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<document id="0E290AD7C5E054ADDB9FA3B6F456D4FA" ID-CLB-Dataset="32293" ID-DOI="10.1206/0003-0090-415.1.1" ID-GBIF-Dataset="22b02e67-634f-4e71-a8fa-3392aff63985" ID-Handle="2246/6723" ID-Zenodo-Dep="851382" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1503785870183" checkinUser="admin" docAuthor="Lauren A. Esposito, Humberto Y. Yamaguti, Cláudio A. Souza, Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" docDate="2017" docId="8F65ED57FFC4B14D3B43CF6EB04F6CFF" docLanguage="en" docName="BulAmeMusNatHis.2017.415.1-63.pdf.imf" docOrigin="Bulletin of the American Museum of Natural History 415" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Heteroctenus Pocock 1893" docType="treatment" docVersion="20" lastPageNumber="25" masterDocId="735C952FFFCAB1543860CC2CB35C6E75" masterDocTitle="Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="14" updateTime="1698467378382" updateUser="ExternalLinkService">
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<mods:title id="7AC00C5B92CC83CBC824F1A2E27E73DD">Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)</mods:title>
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<mods:namePart id="32F77BC2CAC80C8FFF29BA51237ACB48">Lauren A. Esposito</mods:namePart>
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<mods:namePart id="DB5A22730E5223C9F4E4EAFEDAFA16B6">Humberto Y. Yamaguti</mods:namePart>
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<mods:namePart id="59BC8C5426D2497E8329F67CB046A043">Cláudio A. Souza</mods:namePart>
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<mods:namePart id="B16EB5164B1A404616C6CD1798E6939E">Ricardo Pinto-Da-Rocha</mods:namePart>
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<mods:namePart id="E06C227ECCECB9A3EDCA11859C41D60A">Lorenzo Prendini</mods:namePart>
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<treatment id="8F65ED57FFC4B14D3B43CF6EB04F6CFF" ID-DOI="http://doi.org/10.5281/zenodo.4610649" ID-GBIF-Taxon="132953574" ID-Zenodo-Dep="4610649" LSID="urn:lsid:plazi:treatment:8F65ED57FFC4B14D3B43CF6EB04F6CFF" httpUri="http://treatment.plazi.org/id/8F65ED57FFC4B14D3B43CF6EB04F6CFF" lastPageId="25" lastPageNumber="25" pageId="14" pageNumber="14">
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<taxonomicName id="C0CC27C2FFC4B15A3B43CF6EB71A6D2F" ID-CoL="4WDH" authority="Pocock, 1893" authorityName="Pocock" authorityYear="1893" box="[803,1094,834,858]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC4B15A3B43CF6EB0F16D2C" box="[803,941,834,857]" italics="true" pageId="14" pageNumber="14">Heteroctenus</emphasis>
<bibRefCitation id="635D21B0FFC4B15A3BD5CF6EB71A6D2F" author="Pocock" box="[949,1094,834,858]" firstAuthor="Pocock" journalOrPublisher="Journal of the Linnaean Society" pageId="14" pageNumber="14" pagination="374 - 409" part="24" refString="Pocock, R. I. 1893. Contribution to our knowledge of the arthropod fauna of the West Indies. Part I. Scorpiones and Pedipalpi, with a supplementary note upon freshwater Decapoda of St. Vincent. Journal of the Linnaean Society 24: 374 - 409." title="Contribution to our knowledge of the arthropod fauna of the West Indies. Part I. Scorpiones and Pedipalpi, with a supplementary note upon freshwater Decapoda of St. Vincent" type="journal article" year="1893">Pocock, 1893</bibRefCitation>
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<subSubSection id="4FD60FCAFFC4B15A3AA3CF5FB7F86DD9" pageId="14" pageNumber="14" type="description">
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<figureCitation id="9FF740C4FFC4B15A3AA3CF5FB0606DFE" box="[707,828,883,907]" captionStart="FIG" captionStartId="16.[107,147,1517,1538]" captionTargetBox="[188,1135,224,1487]" captionTargetId="figure@16.[142,1179,224,1487]" captionTargetPageId="16" captionText="FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil." httpUri="https://zenodo.org/record/851384/files/figure.png" pageId="14" pageNumber="14">
<figureCitation id="9FF740C4FFC4B15A3AA3CF5FB0796DFE" box="[707,805,883,907]" captionStart="FIG. 1" captionStartId="16.[107,147,1517,1538]" captionTargetBox="[188,1135,224,1487]" captionTargetId="figure@16.[142,1179,224,1487]" captionTargetPageId="16" captionText="FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil." httpUri="https://zenodo.org/record/851384/files/figure.png" pageId="14" pageNumber="14">Figures 1</figureCitation>
A
</figureCitation>
,
<figureCitation id="9FF740C4FFC4B15A3B28CF5FB0DB6DFE" box="[840,903,883,907]" captionStart="FIG" captionStartId="17.[111,153,1486,1507]" captionTargetBox="[155,1166,224,1455]" captionTargetId="figure@17.[111,1214,224,1455]" captionTargetPageId="17" captionText="FIG. 2. Representative habitats of species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971. A. Isla Mona, Puerto Rico, habitat of Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov. B. Sierra de Cubitas, Camagüey, Cuba, habitat of Heteroctenus junceus (Herbst, 1800), comb. nov. C. Guanambí, Bahía, Brazil, habitat of Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., and Jaguajir rochae (Borelli, 1910), comb. nov. D. Exu, Pernambuco, Brazil, habitat of J. rochae and Physoctonus debilis (C.L. Koch, 1840). E. Santarém, Pará, Brazil, habitat of Rhopalurus laticauda Thorell, 1876. F. Normandia, Roraima, Brazil, habitat of R. laticauda. G. Lapa do Bode, Itaeté, Bahía, Brazil, habitat of Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov. H. Gruta do Lapão, Lençóis, Bahía, Brazil, habitat of Troglorhopalurus translucidus Lourenço et al., 2004." httpUri="https://zenodo.org/record/851386/files/figure.png" pageId="14" pageNumber="14">
<figureCitation id="9FF740C4FFC4B15A3B28CF5FB0096DFE" box="[840,853,883,907]" captionStart="FIG. 2" captionStartId="17.[111,153,1486,1507]" captionTargetBox="[155,1166,224,1455]" captionTargetId="figure@17.[111,1214,224,1455]" captionTargetPageId="17" captionText="FIG. 2. Representative habitats of species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971. A. Isla Mona, Puerto Rico, habitat of Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov. B. Sierra de Cubitas, Camagüey, Cuba, habitat of Heteroctenus junceus (Herbst, 1800), comb. nov. C. Guanambí, Bahía, Brazil, habitat of Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., and Jaguajir rochae (Borelli, 1910), comb. nov. D. Exu, Pernambuco, Brazil, habitat of J. rochae and Physoctonus debilis (C. L. Koch, 1840). E. Santarém, Pará, Brazil, habitat of Rhopalurus laticauda Thorell, 1876. F. Normandia, Roraima, Brazil, habitat of R. laticauda. G. Lapa do Bode, Itaeté, Bahía, Brazil, habitat of Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov. H. Gruta do Lapão, Lençóis, Bahía, Brazil, habitat of Troglorhopalurus translucidus Lourenço et al., 2004." httpUri="https://zenodo.org/record/851386/files/figure.png" pageId="14" pageNumber="14">2</figureCitation>
A, B
</figureCitation>
,
<figureCitation id="9FF740C4FFC4B15A3BF3CF5FB0FD6DFE" box="[915,929,883,907]" captionStart="FIG" captionStartId="18.[973,994,1690,1732]" captionTargetBox="[401,930,221,1725]" captionTargetId="figure@18.[382,942,211,1738]" captionTargetPageId="18" captionText="FIG. 3. Map of Cuba and neighboring islets, plotting known locality records of three species of Heteroctenus Pocock, 1893: A. H. junceus (Herbst, 1800) (circles). B. H. garridoi (Armas, 1974), comb. nov. (squares); H. gibarae (Teruel, 2006), comb. nov. (crosses)." httpUri="https://zenodo.org/record/851388/files/figure.png" pageId="14" pageNumber="14">3</figureCitation>
,
<figureCitation id="9FF740C4FFC4B15A3BCDCF5FB0E06DFE" box="[941,956,883,907]" captionStart="FIG" captionStartId="19.[107,149,1161,1182]" captionTargetBox="[120,1211,195,1121]" captionTargetId="figure@19.[107,1211,195,1130]" captionTargetPageId="19" captionText="FIG. 4. Map of Hispaniola and neighboring islets, plotting known locality records of three species of Heteroctenus Pocock, 1893: A. H. abudi (Armas and Marcano Fondeur, 1987), comb. nov. (triangles); H. bonettii (Armas, 1999), comb. nov. (circles). B. H. princeps (Karsch, 1879), comb. nov. (squares)." httpUri="https://zenodo.org/record/851390/files/figure.png" pageId="14" pageNumber="14">4</figureCitation>
,
<figureCitation id="9FF740C4FFC4B15A3BA8CF5FB7486DFE" box="[968,1044,883,907]" captionStart="FIG" captionStartId="28.[108,150,1167,1188]" captionTargetBox="[108,1213,224,1136]" captionTargetId="figure@28.[108,1213,224,1136]" captionTargetPageId="28" captionText="FIG. 11. Rhopalurusinae Bücherl, 1971, dextral pecten, ventral aspect (A, C, E, G) and pectinal teeth, dorsal aspect (B, D, F). A. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♀ (AMNH). C. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP) D. Jaguajir rochae (Borelli, 1910), comb. nov., (AMNH). E. Rhopalurus laticauda Thorell, 1876, ♂ (AMCC [LP 2845]). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (AMCC [LP 3260]). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP)." httpUri="https://zenodo.org/record/851404/files/figure.png" pageId="14" pageNumber="14">11A, B</figureCitation>
,
<figureCitation id="9FF740C4FFC4B15A3C40CF5FB7176DFE" box="[1056,1099,883,907]" captionStart="FIG" captionStartId="29.[107,149,984,1005]" captionTargetBox="[107,1211,224,956]" captionTargetId="figure@29.[107,1211,224,956]" captionTargetPageId="29" captionText="FIG. 12. Rhopalurusinae Bücherl, 1971, pectinal teeth, dorsal aspect, fine structure. A. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., (MZSP). B. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). C. Rhopalurus laticauda Thorell, 1876, (MZSP). D. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (MZSP). E. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). F. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP)." httpUri="https://zenodo.org/record/851406/files/figure.png" pageId="14" pageNumber="14">12B</figureCitation>
,
<figureCitation id="9FF740C4FFC4B15A3C37CF5FB7FF6DFE" box="[1111,1187,883,907]" captionStart="FIG" captionStartId="32.[107,149,1598,1619]" captionTargetBox="[220,1108,224,1568]" captionTargetId="figure@32.[118,1207,224,1568]" captionTargetPageId="32" captionText="FIG. 14. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. Heteroctenus garridoi (Armas, 1974), comb. nov., ♂ (AMNH). D. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). E. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851410/files/figure.png" pageId="14" pageNumber="14">14A–E</figureCitation>
,
<figureCitation id="9FF740C4FFC4B15A3AA5CFB9B1A86DD8" box="[709,756,917,941]" captionStart="FIG" captionStartId="36.[106,149,1153,1174]" captionTargetBox="[106,634,224,1125]" captionTargetId="figure@36.[106,634,224,1125]" captionTargetPageId="36" captionText="FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851416/files/figure.png" pageId="14" pageNumber="14">17A</figureCitation>
,
<figureCitation id="9FF740C4FFC4B15A3B60CFB9B0176DD8" box="[768,843,917,941]" captionStart="FIG" captionStartId="38.[107,149,1554,1575]" captionTargetBox="[108,1212,224,1525]" captionTargetId="figure@38.[108,1212,224,1526]" captionTargetPageId="38" captionText="FIG. 18. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. Heteroctenus junceus (Herbst, 1800), comb. nov., ♂ (AMNH). D. Heteroctenus junceus (Herbst, 1800), ♀ (AMNH). E. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851418/files/figure.png" pageId="14" pageNumber="14">18A–E</figureCitation>
,
<figureCitation id="9FF740C4FFC4B15A3B37CFB9B0FF6DD9" box="[855,931,917,941]" captionStart="FIG" captionStartId="44.[107,149,1352,1373]" captionTargetBox="[108,1212,224,1323]" captionTargetId="figure@44.[108,1212,224,1323]" captionTargetPageId="44" captionText="FIG. 21. Rhopalurusinae Bücherl, 1971, telson, lateral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851424/files/figure.png" pageId="14" pageNumber="14">21A, B</figureCitation>
,
<figureCitation id="9FF740C4FFC4B15A3BCECFB9B0A76DD9" box="[942,1019,917,941]" captionStart="FIG" captionStartId="45.[108,147,1513,1534]" captionTargetBox="[176,1145,224,1484]" captionTargetId="figure@45.[108,1212,224,1484]" captionTargetPageId="45" captionText="FIG. 22. Rhopalurusinae Bücherl, 1971, telson, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH). J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851426/files/figure.png" pageId="14" pageNumber="14">22A, B</figureCitation>
,
<figureCitation id="9FF740C4FFC4B15A3C66CFB9B70A6DD8" box="[1030,1110,917,941]" captionStart="FIG" captionStartId="46.[109,151,1497,1518]" captionTargetBox="[117,1187,233,1462]" captionTargetId="figure@46.[109,1213,228,1467]" captionTargetPageId="46" captionText="FIG. 23. Rhopalurusinae Bücherl, 1971, hemispermatophores, dorsal aspect (E), dorsal aspect, detail (D, L, R), ectal aspect (A, F, H, O, T), ectal aspect, detail (B, K, M, V), ental aspect (G, I, P, S) and ental aspect, detail (C, J, N, Q, U). A–D. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). E–G. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). H–K. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). L–N. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). O–R. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). S–V. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♂ (MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/851428/files/figure.png" pageId="14" pageNumber="14">23A–N</figureCitation>
,
<figureCitation id="9FF740C4FFC4B15A3C02CFB8B7F86DD9" box="[1122,1188,916,940]" captionStart-0="FIG" captionStart-1="FIG" captionStart-10="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStart-5="FIG" captionStart-6="FIG" captionStart-7="FIG" captionStart-8="FIG" captionStart-9="FIG" captionStartId-0="48.[108,150,1256,1277]" captionStartId-1="49.[108,150,1259,1280]" captionStartId-10="60.[107,149,1681,1702]" captionStartId-2="50.[107,149,1654,1675]" captionStartId-3="52.[107,149,1350,1371]" captionStartId-4="53.[108,150,1350,1371]" captionStartId-5="54.[108,150,1344,1365]" captionStartId-6="56.[108,150,1690,1711]" captionStartId-7="57.[107,149,1673,1694]" captionStartId-8="58.[107,149,1715,1736]" captionStartId-9="59.[107,150,1464,1485]" captionTargetBox-0="[108,1212,224,1227]" captionTargetBox-1="[108,1212,224,1229]" captionTargetBox-10="[222,1112,224,1652]" captionTargetBox-2="[108,1212,224,1625]" captionTargetBox-3="[108,1212,224,1322]" captionTargetBox-4="[108,1212,224,1322]" captionTargetBox-5="[108,1212,224,1316]" captionTargetBox-6="[108,1212,224,1661]" captionTargetBox-7="[108,1212,224,1645]" captionTargetBox-8="[191,1129,228,1686]" captionTargetBox-9="[108,1212,224,1435]" captionTargetId-0="figure@48.[108,1212,224,1227]" captionTargetId-1="figure@49.[108,1212,224,1229]" captionTargetId-10="figure@60.[107,1211,224,1652]" captionTargetId-2="figure@50.[108,1212,224,1625]" captionTargetId-3="figure@52.[108,1212,224,1322]" captionTargetId-4="figure@53.[108,1212,224,1322]" captionTargetId-5="figure@54.[108,1212,224,1316]" captionTargetId-6="figure@56.[108,1212,224,1661]" captionTargetId-7="figure@57.[108,1212,224,1645]" captionTargetId-8="figure@58.[108,1212,228,1687]" captionTargetId-9="figure@59.[108,1212,224,1435]" captionTargetPageId-0="48" captionTargetPageId-1="49" captionTargetPageId-10="60" captionTargetPageId-2="50" captionTargetPageId-3="52" captionTargetPageId-4="53" captionTargetPageId-5="54" captionTargetPageId-6="56" captionTargetPageId-7="57" captionTargetPageId-8="58" captionTargetPageId-9="59" captionText-0="FIG. 25. Heteroctenus Pocock, 1893, pedipalp chela, retrolateral aspect. A. H. abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. H. bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. H. garridoi (Armas, 1974), comb. nov., ♂ (AMNH). Scale bars = 2 mm." captionText-1="FIG. 26. Heteroctenus Pocock, 1893, pedipalp chela, retrolateral aspect. A. H. abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♀ (AMNH). B. H. bonettii (Armas and Marcano Fondeur, 1987), comb. nov., ♀ (AMNH). C. H. garridoi (Armas, 1974), comb. nov., ♀ (AMNH). Scale bars = 2 mm." captionText-10="FIG. 35. Heteroctenus princeps (Karsch, 1879), comb. nov., habitus, dorsal (A, C) and ventral (B, D) aspects. A, B. ♂ (AMNH). C, D. ♀ (AMNH). Scale bars = 10 mm." captionText-2="FIG. 27. Heteroctenus Pocock, 1893, pedipalp chela, retrolateral aspect. A. H. junceus (Herbst, 1800), comb. nov., ♂ (AMNH). B. H. princeps (Karsch, 1879), comb. nov., ♂ (AMNH). C. H. junceus (Herbst, 1800), ♀ (AMNH). D. H. princeps (Karsch, 1879), comb. nov., ♀ (AMNH). Scale bars = 2 mm." captionText-3="FIG. 28. Heteroctenus Pocock, 1893, metasoma and telson, dorsal aspect. A. H. abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. H. bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. H. garridoi (Armas, 1974), comb. nov., ♂ (AMNH). D. H. junceus (Herbst, 1800), ♂ (AMNH). E. H. princeps (Karsch, 1879), comb. nov., ♂ (AMNH). Scale bars = 5 mm." captionText-4="FIG. 29. Heteroctenus Pocock, 1893, metasoma and telson, ventral aspect. A. H. abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. H. bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. H. garridoi (Armas, 1974), comb. nov., ♂ (AMNH). D. H. junceus (Herbst, 1800), ♂ (AMNH). E. H. princeps (Karsch, 1879), comb. nov., ♂ (AMNH). Scale bars = 5 mm." captionText-5="FIG. 30. Heteroctenus Pocock, 1893, metasoma and telson, lateral aspect. A. H. abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. H. bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. H. garridoi (Armas, 1974), comb. nov., ♂ (AMNH). D. H. junceus (Herbst, 1800), ♂ (AMNH). E. H. princeps (Karsch, 1879), comb. nov., ♂ (AMNH). Scale bars = 5 mm." captionText-6="FIG. 31. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., habitus, dorsal (A, C) and ventral (B, D) aspects. A, B. ♂ (AMNH). C, D. ♀ (AMNH). Scale bars = 10 mm." captionText-7="FIG. 32. Heteroctenus bonettii (Armas, 1999), comb. nov., habitus, dorsal (A, C) and ventral (B, D) aspects. A, B. ♂ (AMNH). C, D. ♀ (AMNH). Scale bars = 10 mm." captionText-8="FIG. 33. Heteroctenus garridoi (Armas, 1974), comb. nov., habitus, dorsal (A, C) and ventral (B, D) aspects. A, B. ♂ (CAS). C, D. ♀ (CAS). Scale bars = 10 mm." captionText-9="FIG. 34. Heteroctenus junceus (Herbst, 1800), habitus, dorsal (A, C) and ventral (B, D) aspects. A, B. ♂ (AMNH). C, D. ♀ (AMNH). Scale bars = 10 mm." httpUri-0="https://zenodo.org/record/851432/files/figure.png" httpUri-1="https://zenodo.org/record/851434/files/figure.png" httpUri-10="https://zenodo.org/record/851452/files/figure.png" httpUri-2="https://zenodo.org/record/851436/files/figure.png" httpUri-3="https://zenodo.org/record/851438/files/figure.png" httpUri-4="https://zenodo.org/record/851440/files/figure.png" httpUri-5="https://zenodo.org/record/851442/files/figure.png" httpUri-6="https://zenodo.org/record/851444/files/figure.png" httpUri-7="https://zenodo.org/record/851446/files/figure.png" httpUri-8="https://zenodo.org/record/851448/files/figure.png" httpUri-9="https://zenodo.org/record/851450/files/figure.png" pageId="14" pageNumber="14">25–35</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFC4B15B3ACCCFE7B0396F99" lastPageId="15" lastPageNumber="15" pageId="14" pageNumber="14" type="reference_group">
<paragraph id="07735C41FFC4B15A3ACCCFE7B0E56A52" blockId="14.[684,1213,970,1741]" pageId="14" pageNumber="14">
<taxonomicName id="C0CC27C2FFC4B15A3ACCCFE7B0836D96" authority="Herbst, 1800" authorityName="Herbst" authorityYear="1800" box="[684,991,970,995]" class="Arachnida" family="Buthidae" genus="Scorpio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="species" species="junceus">
<emphasis id="35B88053FFC4B15A3ACCCFE7B0136D97" box="[684,847,971,994]" italics="true" pageId="14" pageNumber="14">Scorpio junceus</emphasis>
<bibRefCitation id="635D21B0FFC4B15A3B37CFE6B0836D96" author="Herbst" box="[855,991,970,995]" firstAuthor="Herbst" journalOrPublisher="In Natursystem der ungeflugelten Insekten" pageId="14" pageNumber="14" pagination="1 - 86" part="4" refString="Herbst, J. F. W. 1800. Naturgeschichte der Skorpionen. In Natursystem der ungeflugelten Insekten 4: 1 - 86. Berlin: Gottlieb August Lange." title="Naturgeschichte der Skorpionen" type="journal article" year="1800">Herbst, 1800</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="C0CC27C2FFC4B15A3C67CFE7B0846A70" authority="(Herbst, 1800))" authorityName="" baseAuthorityName="Herbst" baseAuthorityYear="1800" class="Arachnida" family="Scorpionidae" genus="Heteroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="species" species="junceus">
<emphasis id="35B88053FFC4B15A3C67CFE7B0766A70" italics="true" pageId="14" pageNumber="14">Heteroctonus junceus</emphasis>
<bibRefCitation id="635D21B0FFC4B15A3B52CFC1B0846A70" author="Herbst, J. F. W." box="[818,984,1005,1029]" journalOrPublisher="In Natursystem der ungeflugelten Insekten" pageId="14" pageNumber="14" pagination="1 - 86" part="4" refString="Herbst, J. F. W. 1800. Naturgeschichte der Skorpionen. In Natursystem der ungeflugelten Insekten 4: 1 - 86. Berlin: Gottlieb August Lange." title="Naturgeschichte der Skorpionen" type="journal article" year="1800">(Herbst, 1800))</bibRefCitation>
</taxonomicName>
, type species by subsequent designation.
</paragraph>
<paragraph id="07735C41FFC4B15A3ACCC81EB0BE6A19" blockId="14.[684,1213,970,1741]" pageId="14" pageNumber="14">
<treatmentCitationGroup id="27DC7B6FFFC4B15A3ACCC81EB0BE6A19" pageId="14" pageNumber="14">
<taxonomicName id="C0CC27C2FFC4B15A3ACCC81EB1A56A3C" authority="Herbst, 1800" box="[684,761,1074,1097]" class="Arachnida" family="Buthidae" genus="Scorpio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC4B15A3ACCC81EB1A56A3C" box="[684,761,1074,1097]" italics="true" pageId="14" pageNumber="14">Scorpio</emphasis>
</taxonomicName>
(part):
<bibRefCitation id="635D21B0FFC4B15A3B2DC81DB7796A3C" author="Herbst" box="[845,1061,1073,1097]" firstAuthor="Herbst" journalOrPublisher="In Natursystem der ungeflugelten Insekten" pageId="14" pageNumber="14" pagination="1 - 86" part="4" refString="Herbst, J. F. W. 1800. Naturgeschichte der Skorpionen. In Natursystem der ungeflugelten Insekten 4: 1 - 86. Berlin: Gottlieb August Lange." title="Naturgeschichte der Skorpionen" type="journal article" year="1800">
<treatmentCitation id="866D7A50FFC4B15A3B2DC81DB0A16A3C" author="Herbst, J. F. W." box="[845,1021,1073,1097]" page="65" pageId="14" pageNumber="14" year="1800">Herbst, 1800: 65</treatmentCitation>
–67
</bibRefCitation>
, pl. III, fig. 2;
<treatmentCitation id="866D7A50FFC4B15A3ABCC87FB0F56A1E" author="Latreille, P. A." box="[732,937,1107,1132]" page="126" pageId="14" pageNumber="14" year="1804">
<bibRefCitation id="635D21B0FFC4B15A3ABCC87FB0F56A1E" author="Latreille" box="[732,937,1107,1132]" firstAuthor="Latreille" journalOrPublisher="Paris: F. Dufart." pageId="14" pageNumber="14" pagination="110 - 129" refString="Latreille, P. A. 1804. Histoire des scorpions. In Histoire naturelle, generale et particuliere des crustaces et des insectes. Ouvrage faisant suite aux oeuvres de Leclerc de Buffon, et partyie de cours complet d'histoire naturelle redigee par C. S. Sonnini, Tome VII: 110 - 129. Paris: F. Dufart." title="Histoire des scorpions. In Histoire naturelle, generale et particuliere des crustaces et des insectes. Ouvrage faisant suite aux oeuvres de Leclerc de Buffon, et partyie de cours complet d'histoire naturelle redigee par C. S. Sonnini, Tome VII" type="book chapter" year="1804">Latreille, 1804: 126</bibRefCitation>
</treatmentCitation>
, 127.
