripes, Geb., det. Dr. Kaszab&quot;, male and female (MNHN): &quot;Boukoko, Rep. Centr Afric.&quot;,
<pageBreakToken id="3FE3679C7570A99F355F3E7D9D6DC38C" pageId="8" pageNumber="89" start="start">male</pageBreakToken>
and female (MNHN): &quot;La Maboke, Rep. Centr Afric.&quot;, 2 males and female (MNHN):
<normalizedToken id="476EA2A4520C5F223EB8BA5DBB8AC4DD" originalValue="“Lamaboké">&quot;Lamaboke</normalizedToken>
, 10.XI.1965&quot;, male and female (MNHN):
<normalizedToken id="C014CF4B3B8EBBBEF844D50EB951AC84" originalValue="“Lamaboké">&quot;Lamaboke</normalizedToken>
, 20-21.XI.1966&quot;, male (MNHN):
<normalizedToken id="8FAFDFAC77DDF75ABF5ACBD1A6CA7ADB" originalValue="“Lamaboké">&quot;Lamaboke</normalizedToken>
, 18.XI.1965&quot;, 2 females (HNHM): &quot;Cameroon, Dja Reserve, Ekom, 21-26.XII. 1996, G.
<normalizedToken id="48128AF04A76548AFACD9DCE7573C451" originalValue="Hácz">Hacz</normalizedToken>
&amp; G.
<normalizedToken id="30AC73AC183343B4DB3C336E3EA0A08A" originalValue="Köszegi”">Koeszegi&quot;</normalizedToken>
, 3 males (MNB): &quot;Kamerun Buea, 1-7.III.1912, v. Rothkirch S.G.&quot;, female (MNB): &quot;Kamerun, Brea 1000m, 2. - 7. III- 1912; v. Rothkirch S.G.&quot;, female (MNB): &quot;Kamerun, Soppo 25. II. 1912, v. Rothkirch S.G.&quot;, 2 males (MNHN):
<normalizedToken id="FFFC5ECA1D8DB15D9688D20E7B01C78D" originalValue="“Lamaboké">&quot;Lamaboke</normalizedToken>
, Rep. Centrafric., V-1964, J. Carayon&quot;, female (MNHN): &quot;1968-69, La
<normalizedToken id="6C8BAADE33798BD612511924FAE40C30" originalValue="Maboké">Maboke</normalizedToken>
,
<normalizedToken id="F72AB85B2EAFC10CABD2B63B2BA5212F" originalValue="Rép">Rep</normalizedToken>
. Centre Afr., P. Teochhi leg.&quot;, male (MNHN): &quot;21.VIII.1969, La
<normalizedToken id="FB284A0A6A2FF11B2E96EC2328232103" originalValue="Maboké">Maboke</normalizedToken>
,
<normalizedToken id="EACC26F58EEF385D4B19EB216D9066CA" originalValue="Rép">Rep</normalizedToken>
. Centre Afr., P. Teochhi leg.&quot;, female (RBINS): &quot;Coll. R. I. Sc. N. B., congo belge Beni, crottes
<normalizedToken id="955535A312849A5CE6F4C037919B865B" originalValue="d">d'</normalizedToken>
<normalizedToken id="8EE59F28CBA1D8E5F6812A4105C6FB58" originalValue="éléphants">elephants</normalizedToken>
, 18-I-1952 Ch. Verbeke&quot;, female (MNHN):
<normalizedToken id="C279568AA98121FF6A010E170FCA8228" originalValue="“Muséum">&quot;Museum</normalizedToken>
Paris, La
<normalizedToken id="A31B83A4B0DD2E7AC112F6FAFCBB080C" originalValue="Maboké">Maboke</normalizedToken>
,
<normalizedToken id="0602B379F9D76088B05D98FBC52D028A" originalValue="Rép">Rep</normalizedToken>
. Centrafric.&quot;, male (MRAC):
<normalizedToken id="7BFD47AEEB275D6AF2DCF5BCE9A54062" originalValue="“Musée">&quot;Musee</normalizedToken>
du Congo,
<normalizedToken id="27FFB6705D4996169DB98A57D4C5B60F" originalValue="Région">Region</normalizedToken>
de Sassa, 1895-96, Colmant&quot;, male and female (MNHN): &quot;Cameroun, Dr. G. Nonveiller&quot;, female (MNHN):
<normalizedToken id="2C39681680C11ACE0A11DFCC5010ABC6" originalValue="“Muséum">&quot;Museum</normalizedToken>
Paris, Congo Moyen,
<normalizedToken id="3796B3F1353365B5612549A5B81406A4" originalValue="Rég">Reg</normalizedToken>
. de
<normalizedToken id="BE893A6B9753B57B2E0C139C5A4B3468" originalValue="M`Baiki">M'Baiki</normalizedToken>
, (D Fidao), Pitard 1919&quot;, male (MNHN): &quot;Uganda Prot., Mabira Forest, Chagwe., 3,500-3,800 ft.&quot;, female (MNHN): &quot;Uganda, Nimuli to, Murchison Falls&quot;, male (MNB): &quot;Neu-Kamerun, No. 3360-71, Tessmann S.G.&quot;
</paragraph>
</subSubSection>
<subSubSection id="214E0E9A7BF2E509C0E4A2EE0A3A5A83" lastPageId="9" lastPageNumber="90" pageId="8" pageNumber="89" type="redescription">
<paragraph id="2BEB527FFE231641CD440E1C1A38ADA7" pageId="8" pageNumber="89">Redescription.</paragraph>
