513 lines
92 KiB
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513 lines
92 KiB
XML
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<mods:title id="3BCDAAF25D71A62FBC2662EE72DB839A">Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)</mods:title>
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<mods:namePart id="98F677C356DA626259C3D77C653C82F7">Lauren A. Esposito</mods:namePart>
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<mods:namePart id="68D67A469A795152386035809C3928EB">Humberto Y. Yamaguti</mods:namePart>
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<mods:namePart id="F07FD7E2D023D1800525EF5DED8E2313">Cláudio A. Souza</mods:namePart>
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<mods:namePart id="6A6C907CDA1640381A61484B49C2B9BF">Ricardo Pinto-Da-Rocha</mods:namePart>
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<mods:namePart id="D3F263FE8CC08324721EC889EDFF192F">Lorenzo Prendini</mods:namePart>
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<treatment id="8F65ED57FFBEB12338F5CCC8B21C6ADF" ID-DOI="http://doi.org/10.5281/zenodo.4610710" ID-GBIF-Taxon="132953581" ID-Zenodo-Dep="4610710" LSID="urn:lsid:plazi:treatment:8F65ED57FFBEB12338F5CCC8B21C6ADF" httpUri="http://treatment.plazi.org/id/8F65ED57FFBEB12338F5CCC8B21C6ADF" lastPageId="119" lastPageNumber="118" pageId="116" pageNumber="116">
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<taxonomicName id="C0CC27C2FFBEB12038F5CCC8B2946F6B" ID-CoL="82VX" authority="Lourenco, Baptista and Giupponi, 2004" authorityName="Lourenco, Baptista and Giupponi" authorityYear="2004" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
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<emphasis id="35B88053FFBEB12038F5CCC8B2106E89" box="[149,332,228,252]" italics="true" pageId="116" pageNumber="116">Troglorhopalurus</emphasis>
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<bibRefCitation id="635D21B0FFBEB1203935CCC9B2946F6B" author="Lourenco, Baptista and Giupponi" firstAuthor="Lourenco" journalOrPublisher="Comptes Rendus Biologies" pageId="116" pageNumber="116" pagination="1151 - 1156" part="327" refId="ref68662" refString="Lourenco, W. R., R. L. C. Baptista, and A. P. L. Giupponi. 2004. Troglobitic scorpions: a new genus and species from Brazil. Comptes Rendus Biologies 327: 1151 - 1156." title="Troglobitic scorpions: a new genus and species from Brazil" type="journal article" year="2004">Lourenço, Baptista and Giupponi, 2004</bibRefCitation>
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<subSubSection id="4FD60FCAFFBEB12038E1CD14B1096F04" pageId="116" pageNumber="116" type="description">
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<paragraph id="07735C41FFBEB12038E1CD14B1096F04" blockId="116.[129,613,312,369]" pageId="116" pageNumber="116">
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<figureCitation id="9FF740C4FFBEB12038E1CD14B2416F3A" box="[129,285,312,336]" captionStart="FIG" captionStartId="16.[107,147,1517,1538]" captionTargetBox="[188,1135,224,1487]" captionTargetId="figure@16.[142,1179,224,1487]" captionTargetPageId="16" captionText="FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil." httpUri="https://zenodo.org/record/851384/files/figure.png" pageId="116" pageNumber="116">
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<figureCitation id="9FF740C4FFBEB12038E1CD14B3BF6F25" box="[129,227,312,336]" captionStart="FIG. 1" captionStartId="16.[107,147,1517,1538]" captionTargetBox="[188,1135,224,1487]" captionTargetId="figure@16.[142,1179,224,1487]" captionTargetPageId="16" captionText="FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil." httpUri="https://zenodo.org/record/851384/files/figure.png" pageId="116" pageNumber="116">Figures 1</figureCitation>
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G, H
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</figureCitation>
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,