</treatmentCitationGroup>
</paragraph>
<paragraph id="07735C41FFC4B15A3ACCC85AB72A6AFB" blockId="14.[684,1213,970,1741]" box="[684,1142,1142,1166]" pageId="14" pageNumber="14">
<treatmentCitationGroup id="27DC7B6FFFC4B15A3ACCC85AB72A6AFB" box="[684,1142,1142,1166]" pageId="14" pageNumber="14">
<taxonomicName id="C0CC27C2FFC4B15A3ACCC85AB0116AF8" authority="(Atreus" authorityName="Gervais" box="[684,845,1142,1166]" class="Arachnida" family="Buthidae" genus="Scorpio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC4B15A3ACCC85AB1A56AF8" box="[684,761,1142,1165]" italics="true" pageId="14" pageNumber="14">Scorpio</emphasis>
(
<emphasis id="35B88053FFC4B15A3B6AC85AB0116AF8" box="[778,845,1142,1165]" italics="true" pageId="14" pageNumber="14">Atreus</emphasis>
</taxonomicName>
) (part):
<treatmentCitation id="866D7A50FFC4B15A3BCAC85AB72E6AFB" author="Gervais, M. P." box="[938,1138,1142,1166]" page="130" pageId="14" pageNumber="14" year="1843">
<bibRefCitation id="635D21B0FFC4B15A3BCAC85AB72E6AFB" author="Gervais" box="[938,1138,1142,1166]" firstAuthor="Gervais" journalOrPublisher="Societe Philomatique de Paris, Extraits des Proces-Verbaux des Seances 5" pageId="14" pageNumber="14" pagination="129 - 131" part="7" refString="Gervais, M. P. 1843. [Les principaux resultats d'um travail sur le famille des scorpions]. Societe Philomatique de Paris, Extraits des Proces-Verbaux des Seances 5 (7): 129 - 131." title="Les principaux resultats d'um travail sur le famille des scorpions" type="journal article" year="1843">Gervais, 1843: 130</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="07735C41FFC4B15A3ABCC8B4B0CC6A81" blockId="14.[684,1213,970,1741]" pageId="14" pageNumber="14">
<bibRefCitation id="635D21B0FFC4B15A3ABCC8B4B04E6AC5" author="Gervais" box="[732,786,1176,1200]" editor="C. A. Walckenaer" firstAuthor="Gervais" journalOrPublisher="8" pageId="14" pageNumber="14" pagination="14 - 74" part="3" refId="ref66100" refString="Gervais, P. M. 1844 b. Scorpions. In C. A. Walckenaer (editor), Histoire naturelle des Insectes. Apteres. Librarie Encyclopedique de Roret, Paris 3 (8): 14 - 74." title="Scorpions" type="journal article" volumeTitle="Histoire naturelle des Insectes. Apteres" year="1844">1844</bibRefCitation>
a: 218, fig. 18; 1844b: 39, 54;
<bibRefCitation id="635D21B0FFC4B15A3C27C8B4B0656AA7" author="Lucas" firstAuthor="Lucas" journalOrPublisher="Paris: A. Bertrand." pageId="14" pageNumber="14" refString="Lucas, H. 1851. Crustacea et Insecta, Atlas Anim. Artic. In Ramon de la Sagra, Historia fisica, politica y natural de la Isla de Cuba, Volume 5. Paris: A. Bertrand." title="Crustacea et Insecta, Atlas Anim. Artic. In Ramon de la Sagra, Historia fisica, politica y natural de la Isla de Cuba, Volume 5" type="book" year="1851">Lucas, 1851: 70</bibRefCitation>
, pl. V, fig. 5–5c;
<bibRefCitation id="635D21B0FFC4B15A3B8AC896B7F86AA7" author="Gervais" box="[1002,1188,1210,1234]" editor="F. de Castelnau" firstAuthor="Gervais" journalOrPublisher="Paris: P. Bertrand." pageId="14" pageNumber="14" pagination="41 - 43" refString="Gervais, P. M. 1859. Myriapodes et Scorpions. In F. de Castelnau (editor), Animaux nouveaux ou rares recueillis pendant l'expedition dans les parties centrales de l'Amerique du Sud, de Rio de Janeiro, a Lima, et de Lima au Para: 41 - 43. Paris: P. Bertrand." title="Myriapodes et Scorpions" type="book chapter" volumeTitle="Animaux nouveaux ou rares recueillis pendant l'expedition dans les parties centrales de l'Amerique du Sud" year="1859">Gervais, 1859: 41</bibRefCitation>
, pl. I, fig. 2, 2a–b.
</paragraph>
<paragraph id="07735C41FFC4B15B3ACCC8D2B24568FD" blockId="14.[684,1213,970,1741]" lastBlockId="15.[107,634,228,1741]" lastPageId="15" lastPageNumber="15" pageId="14" pageNumber="14">
<treatmentCitationGroup id="27DC7B6FFFC4B15B3ACCC8D2B24568FD" lastPageId="15" lastPageNumber="15" pageId="14" pageNumber="14">
<taxonomicName id="C0CC27C2FFC4B15A3ACCC8D2B07A6B63" ID-CoL="63QFB" authorityName="Thorell" authorityYear="1876" box="[684,806,1278,1302]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC4B15A3ACCC8D2B07A6B63" box="[684,806,1278,1302]" italics="true" pageId="14" pageNumber="14">Rhopalurus</emphasis>
</taxonomicName>
(part):
<treatmentCitation id="866D7A50FFC4B15A3B1AC8D2B7176B63" author="Karsch, F." box="[890,1099,1278,1302]" page="119" pageId="14" pageNumber="14" year="1879">
<bibRefCitation id="635D21B0FFC4B15A3B1AC8D2B7446B63" author="Karsch, F." box="[890,1048,1278,1302]" journalOrPublisher="Mitteilungen des Munchener Entomologischen Vereins" pageId="14" pageNumber="14" pagination="97 - 136" part="3" refString="Karsch, F. 1879 b. Skorpionologische Beitrage. Part II. Mitteilungen des Munchener Entomologischen Vereins 3 (2): 97 - 136." title="Skorpionologische Beitrage. Part II" type="journal article" year="1879" yearSuffix="b">Karsch, 1879b</bibRefCitation>
: 119
</treatmentCitation>
, 121, 122;
<treatmentCitation id="866D7A50FFC4B15A3ABCC90CB0FC6B4D" author="Pocock, R. I." box="[732,928,1312,1336]" page="36" pageId="14" pageNumber="14" year="1902">
<bibRefCitation id="635D21B0FFC4B15A3ABCC90CB0276B4D" author="Pocock, R. I." bookContentInfo="71 pp." box="[732,891,1312,1336]" journalOrPublisher="Biologia Centrali-Americana. London: Taylor and Francis" pageId="14" pageNumber="14" refString="Pocock, R. I. 1902 a. Arachnida. Scorpiones, Pedipalpi, and Solifugae. Biologia Centrali-Americana. London: Taylor and Francis. 71 pp." title="Arachnida. Scorpiones, Pedipalpi, and Solifugae" type="book" year="1902" yearSuffix="a">Pocock, 1902a</bibRefCitation>
: 36
</treatmentCitation>
–38, pl. VIII, fig. 5, 5a, pl. IX, fig. 1, 1a;
<treatmentCitation id="866D7A50FFC4B15A3B0EC96DB7746B2F" author="Banks, N." box="[878,1064,1345,1370]" page="172" pageId="14" pageNumber="14" year="1909">
<bibRefCitation id="635D21B0FFC4B15A3B0EC96DB7746B2F" author="Banks" box="[878,1064,1345,1370]" firstAuthor="Banks" journalOrPublisher="Informe de la Estacion Experimental Agronomica (Santiago de Las Vegas, Cuba) 2" pageId="14" pageNumber="14" pagination="150 - 174" part="2" refString="Banks, N. 1909. Arachnida of Cuba. Informe de la Estacion Experimental Agronomica (Santiago de Las Vegas, Cuba) 2 (2): 150 - 174." title="Arachnida of Cuba" type="journal article" year="1909">Banks, 1909: 172</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC4B15A3C53C96EB01A6B0E" author="Herrera, M." page="271" pageId="14" pageNumber="14" year="1917">
<bibRefCitation id="635D21B0FFC4B15A3C53C96EB01A6B0E" author="Herrera" firstAuthor="Herrera" journalOrPublisher="Boletin de la Direccion de Estudios Biologicos" pageId="14" pageNumber="14" pagination="265 - 275" part="2" refString="Herrera, M. 1917. Los alacranes de Mexico. Boletin de la Direccion de Estudios Biologicos 2 (2): 265 - 275." title="Los alacranes de Mexico" type="journal article" year="1917">Herrera, 1917: 271</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC4B15A3B32C948B74F6B0E" author="Lampe, E." box="[850,1043,1379,1403]" page="197" pageId="14" pageNumber="14" year="1917">
<bibRefCitation id="635D21B0FFC4B15A3B32C948B74F6B0E" author="Lampe" box="[850,1043,1379,1403]" firstAuthor="Lampe" journalOrPublisher="Jahrbucher des Nassauischen Verein fur Naturkunde, Wiesbaden" pageId="14" pageNumber="14" pagination="185 - 208" part="70" refString="Lampe, E. 1917. Katalog der Skorpione, Pedipalpen und Solifugen des Naturhistorischen Museums der Residenzstadt Wiesbaden. Jahrbucher des Nassauischen Verein fur Naturkunde, Wiesbaden 70 (1): 185 - 208." title="Katalog der Skorpione, Pedipalpen und Solifugen des Naturhistorischen Museums der Residenzstadt Wiesbaden" type="journal article" year="1917">Lampe, 1917: 197</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC4B15A3C7FC94FB0196BE8" author="Franganillo, P. B." page="95" pageId="14" pageNumber="14" year="1930">
<bibRefCitation id="635D21B0FFC4B15A3C7FC94FB0436BE8" author="Franganillo, P. B." journalOrPublisher="Memorias del Instituto Nacional de Investigaciones Cientificas, Museo de Historia Natural, La Habana" pageId="14" pageNumber="14" pagination="45 - 97" part="1" refString="Franganillo, P. B. 1930 a. Aracnidos de Cuba. Mas aracnidos nuevos de la Isla de Cuba. Memorias del Instituto Nacional de Investigaciones Cientificas, Museo de Historia Natural, La Habana 1 (1): 45 - 97 (1 - 53 in a separatum)." title="Aracnidos de Cuba. Mas aracnidos nuevos de la Isla de Cuba" type="journal article" year="1930" yearSuffix="a">Franganillo, 1930a</bibRefCitation>
: 95
</treatmentCitation>
, figs. 19–21; 1930b: 119; Mello-
<treatmentCitation id="866D7A50FFC4B15A3ABCC98BB0D56BCA" author="Mello-Leitao, C. F. de" box="[732,905,1447,1471]" page="14" pageId="14" pageNumber="14" year="1932">
<bibRefCitation id="635D21B0FFC4B15A3ABCC98BB03F6BCA" author="Mello-Leitao, C. F. de" box="[732,867,1447,1471]" journalOrPublisher="Arquivos do Museu Nacional" pageId="14" pageNumber="14" pagination="9 - 46" part="34" refString="Mello-Leitao, C. F. de. 1932. Notas sobre escorpioes sulamericanos. Arquivos do Museu Nacional 34: 9 - 46." title="Notas sobre escorpioes sulamericanos" type="journal article" year="1932">Leitão, 1932</bibRefCitation>
: 14
</treatmentCitation>
, 15;
<treatmentCitation id="866D7A50FFC4B15A3BDCC98BB73A6BCA" author="Meise, W." box="[956,1126,1447,1471]" page="29" pageId="14" pageNumber="14" year="1934">
<bibRefCitation id="635D21B0FFC4B15A3BDCC98BB73A6BCA" author="Meise" box="[956,1126,1447,1471]" firstAuthor="Meise" journalOrPublisher="Nyt Magazin for Naturvidenskaberne" pageId="14" pageNumber="14" pagination="25 - 43" part="72" refString="Meise, W. 1934. Scorpiones. Nyt Magazin for Naturvidenskaberne 72: 25 - 43." title="Scorpiones" type="journal article" year="1934">Meise, 1934: 29</bibRefCitation>
</treatmentCitation>
, 32, 38;
<treatmentCitation id="866D7A50FFC4B15A3ABCC9E4B09E6B95" author="Franganillo, P. B." box="[732,962,1480,1504]" page="21" pageId="14" pageNumber="14" year="1935">
<bibRefCitation id="635D21B0FFC4B15A3ABCC9E4B09E6B95" author="Franganillo" box="[732,962,1480,1504]" firstAuthor="Franganillo" journalOrPublisher="Revista &quot; Belen &quot; (La Habana) 9 (49 - 50)" pageId="14" pageNumber="14" pagination="20 - 26" part="1934" refString="Franganillo, P. B. 1935. Estudio de los aracnidos recogidos durante el verano de 1934. Revista &quot; Belen &quot; (La Habana) 9 (49 - 50): 20 - 26." title="Estudio de los aracnidos recogidos durante el verano de" type="journal article" year="1935">Franganillo, 1935: 21</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="635D21B0FFC4B15A3BAEC9E4B7646B95" author="Franganillo" bookContentInfo="179 pp." box="[974,1080,1480,1504]" firstAuthor="Franganillo" journalOrPublisher="Havana, Cuba: Imprenta Cultural, S. A." pageId="14" pageNumber="14" refId="ref65858" refString="Franganillo, P. B. 1936. Los Aracnidos de Cuba hasta 1936. Havana, Cuba: Imprenta Cultural, S. A. 179 pp." title="Los Aracnidos de Cuba hasta 1936" type="book" year="1936">1936: 164</bibRefCitation>
, figs. 86, 87;
<treatmentCitation id="866D7A50FFC4B15A3ABCC9C7B0FA6877" author="Moreno, A." box="[732,934,1514,1538]" page="65" pageId="14" pageNumber="14" year="1939">
<bibRefCitation id="635D21B0FFC4B15A3ABCC9C7B0DD6877" author="Moreno, A." box="[732,897,1514,1538]" journalOrPublisher="Memorias de la Sociedad Cubana de Historia Natural" pageId="14" pageNumber="14" pagination="63 - 75" part="13" refString="Moreno, A. 1939 a. Contribucion al estudio de los escorpionidos cubanos. Parte II. Superfamilia Buthoidea. Memorias de la Sociedad Cubana de Historia Natural 13 (2): 63 - 75." title="Contribucion al estudio de los escorpionidos cubanos. Parte II. Superfamilia Buthoidea" type="journal article" year="1939" yearSuffix="a">Moreno, 1939a</bibRefCitation>
: 65
</treatmentCitation>
–67, pl. 6, fig. 3; 1939b: 124–128, pl. XIX–XXI; 1940b: 129–130, pl. XXII;
<bibRefCitation id="635D21B0FFC4B15A3B40CA01B0AE6830" author="Prado" box="[800,1010,1581,1606]" firstAuthor="Prado" journalOrPublisher="Sinopse das especies de Rhopalurus. Memorias do Instituto Butantan" pageId="14" pageNumber="14" pagination="25 - 36" part="13" refString="Prado, A. 1940. Contribuicao ao conhecimento dos escorpioes sul-americanos. Sinopse das especies de Rhopalurus. Memorias do Instituto Butantan 13: 25 - 36." title="Contribuicao ao conhecimento dos escorpioes sul-americanos" type="journal article" year="1940">
<treatmentCitation id="866D7A50FFC4B15A3B40CA01B0966830" author="Prado, A." box="[800,970,1581,1606]" page="26" pageId="14" pageNumber="14" year="1940">Prado, 1940: 26</treatmentCitation>
–28
</bibRefCitation>
;
<treatmentCitation id="866D7A50FFC4B15A3B9ECA02B0596812" author="Roewer, C. F." page="219" pageId="14" pageNumber="14" year="1943">
<bibRefCitation id="635D21B0FFC4B15A3B9ECA02B0596812" author="Roewer" firstAuthor="Roewer" journalOrPublisher="Senckenbergiana" pageId="14" pageNumber="14" pagination="205 - 244" part="26" refString="Roewer, C. F. 1943. Uber eine neuerworbene Sammlung von Skorpionen des Natur-Museums Senckenberg. Senckenbergiana 26 (4): 205 - 244." title="Uber eine neuerworbene Sammlung von Skorpionen des Natur-Museums Senckenberg" type="journal article" year="1943">Roewer, 1943: 219</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC4B15A3B70CA7CB0846812" author="Jaume, M. L." box="[784,984,1615,1639]" page="1090" pageId="14" pageNumber="14" year="1954">
<bibRefCitation id="635D21B0FFC4B15A3B70CA7CB0846812" author="Jaume" box="[784,984,1615,1639]" firstAuthor="Jaume" journalOrPublisher="Circulares del Museo y Biblioteca de Zoologia de La Habana" pageId="14" pageNumber="14" pagination="1034 - 1092" refString="Jaume, M. L. 1954. Catalogo de la fauna cubana. IV. Catalogo de los Scorpionida de Cuba. Circulares del Museo y Biblioteca de Zoologia de La Habana: 1034 - 1092." title="Catalogo de la fauna cubana. IV. Catalogo de los Scorpionida de Cuba" type="book chapter" year="1954">Jaume, 1954: 1090</bibRefCitation>
</treatmentCitation>
, 1091;
<bibRefCitation id="635D21B0FFC4B15A3C45CA63B0DA68FC" author="Esquivel de Verde, M. A." journalOrPublisher="Acta Biologica Venezuelica" pageId="14" pageNumber="14" pagination="66 - 70" part="6" refId="ref65225" refString="Esquivel de Verde, M. A. 1968. Notas sobre los Scorpionidae de Venezuela. 1. Nuevos registros sobre la distribucion de algunos grupos en Venezuela. Acta Biologica Venezuelica 6 (2): 66 - 70." title="Notas sobre los Scorpionidae de Venezuela. 1. Nuevos registros sobre la distribucion de algunos grupos en Venezuela" type="journal article" year="1968">Esquivel de Verde, 1968: 67</bibRefCitation>
;
<bibRefCitation id="635D21B0FFC4B15A3BF1CA5DB75F68DE" author="Esquivel de Verde, M. A. &amp; C. E. Machado-Allison" bookContentInfo="53 pp" journalOrPublisher="Cuadernos Cientificos: Direccion de Cultura. Caracas: Universidad Central de Venezuela" pageId="14" pageNumber="14" refId="ref65267" refString="Esquivel de Verde, M. A., and C. E. Machado-Allison. 1969. Escorpiones. Cuadernos Cientificos: Direccion de Cultura. Caracas: Universidad Central de Venezuela. 53 pp" title="Escorpiones" type="book" year="1969">Esquivel de Verde and Machado-Allison, 1969: 33</bibRefCitation>
;
<treatmentCitation id="866D7A50FFC4B15A3C6FCABFB05968B9" author="Bucherl, W." page="327" pageId="14" pageNumber="14" year="1971">
<bibRefCitation id="635D21B0FFC4B15A3C6FCABFB05968B9" author="Bucherl" editor="W. Bucherl" firstAuthor="Bucherl" journalOrPublisher="New York: Academic Press" pageId="14" pageNumber="14" pagination="317 - 348" part="3" refString="Bucherl, W. 1971. Classification, biology and venom extraction of scorpions. In W. Bucherl and E. R. Buckley (editors), Venomous animals and their venoms 3: 317 - 348. New York: Academic Press." title="Classification, biology and venom extraction of scorpions" type="journal article" volumeTitle="Venomous animals and their venoms" year="1971">Bücherl, 1971: 327</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC4B15A3B70CA99B0EA68B9" author="Armas, L. F. de" box="[784,950,1716,1741]" page="7" pageId="14" pageNumber="14" year="1973">
<bibRefCitation id="635D21B0FFC4B15A3B70CA99B0EA68B9" author="Armas" box="[784,950,1716,1741]" firstAuthor="Armas" journalOrPublisher="Poeyana" pageId="14" pageNumber="14" pagination="1 - 18" part="101" refString="Armas, L. F. de. 1973. Tipos de las Colecciones Escorpiologicas P. Franganillo y Universidad de La Habana (Arachnida: Scorpionida). Poeyana 101: 1 - 18." title="Tipos de las Colecciones Escorpiologicas P. Franganillo y Universidad de La Habana (Arachnida: Scorpionida)" type="journal article" year="1973">Armas, 1973: 7</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="635D21B0FFC4B15A3BA2CA98B0A568B9" author="Armas" box="[962,1017,1716,1740]" firstAuthor="Armas" journalOrPublisher="Poeyana" pageId="14" pageNumber="14" pagination="1 - 12" part="136" refId="ref63522" refString="Armas, L. F. de. 1974 b. Escorpiones del archipielago cubano. IV. Nueva especie de Rhopalurus (Scorpionidae: Buthidae). Poeyana 136: 1 - 12." title="Escorpiones del archipielago cubano. IV. Nueva especie de Rhopalurus (Scorpionidae: Buthidae)" type="journal article" year="1974">1974</bibRefCitation>
b: 2–6, figs. 1A, B, 2A, B, 3A, B, 4,
<tableCitation id="4A4E69FAFFC5B15B3929CCC8B2CE6E89" box="[329,402,228,252]" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">table 1</tableCitation>
; 1977: 3;
<bibRefCitation id="635D21B0FFC5B15B399BCCC8B2236F6B" author="Stahnke and Calos" firstAuthor="Stahnke" journalOrPublisher="Entomological News" pageId="15" pageNumber="15" pagination="111 - 120" part="88" refString="Stahnke, H. L., and M. Calos. 1977. A key to the species of the genus Centruroides Marx (Buthidae, Scorpionida). Entomological News 88 (5 - 6): 111 - 120." title="A key to the species of the genus Centruroides Marx (Buthidae, Scorpionida)" type="journal article" year="1977">
Stahnke and
<treatmentCitation id="866D7A50FFC5B15B38AACD29B2236F6B" author="Stahnke, H. L. &amp; M. Calos" box="[202,383,261,286]" page="119" pageId="15" pageNumber="15" year="1977">Calos, 1977: 119</treatmentCitation>
</bibRefCitation>
;
<treatmentCitation id="866D7A50FFC5B15B39EACD2AB1356F68" author="Lourenco, W. R." box="[394,617,261,286]" page="215" pageId="15" pageNumber="15" year="1979">
<bibRefCitation id="635D21B0FFC5B15B39EACD2AB1356F68" author="Lourenco" box="[394,617,261,286]" firstAuthor="Lourenco" journalOrPublisher="Revue Arachnologique" pageId="15" pageNumber="15" pagination="213 - 219" part="2" refString="Lourenco, W. R. 1979. A propos de la veritable identite des genres Rhopalurus Thorell, 1879 et Centruroides Marx, 1889 (Scorpiones, Buthidae). Revue Arachnologique 2 (5): 213 - 219." title="A propos de la veritable identite des genres Rhopalurus Thorell, 1879 et Centruroides Marx, 1889 (Scorpiones, Buthidae)" type="journal article" year="1979">Lourenço, 1979: 215</bibRefCitation>
</treatmentCitation>
, 216, fig. 8;
<treatmentCitation id="866D7A50FFC5B15B3972CD0BB28D6F4A" author="Armas, L. F. de" box="[274,465,295,319]" page="52" pageId="15" pageNumber="15" year="1981">
<bibRefCitation id="635D21B0FFC5B15B3972CD0BB2F66F4A" author="Armas, L. F. de" box="[274,426,295,319]" journalOrPublisher="Revista El Yunque, Baracoa 1 - 2 (2 - 3)" pageId="15" pageNumber="15" pagination="51 - 55" refString="Armas, L. F. de. 1981 a. Algunas consideraciones acerca de la fauna cubana de escorpiones. Revista El Yunque, Baracoa 1 - 2 (2 - 3): 51 - 55." title="Algunas consideraciones acerca de la fauna cubana de escorpiones" type="book chapter" year="1981" yearSuffix="a">Armas, 1981a</bibRefCitation>
: 52
</treatmentCitation>
;
<bibRefCitation id="635D21B0FFC5B15B39BDCD0BB14F6F4A" author="Armas" box="[477,531,295,319]" firstAuthor="Armas" journalOrPublisher="Poeyana" pageId="15" pageNumber="15" pagination="1 - 7" part="227" refId="ref63628" refString="Armas, L. F. de. 1981 b. Redescripcion de Rhopalurus princeps (Karsch, 1879) (Scorpionida: Buthidae). Poeyana 227: 1 - 7." title="Redescripcion de Rhopalurus princeps (Karsch, 1879) (Scorpionida: Buthidae)" type="journal article" year="1981">1981</bibRefCitation>
b: 2–5, figs. 1, 2,
<tableCitation id="4A4E69FAFFC5B15B3962CD65B2176F14" box="[258,331,329,353]" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">table 1</tableCitation>
; 1982a: 4; 1982b: 5,
<tableCitation id="4A4E69FAFFC5B15B3A49CD65B12F6F14" box="[553,627,329,353]" captionStart="TABLE 2" captionStartId="7.[610,690,228,252]" captionTargetBox="[235,1058,354,747]" captionText="TABLE 2 Diploid chromosome numbers (2 n) for Brazilian species of Rhopalurusinae from Ubinski et al., (2016) with corresponding genera indicated." httpUri="http://table.plazi.org/id/53B30CC9FFCDB1533A02CCC8B0866F32" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCDB1533A02CCC8B0866F32">table 2</tableCitation>
;
<treatmentCitation id="866D7A50FFC5B15B38FBCD47B2D96FF6" author="Lourenco, W. R." box="[155,389,363,387]" page="108" pageId="15" pageNumber="15" year="1982">