<paragraph id="505D2353629D5C8046557D8C1C554AC9" pageId="8" pageNumber="89">Habitus as in Fig. 45. Body length = 11.5-14.0 mm. Elytra wider and longer than pronotum (width ratio elytra / pronotum = 1.1-1.2; length ratio elytra / the middle of pronotum = 2.4-2.6).</paragraph>
<paragraph id="14E6AF52FF83A6B2385C772BB091E1D8" pageId="8" pageNumber="89">Dorsal side of head dull, with punctures (the intervals between the punctures are smaller than the diameter of the puncture). Frontoclypeal suture coarse. Clypeal emargination relatively deep (clypeal emargination width / depth ratio = 8.0-8.6). Mentum with median part narrow. Submentum with short base. Maxillary palp not widened (width of maxillary palp / length of 3rd antennomere = 1.0-1.1). Length of antennae greater than pronotal length (ratio antenna / pronotum from tip of anterior pronotal angle to tip of posterior pronotal angle = 1.1-1.2). 3rd antennomere relatively long (length ratio of antennomere 3rd / 2nd = 2.8-3.0).</paragraph>
<paragraph id="011CC4A3676F85EABC887B4AA745A23B" pageId="8" pageNumber="89">Pronotal disc transverse (middle of pronotum length / width ratio = 0.4-0.5); dull, with coarse punctures (the intervals between the punctures are smaller than the diameter of the puncture). Anterior pronotal angles sharp and protruding outwards. Lateral margins of pronotal disc sinusoidal. Apophyseal and basal depressions on pronotal disc present; apophyseal depressions rounded. Pronotal hypomera dull; without punctures.</paragraph>
<paragraph id="A85752E30C53C8EF26913CF880DCEFCB" pageId="8" pageNumber="89">Elytra oblong (elytra length / width ratio = 1.1-1.2). Elytral striae with fine punctures; intervals non-convex, with transverse sculpture (Fig. 27). Elytral base slightly rounded. Elytral humeri rounded, not protruding laterad. Wings absent. Scutellum triangular; situated in a depression.</paragraph>
<paragraph id="67D3A98A9D4C4DD72895DDAC93057C54" pageId="8" pageNumber="89">Intercoxal process protruding towards mesoventrite; peaked at the apex. Metaventrite reduced (length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 2). In both sexes abdominal process without tubercles; relatively narrow (process of 1st abdominal ventrite / process of metaventrite = 2.1-2.2). 5th abdominal ventrite without bordering; punctures fine (the intervals between the punctures are greater than the 2 diameters of the puncture).</paragraph>
<paragraph id="2D82501DC470B6B8E229A946185412B5" pageId="9" pageNumber="90">
<pageBreakToken id="DC9795342985EE730F980F5EAF5DA08F" pageId="9" pageNumber="90" start="start">Male</pageBreakToken>
legs. Protarsi slightly narrow. Protibiae as in Fig. 31. Mesotibiae and mesofemorae with large denticle. Metafemorae with an hair fringe. Female legs simple.
</paragraph>
<paragraph id="24BD315EDEB7AA86CE58D6EF0E3B4B8A" pageId="9" pageNumber="90">Male genitalia. Parameres narrowest in the half of their length; length equal to the 0.2 of the rest of aedeagal tegmen (Fig. 20). Clavae hook-shaped (Fig. 20). Female genitalia. Paraproct equal to coxites. Bursa copulatrix with a sclerite in the distal part. Spermatheca with narrow ducts.</paragraph>
</subSubSection>
<subSubSection id="E10E9ACD9C3E5703ABB9C94EEF929F2A" pageId="9" pageNumber="90" type="distribution">
<paragraph id="53B3E6D86FF31988F2C1517B7E00203C" pageId="9" pageNumber="90">Distribution.</paragraph>
<paragraph id="D33D565CC5E0D85D6E3BCCC3C308D2D8" pageId="9" pageNumber="90">This species has been collected in the following ecoregions of Central Africa (Cameroon, Central African Republic, Democratic Republic of the Congo, South Sudan): Atlantic Equatorial coastal forests, East Sudanian savanna, Mount Cameroon and Bioko montane forests, Northeastern Congolian lowland forests, Northwestern Congolian lowland forests (Fig. 41).</paragraph>
</subSubSection>
</treatment>
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