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<figureCitation id="9FF740C4FFBEB1203949CD14B2326F3A" box="[297,366,312,336]" captionStart="FIG" captionStartId="17.[111,153,1486,1507]" captionTargetBox="[155,1166,224,1455]" captionTargetId="figure@17.[111,1214,224,1455]" captionTargetPageId="17" captionText="FIG. 2. Representative habitats of species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971. A. Isla Mona, Puerto Rico, habitat of Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov. B. Sierra de Cubitas, Camagüey, Cuba, habitat of Heteroctenus junceus (Herbst, 1800), comb. nov. C. Guanambí, Bahía, Brazil, habitat of Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., and Jaguajir rochae (Borelli, 1910), comb. nov. D. Exu, Pernambuco, Brazil, habitat of J. rochae and Physoctonus debilis (C.L. Koch, 1840). E. Santarém, Pará, Brazil, habitat of Rhopalurus laticauda Thorell, 1876. F. Normandia, Roraima, Brazil, habitat of R. laticauda. G. Lapa do Bode, Itaeté, Bahía, Brazil, habitat of Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov. H. Gruta do Lapão, Lençóis, Bahía, Brazil, habitat of Troglorhopalurus translucidus Lourenço et al., 2004." httpUri="https://zenodo.org/record/851386/files/figure.png" pageId="116" pageNumber="116">2G, H</figureCitation>
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,
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<figureCitation id="9FF740C4FFBEB120391ACD14B2D56F25" box="[378,393,312,336]" captionStart="FIG" captionStartId="24.[107,149,1416,1437]" captionTargetBox="[138,1207,234,1374]" captionTargetId="figure@24.[107,1211,224,1383]" captionTargetPageId="24" captionText="FIG. 9. Map of northeastern South America, plotting known locality records of six species of Rhopalurusinae Bücherl, 1971: A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov. (star), Ischnotelson peruassu, sp. nov. (triangle), Physoctonus striatus, sp. nov. (circle), Troglorhopalurus lacrau (Lourenço and Pinto-da- Rocha, 1997), comb. nov (diamonds). B. Physoctonus debilis (C.L. Koch, 1840) (squares), Troglorhopalurus translucidus Lourenço et al., 2004 (crosses)." httpUri="https://zenodo.org/record/851400/files/figure.png" pageId="116" pageNumber="116">9</figureCitation>
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,
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<figureCitation id="9FF740C4FFBEB12039F5CD14B29D6F25" box="[405,449,312,336]" captionStart="FIG" captionStartId="26.[109,151,1396,1417]" captionTargetBox="[109,1213,224,1368]" captionTargetId="figure@26.[109,1213,224,1368]" captionTargetPageId="26" captionText="FIG. 10. Rhopalurusinae Bücherl, 1971, sternites III (A–C, E, F), V and VI (D), ventral aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). B. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (MZSP). C, D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E. Physoctonus debilis (C.L. Koch, 1840), ♂ (MZSP). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 1 mm (A, E, F), 2 mm (B, C, D)." httpUri="https://zenodo.org/record/851402/files/figure.png" pageId="116" pageNumber="116">10E</figureCitation>
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,
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<figureCitation id="9FF740C4FFBEB12039ACCD14B2AF6F25" box="[460,499,312,336]" captionStart="FIG" captionStartId="28.[108,150,1167,1188]" captionTargetBox="[108,1213,224,1136]" captionTargetId="figure@28.[108,1213,224,1136]" captionTargetPageId="28" captionText="FIG. 11. Rhopalurusinae Bücherl, 1971, dextral pecten, ventral aspect (A, C, E, G) and pectinal teeth, dorsal aspect (B, D, F). A. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♀ (AMNH). C. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP) D. Jaguajir rochae (Borelli, 1910), comb. nov., (AMNH). E. Rhopalurus laticauda Thorell, 1876, ♂ (AMCC [LP 2845]). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (AMCC [LP 3260]). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP)." httpUri="https://zenodo.org/record/851404/files/figure.png" pageId="116" pageNumber="116">11F</figureCitation>
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,