<bibRefCitation id="635D21B0FFC5B15B38FBCD47B20D6FF6" author="Lourenco, W. R." box="[155,337,363,387]" journalOrPublisher="Revue Arachnologique" pageId="15" pageNumber="15" pagination="107 - 141" part="4" refString="Lourenco, W. R. 1982 a. Revision du genre Rhopalurus Thorell, 1876 (Scorpiones, Buthidae). Revue Arachnologique 4: 107 - 141." title="Revision du genre Rhopalurus Thorell, 1876 (Scorpiones, Buthidae)" type="journal article" year="1982" yearSuffix="a">Lourenço, 1982a</bibRefCitation>
: 108
</treatmentCitation>
, 110–112, 114–116, 134–138, figs. 2–11,
<figureCitation id="9FF740C4FFC5B15B3919CDA1B2E16FD0" box="[377,445,397,421]" captionStart-0="FIG. 14" captionStart-1="FIG. 15" captionStart-10="FIG. 24" captionStart-2="FIG. 16" captionStart-3="FIG. 17" captionStart-4="FIG. 18" captionStart-5="FIG. 19" captionStart-6="FIG. 20" captionStart-7="FIG. 21" captionStart-8="FIG. 22" captionStart-9="FIG. 23" captionStartId-0="32.[107,149,1598,1619]" captionStartId-1="33.[108,150,1627,1648]" captionStartId-10="47.[107,149,1521,1542]" captionStartId-2="34.[109,151,1628,1649]" captionStartId-3="36.[106,149,1153,1174]" captionStartId-4="38.[107,149,1554,1575]" captionStartId-5="39.[108,151,1503,1524]" captionStartId-6="40.[108,151,1490,1511]" captionStartId-7="44.[107,149,1352,1373]" captionStartId-8="45.[108,147,1513,1534]" captionStartId-9="46.[109,151,1497,1518]" captionTargetBox-0="[220,1108,224,1568]" captionTargetBox-1="[235,1082,224,1599]" captionTargetBox-10="[107,1211,228,1487]" captionTargetBox-2="[208,1116,224,1599]" captionTargetBox-3="[106,634,224,1125]" captionTargetBox-4="[108,1212,224,1525]" captionTargetBox-5="[108,1212,227,1475]" captionTargetBox-6="[108,1212,224,1461]" captionTargetBox-7="[108,1212,224,1323]" captionTargetBox-8="[176,1145,224,1484]" captionTargetBox-9="[117,1187,233,1462]" captionTargetId-0="figure@32.[118,1207,224,1568]" captionTargetId-1="figure@33.[108,1209,224,1599]" captionTargetId-10="figure@47.[107,1211,228,1493]" captionTargetId-2="figure@34.[109,1212,224,1599]" captionTargetId-3="figure@36.[106,634,224,1125]" captionTargetId-4="figure@38.[108,1212,224,1526]" captionTargetId-5="figure@39.[108,1212,227,1475]" captionTargetId-6="figure@40.[108,1212,224,1462]" captionTargetId-7="figure@44.[108,1212,224,1323]" captionTargetId-8="figure@45.[108,1212,224,1484]" captionTargetId-9="figure@46.[109,1213,228,1467]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionTargetPageId-10="47" captionTargetPageId-2="34" captionTargetPageId-3="36" captionTargetPageId-4="38" captionTargetPageId-5="39" captionTargetPageId-6="40" captionTargetPageId-7="44" captionTargetPageId-8="45" captionTargetPageId-9="46" captionText-0="FIG. 14. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. Heteroctenus garridoi (Armas, 1974), comb. nov., ♂ (AMNH). D. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). E. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." captionText-1="FIG. 15. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). B. Ischnotelson peruassu, sp. nov., paratype ♂ (MZSP). C. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♂ (MZSP). D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♀ (MZSP). E. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (AMNH). Scale bars = 2 mm." captionText-10="FIG. 24. Rhopalurusinae Bücherl, 1971, hemispermatophores, dorsal aspect (E), dorsal aspect, detail (B, H, M, P), ectal aspect (F, I, N, S), ectal aspect, detail (D, J, R, T, U), ental aspect (A, G, K, O), and ental aspect, detail (C, L, Q). A – D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E – H. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). I – M. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP). N – R. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (MZSP). S – U. Physoctonus debilis (C. L. Koch, 1840), ♂ (MZSP). Scale bars = 1 mm." captionText-2="FIG. 16. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Physoctonus debilis (C. L. Koch, 1840), ♀ (MZSP). B. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). C. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). D. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. E. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Guatire, Venezuela. F. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). Scale bars = 1 mm (A, B), 2 mm (C – F)." captionText-3="FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." captionText-4="FIG. 18. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. Heteroctenus junceus (Herbst, 1800), comb. nov., ♂ (AMNH). D. Heteroctenus junceus (Herbst, 1800), ♀ (AMNH). E. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." captionText-5="FIG. 19. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♀ (MZSP). B. Ischnotelson peruassu, sp. nov., paratype ♂ (MZSP). C. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♂ (MZSP). D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). Scale bars = 2 mm." captionText-6="FIG. 20. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Physoctonus debilis (C. L. Koch, 1840), ♂ (MZSP). B. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). C. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). D. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. E. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Puerto Colombia, Venezuela. F. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). Scale bars = 2 mm." captionText-7="FIG. 21. Rhopalurusinae Bücherl, 1971, telson, lateral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C. L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." captionText-8="FIG. 22. Rhopalurusinae Bücherl, 1971, telson, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C. L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH). J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." captionText-9="FIG. 23. Rhopalurusinae Bücherl, 1971, hemispermatophores, dorsal aspect (E), dorsal aspect, detail (D, L, R), ectal aspect (A, F, H, O, T), ectal aspect, detail (B, K, M, V), ental aspect (G, I, P, S) and ental aspect, detail (C, J, N, Q, U). A – D. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). E – G. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). H – K. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). L – N. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). O – R. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). S – V. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♂ (MZSP). Scale bars = 1 mm." httpUri-0="https://zenodo.org/record/851410/files/figure.png" httpUri-1="https://zenodo.org/record/851412/files/figure.png" httpUri-10="https://zenodo.org/record/851430/files/figure.png" httpUri-2="https://zenodo.org/record/851414/files/figure.png" httpUri-3="https://zenodo.org/record/851416/files/figure.png" httpUri-4="https://zenodo.org/record/851418/files/figure.png" httpUri-5="https://zenodo.org/record/851420/files/figure.png" httpUri-6="https://zenodo.org/record/851422/files/figure.png" httpUri-7="https://zenodo.org/record/851424/files/figure.png" httpUri-8="https://zenodo.org/record/851426/files/figure.png" httpUri-9="https://zenodo.org/record/851428/files/figure.png" pageId="15" pageNumber="15">14–24</figureCitation>
, 78,
<tableCitation id="4A4E69FAFFC5B15B3990CDA1B1646FD0" box="[496,568,397,421]" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">table 1</tableCitation>
;
<treatmentCitation id="866D7A50FFC5B15B38FBCD83B21C6FB2" author="Armas, L. F. de" box="[155,320,431,455]" page="3" pageId="15" pageNumber="15" year="1983">
<bibRefCitation id="635D21B0FFC5B15B38FBCD83B21C6FB2" author="Armas" box="[155,320,431,455]" firstAuthor="Armas" journalOrPublisher="Poeyana" pageId="15" pageNumber="15" pagination="1 - 7" part="257" refString="Armas, L. F. de. 1983. Escorpiofauna de la Isla de la Juventud, Cuba. Composicion, distribucion y origen. Poeyana 257: 1 - 7." title="Escorpiofauna de la Isla de la Juventud, Cuba. Composicion, distribucion y origen" type="journal article" year="1983">Armas, 1983: 3</bibRefCitation>
</treatmentCitation>
, fig. 1; 1984: 8;
<treatmentCitation id="866D7A50FFC5B15B398BCD83B24D6F9D" author="Lourenco, W. R." page="169" pageId="15" pageNumber="15" year="1984">
<bibRefCitation id="635D21B0FFC5B15B398BCD83B3816F9D" author="Lourenco, W. R." journalOrPublisher="Revista Brasileira de Biologia" pageId="15" pageNumber="15" pagination="169 - 170" part="44" refString="Lourenco, W. R. 1984 a. Complementary notes on the systematics of the genus Rhopalurus for the Caribbean area (Scorpiones, Buthidae). Revista Brasileira de Biologia 44 (2): 169 - 170." title="Complementary notes on the systematics of the genus Rhopalurus for the Caribbean area (Scorpiones, Buthidae)" type="journal article" year="1984" yearSuffix="a">Lourenço, 1984a</bibRefCitation>
: 169
</treatmentCitation>
, 170; 1986a: 133, 135, 136,
<figureCitation id="9FF740C4FFC5B15B3A5BCDFCB3EB6C7F" captionStart="FIG. 17" captionStartId="36.[106,149,1153,1174]" captionTargetBox="[106,634,224,1125]" captionTargetId="figure@36.[106,634,224,1125]" captionTargetPageId="36" captionText="FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851416/files/figure.png" pageId="15" pageNumber="15">figs. 17</figureCitation>
, 18; 1986b: 165, fig. 7;
<treatmentCitation id="866D7A50FFC5B15B39D0CDDEB2C76C59" author="Armas, L. F. de &amp; E. J. Marcano Fondeur" page="19" pageId="15" pageNumber="15" year="1987">
<bibRefCitation id="635D21B0FFC5B15B39D0CDDEB2C76C59" author="Armas and Marcano Fondeur" firstAuthor="Armas" journalOrPublisher="Poeyana" pageId="15" pageNumber="15" pagination="1 - 24" part="356" refString="Armas, L. F. de, and E. J. Marcano Fondeur. 1987. Nuevos escorpiones (Arachnida: Scorpiones) de Republica Dominicana. Poeyana 356: 1 - 24." title="Nuevos escorpiones (Arachnida: Scorpiones) de Republica Dominicana" type="journal article" year="1987">Armas and Marcano Fondeur, 1987: 19</bibRefCitation>
</treatmentCitation>
, 20, 23, pl. II, fig. 4, tables 10, 11;
<bibRefCitation id="635D21B0FFC5B15B394FCE1AB1566C3B" author="Armas" bookContentInfo="102 pp." box="[303,522,566,590]" firstAuthor="Armas" journalOrPublisher="Havana, Cuba: Editorial Cientifico-Tecnica" pageId="15" pageNumber="15" refString="Armas, L. F. de. 1988. Sinopsis de los escorpiones antillanos. Havana, Cuba: Editorial Cientifico-Tecnica. 102 pp." title="Sinopsis de los escorpiones antillanos" type="book" year="1988">
<treatmentCitation id="866D7A50FFC5B15B394FCE1AB2BD6C3B" author="Armas, L. F. de" box="[303,481,566,590]" page="68" pageId="15" pageNumber="15" year="1988">Armas, 1988: 68</treatmentCitation>
–71
</bibRefCitation>
, 93, 97, figs. 27, 36;
<treatmentCitation id="866D7A50FFC5B15B397CCE74B2B26C1A" author="Lourenco, W. R." box="[284,494,599,624]" page="55" pageId="15" pageNumber="15" year="1992">
<bibRefCitation id="635D21B0FFC5B15B397CCE74B2B26C1A" author="Lourenco" box="[284,494,599,624]" firstAuthor="Lourenco" journalOrPublisher="Studies of Neotropical Fauna and Environment" pageId="15" pageNumber="15" pagination="43 - 62" part="27" refString="Lourenco, W. R. 1992. Les peuplements des scorpions des Antilles; facteurs historiques et ecologiques en association avec les strategies demographiques. Studies of Neotropical Fauna and Environment 27 (1): 43 - 62." title="Les peuplements des scorpions des Antilles; facteurs historiques et ecologiques en association avec les strategies demographiques" type="journal article" year="1992">Lourenço, 1992: 55</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC5B15B3999CE74B3B66CE7" author="Rudloff, J. - P." page="9" pageId="15" pageNumber="15" year="1994">
<bibRefCitation id="635D21B0FFC5B15B3999CE74B3B66CE7" author="Rudloff" firstAuthor="Rudloff" journalOrPublisher="Arthropoda" pageId="15" pageNumber="15" pagination="3 - 12" part="2" refString="Rudloff, J. - P. 1994. Die Skorpionsfauna der Antillen (Arachnida: Scorpiones). Teil I. Arthropoda 2: 3 - 12." title="Die Skorpionsfauna der Antillen (Arachnida: Scorpiones). Teil I" type="journal article" year="1994">Rudloff, 1994: 9</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC5B15B3897CE56B2656CC6" author="Lourenco, W. R. &amp; J. L. Cloudsley-Thompson" page="424" pageId="15" pageNumber="15" year="1995">
<bibRefCitation id="635D21B0FFC5B15B3897CE56B2656CC6" author="Lourenco and Cloudsley-Thompson" firstAuthor="Lourenco" journalOrPublisher="Journal of Arid Environments" pageId="15" pageNumber="15" pagination="423 - 429" part="31" refString="Lourenco, W. R., and J. L. Cloudsley-Thompson. 1995. Stridulatory apparatus and the evolutionary significance of sound production in Rhopalurus species (Scorpiones: Buthidae). Journal of Arid Environments 31: 423 - 429." title="Stridulatory apparatus and the evolutionary significance of sound production in Rhopalurus species (Scorpiones: Buthidae)" type="journal article" year="1995">Lourenço and Cloudsley-Thompson, 1995: 424</bibRefCitation>
</treatmentCitation>
, 426;
<treatmentCitation id="866D7A50FFC5B15B3919CEB0B13F6CC6" author="Lourenco, W. R." box="[377,611,667,691]" page="590" pageId="15" pageNumber="15" year="1997">
<bibRefCitation id="635D21B0FFC5B15B3919CEB0B1736CC6" author="Lourenco, W. R." box="[377,559,667,691]" journalOrPublisher="Revue Suisse de Zoologie" pageId="15" pageNumber="15" pagination="587 - 604" part="104" refString="Lourenco, W. R. 1997 a. Additions a la faune de scorpions neotropicaux (Arachnida). Revue Suisse de Zoologie 104 (3): 587 - 604." title="Additions a la faune de scorpions neotropicaux (Arachnida)" type="journal article" year="1997" yearSuffix="a">Lourenço, 1997a</bibRefCitation>
: 590
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC5B15B38FBCE92B1006CA0" author="Lourenco, W. R. &amp; R. Pinto-da-Rocha" box="[155,604,701,725]" page="181" pageId="15" pageNumber="15" year="1997">
<bibRefCitation id="635D21B0FFC5B15B38FBCE92B1006CA0" author="Lourenco and Pinto-da-Rocha" box="[155,604,701,725]" firstAuthor="Lourenco" journalOrPublisher="Biogeographica (Paris) 73" pageId="15" pageNumber="15" pagination="181 - 191" part="4" refString="Lourenco, W. R., and R. Pinto-da-Rocha. 1997. A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species. Biogeographica (Paris) 73 (4): 181 - 191." title="A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species" type="journal article" year="1997">Lourenço and Pinto-da-Rocha, 1997: 181</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC5B15B38FBCEF3B2346C82" author="Kovarik, F." box="[155,360,735,759]" page="181" pageId="15" pageNumber="15" year="1997">
<bibRefCitation id="635D21B0FFC5B15B38FBCEF3B2346C82" author="Kovarik" box="[155,360,735,759]" firstAuthor="Kovarik" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="15" pageNumber="15" pagination="177 - 185" part="89" refString="Kovarik, F. 1997. A check-list of scorpions (Arachnida) in the collection of the Hungarian Natural History Museum, Budapest. Annales Historico-Naturales Musei Nationalis Hungarici 89: 177 - 185." title="A check-list of scorpions (Arachnida) in the collection of the Hungarian Natural History Museum, Budapest" type="journal article" year="1997">Kovařík, 1997: 181</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC5B15B3914CEF3B2826C82" author="Kovarik, F." box="[372,478,735,759]" page="118" pageId="15" pageNumber="15" year="1998">
<bibRefCitation id="635D21B0FFC5B15B3914CEF3B2826C82" author="Kovarik" bookContentInfo="175 pp." box="[372,478,735,759]" firstAuthor="Kovarik" journalOrPublisher="Madagaskar: Jihlava" pageId="15" pageNumber="15" refString="Kovarik, F. 1998. Stiri [Scorpions]. Madagaskar: Jihlava. 175 pp. [in Czech]" title="Stiri [Scorpions]" type="book" year="1998">1998: 118</bibRefCitation>
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;
<treatmentCitation id="866D7A50FFC5B15B398ACEF3B3986D6C" author="Armas, L. F. de" page="127" pageId="15" pageNumber="15" year="1999">
<bibRefCitation id="635D21B0FFC5B15B398ACEF3B3986D6C" author="Armas" firstAuthor="Armas" journalOrPublisher="Avicennia 10" pageId="15" pageNumber="15" pagination="101 - 136" part="11" refString="Armas, L. F. de. 1999. Quince nuevos alacranes de La Espanola y Navassa, Antillas Mayores (Arachnida: Scorpiones). Avicennia 10 / 11: 101 - 136." title="Quince nuevos alacranes de La Espanola y Navassa, Antillas Mayores (Arachnida: Scorpiones)" type="journal article" year="1999">Armas, 1999: 127</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="635D21B0FFC5B15B38AFCF2DB2BA6D6C" author="Armas" box="[207,486,769,793]" etAl="et al." firstAuthor="Armas" journalOrPublisher="Avicennia 10" pageId="15" pageNumber="15" pagination="101 - 136" part="11" refString="Armas, L. F. de. 1999. Quince nuevos alacranes de La Espanola y Navassa, Antillas Mayores (Arachnida: Scorpiones). Avicennia 10 / 11: 101 - 136." title="Quince nuevos alacranes de La Espanola y Navassa, Antillas Mayores (Arachnida: Scorpiones)" type="journal article" year="1999">
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–32
</bibRefCitation>
;
<bibRefCitation id="635D21B0FFC5B15B3991CF2DB23C6D4E" author="Lourenco" editor="P. Gajdo" etAl="et al." firstAuthor="Lourenco" journalOrPublisher="Ekologia (Bratislava) 19" pageId="15" pageNumber="15" pagination="141 - 144" part="3" refString="Lourenco, W. R., D. Huber, and J. L. Cloudsley-Thompson. 2000. Description of the stridulatory organ in some species of the genus Rhopalurus Thorell (Scorpiones: Buthidae). In P. Gajdo and S. Pekar (editors), Proceedings of the 18 th European Colloquium of Arachnology, Stara Lesna, 1999. Ekologia (Bratislava) 19 (3): 141 - 144." title="Description of the stridulatory organ in some species of the genus Rhopalurus Thorell (Scorpiones: Buthidae)" type="journal article" volumeTitle="Proceedings of the 18 th European Colloquium of Arachnology, Stara Lesna, 1999" year="2000">
<treatmentCitation id="866D7A50FFC5B15B3991CF2DB2766D4E" author="Lourenco, W. R. &amp; D. Huber &amp; J. L. Cloudsley-Thompson" page="141" pageId="15" pageNumber="15" year="2000">Lourenço et al., 2000: 141</treatmentCitation>
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</bibRefCitation>
;
<treatmentCitation id="866D7A50FFC5B15B390CCF0FB12A6D4E" author="Fet, V." box="[364,630,802,827]" page="217" pageId="15" pageNumber="15" year="2000">
<bibRefCitation id="635D21B0FFC5B15B390CCF0FB12A6D4E" author="Fet and Lowe" box="[364,630,802,827]" editor="V. Fet" firstAuthor="Fet" journalOrPublisher="New York: New York Entomological Society" pageId="15" pageNumber="15" pagination="54 - 286" refString="Fet, V., and G. Lowe. 2000. Family Buthidae C. L. Koch, 1837. In V. Fet, W. D. Sissom, G., Lowe, and M. E. Braunwalder, Catalog of the scorpions of the world (1758 - 1998): 54 - 286. New York: New York Entomological Society." title="Family Buthidae C. L. Koch, 1837" type="book chapter" volumeTitle="G., Lowe, and M. E" year="2000">Fet and Lowe, 2000: 217</bibRefCitation>
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, 219–221;
<treatmentCitation id="866D7A50FFC5B15B3964CF68B2986D29" author="Armas, L. F. de" box="[260,452,836,860]" page="246" pageId="15" pageNumber="15" year="2001">
<bibRefCitation id="635D21B0FFC5B15B3964CF68B2986D29" author="Armas" box="[260,452,836,860]" editor="V. Fet" firstAuthor="Armas" journalOrPublisher="Burnham Beeches, Bucks, UK: British Arachnological Society" pageId="15" pageNumber="15" pagination="245 - 253" part="2001" refString="Armas, L. F. de. 2001. Scorpions of the Greater Antilles, with the description of a new troglobitic species (Scorpiones: Diplocentridae). In V. Fet and P. A. Selden (editors), Scorpions 2001: in memoriam Gary A. Polis: 245 - 253. Burnham Beeches, Bucks, UK: British Arachnological Society." title="Scorpions of the Greater Antilles, with the description of a new troglobitic species (Scorpiones: Diplocentridae)" type="journal article" volumeTitle="Scorpions" year="2001">Armas, 2001: 246</bibRefCitation>
</treatmentCitation>
,
<tableCitation id="4A4E69FAFFC5B15B39AFCF68B1446D29" box="[463,536,836,860]" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">table 1</tableCitation>
;
<treatmentCitation id="866D7A50FFC5B15B3A45CF69B2416D0B" author="Fet, V., B. &amp; Gantenbein., A. V. &amp; Gromov, G. &amp; Lowe &amp; W. R. Lourenco" page="2" pageId="15" pageNumber="15" year="2003">
<bibRefCitation id="635D21B0FFC5B15B3A45CF69B2586D0B" author="Fet, V., B. &amp; Gantenbein., A. V. &amp; Gromov, G. &amp; Lowe &amp; W. R. Lourenco" journalOrPublisher="Euscorpius" pageId="15" pageNumber="15" pagination="1 - 10" part="4" refString="Fet, V., B. Gantenbein., A. V. Gromov, G. Lowe, and W. R. Lourenco. 2003 a. The first molecular phylogeny of Buthidae (Scorpiones). Euscorpius 4: 1 - 10." title="The first molecular phylogeny of Buthidae (Scorpiones)" type="journal article" year="2003" yearSuffix="a">Fet et al., 2003a</bibRefCitation>
: 2
</treatmentCitation>
, 3, 6, 10,
<tableCitation id="4A4E69FAFFC5B15B39E5CF4AB2926D0B" box="[389,462,870,894]" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">table 1</tableCitation>
;