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<figureCitation id="9FF740C4FFBEB120399FCD14B1716F25" box="[511,557,312,336]" captionStart="FIG" captionStartId="29.[107,149,984,1005]" captionTargetBox="[107,1211,224,956]" captionTargetId="figure@29.[107,1211,224,956]" captionTargetPageId="29" captionText="FIG. 12. Rhopalurusinae Bücherl, 1971, pectinal teeth, dorsal aspect, fine structure. A. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., (MZSP). B. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). C. Rhopalurus laticauda Thorell, 1876, (MZSP). D. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (MZSP). E. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). F. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP)." httpUri="https://zenodo.org/record/851406/files/figure.png" pageId="116" pageNumber="116">12D</figureCitation>
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,
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<figureCitation id="9FF740C4FFBEB1203A5ACD14B13D6F25" box="[570,609,312,336]" captionStart="FIG" captionStartId="32.[107,149,1598,1619]" captionTargetBox="[220,1108,224,1568]" captionTargetId="figure@32.[118,1207,224,1568]" captionTargetPageId="32" captionText="FIG. 14. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. Heteroctenus garridoi (Armas, 1974), comb. nov., ♂ (AMNH). D. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). E. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851410/files/figure.png" pageId="116" pageNumber="116">14F</figureCitation>
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,
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<figureCitation id="9FF740C4FFBEB12038F2CD75B3E16F04" box="[146,189,345,369]" captionStart="FIG" captionStartId="33.[108,150,1627,1648]" captionTargetBox="[235,1082,224,1599]" captionTargetId="figure@33.[108,1209,224,1599]" captionTargetPageId="33" captionText="FIG. 15. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). B. Ischnotelson peruassu, sp. nov., paratype ♂ (MZSP). C. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♂ (MZSP). D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♀ (MZSP). E. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (AMNH). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851412/files/figure.png" pageId="116" pageNumber="116">15E</figureCitation>
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,
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<figureCitation id="9FF740C4FFBEB12038A9CD75B3AC6F04" box="[201,240,345,369]" captionStart="FIG" captionStartId="36.[106,149,1153,1174]" captionTargetBox="[106,634,224,1125]" captionTargetId="figure@36.[106,634,224,1125]" captionTargetPageId="36" captionText="FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851416/files/figure.png" pageId="116" pageNumber="116">17F</figureCitation>
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,
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<figureCitation id="9FF740C4FFBEB120389CCD75B27F6F04" box="[252,291,345,369]" captionStart="FIG" captionStartId="38.[107,149,1554,1575]" captionTargetBox="[108,1212,224,1525]" captionTargetId="figure@38.[108,1212,224,1526]" captionTargetPageId="38" captionText="FIG. 18. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. Heteroctenus junceus (Herbst, 1800), comb. nov., ♂ (AMNH). D. Heteroctenus junceus (Herbst, 1800), ♀ (AMNH). E. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851418/files/figure.png" pageId="116" pageNumber="116">18F</figureCitation>
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,
|
||
<figureCitation id="9FF740C4FFBEB120394FCD75B2076F04" box="[303,347,345,369]" captionStart="FIG" captionStartId="39.[108,151,1503,1524]" captionTargetBox="[108,1212,227,1475]" captionTargetId="figure@39.[108,1212,227,1475]" captionTargetPageId="39" captionText="FIG. 19. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♀ (MZSP). B. Ischnotelson peruassu, sp. nov., paratype ♂ (MZSP). C. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♂ (MZSP). D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851420/files/figure.png" pageId="116" pageNumber="116">19E</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FFBEB1203906CD75B2ED6F04" box="[358,433,345,369]" captionStart="FIG" captionStartId="44.[107,149,1352,1373]" captionTargetBox="[108,1212,224,1323]" captionTargetId="figure@44.[108,1212,224,1323]" captionTargetPageId="44" captionText="FIG. 21. Rhopalurusinae Bücherl, 1971, telson, lateral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851424/files/figure.png" pageId="116" pageNumber="116">21K, L</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FFBEB12039DCCD75B15B6F04" box="[444,519,345,369]" captionStart="FIG" captionStartId="45.