<treatmentCitation id="866D7A50FFC5B15B39BBCF4AB3986DD5" author="Teruel, R." page="149" pageId="15" pageNumber="15" year="2003">
<bibRefCitation id="635D21B0FFC5B15B39BBCF4AB3986DD5" author="Teruel" firstAuthor="Teruel" journalOrPublisher="Revista Iberica de Aracnologia" pageId="15" pageNumber="15" pagination="149 - 150" part="7" refString="Teruel, R. 2003. Registro de maxima altitud para Rhopalurus junceus (Herbst 1800) (Scorpiones: Buthidae). Revista Iberica de Aracnologia 7: 149 - 150." title="Registro de maxima altitud para Rhopalurus junceus (Herbst 1800) (Scorpiones: Buthidae)" type="journal article" year="2003">Teruel, 2003: 149</bibRefCitation>
</treatmentCitation>
, 150, figs. 1, 2; 2005: 165;
<bibRefCitation id="635D21B0FFC5B15B3981CFA4B2F66DB7" author="Teruel and Montano" firstAuthor="Teruel" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="15" pageNumber="15" pagination="219 - 228" part="37" refString="Teruel, R., and L. Montano. 2005. Los escorpiones (Arachnida: Scorpiones) del Parque Nacional &quot; Desembarco del Granma, &quot; Cuba. Boletin de la Sociedad Entomologica Aragonesa 37: 219 - 228." title="Los escorpiones (Arachnida: Scorpiones) del Parque Nacional &quot; Desembarco del Granma, &quot; Cuba" type="journal article" year="2005">
<treatmentCitation id="866D7A50FFC5B15B3981CFA4B22F6DB7" author="Teruel, R. &amp; L. Montano" page="221" pageId="15" pageNumber="15" year="2005">Teruel and Montano, 2005: 221</treatmentCitation>
–223
</bibRefCitation>
, 225–227, figs. 10, 14,
<tableCitation id="4A4E69FAFFC5B15B38A2CFE7B24A6D91" box="[194,278,971,996]" captionStart="TABLE 2" captionStartId="7.[610,690,228,252]" captionTargetBox="[235,1058,354,747]" captionText="TABLE 2 Diploid chromosome numbers (2 n) for Brazilian species of Rhopalurusinae from Ubinski et al., (2016) with corresponding genera indicated." httpUri="http://table.plazi.org/id/53B30CC9FFCDB1533A02CCC8B0866F32" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCDB1533A02CCC8B0866F32">tables 2</tableCitation>
,
<tableCitation id="4A4E69FAFFC5B15B3942CFE0B26D6D91" box="[290,305,972,996]" captionStart="TABLE 4" captionStartId="112.[610,690,228,252]" captionTargetBox="[124,1197,381,1724]" captionText="TABLE 4 Meristic data for type material of Rhopalurus ochoai, sp. n. Measurements follow Prendini (2000, 2003, 2004 a). 1 Sum of carapace, tergites I – VII, metasomal segments I – V, and telson; 2 sum of metasomal segments I – V and telson; 3 measured from base of condyle to tip of fixed finger." httpUri="http://table.plazi.org/id/53B30CC9FFBAB1243A02CCC8B7F16F11" pageId="15" pageNumber="15" tableUuid="53B30CC9FFBAB1243A02CCC8B7F16F11">4</tableCitation>
;
<treatmentCitation id="866D7A50FFC5B15B395CCFE0B2BF6D96" author="Armas, L. F. de" box="[316,483,971,996]" page="6" pageId="15" pageNumber="15" year="2006">
<bibRefCitation id="635D21B0FFC5B15B395CCFE0B2BF6D96" author="Armas" box="[316,483,971,996]" firstAuthor="Armas" journalOrPublisher="Euscorpius" pageId="15" pageNumber="15" pagination="1 - 14" part="33" refString="Armas, L. F. de. 2006. Name-bearing types of scorpions deposited at the Institute of Ecology and Systematics, Havana, Cuba (Arachnida: Scorpiones). Euscorpius 33: 1 - 14." title="Name-bearing types of scorpions deposited at the Institute of Ecology and Systematics, Havana, Cuba (Arachnida: Scorpiones)" type="journal article" year="2006">Armas, 2006: 6</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="635D21B0FFC5B15B398ECFE7B3826A70" author="Teruel" firstAuthor="Teruel" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="15" pageNumber="15" pagination="43 - 56" part="38" refString="Teruel, R. 2006. Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba. Boletin de la Sociedad Entomologica Aragonesa 38: 43 - 56." title="Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba" type="journal article" year="2006">
<treatmentCitation id="866D7A50FFC5B15B398ECFE7B3EA6A70" author="Teruel, R." page="43" pageId="15" pageNumber="15" year="2006">Teruel, 2006: 43</treatmentCitation>
–53
</bibRefCitation>
, figs. 1–10, 12A, E,
<tableCitation id="4A4E69FAFFC5B15B39D6CFC1B1546A70" box="[438,520,1005,1029]" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">tables 1</tableCitation>
,
<tableCitation id="4A4E69FAFFC5B15B3A76CFC1B1796A70" box="[534,549,1005,1029]" captionStart="TABLE 2" captionStartId="7.[610,690,228,252]" captionTargetBox="[235,1058,354,747]" captionText="TABLE 2 Diploid chromosome numbers (2 n) for Brazilian species of Rhopalurusinae from Ubinski et al., (2016) with corresponding genera indicated." httpUri="http://table.plazi.org/id/53B30CC9FFCDB1533A02CCC8B0866F32" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCDB1533A02CCC8B0866F32">2</tableCitation>
;
<bibRefCitation id="635D21B0FFC5B15B3A50CFC1B2E36A52" author="Teruel and Armas" firstAuthor="Teruel" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="15" pageNumber="15" pagination="175 - 179" part="39" refString="Teruel, R., and L. F. de. Armas. 2006. Un nuevo Rhopalurus Thorell 1876 (Scorpiones: Buthidae) de Cuba oriental. Boletin de la Sociedad Entomologica Aragonesa 39: 175 - 179." title="Un nuevo Rhopalurus Thorell 1876 (Scorpiones: Buthidae) de Cuba oriental" type="journal article" year="2006">
<treatmentCitation id="866D7A50FFC5B15B3A50CFC1B2D46A52" author="Teruel, R. &amp; L. F. de. Armas" page="175" pageId="15" pageNumber="15" year="2006">Teruel and Armas, 2006: 175</treatmentCitation>
–179
</bibRefCitation>
, figs. 1–4,
<tableCitation id="4A4E69FAFFC5B15B3A53C823B3F46A3C" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">tables 1</tableCitation>
,
<tableCitation id="4A4E69FAFFC5B15B38D5C81DB3986A3C" box="[181,196,1073,1097]" captionStart="TABLE 2" captionStartId="7.[610,690,228,252]" captionTargetBox="[235,1058,354,747]" captionText="TABLE 2 Diploid chromosome numbers (2 n) for Brazilian species of Rhopalurusinae from Ubinski et al., (2016) with corresponding genera indicated." httpUri="http://table.plazi.org/id/53B30CC9FFCDB1533A02CCC8B0866F32" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCDB1533A02CCC8B0866F32">2</tableCitation>
;
<bibRefCitation id="635D21B0FFC5B15B38B0C81DB2A16A3C" author="Teruel" box="[208,509,1073,1097]" etAl="et al." firstAuthor="Teruel" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="15" pageNumber="15" pagination="43 - 56" part="38" refString="Teruel, R. 2006. Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba. Boletin de la Sociedad Entomologica Aragonesa 38: 43 - 56." title="Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba" type="journal article" year="2006">
<treatmentCitation id="866D7A50FFC5B15B38B0C81DB29A6A3C" author="Teruel, R." box="[208,454,1073,1097]" page="219" pageId="15" pageNumber="15" year="2006">Teruel et al., 2006: 219</treatmentCitation>
–223
</bibRefCitation>
, fig. 1;
<treatmentCitation id="866D7A50FFC5B15B38FBC87FB2256A1E" author="Lourenco, W. R." box="[155,377,1107,1131]" page="359" pageId="15" pageNumber="15" year="2007">
<bibRefCitation id="635D21B0FFC5B15B38FBC87FB2256A1E" author="Lourenco" box="[155,377,1107,1131]" firstAuthor="Lourenco" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="15" pageNumber="15" pagination="359 - 365" part="40" refString="Lourenco, W. R. 2007. New considerations on the taxonomic status of the genus Physoctonus Mello-Leitao, 1934 (Scorpiones, Buthidae). Boletin de la Sociedad Entomologica Aragonesa 40: 359 - 365." title="New considerations on the taxonomic status of the genus Physoctonus Mello-Leitao, 1934 (Scorpiones, Buthidae)" type="journal article" year="2007">Lourenço, 2007: 359</bibRefCitation>
</treatmentCitation>
, 361, 362;
<treatmentCitation id="866D7A50FFC5B15B398DC87FB20A6AF8" author="Kamenz, C. &amp; L. Prendini" page="9" pageId="15" pageNumber="15" year="2008">
<bibRefCitation id="635D21B0FFC5B15B398DC87FB20A6AF8" author="Kamenz and Prendini" firstAuthor="Kamenz" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="15" pageNumber="15" pagination="1 - 359" part="316" refString="Kamenz, C., and L. Prendini. 2008. An atlas of book lung fine structure in the order Scorpiones (Arachnida). Bulletin of the American Museum of Natural History 316: 1 - 359." title="An atlas of book lung fine structure in the order Scorpiones (Arachnida)" type="journal article" year="2008">Kamenz and Prendini, 2008: 9</bibRefCitation>
</treatmentCitation>
,
<tableCitation id="4A4E69FAFFC5B15B3902C859B2F06AF8" box="[354,428,1141,1165]" captionStart="TABLE 2" captionStartId="7.[610,690,228,252]" captionTargetBox="[235,1058,354,747]" captionText="TABLE 2 Diploid chromosome numbers (2 n) for Brazilian species of Rhopalurusinae from Ubinski et al., (2016) with corresponding genera indicated." httpUri="http://table.plazi.org/id/53B30CC9FFCDB1533A02CCC8B0866F32" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCDB1533A02CCC8B0866F32">table 2</tableCitation>
, pl. 41, 43; Perez-
<treatmentCitation id="866D7A50FFC5B15B38FBC8BAB23F6ADB" author="Perez-Gelabert, D. E." box="[155,355,1174,1199]" page="68" pageId="15" pageNumber="15" year="2008">
<bibRefCitation id="635D21B0FFC5B15B38FBC8BAB2616ADA" author="Perez-Gelabert, D. E." box="[155,317,1174,1199]" journalOrPublisher="Zootaxa" pageId="15" pageNumber="15" pagination="1 - 530" part="1831" refString="Perez-Gelabert, D. E. 2008. Arthropods of Hispaniola (Dominican Republic and Haiti): A checklist and bibliography. Zootaxa 1831: 1 - 530." title="Arthropods of Hispaniola (Dominican Republic and Haiti): A checklist and bibliography" type="journal article" year="2008">Gelabert, 2008</bibRefCitation>
: 68
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC5B15B390EC8BAB3986AA5" author="Volschenk, E. S. &amp; L. Prendini" page="654" pageId="15" pageNumber="15" year="2008">
<bibRefCitation id="635D21B0FFC5B15B390EC8BAB3986AA5" author="Volschenk" etAl="et al." firstAuthor="Volschenk" journalOrPublisher="Invertebrate Systematics" pageId="15" pageNumber="15" pagination="235 - 257" part="22" refString="Volschenk, E. S., and L. Prendini. 2008. Aops oncodactylus, gen. et n. sp., the first troglobitic urodacid (Urodacidae: Scorpiones), with a re-assessment of cavernicolous, troglobitic and troglomorphic scorpions. Invertebrate Systematics 22: 235 - 257." title="Aops oncodactylus, gen. et n. sp., the first troglobitic urodacid (Urodacidae: Scorpiones), with a re-assessment of cavernicolous, troglobitic and troglomorphic scorpions" type="journal article" year="2008">Volschenk et al., 2008: 654</bibRefCitation>
</treatmentCitation>
, 658, 659, 663, 664, 674, fig. 1B,
<tableCitation id="4A4E69FAFFC5B15B3A48C894B3F46A87" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">tables 1</tableCitation>
,
<tableCitation id="4A4E69FAFFC5B15B38D5C8F6B3986A87" box="[181,196,1242,1266]" captionStart="TABLE 2" captionStartId="7.[610,690,228,252]" captionTargetBox="[235,1058,354,747]" captionText="TABLE 2 Diploid chromosome numbers (2 n) for Brazilian species of Rhopalurusinae from Ubinski et al., (2016) with corresponding genera indicated." httpUri="http://table.plazi.org/id/53B30CC9FFCDB1533A02CCC8B0866F32" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCDB1533A02CCC8B0866F32">2</tableCitation>
;
<bibRefCitation id="635D21B0FFC5B15B38B0C8F6B14A6A87" author="Prendini" box="[208,534,1242,1266]" etAl="et al." firstAuthor="Prendini" journalOrPublisher="Journal of Arachnology" pageId="15" pageNumber="15" pagination="206 - 224" part="37" refString="Prendini, L., L. A. Esposito, J. Huff, and E. S. Volschenk. 2009. Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola. Journal of Arachnology 37: 206 - 224." title="Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola" type="journal article" year="2009">
<treatmentCitation id="866D7A50FFC5B15B38B0C8F6B2BC6A87" author="Prendini, L. &amp; L. A. Esposito &amp; J. Huff &amp; E. S. Volschenk" box="[208,480,1242,1266]" page="206" pageId="15" pageNumber="15" year="2009">Prendini et al., 2009: 206</treatmentCitation>
–223
</bibRefCitation>
, figs. 1, 2, 4, 5A, B, E, F, 6A, C, 7A, C, 8, 10, 11,
<tableCitation id="4A4E69FAFFC5B15B38FBC932B3B16B43" box="[155,237,1310,1334]" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">tables 1</tableCitation>
,
<tableCitation id="4A4E69FAFFC5B15B389AC932B2546B43" box="[250,264,1310,1334]" captionStart="TABLE 3" captionStartId="69.[610,690,228,252]" captionTargetBox="[131,1171,439,1728]" captionText="TABLE 3 Meristic data for type material of Ischnotelson peruassu, sp. n., Physoctonus striatus, sp. n., and Physoctonus debilis (C. L. Koch, 1840). Measurements follow Prendini (2000, 2003, 2004). 1 Sum of carapace, tergites I – VII, metasomal segments I – V, and telson; 2 sum of metasomal segments I – V and telson; 3 measured from base of condyle to tip of fixed finger." httpUri="http://table.plazi.org/id/53B30CC9FF8FB1113A02CCC8B0CF6FEA" pageId="15" pageNumber="15" tableUuid="53B30CC9FF8FB1113A02CCC8B0CF6FEA">3</tableCitation>
;
<treatmentCitation id="866D7A50FFC5B15B3975C932B1436B40" author="Santiago-Blay, J. A." box="[277,543,1309,1334]" page="115" pageId="15" pageNumber="15" year="2009">
<bibRefCitation id="635D21B0FFC5B15B3975C932B1436B40" author="Santiago-Blay" box="[277,543,1309,1334]" firstAuthor="Santiago-Blay" journalOrPublisher="Entomological News" pageId="15" pageNumber="15" pagination="109 - 124" part="120" refString="Santiago-Blay, J. A. 2009. Systematics and some aspects of the biology of the scorpions (Arachnida) of the greater Puerto Rico region: a biosystematic synopsis. Entomological News 120 (1): 109 - 124." title="Systematics and some aspects of the biology of the scorpions (Arachnida) of the greater Puerto Rico region: a biosystematic synopsis" type="journal article" year="2009">Santiago-Blay, 2009: 115</bibRefCitation>
</treatmentCitation>
, 116, 119, 122, 125, figs. 10, 31;
<treatmentCitation id="866D7A50FFC5B15B39D9C913B2396B0C" author="Teruel, R. &amp; L. F. de Armas" page="153" pageId="15" pageNumber="15" year="2012">
<bibRefCitation id="635D21B0FFC5B15B39D9C913B2746B0C" author="Teruel and Armas" firstAuthor="Teruel" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="15" pageNumber="15" pagination="157 - 174" part="50" refString="Teruel, R., and L. F. de Armas. 2012 a. Redescripcion de Rhopalurus junceus (Herbst 1800) (Scorpiones: Buthidae). Boletin de la Sociedad Entomologica Aragonesa 50: 157 - 174." title="Redescripcion de Rhopalurus junceus (Herbst 1800) (Scorpiones: Buthidae)" type="journal article" year="2012" yearSuffix="a">
Teruel and
<bibRefCitation id="635D21B0FFC5B15B38FBC94DB2746B0C" author="Teruel, R. &amp; L. F. de Armas" box="[155,296,1377,1401]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="15" pageNumber="15" pagination="157 - 174" part="50" refString="Teruel, R., and L. F. de Armas. 2012 a. Redescripcion de Rhopalurus junceus (Herbst 1800) (Scorpiones: Buthidae). Boletin de la Sociedad Entomologica Aragonesa 50: 157 - 174." title="Redescripcion de Rhopalurus junceus (Herbst 1800) (Scorpiones: Buthidae)" type="journal article" year="2012" yearSuffix="a">Armas, 2012</bibRefCitation>
</bibRefCitation>
a: 153
</treatmentCitation>
–167, figs. 1–16,
<tableCitation id="4A4E69FAFFC5B15B3A7CC94DB3F96BEE" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">tables 1</tableCitation>
–7; 2012b: 209, 210, 212, 214–217, figs. 1–7,
<tableCitation id="4A4E69FAFFC5B15B38B0C989B2636BC8" box="[208,319,1445,1469]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="6.[609,689,226,250]" captionStartId-1="7.[610,690,228,252]" captionStartId-2="69.[610,690,228,252]" captionTargetBox-0="[115,1198,353,998]" captionTargetBox-1="[235,1058,354,747]" captionTargetBox-2="[131,1171,439,1728]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." captionText-1="TABLE 2 Diploid chromosome numbers (2 n) for Brazilian species of Rhopalurusinae from Ubinski et al., (2016) with corresponding genera indicated." captionText-2="TABLE 3 Meristic data for type material of Ischnotelson peruassu, sp. n., Physoctonus striatus, sp. n., and Physoctonus debilis (C. L. Koch, 1840). Measurements follow Prendini (2000, 2003, 2004). 1 Sum of carapace, tergites I – VII, metasomal segments I – V, and telson; 2 sum of metasomal segments I – V and telson; 3 measured from base of condyle to tip of fixed finger." httpUri-0="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" httpUri-1="http://table.plazi.org/id/53B30CC9FFCDB1533A02CCC8B0866F32" httpUri-2="http://table.plazi.org/id/53B30CC9FF8FB1113A02CCC8B0CF6FEA" pageId="15" pageNumber="15" tableUuid-0="53B30CC9FFCCB1523A01CCCEB0B66F30" tableUuid-1="53B30CC9FFCDB1533A02CCC8B0866F32" tableUuid-2="53B30CC9FF8FB1113A02CCC8B0CF6FEA">tables 1–3</tableCitation>
;
<bibRefCitation id="635D21B0FFC5B15B392BC989B3A46BAA" author="Teruel and Kovarik" bookContentInfo="229 pp." firstAuthor="Teruel" journalOrPublisher="Prague, Czech Republic: Jakub Rolcik - Clarion Productions" pageId="15" pageNumber="15" refString="Teruel, R., and F. Kovarik. 2012. Scorpions of Cuba. Prague, Czech Republic: Jakub Rolcik - Clarion Productions. 229 pp." title="Scorpions of Cuba" type="book" year="2012">
<treatmentCitation id="866D7A50FFC5B15B392BC989B39F6BAA" author="Teruel, R. &amp; F. Kovarik" page="116" pageId="15" pageNumber="15" year="2012">Teruel and Kovařík, 2012: 116</treatmentCitation>
–141
</bibRefCitation>
, figs. 29, 40, 41, 48, 253–305, 534– 565, 636;
<treatmentCitation id="866D7A50FFC5B15B3963C9C4B2596857" author="Rodriguez-Cabrera, T. M. &amp; R. Teruel" page="121" pageId="15" pageNumber="15" year="2014">
<bibRefCitation id="635D21B0FFC5B15B3963C9C4B2596857" author="Rodriguez-Cabrera and Teruel" firstAuthor="Rodriguez-Cabrera" journalOrPublisher="Revista Iberica de Aracnologia" pageId="15" pageNumber="15" pagination="119 - 122" part="24" refString="Rodriguez-Cabrera, T. M., and R. Teruel. 2014. On the highest altitudinal occurrences of scorpions in Cuba (Arachnida: Scorpiones). Revista Iberica de Aracnologia 24: 119 - 122." title="On the highest altitudinal occurrences of scorpions in Cuba (Arachnida: Scorpiones)" type="journal article" year="2014">Rodríguez-Cabrera and Teruel, 2014: 121</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC5B15B3970CA27B1336857" author="Lourenco, W. R. &amp; L. F. de Armas" box="[272,623,1546,1571]" page="228" pageId="15" pageNumber="15" year="2015">
<bibRefCitation id="635D21B0FFC5B15B3970CA27B1336857" author="Lourenco and Armas" box="[272,623,1546,1571]" firstAuthor="Lourenco" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" pageId="15" pageNumber="15" pagination="225 - 232" part="17" refString="Lourenco, W. R., and L. F. de Armas. 2015. New records of scorpions from Haiti (Scorpiones: Buthidae, Diplocentridae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 17 (194): 225 - 232." title="New records of scorpions from Haiti (Scorpiones: Buthidae, Diplocentridae)" type="journal article" year="2015">Lourenço and Armas, 2015: 228</bibRefCitation>
</treatmentCitation>
, 229;
<treatmentCitation id="866D7A50FFC5B15B38AFCA00B1196831" author="Rodriguez-Cabrera, T. M. &amp; C. A. Martinez-Munoz &amp; R. Teruel" box="[207,581,1580,1604]" page="85" pageId="15" pageNumber="15" year="2015">
<bibRefCitation id="635D21B0FFC5B15B38AFCA00B1196831" author="Rodriguez-Cabrera" box="[207,581,1580,1604]" etAl="et al." firstAuthor="Rodriguez-Cabrera" journalOrPublisher="Revista Iberica de Aracnologia" pageId="15" pageNumber="15" pagination="85 - 86" part="26" refString="Rodriguez-Cabrera, T. M., C. A. Martinez-Munoz, and R. Teruel. 2015. Predation by the scorpion Rhopalurus junceus (Scorpiones: Buthidae) on the centipede Scolopocryptops ferrugineus (Scolopendromorpha: Scolopocryptopidae). Revista Iberica de Aracnologia 26: 85 - 86." title="Predation by the scorpion Rhopalurus junceus (Scorpiones: Buthidae) on the centipede Scolopocryptops ferrugineus (Scolopendromorpha: Scolopocryptopidae)" type="journal article" year="2015">Rodríguez-Cabrera et al., 2015: 85</bibRefCitation>
</treatmentCitation>
, 86, fig. 1;
<treatmentCitation id="866D7A50FFC5B15B38BECA62B2E16813" author="Santos, G. de" box="[222,445,1614,1638]" page="3" pageId="15" pageNumber="15" year="2016">
<bibRefCitation id="635D21B0FFC5B15B38BECA62B2E16813" author="Santos" box="[222,445,1614,1638]" etAl="et al." firstAuthor="Santos" journalOrPublisher="Novitates Caribaea" pageId="15" pageNumber="15" pagination="1 - 22" part="10" refString="Santos, G. de los, L. F. de Armas, and R. Teruel. 2016. Lista anotada de los escorpiones (Arachnida: Scorpiones) de la Espanola (Republica Dominicana y Haiti). Novitates Caribaea 10: 1 - 22." title="Lista anotada de los escorpiones (Arachnida: Scorpiones) de la Espanola (Republica Dominicana y Haiti)" type="journal article" year="2016">Santos et al., 2016: 3</bibRefCitation>
</treatmentCitation>
, 9,
<tableCitation id="4A4E69FAFFC5B15B3984CA62B16A6813" box="[484,566,1614,1638]" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="15" pageNumber="15" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">tables 1</tableCitation>
, figs. 1Ñ, 2B, 3D.