[108,147,1513,1534]" captionTargetBox="[176,1145,224,1484]" captionTargetId="figure@45.[108,1212,224,1484]" captionTargetPageId="45" captionText="FIG. 22. Rhopalurusinae Bücherl, 1971, telson, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH). J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851426/files/figure.png" pageId="116" pageNumber="116">22K, L</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FFBEB1203A73CD75B1096F04" box="[531,597,345,369]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="114.[108,150,859,880]" captionStartId-1="115.[108,150,1567,1588]" captionStartId-2="117.[108,150,1678,1699]" captionTargetBox-0="[108,1212,224,828]" captionTargetBox-1="[108,1212,224,1539]" captionTargetBox-2="[108,1212,224,1650]" captionTargetId-0="figure@114.[108,1212,224,828]" captionTargetId-1="figure@115.[108,1212,224,1539]" captionTargetId-2="figure@117.[108,1212,224,1650]" captionTargetPageId-0="114" captionTargetPageId-1="115" captionTargetPageId-2="117" captionText-0="FIG. 61. Troglorhopalurus Lourenço et al., 2004, pedipalp chela, retrolateral aspect. A. T. lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). B. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." captionText-1="FIG. 62. Troglorhopalurus Lourenço et al., 2004, metasoma and telson, dorsal (A, C), ventral (B, D) and lateral (E, F) aspects. A, B, E. T. lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). C, D, F. T. translucidus Lourenço et al., 2004, ♀ (MZSP)." captionText-2="FIG. 63. Troglorhopalurus Lourenço et al., 2004, habitus, dorsal (A, C) and ventral (B, D) aspects. A, B. T. lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). C, D. T. translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 1 mm." httpUri-0="https://zenodo.org/record/851504/files/figure.png" httpUri-1="https://zenodo.org/record/851506/files/figure.png" httpUri-2="https://zenodo.org/record/851508/files/figure.png" pageId="116" pageNumber="116">61–63</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFFBEB120380BCDBCB2326AD4" pageId="116" pageNumber="116" type="reference_group">
|
||
<paragraph id="07735C41FFBEB120380BCDBCB13D6FBF" blockId="116.[107,634,399,1185]" pageId="116" pageNumber="116">
|
||
<taxonomicName id="C0CC27C2FFBEB120380BCDBCB38C6FBF" ID-CoL="5995L" authority="Lourenco et al., 2004" authorityName="Lourenco et al." authorityYear="2004" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBEB120380BCDBCB2F96FDD" box="[107,421,400,424]" italics="true" pageId="116" pageNumber="116">Troglorhopalurus translucidus</emphasis>
|
||
<bibRefCitation id="635D21B0FFBEB12039CDCDBCB38C6FBF" author="Lourenco" etAl="et al." firstAuthor="Lourenco" journalOrPublisher="Comptes Rendus Biologies" pageId="116" pageNumber="116" pagination="1151 - 1156" part="327" refId="ref68662" refString="Lourenco, W. R., R. L. C. Baptista, and A. P. L. Giupponi. 2004. Troglobitic scorpions: a new genus and species from Brazil. Comptes Rendus Biologies 327: 1151 - 1156." title="Troglobitic scorpions: a new genus and species from Brazil" type="journal article" year="2004">Lourenço et al., 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
, type species by original designation.
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBEB120380BCDF9B1506CEF" blockId="116.[107,634,399,1185]" pageId="116" pageNumber="116">
|
||
<taxonomicName id="C0CC27C2FFBEB120380BCDF9B38C6C7A" ID-CoL="82VX" authority="Lourenco et al., 2004: 1151" authorityName="Lourenco et al., 2004:" authorityPageNumber="1151" authorityYear="1151" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB120380BCDF9B2436F98" box="[107,287,469,493]" italics="true" pageId="116" pageNumber="116">Troglorhopalurus</emphasis>
|
||
<bibRefCitation id="635D21B0FFBEB1203946CDF9B38C6C7A" author="Lourenco" etAl="et al." firstAuthor="Lourenco" journalOrPublisher="Comptes Rendus Biologies" pageId="116" pageNumber="116" pagination="1151 - 1156" part="327" refId="ref68662" refString="Lourenco, W. R., R. L. C. Baptista, and A. P. L. Giupponi. 2004. Troglobitic scorpions: a new genus and species from Brazil. Comptes Rendus Biologies 327: 1151 - 1156." title="Troglobitic scorpions: a new genus and species from Brazil" type="journal article" year="2004">Lourenço et al., 2004: 1151– 1156</bibRefCitation>