</treatmentCitationGroup>
</paragraph>
<paragraph id="07735C41FFC5B15B380BCABFB0166F17" blockId="15.[107,634,228,1741]" lastBlockId="15.[684,1212,227,492]" pageId="15" pageNumber="15">
<treatmentCitationGroup id="27DC7B6FFFC5B15B380BCABFB0166F17" pageId="15" pageNumber="15">
<taxonomicName id="C0CC27C2FFC5B15B380BCABFB38A68DF" authorityName="C. L. Koch" authorityYear="1838" box="[107,214,1683,1706]" class="Arachnida" family="Buthidae" genus="Centrurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B380BCABFB38A68DF" box="[107,214,1683,1706]" italics="true" pageId="15" pageNumber="15">Centrurus</emphasis>
</taxonomicName>
(part):
<treatmentCitation id="866D7A50FFC5B15B3948CABEB2AA68DF" author="Karsch, F." box="[296,502,1682,1706]" page="121" pageId="15" pageNumber="15" year="1879">
<bibRefCitation id="635D21B0FFC5B15B3948CABEB29F68DF" author="Karsch, F." box="[296,451,1682,1706]" journalOrPublisher="Mitteilungen des Munchener Entomologischen Vereins" pageId="15" pageNumber="15" pagination="97 - 136" part="3" refString="Karsch, F. 1879 b. Skorpionologische Beitrage. Part II. Mitteilungen des Munchener Entomologischen Vereins 3 (2): 97 - 136." title="Skorpionologische Beitrage. Part II" type="journal article" year="1879" yearSuffix="b">Karsch, 1879b</bibRefCitation>
: 121
</treatmentCitation>
, 122;
<treatmentCitation id="866D7A50FFC5B15B3A55CABFB21A68B8" author="Kraepelin, K." page="123" pageId="15" pageNumber="15" year="1891">
<bibRefCitation id="635D21B0FFC5B15B3A55CABFB21A68B8" author="Kraepelin" firstAuthor="Kraepelin" journalOrPublisher="Jahrbuch des Hamburgische Wissenschaftlichen Anstalten" pageId="15" pageNumber="15" pagination="1 - 144" part="8" refString="Kraepelin, K. 1891. Revision der Skorpione. I. Die Familie Androctonidae. Jahrbuch des Hamburgische Wissenschaftlichen Anstalten 8: 1 - 144." title="Revision der Skorpione. I. Die Familie Androctonidae" type="journal article" year="1891">Kraepelin, 1891: 123</bibRefCitation>
</treatmentCitation>
, 135–137, 139, figs. 30, 33;
<treatmentCitation id="866D7A50FFC5B15B3ABCCCC8B0C36E8E" author="Pocock, R. I." box="[732,927,227,252]" page="385" pageId="15" pageNumber="15" year="1893">
<bibRefCitation id="635D21B0FFC5B15B3ABCCCC8B0C36E8E" author="Pocock" box="[732,927,227,252]" firstAuthor="Pocock" journalOrPublisher="Journal of the Linnaean Society" pageId="15" pageNumber="15" pagination="374 - 409" part="24" refString="Pocock, R. I. 1893. Contribution to our knowledge of the arthropod fauna of the West Indies. Part I. Scorpiones and Pedipalpi, with a supplementary note upon freshwater Decapoda of St. Vincent. Journal of the Linnaean Society 24: 374 - 409." title="Contribution to our knowledge of the arthropod fauna of the West Indies. Part I. Scorpiones and Pedipalpi, with a supplementary note upon freshwater Decapoda of St. Vincent" type="journal article" year="1893">Pocock, 1893: 385</bibRefCitation>
</treatmentCitation>
, 391;
<treatmentCitation id="866D7A50FFC5B15B3BBFCCC8B7FE6E89" author="Thorell, T." box="[991,1186,228,252]" page="372" pageId="15" pageNumber="15" year="1893">
<bibRefCitation id="635D21B0FFC5B15B3BBFCCC8B7FE6E89" author="Thorell" box="[991,1186,228,252]" firstAuthor="Thorell" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="15" pageNumber="15" pagination="356 - 387" part="25" refString="Thorell, T. 1893. Scorpiones exotici R. Musei Historiea Naturalis Florentini. Bollettino della Societa Entomologica Italiana 25 (4): 356 - 387." title="Scorpiones exotici R. Musei Historiea Naturalis Florentini" type="journal article" year="1893">Thorell, 1893: 372</bibRefCitation>
</treatmentCitation>
, 373;
<treatmentCitation id="866D7A50FFC5B15B3B70CD2AB0BD6F6B" author="Kraepelin, K." box="[784,993,262,286]" page="95" pageId="15" pageNumber="15" year="1895">
<bibRefCitation id="635D21B0FFC5B15B3B70CD2AB0BD6F6B" author="Kraepelin" box="[784,993,262,286]" firstAuthor="Kraepelin" journalOrPublisher="Jahrbuch des Hamburgische Wissenschaftlichen Anstalten" pageId="15" pageNumber="15" pagination="75 - 96" part="12" refString="Kraepelin, K. 1895. Nachtrag zu Theil 1 der Revision der Skorpione. Jahrbuch des Hamburgische Wissenschaftlichen Anstalten 12: 75 - 96." title="Nachtrag zu Theil 1 der Revision der Skorpione" type="journal article" year="1895">Kraepelin, 1895: 95</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="635D21B0FFC5B15B3B8DCD2AB7156F6B" author="Kraepelin" bookContentInfo="265 pp." box="[1005,1097,262,286]" editor="F. Dahl" firstAuthor="Kraepelin" journalOrPublisher="Friedlander and Sohn Verlag" pageId="15" pageNumber="15" refId="ref66921" refString="Kraepelin, K. 1899. Scorpiones und Pedipalpi. In F. Dahl (editor), Das Tierreich. Berlin: R. Friedlander and Sohn Verlag. 265 pp." title="Scorpiones und Pedipalpi" type="book" volumeTitle="Das Tierreich. Berlin: R" year="1899">1899: 89</bibRefCitation>
, 94, 95;
<bibRefCitation id="635D21B0FFC5B15B3ABCCD04B01A6F35" author="Kraepelin" box="[732,838,296,320]" firstAuthor="Kraepelin" journalOrPublisher="Bulletin du Museum National d'Histoire Naturelle, Paris" pageId="15" pageNumber="15" pagination="265 - 274" part="7" refId="ref66954" refString="Kraepelin, K. 1901. Catalogue des scorpions des collections du Museum d'histoire naturelle de Paris. Bulletin du Museum National d'Histoire Naturelle, Paris 7 (6): 265 - 274." title="Catalogue des scorpions des collections du Museum d'histoire naturelle de Paris" type="journal article" year="1901">1901: 270</bibRefCitation>
;
<bibRefCitation id="635D21B0FFC5B15B3B30CD04B0E56F35" author="Kraepelin" box="[848,953,296,320]" firstAuthor="Kraepelin" journalOrPublisher="Jahrbuch des Hamburgische Wissenschaftlichen Anstalten" pageId="15" pageNumber="15" pagination="181 - 203" part="25" refId="ref66989" refString="Kraepelin, K. 1908. Die sekundaren Geschlechtscharaktere der Skorpione, Pedipalpen und Solifugen. I. Skorpione. Jahrbuch des Hamburgische Wissenschaftlichen Anstalten 25 (2): 181 - 203." title="Die sekundaren Geschlechtscharaktere der Skorpione, Pedipalpen und Solifugen. I. Skorpione" type="journal article" year="1908">1908: 187</bibRefCitation>
, 190, 193, 194;
<treatmentCitation id="866D7A50FFC5B15B3C00CD05B01A6F17" author="Werner, F." page="357" pageId="15" pageNumber="15" year="1927">
<bibRefCitation id="635D21B0FFC5B15B3C00CD05B01A6F17" author="Werner" firstAuthor="Werner" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="15" pageNumber="15" pagination="355 - 358" part="40" refString="Werner, F. 1927. Uber einige Skorpione aus Brasilien. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 40 (3): 355 - 358." title="Uber einige Skorpione aus Brasilien" type="journal article" year="1927">Werner, 1927: 357</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="07735C41FFC5B15B3ACCCD42B73B6FD2" blockId="15.[684,1212,227,492]" pageId="15" pageNumber="15">
<treatmentCitationGroup id="27DC7B6FFFC5B15B3ACCCD42B73B6FD2" pageId="15" pageNumber="15">
<treatmentCitation id="866D7A50FFC5B15B3ACCCD42B0A26FF0" author="Pocock, R. I." box="[684,1022,365,389]" page="375" pageId="15" pageNumber="15" year="1893">
<taxonomicName id="C0CC27C2FFC5B15B3ACCCD42B0A26FF0" ID-CoL="4WDH" authority="Pocock, 1893: 375" authorityName="Pocock" authorityPageNumber="375" authorityYear="1893" box="[684,1022,365,389]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3ACCCD42B06F6FF0" box="[684,819,366,389]" italics="true" pageId="15" pageNumber="15">Heteroctenus</emphasis>
<bibRefCitation id="635D21B0FFC5B15B3B5BCD41B0A26FF0" author="Pocock" box="[827,1022,365,389]" firstAuthor="Pocock" journalOrPublisher="Journal of the Linnaean Society" pageId="15" pageNumber="15" pagination="374 - 409" part="24" refString="Pocock, R. I. 1893. Contribution to our knowledge of the arthropod fauna of the West Indies. Part I. Scorpiones and Pedipalpi, with a supplementary note upon freshwater Decapoda of St. Vincent. Journal of the Linnaean Society 24: 374 - 409." title="Contribution to our knowledge of the arthropod fauna of the West Indies. Part I. Scorpiones and Pedipalpi, with a supplementary note upon freshwater Decapoda of St. Vincent" type="journal article" year="1893">Pocock, 1893: 375</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 391, 392;
<treatmentCitation id="866D7A50FFC5B15B3C11CD41B0196FD2" author="Laurie, M." page="131" pageId="15" pageNumber="15" year="1896">
<bibRefCitation id="635D21B0FFC5B15B3C11CD41B0196FD2" author="Laurie" firstAuthor="Laurie" journalOrPublisher="Annals and Magazine of Natural History" pageId="15" pageNumber="15" pagination="121 - 133" part="6" refString="Laurie, M. 1896. Further notes on the anatomy and development of scorpions, and their bearing on the classification of the order. Annals and Magazine of Natural History (6) 18 (104): 121 - 133." title="Further notes on the anatomy and development of scorpions, and their bearing on the classification of the order" type="journal article" year="1896">Laurie, 1896: 131</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="866D7A50FFC5B15B3B30CDA3B7726FD2" author="Lonnberg, E." box="[848,1070,399,423]" page="197" pageId="15" pageNumber="15" year="1897">
<bibRefCitation id="635D21B0FFC5B15B3B30CDA3B7726FD2" author="Lonnberg" box="[848,1070,399,423]" firstAuthor="Lonnberg" journalOrPublisher="Entomologisk Tidskrift" pageId="15" pageNumber="15" pagination="193 - 211" part="18" refString="Lonnberg, E. 1897. Om skorpionernas och pedipalpernas geografiska utbredning. Entomologisk Tidskrift 18 (1 - 4): 193 - 211." title="Om skorpionernas och pedipalpernas geografiska utbredning" type="journal article" year="1897">Lönnberg, 1897: 197</bibRefCitation>
</treatmentCitation>
, 208.
</treatmentCitationGroup>
</paragraph>
<paragraph id="07735C41FFC5B15B3ACCCD9EB0396F99" blockId="15.[684,1212,227,492]" pageId="15" pageNumber="15">
<treatmentCitationGroup id="27DC7B6FFFC5B15B3ACCCD9EB0396F99" pageId="15" pageNumber="15">
<taxonomicName id="C0CC27C2FFC5B15B3ACCCD9EB0996FBF" ID-CoL="3KC2" authority="(Rhopalurus)" authorityName="Marx" authorityYear="1890" baseAuthorityName="Rhopalurus" box="[684,965,434,458]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3ACCCD9EB06F6FBF" box="[684,819,434,458]" italics="true" pageId="15" pageNumber="15">Centruroides</emphasis>
(
<emphasis id="35B88053FFC5B15B3B23CD9EB0E06FBF" box="[835,956,434,458]" italics="true" pageId="15" pageNumber="15">Rhopalurus</emphasis>
)
</taxonomicName>
(part):
<treatmentCitation id="866D7A50FFC5B15B3C78CD9EB0596F99" author="Werner, F." page="274" pageId="15" pageNumber="15" year="1934">
<bibRefCitation id="635D21B0FFC5B15B3C78CD9EB0596F99" author="Werner" editor="H. G. Bronns Klassen" firstAuthor="Werner" journalOrPublisher="Lief. 1" pageId="15" pageNumber="15" pagination="1 - 316" part="2" refString="Werner, F. 1934. Scorpiones, Pedipalpi. In H. G. Bronns Klassen und Ordnungen des Tierreichs: 5, IV, 8, Lief. 1, 2 (Scorpiones): 1 - 316. Leipzig: Akademische Verlaggesellschaft." title="Scorpiones, Pedipalpi" type="journal article" volumeTitle="und Ordnungen des Tierreichs: 5, IV, 8" year="1934">Werner, 1934: 274</bibRefCitation>
</treatmentCitation>
, fig. 33b.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFC5B1473AACCE24B2636B4E" lastPageId="19" lastPageNumber="19" pageId="15" pageNumber="15" type="diagnosis">
<paragraph id="07735C41FFC5B1473AACCE24B2636B4E" blockId="15.[684,1214,520,1741]" lastBlockId="19.[107,637,1282,1737]" lastPageId="19" lastPageNumber="19" pageId="15" pageNumber="15">
DIAGNOSIS:
<emphasis id="35B88053FFC5B15B3B2FCE25B0856C55" box="[847,985,521,544]" italics="true" pageId="15" pageNumber="15">Heteroctonus</emphasis>
differs all from other rhopalurusine genera by the presence of a single, deep, median depression in the male pectinal plate. It differs further from
<taxonomicName id="C0CC27C2FFC5B15B3B9ACE40B7226CF1" authority="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[1018,1150,620,644]" class="Arachnida" family="Buthidae" genus="Ischnotelson" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="35B88053FFC5B15B3B9ACE40B7226CF1" box="[1018,1150,620,644]" italics="true" pageId="15" pageNumber="15">Ischnotelson</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFC5B15B3CF0CE40B1876CD1" pageId="15" pageNumber="15" rank="genus">gen. nov.</taxonomicNameLabel>
,
<taxonomicName id="C0CC27C2FFC5B15B3A85CEA1B06B6CD1" authority="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[741,823,653,676]" class="Arachnida" family="Buthidae" genus="Jaguajir" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="35B88053FFC5B15B3A85CEA1B06B6CD1" box="[741,823,653,676]" italics="true" pageId="15" pageNumber="15">Jaguajir</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFC5B15B3B24CEA1B0F96CD1" box="[836,933,653,677]" pageId="15" pageNumber="15" rank="genus">gen. nov.</taxonomicNameLabel>
,
<taxonomicName id="C0CC27C2FFC5B15B3BCFCEA1B76C6CD0" authorityName="Mello-Leitao" authorityYear="1934" box="[943,1072,653,677]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3BCFCEA1B76C6CD0" box="[943,1072,653,677]" italics="true" pageId="15" pageNumber="15">Physoctonus</emphasis>
</taxonomicName>
,
<taxonomicName id="C0CC27C2FFC5B15B3C5DCEA1B7EB6CD0" authorityName="Thorell" authorityYear="1876" box="[1085,1207,653,677]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3C5DCEA1B7EB6CD0" box="[1085,1207,653,677]" italics="true" pageId="15" pageNumber="15">Rhopalurus</emphasis>
</taxonomicName>
,
<taxonomicName id="C0CC27C2FFC5B15B3ACCCE82B03E6CB3" box="[684,866,686,710]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3ACCCE82B03E6CB3" box="[684,866,686,710]" italics="true" pageId="15" pageNumber="15">Troglorhopalurus</emphasis>
</taxonomicName>
, and many species of
<taxonomicName id="C0CC27C2FFC5B15B3C2DCE82B1916C92" authorityName="Marx" authorityYear="1890" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3C2DCE82B1916C92" italics="true" pageId="15" pageNumber="15">Centruroides</emphasis>
</taxonomicName>
by the absence of a subaculear tubercle on the telson; from
<taxonomicName id="C0CC27C2FFC5B15B3B55CEDCB7176D7D" authority=", Ischnotelson" authorityName="Ischnotelson" box="[821,1099,752,776]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3B55CEDCB0E26D7D" box="[821,958,752,776]" italics="true" pageId="15" pageNumber="15">Centruroides</emphasis>
,
<emphasis id="35B88053FFC5B15B3BAACEDCB7176D7D" box="[970,1099,752,776]" italics="true" pageId="15" pageNumber="15">Ischnotelson</emphasis>
</taxonomicName>
,
<taxonomicName id="C0CC27C2FFC5B15B3C37CEDCB18E6D5F" authorityName="Mello-Leitao" authorityYear="1934" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3C37CEDCB18E6D5F" italics="true" pageId="15" pageNumber="15">Physoctonus</emphasis>
</taxonomicName>
, and
<taxonomicName id="C0CC27C2FFC5B15B3B75CF3EB0936D5F" box="[789,975,786,810]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3B75CF3EB0936D5F" box="[789,975,786,810]" italics="true" pageId="15" pageNumber="15">Troglorhopalurus</emphasis>
</taxonomicName>
by the presence of a pecten-sternite stridulatory organ (proximal pectinal teeth often enlarged, dorsal surfaces with multiple nodules and regular striations, sternite III, lateral margins forming smooth, raised carina, ventromedian carina elevated anteriorly, ventrosubmedian surfaces forming paired depressions, finely and irregularly granular); from
<taxonomicName id="C0CC27C2FFC5B15B3A8BC830B0316A41" authority="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[747,877,1052,1076]" class="Arachnida" family="Buthidae" genus="Ischnotelson" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="35B88053FFC5B15B3A8BC830B0316A41" box="[747,877,1052,1076]" italics="true" pageId="15" pageNumber="15">Ischnotelson</emphasis>
</taxonomicName>
and
<taxonomicName id="C0CC27C2FFC5B15B3BC9C830B7786A41" authorityName="Thorell" authorityYear="1876" box="[937,1060,1052,1076]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3BC9C830B7786A41" box="[937,1060,1052,1076]" italics="true" pageId="15" pageNumber="15">Rhopalurus</emphasis>
</taxonomicName>
, by the separate (unfused) central lateral and posterior central submedian carinae of the carapace; from
<taxonomicName id="C0CC27C2FFC5B15B3ACCC853B06B6AE2" box="[684,823,1151,1175]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3ACCC853B06B6AE2" box="[684,823,1151,1175]" italics="true" pageId="15" pageNumber="15">Centruroides</emphasis>
</taxonomicName>
by the presence of macrosetae on the dorsobasal surface of the pectinal teeth; from
<taxonomicName id="C0CC27C2FFC5B15B3ACCC8EEB06D6AAF" authority="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[684,817,1218,1242]" class="Arachnida" family="Buthidae" genus="Ischnotelson" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="35B88053FFC5B15B3ACCC8EEB06D6AAF" box="[684,817,1218,1242]" italics="true" pageId="15" pageNumber="15">Ischnotelson</emphasis>
</taxonomicName>
, by the separate (unfused) lateral ocular and central lateral carinae of the carapace, and the telson vesicle not being laterally compressed; from
<taxonomicName id="C0CC27C2FFC5B15B3B29C90AB0C06B48" authority="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[841,924,1318,1341]" class="Arachnida" family="Buthidae" genus="Jaguajir" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="35B88053FFC5B15B3B29C90AB0C06B48" box="[841,924,1318,1341]" italics="true" pageId="15" pageNumber="15">Jaguajir</emphasis>
</taxonomicName>
by the separate (unfused) lateral ocular and anterior central submedian carinae of the carapace; from
<taxonomicName id="C0CC27C2FFC5B15B3B90C944B72F6BF5" authorityName="Mello-Leitao" authorityYear="1934" box="[1008,1139,1384,1408]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3B90C944B72F6BF5" box="[1008,1139,1384,1408]" italics="true" pageId="15" pageNumber="15">Physoctonus</emphasis>
</taxonomicName>
by the larger size (30–70 mm), the more distinct carapacial carinae, the setose proximal dorsal fulcra of the pectines, the incrassate pedipalp chela manus of the adult male, the bifurcate prolateral pedal spur of leg I, and the oblique subrows of primary denticles on the pedipalp chela fingers flanked closely by pro- and retrolateral accessory (supernumerary) denticles; and from
<taxonomicName id="C0CC27C2FFC5B15B3C2DCA5EB1A168DE" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFC5B15B3C2DCA5EB1A168DE" italics="true" pageId="15" pageNumber="15">Troglorhopalurus</emphasis>
</taxonomicName>
by the proximal dentate margin of the chela fixed and movable fingers of the adult male emarginate, with a distinct gap evident between them, when closed.
</paragraph>
</subSubSection>
<caption id="53B30CC9FFDAB144380BC9C1B09868A4" httpUri="https://zenodo.org/record/851384/files/figure.png" pageId="16" pageNumber="16" targetBox="[188,1135,224,1487]" targetPageId="16">
<paragraph id="07735C41FFDAB144380BC9C1B09868A4" blockId="16.[107,1214,1516,1745]" pageId="16" pageNumber="16">
FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae
<bibRefCitation id="635D21B0FFDAB1443B81C9C0B7366871" author="Bucherl" box="[993,1130,1516,1540]" editor="W. Bucherl &amp; E. R. Buckley" firstAuthor="Bucherl" journalOrPublisher="New York: Academic Press" pageId="16" pageNumber="16" pagination="317 - 348" part="3" refId="ref64808" refString="Bucherl, W. 1971. Classification, biology and venom extraction of scorpions. In W. Bucherl and E. R. Buckley (editors), Venomous animals and their venoms 3: 317 - 348. New York: Academic Press." title="Classification, biology and venom extraction of scorpions" type="journal article" volumeTitle="Venomous animals and their venoms" year="1971">Bücherl, 1971</bibRefCitation>
, habitus in life.
<emphasis id="35B88053FFDAB14438D1CA25B3956854" bold="true" box="[177,201,1545,1569]" pageId="16" pageNumber="16">A.</emphasis>
<taxonomicName id="C0CC27C2FFDAB14438B0CA26B0516854" authority="(Armas and Marcano Fondeur, 1987)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas and Marcano Fondeur" baseAuthorityYear="1987" box="[208,781,1545,1569]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="abudi" status="comb. nov.">
<emphasis id="35B88053FFDAB14438B0CA26B2D06854" box="[208,396,1545,1569]" italics="true" pageId="16" pageNumber="16">Heteroctenus abudi</emphasis>
(
<bibRefCitation id="635D21B0FFDAB14439FBCA25B05A6854" author="Armas and Marcano Fondeur" box="[411,774,1545,1569]" firstAuthor="Armas" journalOrPublisher="Poeyana" pageId="16" pageNumber="16" pagination="1 - 24" part="356" refId="ref64021" refString="Armas, L. F. de, and E. J. Marcano Fondeur. 1987. Nuevos escorpiones (Arachnida: Scorpiones) de Republica Dominicana. Poeyana 356: 1 - 24." title="Nuevos escorpiones (Arachnida: Scorpiones) de Republica Dominicana" type="journal article" year="1987">Armas and Marcano Fondeur, 1987</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFDAB1443B78CA25B0DB6855" box="[792,903,1545,1569]" pageId="16" pageNumber="16">comb. nov.</taxonomicNameLabel>
, ♂, Isla Mona, Puerto
<taxonomicName id="C0CC27C2FFDAB1443C0ECA25B1D0684B" authority="(Lenarducci et al., 2005)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lenarducci et al." baseAuthorityYear="2005" pageId="16" pageNumber="16" rank="subSpecies" status="comb. nov." subSpecies="guanambiensis">
Rico.
<emphasis id="35B88053FFDAB1443CC7CA25B7E16854" bold="true" box="[1191,1213,1545,1569]" pageId="16" pageNumber="16">B.</emphasis>
<emphasis id="35B88053FFDAB144380BCA0AB2DC684B" box="[107,384,1574,1598]" italics="true" pageId="16" pageNumber="16">Ischnotelson guanambiensis</emphasis>
(
<bibRefCitation id="635D21B0FFDAB14439F1CA0AB1D9684B" author="Lenarducci" box="[401,645,1574,1598]" etAl="et al." firstAuthor="Lenarducci" journalOrPublisher="Biota Neotropica" pageId="16" pageNumber="16" pagination="173 - 180" part="5" refId="ref67356" refString="Lenarducci, A. R. I., R. Pinto-da-Rocha, and S. M. Lucas. 2005. Descricao de uma nova especie de Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) do nordeste brasileiro. Biota Neotropica 5 (1 A): 173 - 180." title="Descricao de uma nova especie de Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) do nordeste brasileiro" type="journal article" year="2005">Lenarducci et al., 2005</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFDAB1443AFACA0AB052684B" box="[666,782,1574,1598]" pageId="16" pageNumber="16">comb. nov.</taxonomicNameLabel>
, ♂, Guanambí, Bahía, Brazil.
<taxonomicName id="C0CC27C2FFDAB1443C2ECA0AB2C76829" authority="(C. L. Koch, 1839)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="C. L. Koch" baseAuthorityYear="1839" pageId="16" pageNumber="16" rank="subSpecies" status="comb. nov." subSpecies="agamemnon">
<emphasis id="35B88053FFDAB1443C2ECA0AB73A684B" bold="true" box="[1102,1126,1574,1598]" pageId="16" pageNumber="16">C.</emphasis>
<emphasis id="35B88053FFDAB1443C10CA0BB3BF682E" italics="true" pageId="16" pageNumber="16">Jaguajir agamemnon</emphasis>
(C.L.
<bibRefCitation id="635D21B0FFDAB1443941CA68B2C86829" author="Koch" box="[289,404,1604,1628]" firstAuthor="Koch" journalOrPublisher="Nurnberg: C. H. Zeh'sche Buchhandlung" pageId="16" pageNumber="16" pagination="1 - 156" part="6" refId="ref66709" refString="Koch, C. L. 1839. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung 6 (1 - 6): 1 - 156." title="Die Arachniden" type="journal article" year="1839">Koch, 1839</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFDAB14439C5CA68B14F682E" box="[421,531,1604,1628]" pageId="16" pageNumber="16">comb. nov.</taxonomicNameLabel>
, ♀, Parque Zoobotânico de Teresina, Piauí, Brazil.
<taxonomicName id="C0CC27C2FFDAB1443C6DCA68B3A7680C" authority="(Borelli, 1910)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Borelli" baseAuthorityYear="1910" pageId="16" pageNumber="16" rank="subSpecies" status="comb. nov." subSpecies="rochae">
<emphasis id="35B88053FFDAB1443C6DCA68B7796829" bold="true" box="[1037,1061,1604,1628]" pageId="16" pageNumber="16">D.</emphasis>
<emphasis id="35B88053FFDAB1443C4BCA68B7E16829" box="[1067,1213,1604,1628]" italics="true" pageId="16" pageNumber="16">Jaguajir rochae</emphasis>
(
<bibRefCitation id="635D21B0FFDAB1443811CA4DB3A9680C" author="Borelli" box="[113,245,1633,1657]" firstAuthor="Borelli" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Realle Universita di Torino" pageId="16" pageNumber="16" pagination="1 - 8" part="25" refId="ref64441" refString="Borelli, A. 1910. Scorpioni nuovi e poco noti del Brasile. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Realle Universita di Torino 25 (629): 1 - 8." title="Scorpioni nuovi e poco noti del Brasile" type="journal article" year="1910">Borelli, 1910</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFDAB1443967CA4DB22A680D" box="[263,374,1633,1657]" pageId="16" pageNumber="16">comb. nov.</taxonomicNameLabel>
, ♂, Jeremoabo, Bahía, Brazil.
<emphasis id="35B88053FFDAB1443AC5CA4DB1E5680C" bold="true" box="[677,697,1633,1657]" pageId="16" pageNumber="16">E.</emphasis>
<taxonomicName id="C0CC27C2FFDAB1443AA0CA4DB020680C" baseAuthorityName="C. L. Koch" baseAuthorityYear="1840" box="[704,892,1633,1657]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="35B88053FFDAB1443AA0CA4DB020680C" box="[704,892,1633,1657]" italics="true" pageId="16" pageNumber="16">Physoctonus debilis</emphasis>
</taxonomicName>
, ♀, Guanambí, Bahía, Brazil.
<emphasis id="35B88053FFDAB1443CCCCA4DB7E1680C" bold="true" box="[1196,1213,1633,1657]" pageId="16" pageNumber="16">F.</emphasis>
<taxonomicName id="C0CC27C2FFDAB144380BCA52B29468E3" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[107,456,1662,1686]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="laticauda">
<emphasis id="35B88053FFDAB144380BCA52B26768E3" box="[107,315,1662,1686]" italics="true" pageId="16" pageNumber="16">Rhopalurus laticauda</emphasis>
<bibRefCitation id="635D21B0FFDAB1443922CA52B29468E3" author="Thorell" box="[322,456,1662,1686]" firstAuthor="Thorell" journalOrPublisher="Annals and Magazine of Natural History" pageId="16" pageNumber="16" pagination="1 - 15" part="4" refId="ref72155" refString="Thorell, T. 1876 a. On the classification of scorpions. Annals and Magazine of Natural History 4 (17): 1 - 15." title="On the classification of scorpions" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
</taxonomicName>
., ♀, Alter do Chão, Santarem, Pará, Brazil.
<emphasis id="35B88053FFDAB1443B15CA52B0D268E3" bold="true" box="[885,910,1662,1686]" pageId="16" pageNumber="16">G.</emphasis>
<taxonomicName id="C0CC27C2FFDAB1443BF4CA52B72268E3" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[916,1150,1662,1686]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFDAB1443BF4CA52B72268E3" box="[916,1150,1662,1686]" italics="true" pageId="16" pageNumber="16">Troglorhopalurus lacrau</emphasis>
</taxonomicName>
(
<bibRefCitation id="635D21B0FFDAB1443CEBCA52B2EC68C1" author="Lourenco and Pinto-da-Rocha" firstAuthor="Lourenco" journalOrPublisher="Biogeographica (Paris) 73" pageId="16" pageNumber="16" pagination="181 - 191" part="4" refId="ref68476" refString="Lourenco, W. R., and R. Pinto-da-Rocha. 1997. A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species. Biogeographica (Paris) 73 (4): 181 - 191." title="A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species" type="journal article" year="1997">Lourenço and Pinto-da-Rocha, 1997</bibRefCitation>
), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil.
<emphasis id="35B88053FFDAB1443BD7CAB0B08D68C1" bold="true" box="[951,977,1692,1716]" pageId="16" pageNumber="16">H.</emphasis>
<taxonomicName id="C0CC27C2FFDAB1443BB7CAB0B2D568A4" authority="Lourenco et al., 2004" authorityName="Lourenco et al." authorityYear="2004" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="16" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFDAB1443BB7CAB0B3F368A4" italics="true" pageId="16" pageNumber="16">Troglorhopalurus translucidus</emphasis>
<bibRefCitation id="635D21B0FFDAB14438D6CA95B2D568A4" author="Lourenco" box="[182,393,1721,1745]" etAl="et al." firstAuthor="Lourenco" journalOrPublisher="Comptes Rendus Biologies" pageId="16" pageNumber="16" pagination="1151 - 1156" part="327" refId="ref68662" refString="Lourenco, W. R., R. L. C. Baptista, and A. P. L. Giupponi. 2004. Troglobitic scorpions: a new genus and species from Brazil. Comptes Rendus Biologies 327: 1151 - 1156." title="Troglobitic scorpions: a new genus and species from Brazil" type="journal article" year="2004">Lourenço et al., 2004</bibRefCitation>
</taxonomicName>
, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil.
</paragraph>
</caption>
<caption id="53B30CC9FFDBB145380FC9E2B1C468BA" httpUri="https://zenodo.org/record/851386/files/figure.png" pageId="17" pageNumber="17" targetBox="[155,1166,224,1455]" targetPageId="17">
<paragraph id="07735C41FFDBB145380FC9E2B1C468BA" blockId="17.[111,1212,1485,1743]" pageId="17" pageNumber="17">
FIG. 2. Representative habitats of species in the New World buthid scorpion family Rhopalurusinae
<bibRefCitation id="635D21B0FFDBB1453C06C9E1B3FE6877" author="Bucherl" editor="W. Bucherl &amp; E. R. Buckley" firstAuthor="Bucherl" journalOrPublisher="New York: Academic Press" pageId="17" pageNumber="17" pagination="317 - 348" part="3" refId="ref64808" refString="Bucherl, W. 1971. Classification, biology and venom extraction of scorpions. In W. Bucherl and E. R. Buckley (editors), Venomous animals and their venoms 3: 317 - 348. New York: Academic Press." title="Classification, biology and venom extraction of scorpions" type="journal article" volumeTitle="Venomous animals and their venoms" year="1971">Bücherl, 1971</bibRefCitation>
.
<emphasis id="35B88053FFDBB14538CCC9C6B3996877" bold="true" box="[172,197,1514,1538]" pageId="17" pageNumber="17">A.</emphasis>
Isla Mona, Puerto Rico, habitat of
<taxonomicName id="C0CC27C2FFDBB1453A4BC9C7B72E6877" authority="(Armas and Marcano Fondeur, 1987)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas and Marcano Fondeur" baseAuthorityYear="1987" box="[555,1138,1514,1538]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="17" phylum="Arthropoda" rank="species" species="abudi" status="comb. nov.">
<emphasis id="35B88053FFDBB1453A4BC9C7B1B06877" box="[555,748,1514,1538]" italics="true" pageId="17" pageNumber="17">Heteroctenus abudi</emphasis>
(
<bibRefCitation id="635D21B0FFDBB1453A9AC9C6B7376877" author="Armas and Marcano Fondeur" box="[762,1131,1514,1538]" firstAuthor="Armas" journalOrPublisher="Poeyana" pageId="17" pageNumber="17" pagination="1 - 24" part="356" refId="ref64021" refString="Armas, L. F. de, and E. J. Marcano Fondeur. 1987. Nuevos escorpiones (Arachnida: Scorpiones) de Republica Dominicana. Poeyana 356: 1 - 24." title="Nuevos escorpiones (Arachnida: Scorpiones) de Republica Dominicana" type="journal article" year="1987">Armas and Marcano Fondeur, 1987</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFDBB1453C1DC9C6B3C6686A" pageId="17" pageNumber="17">comb. nov.</taxonomicNameLabel>
<emphasis id="35B88053FFDBB14538C1CA24B3E46855" bold="true" box="[161,184,1544,1568]" pageId="17" pageNumber="17">B.</emphasis>
Sierra de Cubitas, Camagüey, Cuba, habitat of
<taxonomicName id="C0CC27C2FFDBB1453AC9CA24B742686A" authority="(Herbst, 1800)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Herbst" baseAuthorityYear="1800" box="[681,1054,1543,1567]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="17" phylum="Arthropoda" rank="species" species="junceus" status="comb. nov.">
<emphasis id="35B88053FFDBB1453AC9CA24B022686A" box="[681,894,1544,1567]" italics="true" pageId="17" pageNumber="17">Heteroctenus junceus</emphasis>
(
<bibRefCitation id="635D21B0FFDBB1453BEDCA2BB74A686A" author="Herbst" box="[909,1046,1543,1567]" firstAuthor="Herbst" journalOrPublisher="In Natursystem der ungeflugelten Insekten" pageId="17" pageNumber="17" pagination="1 - 86" part="4" refId="ref66286" refString="Herbst, J. F. W. 1800. Naturgeschichte der Skorpionen. In Natursystem der ungeflugelten Insekten 4: 1 - 86. Berlin: Gottlieb August Lange." title="Naturgeschichte der Skorpionen" type="journal article" year="1800">Herbst, 1800</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFDBB1453C4ACA2BB7C0686A" box="[1066,1180,1543,1567]" pageId="17" pageNumber="17">comb. nov.</taxonomicNameLabel>
<emphasis id="35B88053FFDBB1453CC3CA2BB7E7686A" bold="true" box="[1187,1211,1543,1567]" pageId="17" pageNumber="17">C.</emphasis>
Guanambí, Bahía, Brazil, habitat of
<taxonomicName id="C0CC27C2FFDBB1453987CA09B7546849" authority="(Lenarducci et al., 2005)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2005" box="[487,1032,1572,1597]" class="Arachnida" family="Buthidae" genus="Ischnotelson" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="17" phylum="Arthropoda" rank="species" species="guanambiensis" status="comb. nov.">
<emphasis id="35B88053FFDBB1453987CA09B1A26848" box="[487,766,1573,1597]" italics="true" pageId="17" pageNumber="17">Ischnotelson guanambiensis</emphasis>
(
<bibRefCitation id="635D21B0FFDBB1453B6ECA09B75D6849" author="Lenarducci" box="[782,1025,1572,1597]" etAl="et al." firstAuthor="Lenarducci" journalOrPublisher="Biota Neotropica" pageId="17" pageNumber="17" pagination="173 - 180" part="5" refId="ref67356" refString="Lenarducci, A. R. I., R. Pinto-da-Rocha, and S. M. Lucas. 2005. Descricao de uma nova especie de Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) do nordeste brasileiro. Biota Neotropica 5 (1 A): 173 - 180." title="Descricao de uma nova especie de Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) do nordeste brasileiro" type="journal article" year="2005">Lenarducci et al., 2005</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFDBB1453C75CA09B7D56849" box="[1045,1161,1573,1597]" pageId="17" pageNumber="17">comb. nov.</taxonomicNameLabel>
, and
<taxonomicName id="C0CC27C2FFDBB145380FCA6FB2FD682F" authority="(Borelli, 1910)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Borelli" baseAuthorityYear="1910" box="[111,417,1602,1626]" class="Arachnida" family="Buthidae" genus="Jaguajir" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="17" phylum="Arthropoda" rank="species" species="rochae" status="comb. nov.">
<emphasis id="35B88053FFDBB145380FCA6FB259682F" box="[111,261,1602,1626]" italics="true" pageId="17" pageNumber="17">Jaguajir rochae</emphasis>
(
<bibRefCitation id="635D21B0FFDBB1453973CA6EB2C6682F" author="Borelli" box="[275,410,1602,1626]" firstAuthor="Borelli" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Realle Universita di Torino" pageId="17" pageNumber="17" pagination="1 - 8" part="25" refId="ref64441" refString="Borelli, A. 1910. Scorpioni nuovi e poco noti del Brasile. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Realle Universita di Torino 25 (629): 1 - 8." title="Scorpioni nuovi e poco noti del Brasile" type="journal article" year="1910">Borelli, 1910</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFDBB14539CCCA6EB140682C" box="[428,540,1602,1626]" pageId="17" pageNumber="17">comb. nov.</taxonomicNameLabel>
<emphasis id="35B88053FFDBB1453A43CA6EB167682F" bold="true" box="[547,571,1602,1626]" pageId="17" pageNumber="17">D.</emphasis>
Exu, Pernambuco, Brazil, habitat of
<taxonomicName id="C0CC27C2FFDBB1453BD6CA6FB750682F" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Borelli" baseAuthorityYear="1910" box="[950,1036,1602,1626]" class="Arachnida" family="Buthidae" genus="Jaguajir" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="17" phylum="Arthropoda" rank="species" species="rochae">
<emphasis id="35B88053FFDBB1453BD6CA6FB750682F" box="[950,1036,1602,1626]" italics="true" pageId="17" pageNumber="17">J. rochae</emphasis>
</taxonomicName>
and
<taxonomicName id="C0CC27C2FFDBB1453C21CA6EB2346802" authority="(C. L. Koch, 1840)" authorityName="C.L. Koch" authorityYear="1840" baseAuthorityName="C. L. Koch" baseAuthorityYear="1840" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="17" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="35B88053FFDBB1453C21CA6EB3F26802" italics="true" pageId="17" pageNumber="17">Physoctonus debilis</emphasis>
(C.L.
<bibRefCitation id="635D21B0FFDBB145388BCA73B23E6802" author="Koch" box="[235,354,1631,1655]" firstAuthor="Koch" journalOrPublisher="Nurnberg: C. H. Zeh'sche Buchhandlung" pageId="17" pageNumber="17" pagination="1 - 114" part="8" refId="ref66739" refString="Koch, C. L. 1840. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung 8: 1 - 114 (pts. 1, 2, 1840; pts. 2, 3, 1841)." title="Die Arachniden" type="journal article" year="1840">Koch, 1840</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="35B88053FFDBB1453911CA4CB2DA680D" bold="true" box="[369,390,1632,1656]" pageId="17" pageNumber="17">E.</emphasis>
Santarém, Pará, Brazil, habitat of
<taxonomicName id="C0CC27C2FFDBB1453ABBCA73B7606802" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[731,1084,1631,1655]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="17" phylum="Arthropoda" rank="species" species="laticauda">
<emphasis id="35B88053FFDBB1453ABBCA73B0F36802" box="[731,943,1631,1655]" italics="true" pageId="17" pageNumber="17">Rhopalurus laticauda</emphasis>
<bibRefCitation id="635D21B0FFDBB1453BD5CA73B7606802" author="Thorell" box="[949,1084,1631,1655]" firstAuthor="Thorell" journalOrPublisher="Annals and Magazine of Natural History" pageId="17" pageNumber="17" pagination="1 - 15" part="4" refId="ref72155" refString="Thorell, T. 1876 a. On the classification of scorpions. Annals and Magazine of Natural History 4 (17): 1 - 15." title="On the classification of scorpions" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
</taxonomicName>
.
<emphasis id="35B88053FFDBB1453C25CA4CB708680D" bold="true" box="[1093,1108,1632,1656]" pageId="17" pageNumber="17">F.</emphasis>
Normandia, Roraima, Brazil, habitat of
<taxonomicName id="C0CC27C2FFDBB14539C7CA51B14268E0" authorityName="Thorell" authorityYear="1876" box="[423,542,1661,1685]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="17" phylum="Arthropoda" rank="species" species="laticauda">
<emphasis id="35B88053FFDBB14539C7CA51B14268E0" box="[423,542,1661,1685]" italics="true" pageId="17" pageNumber="17">R. laticauda</emphasis>
</taxonomicName>
.
<emphasis id="35B88053FFDBB1453A49CA51B11E68E0" bold="true" box="[553,578,1661,1685]" pageId="17" pageNumber="17">G.</emphasis>
Lapa do Bode, Itaeté, Bahía, Brazil, habitat of
<taxonomicName id="C0CC27C2FFDBB1453C70CA51B3F168C7" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="17" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFDBB1453C70CA51B3F168C7" italics="true" pageId="17" pageNumber="17">Troglorhopalurus lacrau</emphasis>
</taxonomicName>
(
<bibRefCitation id="635D21B0FFDBB14538DBCAB6B16868C7" author="Lourenco and Pinto-da-Rocha" box="[187,564,1690,1714]" firstAuthor="Lourenco" journalOrPublisher="Biogeographica (Paris) 73" pageId="17" pageNumber="17" pagination="181 - 191" part="4" refId="ref68476" refString="Lourenco, W. R., and R. Pinto-da-Rocha. 1997. A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species. Biogeographica (Paris) 73 (4): 181 - 191." title="A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species" type="journal article" year="1997">Lourenço and Pinto-da-Rocha, 1997</bibRefCitation>
), comb. nov.
<emphasis id="35B88053FFDBB1453ADCCAB6B18B68C7" bold="true" box="[700,727,1690,1714]" pageId="17" pageNumber="17">H.</emphasis>
Gruta do Lapão, Lençóis, Bahía, Brazil, habitat of
<taxonomicName id="C0CC27C2FFDBB14538EACA9BB1C868BA" authority="Lourenco et al., 2004" authorityName="Lourenco et al." authorityYear="2004" box="[138,660,1719,1743]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="17" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFDBB14538EACA9BB2E968BA" box="[138,437,1719,1743]" italics="true" pageId="17" pageNumber="17">Troglorhopalurus translucidus</emphasis>
<bibRefCitation id="635D21B0FFDBB14539DBCA94B1C868BA" author="Lourenco" box="[443,660,1719,1743]" etAl="et al." firstAuthor="Lourenco" journalOrPublisher="Comptes Rendus Biologies" pageId="17" pageNumber="17" pagination="1151 - 1156" part="327" refId="ref68662" refString="Lourenco, W. R., R. L. C. Baptista, and A. P. L. Giupponi. 2004. Troglobitic scorpions: a new genus and species from Brazil. Comptes Rendus Biologies 327: 1151 - 1156." title="Troglobitic scorpions: a new genus and species from Brazil" type="journal article" year="2004">Lourenço et al., 2004</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="53B30CC9FFD8B1463BADCAB6B75D6C4D" httpUri="https://zenodo.org/record/851388/files/figure.png" pageId="18" pageNumber="18" targetBox="[401,930,221,1725]" targetPageId="18">
<paragraph id="07735C41FFD8B1463BADCAB6B75D6C4D" blockId="18.[971,1025,224,1732]" pageId="18" pageNumber="18">
FIG. 3. Map of Cuba and neighboring islets, plotting known locality records of three species of
<taxonomicName id="C0CC27C2FFD8B1463BACCE49B0BF6C70" authority="Pocock , 1893" authorityName="Pocock" authorityYear="1893" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFD8B1463BACCE49B0BF6C93" box="[972,995,613,742]" italics="true" pageId="18" pageNumber="18">Heteroctenus</emphasis>
<bibRefCitation id="635D21B0FFD8B1463BABCE3DB0BF6C70" author="Pocock" box="[971,995,466,605]" firstAuthor="Pocock" journalOrPublisher="Journal of the Linnaean Society" pageId="18" pageNumber="18" pagination="374 - 409" part="24" refId="ref69886" refString="Pocock, R. I. 1893. Contribution to our knowledge of the arthropod fauna of the West Indies. Part I. Scorpiones and Pedipalpi, with a supplementary note upon freshwater Decapoda of St. Vincent. Journal of the Linnaean Society 24: 374 - 409." title="Contribution to our knowledge of the arthropod fauna of the West Indies. Part I. Scorpiones and Pedipalpi, with a supplementary note upon freshwater Decapoda of St. Vincent" type="journal article" year="1893">Pocock, 1893</bibRefCitation>
</taxonomicName>
:
<emphasis id="35B88053FFD8B1463BABCD9FB0BF6FC1" bold="true" box="[971,995,431,454]" pageId="18" pageNumber="18">A.</emphasis>
<emphasis id="35B88053FFD8B1463BACCDBDB0BF6FE4" box="[972,995,398,423]" italics="true" pageId="18" pageNumber="18">H.</emphasis>
<taxonomicName id="C0CC27C2FFD8B1463BACCD17B75D68E7" authority="( Herbst , 1800 )" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Herbst" baseAuthorityYear="1800" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="18" phylum="Arthropoda" rank="species" species="junceus">
<emphasis id="35B88053FFD8B1463BACCD17B0BF6FF3" box="[972,995,315,390]" italics="true" pageId="18" pageNumber="18">junceus</emphasis>
(
<bibRefCitation id="635D21B0FFD8B1463BABCCC8B75D68B1" author="Herbst" box="[971,1025,224,1732]" firstAuthor="Herbst" journalOrPublisher="In Natursystem der ungeflugelten Insekten" pageId="18" pageNumber="18" pagination="1 - 86" part="4" refId="ref66286" refString="Herbst, J. F. W. 1800. Naturgeschichte der Skorpionen. In Natursystem der ungeflugelten Insekten 4: 1 - 86. Berlin: Gottlieb August Lange." title="Naturgeschichte der Skorpionen" type="journal article" year="1800">Herbst, 1800</bibRefCitation>
)
</taxonomicName>
(circles).
<emphasis id="35B88053FFD8B1463B89CA39B75D6860" bold="true" box="[1001,1025,1552,1574]" pageId="18" pageNumber="18">B.</emphasis>
H.
<taxonomicName id="C0CC27C2FFD8B1463B89C9B5B75D6B70" authority="(Armas , 1974 )" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas" baseAuthorityYear="1974" box="[1001,1025,1278,1513]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="18" phylum="Arthropoda" rank="species" species="garridoi">
<emphasis id="35B88053FFD8B1463B89C9B5B75D6B9C" box="[1001,1025,1433,1513]" italics="true" pageId="18" pageNumber="18">garridoi</emphasis>
(
<bibRefCitation id="635D21B0FFD8B1463B89C968B75D6B4C" author="Armas" box="[1001,1025,1285,1420]" firstAuthor="Armas" journalOrPublisher="Poeyana" pageId="18" pageNumber="18" pagination="1 - 26" part="132" refId="ref63476" refString="Armas, L. F. de. 1974 a. Escorpiones del archipielago cubano. II. Hallazgo de genero Microtityus (Scorpionida: Buthidae) con las descripciones de un nuevo subgenero y tres nuevas especies. Poeyana 132: 1 - 26." title="Escorpiones del archipielago cubano. II. Hallazgo de genero Microtityus (Scorpionida: Buthidae) con las descripciones de un nuevo subgenero y tres nuevas especies" type="journal article" year="1974">Armas, 1974</bibRefCitation>
)
</taxonomicName>
, comb. nov. (squares);
<emphasis id="35B88053FFD8B1463B89CFD2B75C6D8B" box="[1001,1024,1019,1044]" italics="true" pageId="18" pageNumber="18">H.</emphasis>
<taxonomicName id="C0CC27C2FFD8B1463B89CF87B75C6D6F" authority="( Teruel, 2006 )" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Teruel" baseAuthorityYear="2006" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="18" phylum="Arthropoda" rank="species" species="gibarae">
<emphasis id="35B88053FFD8B1463B89CF87B75D6D81" box="[1001,1025,939,1012]" italics="true" pageId="18" pageNumber="18">gibarae</emphasis>
(
<bibRefCitation id="635D21B0FFD8B1463B88CF75B75C6D3B" author="Teruel" box="[1000,1024,794,925]" firstAuthor="Teruel" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="18" pageNumber="18" pagination="43 - 56" part="38" refId="ref71569" refString="Teruel, R. 2006. Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba. Boletin de la Sociedad Entomologica Aragonesa 38: 43 - 56." title="Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba" type="journal article" year="2006">Teruel, 2006</bibRefCitation>
)
</taxonomicName>
, comb. nov. (crosses).
</paragraph>
</caption>
<caption id="53B30CC9FFD9B147380BC8A5B0B26AAF" httpUri="https://zenodo.org/record/851390/files/figure.png" pageId="19" pageNumber="19" targetBox="[120,1211,195,1121]" targetPageId="19">
<paragraph id="07735C41FFD9B147380BC8A5B0B26AAF" blockId="19.[107,1211,1159,1242]" pageId="19" pageNumber="19">
FIG. 4. Map of Hispaniola and neighboring islets, plotting known locality records of three species of
<taxonomicName id="C0CC27C2FFD9B1473C06C8A4B2696AC9" authority="Pocock, 1893" authorityName="Pocock" authorityYear="1893" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFD9B1473C06C8A4B3FC6AC9" italics="true" pageId="19" pageNumber="19">Heteroctenus</emphasis>
<bibRefCitation id="635D21B0FFD9B14738C9C888B2696AC9" author="Pocock" box="[169,309,1188,1212]" firstAuthor="Pocock" journalOrPublisher="Journal of the Linnaean Society" pageId="19" pageNumber="19" pagination="374 - 409" part="24" refId="ref69886" refString="Pocock, R. I. 1893. Contribution to our knowledge of the arthropod fauna of the West Indies. Part I. Scorpiones and Pedipalpi, with a supplementary note upon freshwater Decapoda of St. Vincent. Journal of the Linnaean Society 24: 374 - 409." title="Contribution to our knowledge of the arthropod fauna of the West Indies. Part I. Scorpiones and Pedipalpi, with a supplementary note upon freshwater Decapoda of St. Vincent" type="journal article" year="1893">Pocock, 1893</bibRefCitation>
</taxonomicName>
:
<emphasis id="35B88053FFD9B1473921C888B2056AC9" bold="true" box="[321,345,1188,1212]" pageId="19" pageNumber="19">A.</emphasis>
<taxonomicName id="C0CC27C2FFD9B1473902C889B0116AC9" authority="(Armas and Marcano Fondeur, 1987)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas and Marcano Fondeur" baseAuthorityYear="1987" box="[354,845,1188,1212]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="19" phylum="Arthropoda" rank="species" species="abudi" status="comb. nov.">
<emphasis id="35B88053FFD9B1473902C889B2E16AC9" box="[354,445,1188,1212]" italics="true" pageId="19" pageNumber="19">H. abudi</emphasis>
(
<bibRefCitation id="635D21B0FFD9B14739ADC888B01A6AC9" author="Armas and Marcano Fondeur" box="[461,838,1188,1212]" firstAuthor="Armas" journalOrPublisher="Poeyana" pageId="19" pageNumber="19" pagination="1 - 24" part="356" refId="ref64021" refString="Armas, L. F. de, and E. J. Marcano Fondeur. 1987. Nuevos escorpiones (Arachnida: Scorpiones) de Republica Dominicana. Poeyana 356: 1 - 24." title="Nuevos escorpiones (Arachnida: Scorpiones) de Republica Dominicana" type="journal article" year="1987">Armas and Marcano Fondeur, 1987</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFD9B1473B3AC888B0906AC9" box="[858,972,1188,1212]" pageId="19" pageNumber="19">comb. nov.</taxonomicNameLabel>
(triangles);
<taxonomicName id="C0CC27C2FFD9B1473C2CC889B3A36AAF" authority="(Armas, 1999)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas" baseAuthorityYear="1999" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="19" phylum="Arthropoda" rank="species" species="bonettii" status="comb. nov.">
<emphasis id="35B88053FFD9B1473C2CC889B7E76AC9" box="[1100,1211,1188,1212]" italics="true" pageId="19" pageNumber="19">H. bonettii</emphasis>
(
<bibRefCitation id="635D21B0FFD9B1473812C8EEB3A46AAF" author="Armas" box="[114,248,1218,1242]" firstAuthor="Armas" journalOrPublisher="Avicennia 10" pageId="19" pageNumber="19" pagination="101 - 136" part="11" refId="ref63877" refString="Armas, L. F. de. 1999. Quince nuevos alacranes de La Espanola y Navassa, Antillas Mayores (Arachnida: Scorpiones). Avicennia 10 / 11: 101 - 136." title="Quince nuevos alacranes de La Espanola y Navassa, Antillas Mayores (Arachnida: Scorpiones)" type="journal article" year="1999">Armas, 1999</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFD9B147396AC8EDB2266AAC" box="[266,378,1217,1241]" pageId="19" pageNumber="19">comb. nov.</taxonomicNameLabel>
(circles).