|
||
</taxonomicName>
|
||
, figs. 1–10;
|
||
<bibRefCitation id="635D21B0FFBEB120392FCDDBB25F6C47" author="Prendini and Wheeler" firstAuthor="Prendini" journalOrPublisher="Cladistics" pageId="116" pageNumber="116" pagination="446 - 494" part="21" refId="ref70578" refString="Prendini, L., and W. C. Wheeler. 2005. Scorpion higher phylogeny and classification, taxonomic anarchy, and standards for peer review in online publishing. Cladistics 21 (5): 446 - 494." title="Scorpion higher phylogeny and classification, taxonomic anarchy, and standards for peer review in online publishing" type="journal article" year="2005">Prendini and Wheeler, 2005: 481</bibRefCitation>
|
||
, table 10;
|
||
<bibRefCitation id="635D21B0FFBEB1203910CE36B2586C20" author="Volschenk and Prendini" firstAuthor="Volschenk" journalOrPublisher="Invertebrate Systematics" pageId="116" pageNumber="116" pagination="235 - 257" part="22" refId="ref72705" refString="Volschenk, E. S., and L. Prendini. 2008. Aops oncodactylus, gen. et n. sp., the first troglobitic urodacid (Urodacidae: Scorpiones), with a re-assessment of cavernicolous, troglobitic and troglomorphic scorpions. Invertebrate Systematics 22: 235 - 257." title="Aops oncodactylus, gen. et n. sp., the first troglobitic urodacid (Urodacidae: Scorpiones), with a re-assessment of cavernicolous, troglobitic and troglomorphic scorpions" type="journal article" year="2008">Volschenk and Prendini, 2008: 236</bibRefCitation>
|
||
, 249;
|
||
<bibRefCitation id="635D21B0FFBEB1203923CE11B10D6C20" author="Prendini" box="[323,593,573,597]" etAl="et al." firstAuthor="Prendini" journalOrPublisher="Journal of Arachnology" pageId="116" pageNumber="116" pagination="206 - 224" part="37" refId="ref70522" refString="Prendini, L., L. A. Esposito, J. Huff, and E. S. Volschenk. 2009. Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola. Journal of Arachnology 37: 206 - 224." title="Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola" type="journal article" year="2009">Prendini et al., 2009: 206</bibRefCitation>
|
||
, 222;
|
||
<bibRefCitation id="635D21B0FFBEB12038AECE73B2BB6C02" author="Brazil and Porto" bookContentInfo="84 pp." box="[206,487,607,631]" firstAuthor="Brazil" journalOrPublisher="Salvador, BA, Brazil: EDUFBA" pageId="116" pageNumber="116" refId="ref64569" refString="Brazil, T. K., and Porto, T. J. 2010. Os escorpioes. Salvador, BA, Brazil: EDUFBA. 84 pp." title="Os escorpioes" type="book" year="2010">Brazil and Porto, 2010: 57</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FFBEB1203992CE73B3AA6CEF" author="Ochoa" etAl="et al." firstAuthor="Ochoa" journalOrPublisher="American Museum Novitates" pageId="116" pageNumber="116" pagination="1 - 19" part="3691" refId="ref69663" refString="Ochoa, J. A., R. Botero-Trujillo, and L. Prendini. 2010. On the troglomorphic scorpion Troglotayosicus humiculum (Scorpiones, Troglotayosicidae), with first description of the adults. American Museum Novitates 3691: 1 - 19." title="On the troglomorphic scorpion Troglotayosicus humiculum (Scorpiones, Troglotayosicidae), with first description of the adults" type="journal article" year="2010">Ochoa et al., 2010: 17</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FFBEB1203962CEAEB1546CEF" author="Ubinski" box="[258,520,642,666]" etAl="et al." firstAuthor="Ubinski" journalOrPublisher="Cytogenetic and Genome Research" pageId="116" pageNumber="116" pagination="83 - 155" part="148" refId="ref72292" refString="Ubinski, C. V., L. S. Carvalho, and M. C. Schneider. 2016. Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil. Cytogenetic and Genome Research 148: 83 - 155. [doi: 10.1159 / 000446523]" title="Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil" type="journal article" year="2016">Ubinski et al., 2016: 122</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBEB120380BCE88B2326AD4" blockId="116.[107,634,399,1185]" pageId="116" pageNumber="116">
|
||
<taxonomicName id="C0CC27C2FFBEB120380BCE88B3B86CC9" ID-CoL="63QFB" authorityName="Thorell" authorityYear="1876" box="[107,228,676,700]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB120380BCE88B3B86CC9" box="[107,228,676,700]" italics="true" pageId="116" pageNumber="116">Rhopalurus</emphasis>
|