<emphasis id="35B88053FFD9B14739BDC8EEB2A86AAF" bold="true" box="[477,500,1218,1242]" pageId="19" pageNumber="19">B.</emphasis>
<taxonomicName id="C0CC27C2FFD9B147399BC8EEB0566AAF" authority="(Karsch, 1879)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[507,778,1217,1242]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="19" phylum="Arthropoda" rank="species" species="princeps" status="comb. nov.">
<emphasis id="35B88053FFD9B147399BC8EEB1376AAC" box="[507,619,1218,1241]" italics="true" pageId="19" pageNumber="19">H. princeps</emphasis>
(
<bibRefCitation id="635D21B0FFD9B1473A1AC8EDB05F6AAF" author="Karsch" box="[634,771,1217,1242]" firstAuthor="Karsch" journalOrPublisher="Mitteilungen des Munchener Entomologischen Vereins" pageId="19" pageNumber="19" pagination="6 - 22" part="3" refId="ref66627" refString="Karsch, F. 1879 a. Scorpionologische Beitrage. Part I. Mitteilungen des Munchener Entomologischen Vereins 3 (2): 6 - 22." title="Scorpionologische Beitrage. Part I" type="journal article" year="1879">Karsch, 1879</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFD9B1473B75C8EDB0D86AAC" box="[789,900,1217,1241]" pageId="19" pageNumber="19">comb. nov.</taxonomicNameLabel>
(squares).
</paragraph>
</caption>
<subSubSection id="4FD60FCAFFD9B14D38EBC969B3A86A42" lastPageId="25" lastPageNumber="25" pageId="19" pageNumber="19" type="description">
<paragraph id="07735C41FFD9B14738EBC969B2096B95" blockId="19.[107,637,1282,1737]" pageId="19" pageNumber="19">
DESCRIPTION: The following general description outlines characters common to the species of
<taxonomicName id="C0CC27C2FFD9B14738E9C9A4B24F6BEA" authorityName="Pocock" authorityYear="1893" box="[137,275,1416,1439]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFD9B14738E9C9A4B24F6BEA" box="[137,275,1416,1439]" italics="true" pageId="19" pageNumber="19">Heteroctenus</emphasis>
</taxonomicName>
. Descriptions of hemispermatophores are based on
<taxonomicName id="C0CC27C2FFD9B1473927C985B2F96BB5" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas and Marcano Fondeur" baseAuthorityYear="1987" box="[327,421,1448,1472]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="19" phylum="Arthropoda" rank="species" species="abudi">
<emphasis id="35B88053FFD9B1473927C985B2F96BB5" box="[327,421,1448,1472]" italics="true" pageId="19" pageNumber="19">H. abudi</emphasis>
</taxonomicName>
,
<taxonomicName id="C0CC27C2FFD9B14739D1C985B17E6BB5" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas" baseAuthorityYear="1999" box="[433,546,1448,1472]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="19" phylum="Arthropoda" rank="species" species="bonettii">
<emphasis id="35B88053FFD9B14739D1C985B17E6BB5" box="[433,546,1448,1472]" italics="true" pageId="19" pageNumber="19">H. bonettii</emphasis>
</taxonomicName>
,
<taxonomicName id="C0CC27C2FFD9B1473A4FC985B3C46B94" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Herbst" baseAuthorityYear="1800" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="19" phylum="Arthropoda" rank="species" species="junceus">
<emphasis id="35B88053FFD9B1473A4FC985B3C46B94" italics="true" pageId="19" pageNumber="19">H. junceus</emphasis>
</taxonomicName>
, and
<taxonomicName id="C0CC27C2FFD9B14738B5C9E6B2136B94" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[213,335,1482,1505]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="19" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="35B88053FFD9B14738B5C9E6B2136B94" box="[213,335,1482,1505]" italics="true" pageId="19" pageNumber="19">H. princeps</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="07735C41FFD9B14738EBC9C6B2186856" blockId="19.[107,637,1282,1737]" pageId="19" pageNumber="19">
<emphasis id="35B88053FFD9B14738EBC9C6B24D6877" box="[139,273,1514,1538]" italics="true" pageId="19" pageNumber="19">Total length</emphasis>
: Large, robust scorpions (total length, 50–70 mm).
</paragraph>
<paragraph id="07735C41FFD9B14738EBCA00B0FC6BEA" blockId="19.[107,637,1282,1737]" lastBlockId="19.[684,1212,1282,1736]" pageId="19" pageNumber="19">
<emphasis id="35B88053FFD9B14738EBCA00B3956831" box="[139,201,1580,1604]" italics="true" pageId="19" pageNumber="19">Color:</emphasis>
Carapace and tergites I–VI light brown, tergite VII yellowish (fig. 1A). Coxosternal region, pectines and sternites pale yellow. Metasomal segments, dorsal surfaces yellow (segments I–III) to brown (IV and V); ventral surfaces darker; segments IV and V darker than preceding segments, with V darker than IV. Telson reddish brown, aculeus almost black. Chelicerae and legs yellowish. Pedipalps yellow with chela fingers darker than manus, reddish-brown.
</paragraph>
<paragraph id="07735C41FFD9B1473AACC984B7D86B94" blockId="19.[684,1212,1282,1736]" pageId="19" pageNumber="19">
<emphasis id="35B88053FFD9B1473AACC984B0676BB5" box="[716,827,1448,1472]" italics="true" pageId="19" pageNumber="19">Chelicerae</emphasis>
: Base, dorsal surface with medial transverse row of well-developed tubercles.
</paragraph>
<paragraph id="07735C41FFD9B1403AACC9C6B1126AAF" blockId="19.[684,1212,1282,1736]" lastBlockId="20.[107,637,1185,1738]" lastPageId="20" lastPageNumber="20" pageId="19" pageNumber="19">
<emphasis id="35B88053FFD9B1473AACC9C6B0716874" box="[716,813,1514,1537]" italics="true" pageId="19" pageNumber="19">Carapace</emphasis>
: Median ocular tubercle raised (fig. 14A–E); two median ocelli; three pairs of lateral macroocelli; one or two pairs of lateral microocelli. Anteromedian, median ocular, and posteromedian sulci well developed, forming single, almost continuous, longitudinal sulcus. Lateral ocular, central lateral, anterior central submedian, and posterior central submedian carinae distinct, coarsely granular to costate-granular and separate (unfused).
</paragraph>
<caption id="53B30CC9FFDEB140380BC81DB3936A10" httpUri="https://zenodo.org/record/851392/files/figure.png" pageId="20" pageNumber="20" targetBox="[129,1215,225,1032]" targetPageId="20">
<paragraph id="07735C41FFDEB140380BC81DB3936A10" blockId="20.[107,1213,1071,1125]" pageId="20" pageNumber="20">
FIG. 5. Map of northern South America, plotting known locality records of
<taxonomicName id="C0CC27C2FFDEB1403B00C803B7E16A32" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[864,1213,1071,1095]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="20" phylum="Arthropoda" rank="species" species="laticauda">
<emphasis id="35B88053FFDEB1403B00C803B7686A32" box="[864,1076,1071,1095]" italics="true" pageId="20" pageNumber="20">Rhopalurus laticauda</emphasis>
<bibRefCitation id="635D21B0FFDEB1403C5AC803B7E16A32" author="Thorell" box="[1082,1213,1071,1095]" firstAuthor="Thorell" journalOrPublisher="Annals and Magazine of Natural History" pageId="20" pageNumber="20" pagination="1 - 15" part="4" refId="ref72155" refString="Thorell, T. 1876 a. On the classification of scorpions. Annals and Magazine of Natural History 4 (17): 1 - 15." title="On the classification of scorpions" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
</taxonomicName>
(squares).
</paragraph>
</caption>
<paragraph id="07735C41FFDEB14038EBC8CFB76B6B97" blockId="20.[107,637,1185,1738]" lastBlockId="20.[684,1214,1185,1738]" pageId="20" pageNumber="20">
<emphasis id="35B88053FFDEB14038EBC8CFB3AE6A8E" box="[139,242,1251,1275]" italics="true" pageId="20" pageNumber="20">Pedipalps</emphasis>
: Pedipalp femur retrolateral accessory carinae absent. Pedipalp chela manus of adult male incrassate, fixed and movable fingers curved proximally (fixed finger curved dorsally, movable finger curved ventrally), such that proximal dentate margin emarginate, distinct gap present between fingers proximally, when closed (figs. 25–27), manus of female not incrassate, fixed and movable fingers not curved proximally, such that proximal dentate margin sublinear, little or no gap present between them proximally, when closed; manus, proventral and promedian (except in
<taxonomicName id="C0CC27C2FFDEB14039E8CA5DB15168FD" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas" baseAuthorityYear="1999" box="[392,525,1648,1672]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="20" phylum="Arthropoda" rank="species" species="bonettii">
<emphasis id="35B88053FFDEB14039E8CA5DB15168FD" box="[392,525,1648,1672]" italics="true" pageId="20" pageNumber="20">H. bonettii</emphasis>
</taxonomicName>
) carinae absent; fixed and movable fingers, median denticle rows each comprising eight or nine oblique subrows of primary denticles flanked closely by pro- and retrolateral accessory (supernumerary) denticles; movable finger with proximal lobe (
<figureCitation id="9FF740C4FFDEB1403A93C928B01E6B69" box="[755,834,1284,1308]" captionStart="FIG. 17" captionStartId="36.[106,149,1153,1174]" captionTargetBox="[106,634,224,1125]" captionTargetId="figure@36.[106,634,224,1125]" captionTargetPageId="36" captionText="FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851416/files/figure.png" pageId="20" pageNumber="20">fig. 17</figureCitation>
A). Pedipalps orthobothriotaxic Type A, α configuration; femur with five dorsal trichobothria, trichobothrium
<emphasis id="35B88053FFDEB1403C77C96AB7796B2B" box="[1047,1061,1350,1374]" italics="true" pageId="20" pageNumber="20">d</emphasis>
<subScript id="9B485E04FFDEB1403C45C97DB7726B17" attach="left" box="[1061,1070,1361,1378]" fontSize="7" pageId="20" pageNumber="20">2</subScript>
situated on prolateral surface; patella trichobothrium
<emphasis id="35B88053FFDEB1403CC6C94BB7E86B0A" box="[1190,1204,1383,1407]" italics="true" pageId="20" pageNumber="20">d</emphasis>
<subScript id="9B485E04FFDEB1403CD5C95EB7E26BF6" attach="right" box="[1205,1214,1394,1411]" fontSize="7" pageId="20" pageNumber="20">3</subScript>
situated retrolateral to dorsomedian carina; chela fixed finger trichobothrium
<emphasis id="35B88053FFDEB1403C22C985B7016BB4" box="[1090,1117,1449,1473]" italics="true" pageId="20" pageNumber="20">db</emphasis>
aligned with or distal to trichobothrium
<emphasis id="35B88053FFDEB1403C7DC9E7B76C6B97" box="[1053,1072,1483,1506]" italics="true" pageId="20" pageNumber="20">et</emphasis>
.
</paragraph>
<paragraph id="07735C41FFDEB1403AACC9C0B0FB6813" blockId="20.[684,1214,1185,1738]" pageId="20" pageNumber="20">
<emphasis id="35B88053FFDEB1403AACC9C0B1A66876" box="[716,762,1516,1539]" italics="true" pageId="20" pageNumber="20">Legs</emphasis>
: Legs III and IV, tibial spurs absent; I–IV, basitarsi each with bifurcate prolateral pedal spur; telotarsi each with irregular tufts of fine, acuminate macrosetae.
</paragraph>
<paragraph id="07735C41FFDEB1413AACCA5DB2EC6B0A" blockId="20.[684,1214,1185,1738]" lastBlockId="21.[109,637,1185,1738]" lastPageId="21" lastPageNumber="21" pageId="20" pageNumber="20">
<emphasis id="35B88053FFDEB1403AACCA5DB07F68FD" box="[716,803,1649,1672]" italics="true" pageId="20" pageNumber="20">Pectines</emphasis>
: Pectinal plate with single median depression (male), anterior margin with sulcus (fig. 18A–E). Pectines often proximally expanded, at least 1.5× wider proximally than medially; proximal dorsal fulcra setose; pectinal teeth almost straight, slightly curved laterally, proximal teeth often enlarged, dorsal surfaces with multiple nodules and regular striations (figs. 11B, 12B), dorsobasal surfaces with macrosetae; pectinal sensillae peg shaped.
</paragraph>
<caption id="53B30CC9FFDFB141380BC802B2276A0A" httpUri="https://zenodo.org/record/851394/files/figure.png" pageId="21" pageNumber="21" targetBox="[112,1197,224,1032]" targetPageId="21">
<paragraph id="07735C41FFDFB141380BC802B2276A0A" blockId="21.[107,1208,1068,1151]" pageId="21" pageNumber="21">
FIG. 6. Map of northern South America, plotting known locality records of
<taxonomicName id="C0CC27C2FFDFB1413BECC801B3FA6A17" authority="(Mello-Leitao, 1932)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1932" class="Arachnida" family="Buthidae" genus="Jaguajir" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="21" phylum="Arthropoda" rank="species" species="pintoi" status="comb. nov.">
<emphasis id="35B88053FFDFB1413BECC801B7426A31" box="[908,1054,1069,1092]" italics="true" pageId="21" pageNumber="21">Jaguajir pintoi</emphasis>
(
<bibRefCitation id="635D21B0FFDFB1413C4DC800B3C36A17" author="Mello-Leitao" firstAuthor="Mello-Leitao" journalOrPublisher="Arquivos do Museu Nacional" pageId="21" pageNumber="21" pagination="9 - 46" part="34" refId="ref69227" refString="Mello-Leitao, C. F. de. 1932. Notas sobre escorpioes sulamericanos. Arquivos do Museu Nacional 34: 9 - 46." title="Notas sobre escorpioes sulamericanos" type="journal article" year="1932">Mello-Leitão, 1932</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFDFB14138D3C866B2796A14" box="[179,293,1098,1122]" pageId="21" pageNumber="21">comb. nov.</taxonomicNameLabel>
(circles),
<taxonomicName id="C0CC27C2FFDFB14139EDC866B0D96A17" authority="Teruel and Roncallo, 2008" authorityName="Teruel and Roncallo" authorityYear="2008" box="[397,901,1098,1122]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="21" phylum="Arthropoda" rank="species" species="caribensis">
<emphasis id="35B88053FFDFB14139EDC866B1376A17" box="[397,619,1098,1122]" italics="true" pageId="21" pageNumber="21">Rhopalurus caribensis</emphasis>
<bibRefCitation id="635D21B0FFDFB1413A13C866B0D96A17" author="Teruel and Roncallo" box="[627,901,1098,1122]" firstAuthor="Teruel" journalOrPublisher="Euscorpius" pageId="21" pageNumber="21" pagination="1 - 12" part="68" refId="ref71905" refString="Teruel, R., and C. A. Roncallo. 2008. Rare or poorly known scorpions from Colombia. III. On the taxonomy and distribution of Rhopalurus laticauda Thorell, 1876 (Scorpiones: Buthidae), with description of a new species of the genus. Euscorpius 68: 1 - 12." title="Rare or poorly known scorpions from Colombia. III. On the taxonomy and distribution of Rhopalurus laticauda Thorell, 1876 (Scorpiones: Buthidae), with description of a new species of the genus" type="journal article" year="2008">Teruel and Roncallo, 2008</bibRefCitation>
</taxonomicName>
(pentagons), and
<taxonomicName id="C0CC27C2FFDFB1413C25C866B3F06A0A" authority="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="21" phylum="Arthropoda" rank="species" species="ochoai" status="sp. nov.">
<emphasis id="35B88053FFDFB1413C25C866B3F06A0A" italics="true" pageId="21" pageNumber="21">Rhopalurus ochoai</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFDFB14138D8C84BB2596A0B" box="[184,261,1127,1150]" pageId="21" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
(triangles).
</paragraph>
</caption>
<paragraph id="07735C41FFDFB14138EDC9A5B0276B69" blockId="21.[109,637,1185,1738]" lastBlockId="21.[684,1211,1185,1738]" pageId="21" pageNumber="21">
<emphasis id="35B88053FFDFB14138EDC9A5B3A46BD5" box="[141,248,1417,1440]" italics="true" pageId="21" pageNumber="21">Mesosoma</emphasis>
: Tergites IV–VI wider than I–III and VII (figs. 31–35); I–VI tricarinate, dorsomedian and dorsosubmedian carinae granular to costategranular, restricted to posterior half on I–VI, dorsosubmedian carinae more prominent on IV–VI. Tergite VII pentacarinate, dorsomedian carina restricted to anterior two thirds of segment. Sternites smooth, carinate obsolete, more developed on VI and VII; sternite III, lateral margins forming smooth, raised carina, ventromedian carina elevated anteriorly, ventrosubmedian surfaces forming paired depressions, finely and irregularly granular; respiratory spiracles (stigmata) width more than 5× length (
<figureCitation id="9FF740C4FFDFB1413A9CC928B0606B69" box="[764,828,1284,1308]" captionStart="FIG. 11" captionStartId="28.[108,150,1167,1188]" captionTargetBox="[108,1213,224,1136]" captionTargetId="figure@28.[108,1213,224,1136]" captionTargetPageId="28" captionText="FIG. 11. Rhopalurusinae Bücherl, 1971, dextral pecten, ventral aspect (A, C, E, G) and pectinal teeth, dorsal aspect (B, D, F). A. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♀ (AMNH). C. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP) D. Jaguajir rochae (Borelli, 1910), comb. nov., (AMNH). E. Rhopalurus laticauda Thorell, 1876, ♂ (AMCC [LP 2845]). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (AMCC [LP 3260]). G. Physoctonus debilis (C. L. Koch, 1840), ♀ (MZSP)." httpUri="https://zenodo.org/record/851404/files/figure.png" pageId="21" pageNumber="21">fig. 11</figureCitation>
A, B).
</paragraph>
<paragraph id="07735C41FFDFB14D3AACC90AB3816F96" blockId="21.[684,1211,1185,1738]" lastBlockId="25.[107,637,228,1740]" lastPageId="25" lastPageNumber="25" pageId="21" pageNumber="21">
<emphasis id="35B88053FFDFB1413AACC90AB0696B48" box="[716,821,1318,1341]" italics="true" pageId="21" pageNumber="21">Metasoma</emphasis>
: Metasoma robust, but not increasing markedly in width posteriorly, segment V slightly wider than I in adult male, I and V usually similar width in adult female (figs. 28–30). Segments I and II each with 10 distinct, costate-granular carinae, III and IV each with eight distinct, costate-granular carinae, V with seven distinct but less pronounced, granular carinae; dorsosubmedian carinae obsolete, reduced to rows of granules on dorsal surfaces of segments I–IV, more pronounced on segment I; dorsolateral carinae complete on segments I–IV, and terminating in prominent, spiniform granules posteriorly on III and IV, absent on V; lateral supramedian carinae complete on segments I–V; lateral inframedian carinae complete on segment I, partial on II, absent on III–V; ventrosubmedian carinae complete on segments I–IV, restricted to anterior third of V; ventromedian carina absent on segments I–IV, complete on V. Intercarinal surfaces finely to coarsely granular, less so on dorsal surfaces, especially on V.
</paragraph>
<caption id="53B30CC9FFDCB142380CC9D4B292685E" httpUri="https://zenodo.org/record/851396/files/figure.png" pageId="22" pageNumber="22" targetBox="[120,1208,225,1490]" targetPageId="22">
<paragraph id="07735C41FFDCB142380CC9D4B292685E" blockId="22.[108,1214,1526,1579]" pageId="22" pageNumber="22">
FIG. 7. Map of northeastern South America, plotting known locality records of
<taxonomicName id="C0CC27C2FFDCB1423BD0C9DBB3B5685E" authority="(C. L. Koch, 1839)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="C. L. Koch" baseAuthorityYear="1839" class="Arachnida" family="Buthidae" genus="Jaguajir" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="22" phylum="Arthropoda" rank="species" species="agamemnon" status="comb. nov.">
<emphasis id="35B88053FFDCB1423BD0C9DBB7DD687B" box="[944,1153,1527,1550]" italics="true" pageId="22" pageNumber="22">Jaguajir agamemnon</emphasis>
(C.L.
<bibRefCitation id="635D21B0FFDCB142380CCA3FB3BE685E" author="Koch" box="[108,226,1555,1579]" firstAuthor="Koch" journalOrPublisher="Nurnberg: C. H. Zeh'sche Buchhandlung" pageId="22" pageNumber="22" pagination="1 - 156" part="6" refId="ref66709" refString="Koch, C. L. 1839. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung 6 (1 - 6): 1 - 156." title="Die Arachniden" type="journal article" year="1839">Koch, 1839</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFDCB1423894CA3FB238685E" box="[244,356,1555,1579]" pageId="22" pageNumber="22">comb. nov.</taxonomicNameLabel>
(squares).
</paragraph>
</caption>
<caption id="53B30CC9FFDDB143380BC9D4B2646859" httpUri="https://zenodo.org/record/851398/files/figure.png" pageId="23" pageNumber="23" targetBox="[117,1207,234,1492]" targetPageId="23">
<paragraph id="07735C41FFDDB143380BC9D4B2646859" blockId="23.[107,1211,1526,1580]" pageId="23" pageNumber="23">
FIG. 8. Map of northeastern South America, plotting known locality records of
<taxonomicName id="C0CC27C2FFDDB1433BEAC9DBB7EB687A" authority="(Borelli, 1910)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Borelli" baseAuthorityYear="1910" box="[906,1207,1526,1551]" class="Arachnida" family="Buthidae" genus="Jaguajir" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="23" phylum="Arthropoda" rank="species" species="rochae" status="comb. nov.">
<emphasis id="35B88053FFDDB1433BEAC9DBB743687B" box="[906,1055,1526,1550]" italics="true" pageId="23" pageNumber="23">Jaguajir rochae</emphasis>
(
<bibRefCitation id="635D21B0FFDDB1433C4BC9DAB7EC687A" author="Borelli" box="[1067,1200,1526,1551]" firstAuthor="Borelli" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Realle Universita di Torino" pageId="23" pageNumber="23" pagination="1 - 8" part="25" refId="ref64441" refString="Borelli, A. 1910. Scorpioni nuovi e poco noti del Brasile. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Realle Universita di Torino 25 (629): 1 - 8." title="Scorpioni nuovi e poco noti del Brasile" type="journal article" year="1910">Borelli, 1910</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFDDB143380BCA38B386685E" box="[107,218,1556,1580]" pageId="23" pageNumber="23">comb. nov.</taxonomicNameLabel>
(circles).
</paragraph>
</caption>
<caption id="53B30CC9FFD2B14C380BC9A4B17B6866" httpUri="https://zenodo.org/record/851400/files/figure.png" pageId="24" pageNumber="24" targetBox="[138,1207,234,1374]" targetPageId="24">
<paragraph id="07735C41FFD2B14C380BC9A4B17B6866" blockId="24.[107,1214,1414,1555]" pageId="24" pageNumber="24">
FIG. 9. Map of northeastern South America, plotting known locality records of six species of Rhopalurusinae
<bibRefCitation id="635D21B0FFD2B14C380BC98FB3AB6BCE" author="Bucherl" box="[107,247,1443,1467]" editor="W. Bucherl &amp; E. R. Buckley" firstAuthor="Bucherl" journalOrPublisher="New York: Academic Press" pageId="24" pageNumber="24" pagination="317 - 348" part="3" refId="ref64808" refString="Bucherl, W. 1971. Classification, biology and venom extraction of scorpions. In W. Bucherl and E. R. Buckley (editors), Venomous animals and their venoms 3: 317 - 348. New York: Academic Press." title="Classification, biology and venom extraction of scorpions" type="journal article" volumeTitle="Venomous animals and their venoms" year="1971">Bücherl, 1971</bibRefCitation>
:
<taxonomicName id="C0CC27C2FFD2B14C389FC98FB07A6BCE" authority="(Lenarducci et al., 2005)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lenarducci et al." baseAuthorityYear="2005" box="[255,806,1443,1467]" pageId="24" pageNumber="24" rank="subSpecies" status="comb. nov." subSpecies="guanambiensis">
<emphasis id="35B88053FFD2B14C389FC98FB2446BCE" bold="true" box="[255,280,1443,1467]" pageId="24" pageNumber="24">A.</emphasis>
<emphasis id="35B88053FFD2B14C397DC98FB1716BCE" box="[285,557,1443,1467]" italics="true" pageId="24" pageNumber="24">Ischnotelson guanambiensis</emphasis>
(
<bibRefCitation id="635D21B0FFD2B14C3A59C98FB0436BCE" author="Lenarducci" box="[569,799,1443,1467]" etAl="et al." firstAuthor="Lenarducci" journalOrPublisher="Biota Neotropica" pageId="24" pageNumber="24" pagination="173 - 180" part="5" refId="ref67356" refString="Lenarducci, A. R. I., R. Pinto-da-Rocha, and S. M. Lucas. 2005. Descricao de uma nova especie de Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) do nordeste brasileiro. Biota Neotropica 5 (1 A): 173 - 180." title="Descricao de uma nova especie de Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) do nordeste brasileiro" type="journal article" year="2005">Lenarducci et al., 2005</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFD2B14C3B4FC98FB0C16BCE" box="[815,925,1443,1467]" pageId="24" pageNumber="24">comb. nov.</taxonomicNameLabel>
(star),
<taxonomicName id="C0CC27C2FFD2B14C3B83C98FB7EB6BCE" authority="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[995,1207,1443,1467]" class="Arachnida" family="Buthidae" genus="Ischnotelson" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="peruassu" status="sp. nov.">
<emphasis id="35B88053FFD2B14C3B83C98FB7EB6BCE" box="[995,1207,1443,1467]" italics="true" pageId="24" pageNumber="24">Ischnotelson peruassu</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFD2B14C380BC9EDB3E66BAD" box="[107,186,1473,1496]" pageId="24" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
(triangle),
<taxonomicName id="C0CC27C2FFD2B14C3951C9ECB2A36BAD" authority="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[305,511,1472,1496]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
<emphasis id="35B88053FFD2B14C3951C9ECB2A36BAD" box="[305,511,1472,1496]" italics="true" pageId="24" pageNumber="24">Physoctonus striatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFD2B14C3A6EC9EDB1016BAD" box="[526,605,1473,1496]" pageId="24" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
(circle),
<taxonomicName id="C0CC27C2FFD2B14C3ADCC9ECB0EC6BAD" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[700,944,1472,1496]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFD2B14C3ADCC9ECB0EC6BAD" box="[700,944,1472,1496]" italics="true" pageId="24" pageNumber="24">Troglorhopalurus lacrau</emphasis>
</taxonomicName>
(Lourenço and Pinto-da- Rocha, 1997), comb. nov (diamonds).
<emphasis id="35B88053FFD2B14C399BC9F2B14E6B83" bold="true" box="[507,530,1502,1526]" pageId="24" pageNumber="24">B.</emphasis>
<taxonomicName id="C0CC27C2FFD2B14C3A7AC9F2B0C16B83" authority="(C. L. Koch, 1840)" authorityName="C.L. Koch" authorityYear="1840" baseAuthorityName="C. L. Koch" baseAuthorityYear="1840" box="[538,925,1502,1526]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="35B88053FFD2B14C3A7AC9F2B1806B83" box="[538,732,1502,1526]" italics="true" pageId="24" pageNumber="24">Physoctonus debilis</emphasis>
(C.L.
<bibRefCitation id="635D21B0FFD2B14C3B40C9F2B0CB6B83" author="Koch" box="[800,919,1502,1526]" firstAuthor="Koch" journalOrPublisher="Nurnberg: C. H. Zeh'sche Buchhandlung" pageId="24" pageNumber="24" pagination="1 - 114" part="8" refId="ref66739" refString="Koch, C. L. 1840. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung 8: 1 - 114 (pts. 1, 2, 1840; pts. 2, 3, 1841)." title="Die Arachniden" type="journal article" year="1840">Koch, 1840</bibRefCitation>
)
</taxonomicName>
(squares),
<taxonomicName id="C0CC27C2FFD2B14C3C72C9F2B29D6866" authority="Lourenco et al., 2004" authorityName="Lourenco et al." authorityYear="2004" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="24" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFD2B14C3C72C9F2B3B86866" italics="true" pageId="24" pageNumber="24">Troglorhopalurus translucidus</emphasis>
<bibRefCitation id="635D21B0FFD2B14C388AC9D0B29D6866" author="Lourenco" box="[234,449,1531,1555]" etAl="et al." firstAuthor="Lourenco" journalOrPublisher="Comptes Rendus Biologies" pageId="24" pageNumber="24" pagination="1151 - 1156" part="327" refId="ref68662" refString="Lourenco, W. R., R. L. C. Baptista, and A. P. L. Giupponi. 2004. Troglobitic scorpions: a new genus and species from Brazil. Comptes Rendus Biologies 327: 1151 - 1156." title="Troglobitic scorpions: a new genus and species from Brazil" type="journal article" year="2004">Lourenço et al., 2004</bibRefCitation>
</taxonomicName>
(crosses).
</paragraph>
</caption>
<paragraph id="07735C41FFD3B14D38EBCDC1B2866CDF" blockId="25.[107,637,228,1740]" pageId="25" pageNumber="25">
<emphasis id="35B88053FFD3B14D38EBCDC1B3966C70" box="[139,202,493,517]" italics="true" pageId="25" pageNumber="25">Telson</emphasis>
: Vesicle slightly elongate, length ca. 1.5× width, not laterally compressed, similar in width or slightly narrower than metasoma V; anterodorsal lateral lobes reduced or absent; lateral and ventral surfaces granular, with distinct ventromedian carina; subaculear tubercle absent.
</paragraph>
<paragraph id="07735C41FFD3B14D38EBCE9FB3A86A42" blockId="25.[107,637,228,1740]" pageId="25" pageNumber="25">
<emphasis id="35B88053FFD3B14D38EBCE9FB23B6CBE" box="[139,359,691,715]" italics="true" pageId="25" pageNumber="25">Hemispermatophore</emphasis>
: Flagelliform; flagellum, elongate and narrow (fig. 23A–N); trunk markedly concave; three lobules, ental (LI), ectal (LE), and basal (LB); LI continuous until flagellar base; flagellar base narrow, half (
<taxonomicName id="C0CC27C2FFD3B14D3A77CF14B1206D3A" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas and Marcano Fondeur" baseAuthorityYear="1987" box="[535,636,823,847]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="abudi">
<emphasis id="35B88053FFD3B14D3A77CF14B1206D3A" box="[535,636,823,847]" italics="true" pageId="25" pageNumber="25">H. abudi</emphasis>
</taxonomicName>
and
<taxonomicName id="C0CC27C2FFD3B14D38FBCF75B24C6D05" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas" baseAuthorityYear="1999" box="[155,272,856,880]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="bonettii">
<emphasis id="35B88053FFD3B14D38FBCF75B24C6D05" box="[155,272,856,880]" italics="true" pageId="25" pageNumber="25">H. bonettii</emphasis>
</taxonomicName>
) to one-third (
<taxonomicName id="C0CC27C2FFD3B14D39D7CF75B1776D05" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Herbst" baseAuthorityYear="1800" box="[439,555,857,880]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="junceus">
<emphasis id="35B88053FFD3B14D39D7CF75B1776D05" box="[439,555,857,880]" italics="true" pageId="25" pageNumber="25">H. junceus</emphasis>
</taxonomicName>
and
<taxonomicName id="C0CC27C2FFD3B14D3A01CF75B3996DE4" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Karsch" baseAuthorityYear="1879" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="35B88053FFD3B14D3A01CF75B3996DE4" italics="true" pageId="25" pageNumber="25">H. princeps</emphasis>
</taxonomicName>
) the maximum width of trunk; LE ca. half (
<emphasis id="35B88053FFD3B14D38C9CFB7B2C96DC7" box="[169,405,923,947]" italics="true" pageId="25" pageNumber="25">
<taxonomicName id="C0CC27C2FFD3B14D38C9CFB7B2536DC6" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas and Marcano Fondeur" baseAuthorityYear="1987" box="[169,271,923,947]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="abudi">H. abudi</taxonomicName>
,
<taxonomicName id="C0CC27C2FFD3B14D397DCFB7B2C96DC7" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Herbst" baseAuthorityYear="1800" box="[285,405,923,946]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="junceus">H. junceus</taxonomicName>
</emphasis>
and
<taxonomicName id="C0CC27C2FFD3B14D39B2CFB7B10E6DC7" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[466,594,923,946]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="35B88053FFD3B14D39B2CFB7B10E6DC7" box="[466,594,923,946]" italics="true" pageId="25" pageNumber="25">H. princeps</emphasis>
</taxonomicName>
) to one third (
<taxonomicName id="C0CC27C2FFD3B14D3883CF90B2056DA1" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas" baseAuthorityYear="1999" box="[227,345,956,980]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="bonettii">
<emphasis id="35B88053FFD3B14D3883CF90B2056DA1" box="[227,345,956,980]" italics="true" pageId="25" pageNumber="25">H. bonettii</emphasis>
</taxonomicName>
) the length of LI and may be spiniform (
<taxonomicName id="C0CC27C2FFD3B14D3970CFF1B2D56D80" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas" baseAuthorityYear="1999" box="[272,393,989,1013]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="bonettii">
<emphasis id="35B88053FFD3B14D3970CFF1B2D56D80" box="[272,393,989,1013]" italics="true" pageId="25" pageNumber="25">H. bonettii</emphasis>
</taxonomicName>
); LB very short with sharp (
<taxonomicName id="C0CC27C2FFD3B14D38DFCFD3B27B6A63" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas and Marcano Fondeur" baseAuthorityYear="1987" box="[191,295,1022,1046]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="abudi">
<emphasis id="35B88053FFD3B14D38DFCFD3B27B6A63" box="[191,295,1022,1046]" italics="true" pageId="25" pageNumber="25">H. abudi</emphasis>
</taxonomicName>
) or setalike (acuminate) (
<taxonomicName id="C0CC27C2FFD3B14D3A01CFD3B3E36A42" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas" baseAuthorityYear="1999" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="bonettii">
<emphasis id="35B88053FFD3B14D3A01CFD3B3E36A42" italics="true" pageId="25" pageNumber="25">H. bonettii</emphasis>
</taxonomicName>
) tip.
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFD3B14D38EBC86CB2D56B6B" pageId="25" pageNumber="25" type="discussion">
<paragraph id="07735C41FFD3B14D38EBC86CB2D56B6B" blockId="25.[107,637,228,1740]" pageId="25" pageNumber="25">
INCLUDED SPECIES:
<taxonomicName id="C0CC27C2FFD3B14D393DC86DB2C26A0C" authority="(Armas and Marcano Fondeur, 1987)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas and Marcano Fondeur" baseAuthorityYear="1987" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="abudi" status="comb. nov.">
<emphasis id="35B88053FFD3B14D393DC86DB1796A2D" box="[349,549,1088,1112]" italics="true" pageId="25" pageNumber="25">Heteroctenus abudi</emphasis>
(
<bibRefCitation id="635D21B0FFD3B14D3A52C86CB2CA6A0C" author="Armas and Marcano Fondeur" firstAuthor="Armas" journalOrPublisher="Poeyana" pageId="25" pageNumber="25" pagination="1 - 24" part="356" refString="Armas, L. F. de, and E. J. Marcano Fondeur. 1987. Nuevos escorpiones (Arachnida: Scorpiones) de Republica Dominicana. Poeyana 356: 1 - 24." title="Nuevos escorpiones (Arachnida: Scorpiones) de Republica Dominicana" type="journal article" year="1987">Armas and Marcano Fondeur, 1987</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFD3B14D39C7C84DB1476A0D" box="[423,539,1121,1145]" pageId="25" pageNumber="25">comb. nov.</taxonomicNameLabel>
;
<taxonomicName id="C0CC27C2FFD3B14D3A44C84EB2C66AEF" authority="(Armas, 1999)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas" baseAuthorityYear="1999" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="bonettii" status="comb. nov.">
<emphasis id="35B88053FFD3B14D3A44C84EB3AB6AEF" italics="true" pageId="25" pageNumber="25">Heteroctenus bonettii</emphasis>
(
<bibRefCitation id="635D21B0FFD3B14D3966C8AEB2CE6AEF" author="Armas" box="[262,402,1154,1178]" firstAuthor="Armas" journalOrPublisher="Avicennia 10" pageId="25" pageNumber="25" pagination="101 - 136" part="11" refString="Armas, L. F. de. 1999. Quince nuevos alacranes de La Espanola y Navassa, Antillas Mayores (Arachnida: Scorpiones). Avicennia 10 / 11: 101 - 136." title="Quince nuevos alacranes de La Espanola y Navassa, Antillas Mayores (Arachnida: Scorpiones)" type="journal article" year="1999">Armas, 1999</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFD3B14D39C4C8AEB1466AEC" box="[420,538,1154,1178]" pageId="25" pageNumber="25">comb. nov.</taxonomicNameLabel>
;
<taxonomicName id="C0CC27C2FFD3B14D3A43C8AFB2CF6ACE" authority="(Herbst, 1800)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Herbst" baseAuthorityYear="1800" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="junceus">
<emphasis id="35B88053FFD3B14D3A43C8AFB3A86ACE" italics="true" pageId="25" pageNumber="25">Heteroctenus junceus</emphasis>
(
<bibRefCitation id="635D21B0FFD3B14D3962C88FB2D06ACE" author="Herbst" box="[258,396,1187,1211]" firstAuthor="Herbst" journalOrPublisher="In Natursystem der ungeflugelten Insekten" pageId="25" pageNumber="25" pagination="1 - 86" part="4" refString="Herbst, J. F. W. 1800. Naturgeschichte der Skorpionen. In Natursystem der ungeflugelten Insekten 4: 1 - 86. Berlin: Gottlieb August Lange." title="Naturgeschichte der Skorpionen" type="journal article" year="1800">Herbst, 1800</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="C0CC27C2FFD3B14D39FEC888B2566AA9" authority="(Armas, 1974)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Armas" baseAuthorityYear="1974" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="garridoi" status="comb. nov.">
<emphasis id="35B88053FFD3B14D39FEC888B1206ACE" box="[414,636,1187,1211]" italics="true" pageId="25" pageNumber="25">Heteroctenus garridoi</emphasis>
(
<bibRefCitation id="635D21B0FFD3B14D3812C8E8B25E6AA9" author="Armas" box="[114,258,1220,1244]" firstAuthor="Armas" journalOrPublisher="Poeyana" pageId="25" pageNumber="25" pagination="1 - 26" part="132" refString="Armas, L. F. de. 1974 a. Escorpiones del archipielago cubano. II. Hallazgo de genero Microtityus (Scorpionida: Buthidae) con las descripciones de un nuevo subgenero y tres nuevas especies. Poeyana 132: 1 - 26." title="Escorpiones del archipielago cubano. II. Hallazgo de genero Microtityus (Scorpionida: Buthidae) con las descripciones de un nuevo subgenero y tres nuevas especies" type="journal article" year="1974">Armas, 1974</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFD3B14D3977C8E8B2CD6AA9" box="[279,401,1220,1244]" pageId="25" pageNumber="25">comb. nov.</taxonomicNameLabel>
;
<taxonomicName id="C0CC27C2FFD3B14D39FDC8E9B2586A88" authority="(Teruel, 2006)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Teruel" baseAuthorityYear="2006" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="gibarae" status="comb. nov.">
<emphasis id="35B88053FFD3B14D39FDC8E9B1206AA9" box="[413,636,1220,1244]" italics="true" pageId="25" pageNumber="25">Heteroctenus gibarae</emphasis>
(
<bibRefCitation id="635D21B0FFD3B14D3812C8C9B3A16A88" author="Teruel" box="[114,253,1253,1277]" firstAuthor="Teruel" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="25" pageNumber="25" pagination="43 - 56" part="38" refString="Teruel, R. 2006. Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba. Boletin de la Sociedad Entomologica Aragonesa 38: 43 - 56." title="Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba" type="journal article" year="2006">Teruel, 2006</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFD3B14D3971C8C9B2D56A88" box="[273,393,1253,1277]" pageId="25" pageNumber="25">comb. nov.</taxonomicNameLabel>
;
<taxonomicName id="C0CC27C2FFD3B14D39F6C8CAB2556B6B" authority="(Karsch, 1879)" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Karsch" baseAuthorityYear="1879" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="princeps" status="comb. nov.">
<emphasis id="35B88053FFD3B14D39F6C8CAB1276A88" box="[406,635,1254,1277]" italics="true" pageId="25" pageNumber="25">Heteroctenus princeps</emphasis>
(
<bibRefCitation id="635D21B0FFD3B14D3812C92AB25E6B6B" author="Karsch" box="[114,258,1286,1310]" firstAuthor="Karsch" journalOrPublisher="Mitteilungen des Munchener Entomologischen Vereins" pageId="25" pageNumber="25" pagination="6 - 22" part="3" refString="Karsch, F. 1879 a. Scorpionologische Beitrage. Part I. Mitteilungen des Munchener Entomologischen Vereins 3 (2): 6 - 22." title="Scorpionologische Beitrage. Part I" type="journal article" year="1879">Karsch, 1879</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2E8B3D28FFD3B14D3974C92AB2D56B6B" box="[276,393,1286,1310]" pageId="25" pageNumber="25">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFD3B14D38EBC904B2906873" pageId="25" pageNumber="25" type="distribution">
<paragraph id="07735C41FFD3B14D38EBC904B2906873" blockId="25.[107,637,228,1740]" pageId="25" pageNumber="25">
DISTRIBUTION: The species of
<taxonomicName id="C0CC27C2FFD3B14D398FC904B1206B4A" authorityName="Pocock" authorityYear="1893" box="[495,636,1320,1343]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFD3B14D398FC904B1206B4A" box="[495,636,1320,1343]" italics="true" pageId="25" pageNumber="25">Heteroctenus</emphasis>
</taxonomicName>
are endemic to the Greater Antilles of the Caribbean (figs. 3, 4): Cuba, Hispaniola (Haiti and the Dominican Republic), and Puerto Rico (Isla Mona). The known locality records range in altitudes from below sea level to 1200 m (
<bibRefCitation id="635D21B0FFD3B14D3A50C9E1B29C6873" author="Rodriguez-Cabrera and Teruel" firstAuthor="Rodriguez-Cabrera" journalOrPublisher="Revista Iberica de Aracnologia" pageId="25" pageNumber="25" pagination="119 - 122" part="24" refString="Rodriguez-Cabrera, T. M., and R. Teruel. 2014. On the highest altitudinal occurrences of scorpions in Cuba (Arachnida: Scorpiones). Revista Iberica de Aracnologia 24: 119 - 122." title="On the highest altitudinal occurrences of scorpions in Cuba (Arachnida: Scorpiones)" type="journal article" year="2014">Rodríguez-Cabrera and Teruel, 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFD3B14D38EBCA23B7736F68" pageId="25" pageNumber="25" type="biology_ecology">
<paragraph id="07735C41FFD3B14D38EBCA23B7736F68" blockId="25.[107,637,228,1740]" lastBlockId="25.[684,1214,228,650]" pageId="25" pageNumber="25">
ECOLOGY: Whereas
<taxonomicName id="C0CC27C2FFD3B14D3919CA3CB1596852" authorityName="Pocock" authorityYear="1893" box="[377,517,1552,1575]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFD3B14D3919CA3CB1596852" box="[377,517,1552,1575]" italics="true" pageId="25" pageNumber="25">Heteroctenus</emphasis>
</taxonomicName>
primarily inhabit open vegetation formations (fig. 2A, B), in common with
<taxonomicName id="C0CC27C2FFD3B14D396BCA7DB2DE681C" authorityName="Thorell" authorityYear="1876" box="[267,386,1617,1641]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFD3B14D396BCA7DB2DE681C" box="[267,386,1617,1641]" italics="true" pageId="25" pageNumber="25">Rhopalurus</emphasis>
</taxonomicName>
and the South American genera (
<bibRefCitation id="635D21B0FFD3B14D3899CA5EB2EA68FF" author="Lourenco, W. R." box="[249,438,1650,1674]" journalOrPublisher="Memoires de la Societe Royale Belge d'Entomologie" pageId="25" pageNumber="25" pagination="129 - 137" part="33" refString="Lourenco, W. R. 1986 a. Biogeographie et phylogenie des scorpions du genre Rhopalurus Thorell, 1876 (Scorpiones, Buthidae). Memoires de la Societe Royale Belge d'Entomologie 33: 129 - 137." title="Biogeographie et phylogenie des scorpions du genre Rhopalurus Thorell, 1876 (Scorpiones, Buthidae)" type="journal article" year="1986" yearSuffix="a">Lourenço, 1986a</bibRefCitation>
,
<bibRefCitation id="635D21B0FFD3B14D39A6CA5EB2A268FF" author="Lourenco" box="[454,510,1650,1674]" firstAuthor="Lourenco" pageId="25" pageNumber="25" refString="2008: 9, table 2, pl. 42; Lourenco, 2008: 3, 4, 5, 7, 12, table 1, fig. 4; Teruel and Roncallo, 2008: 2 - 5, 11, fig. 2; Prendini et al., 2009: 222, 223; Florez, 2012 (part): 1, 2, figs. 1, 2, table 1; Teruel and Roncallo, 2013: 112, 113." type="journal article" year="2008">2008</bibRefCitation>
), they also occur in semideciduous forests (
<bibRefCitation id="635D21B0FFD3B14D39A5CABFB10E68DE" author="Armas" box="[453,594,1683,1707]" editor="V. Fet" firstAuthor="Armas" journalOrPublisher="Burnham Beeches, Bucks, UK: British Arachnological Society" pageId="25" pageNumber="25" pagination="245 - 253" part="2001" refString="Armas, L. F. de. 2001. Scorpions of the Greater Antilles, with the description of a new troglobitic species (Scorpiones: Diplocentridae). In V. Fet and P. A. Selden (editors), Scorpions 2001: in memoriam Gary A. Polis: 245 - 253. Burnham Beeches, Bucks, UK: British Arachnological Society." title="Scorpions of the Greater Antilles, with the description of a new troglobitic species (Scorpiones: Diplocentridae)" type="journal article" volumeTitle="Scorpions" year="2001">Armas, 2001</bibRefCitation>
). In the karst limestone landscapes where they occur, these lapidicolous scorpions shelter under stones or any other available surface debris.
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFD3B14D3AACCD0BB04F6CFF" pageId="25" pageNumber="25" type="discussion">
<paragraph id="07735C41FFD3B14D3AACCD0BB04F6CFF" blockId="25.[684,1214,228,650]" pageId="25" pageNumber="25">
REMARKS: This genus accommodates species previously assigned to
<taxonomicName id="C0CC27C2FFD3B14D3BA5CD64B7196F15" authorityName="Thorell" authorityYear="1876" box="[965,1093,328,352]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FFD3B14D3BA5CD64B7196F15" box="[965,1093,328,352]" italics="true" pageId="25" pageNumber="25">Rhopalurus</emphasis>
</taxonomicName>
from the Greater Antilles, the monophyly of which was consistently recovered in the analyses by Esposito et al. (in review), resulting in four new combinations.
<taxonomicName id="C0CC27C2FFD3B14D3A92CDE0B08B6F91" baseAuthorityName="Teruel and Armas" baseAuthorityYear="2012" box="[754,983,460,484]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="aridicola">
<emphasis id="35B88053FFD3B14D3A92CDE0B08B6F91" box="[754,983,460,484]" italics="true" pageId="25" pageNumber="25">Rhopalurus aridicola</emphasis>
</taxonomicName>
,
<taxonomicName id="C0CC27C2FFD3B14D3B87CDE1B7D86F91" authorityName="Teruel and Armas" authorityYear="2006" box="[999,1156,460,484]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="melloleitaoi">
<emphasis id="35B88053FFD3B14D3B87CDE1B7D86F91" box="[999,1156,460,484]" italics="true" pageId="25" pageNumber="25">R. melloleitaoi</emphasis>
</taxonomicName>
, and
<taxonomicName id="C0CC27C2FFD3B14D3ACCCDC2B0526C70" authorityName="Santiago-Blay" authorityYear="2009" box="[684,782,493,517]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="virkkii">
<emphasis id="35B88053FFD3B14D3ACCCDC2B0526C70" box="[684,782,493,517]" italics="true" pageId="25" pageNumber="25">R. virkkii</emphasis>
</taxonomicName>
are newly synonymized based on morphological and, in the case of
<taxonomicName id="C0CC27C2FFD3B14D3B93CE23B7096C52" authorityName="Santiago-Blay" authorityYear="2009" box="[1011,1109,527,551]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="virkkii">
<emphasis id="35B88053FFD3B14D3B93CE23B7096C52" box="[1011,1109,527,551]" italics="true" pageId="25" pageNumber="25">R. virkkii</emphasis>
</taxonomicName>
, molecular evidence. The validity of
<taxonomicName id="C0CC27C2FFD3B14D3B82CE1DB70E6C3D" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Teruel" baseAuthorityYear="2006" box="[994,1106,560,584]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="25" phylum="Arthropoda" rank="species" species="gibarae">
<emphasis id="35B88053FFD3B14D3B82CE1DB70E6C3D" box="[994,1106,560,584]" italics="true" pageId="25" pageNumber="25">H. gibarae</emphasis>
</taxonomicName>
will need to be reassessed when material becomes available for study.
</paragraph>
</subSubSection>
</treatment>
</document>
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