||
</taxonomicName>
|
||
(part): Lourenço and Pinto-da- Rocha, 1997: 182–188, 191, figs. 1–3, 5, 7, 9, 11, 14;
|
||
<bibRefCitation id="635D21B0FFBEB1203889CEC6B2EF6D77" author="Kovarik" bookContentInfo="175 pp." box="[233,435,746,770]" firstAuthor="Kovarik" journalOrPublisher="Madagaskar: Jihlava" pageId="116" pageNumber="116" refId="ref66846" refString="Kovarik, F. 1998. Stiri [Scorpions]. Madagaskar: Jihlava. 175 pp. [in Czech]" title="Stiri [Scorpions]" type="book" year="1998">Kovařík, 1998: 118</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FFBEB12039DFCEC6B2586D51" author="Fet and Lowe" editor="V. Fet & W. D. Sissom" firstAuthor="Fet" journalOrPublisher="New York: New York Entomological Society" pageId="116" pageNumber="116" pagination="54 - 286" refId="ref65304" refString="Fet, V., and G. Lowe. 2000. Family Buthidae C. L. Koch, 1837. In V. Fet, W. D. Sissom, G., Lowe, and M. E. Braunwalder, Catalog of the scorpions of the world (1758 - 1998): 54 - 286. New York: New York Entomological Society." title="Family Buthidae C. L. Koch, 1837" type="book chapter" volumeTitle="G., Lowe, and M. E" year="2000">Fet and Lowe, 2000: 220</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FFBEB120396FCF21B2B66D51" author="Lourenco" bookContentInfo="306 pp." box="[271,490,780,804]" firstAuthor="Lourenco" journalOrPublisher="Paris: Les Editions de l'If" pageId="116" pageNumber="116" refId="ref68189" refString="Lourenco, W. R. 2002. Scorpions of Brazil. Paris: Les Editions de l'If. 306 pp." title="Scorpions of Brazil" type="book" year="2002">Lourenço, 2002: 102</bibRefCitation>
|
||
, 111, figs. 228–231;
|
||
<bibRefCitation id="635D21B0FFBEB1203963CF03B1166D32" author="Manzanilla and Sousa" box="[259,586,815,839]" firstAuthor="Manzanilla" journalOrPublisher="Saber" pageId="116" pageNumber="116" pagination="3 - 14" part="15" refId="ref68993" refString="Manzanilla, J., and L. Sousa. 2003. Ecologia y distribucion de Rhopalurus laticauda Thorell, 1876 (Scorpiones: Buthidae) en Venezuela. Saber 15 (1): 3 - 14." title="Ecologia y distribucion de Rhopalurus laticauda Thorell, 1876 (Scorpiones: Buthidae) en Venezuela" type="journal article" year="2003">Manzanilla and Sousa, 2003: 3</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FFBEB12038FBCF7EB2E26D1C" author="Lourenco" box="[155,446,849,874]" etAl="et al." firstAuthor="Lourenco" journalOrPublisher="Comptes Rendus Biologies" pageId="116" pageNumber="116" pagination="1151 - 1156" part="327" refId="ref68662" refString="Lourenco, W. R., R. L. C. Baptista, and A. P. L. Giupponi. 2004. Troglobitic scorpions: a new genus and species from Brazil. Comptes Rendus Biologies 327: 1151 - 1156." title="Troglobitic scorpions: a new genus and species from Brazil" type="journal article" year="2004">Lourenço et al., 2004: 1152</bibRefCitation>
|
||
, 1156; Lenar- ducci et al., 2005: 1, 2, 7;
|
||
<bibRefCitation id="635D21B0FFBEB12039CBCF58B3B56DDA" author="Lira-da-Silva" etAl="et al." firstAuthor="Lira-da-Silva" journalOrPublisher="Biota Neotropica" pageId="116" pageNumber="116" pagination="1 - 3" part="5" refId="ref67409" refString="Lira-da-Silva, R. M., G. M. Jordao, T. F. Silva, D. M. Candido, and T. K. Brazil. 2005. Ocorrencia de Rhopalurus debilis (C. L. Koch, 1840) (Scorpiones, Buthidae) no estado da Bahia, Brasil. Biota Neotropica 5 (1 A): 1 - 3." title="Ocorrencia de Rhopalurus debilis (C. L. Koch, 1840) (Scorpiones, Buthidae) no estado da Bahia, Brasil" type="journal article" year="2005">Lira-da-Silva et al., 2005: 2</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FFBEB1203895CFBBB2FF6DDA" author="Teruel" box="[245,419,919,943]" firstAuthor="Teruel" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="116" pageNumber="116" pagination="43 - 56" part="38" refId="ref71569" refString="Teruel, R. 2006. Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba. Boletin de la Sociedad Entomologica Aragonesa 38: 43 - 56." title="Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba" type="journal article" year="2006">Teruel, 2006: 52</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FFBEB12039CDCFBBB39F6DA4" author="Lourenco" firstAuthor="Lourenco" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="116" pageNumber="116" pagination="359 - 365" part="40" refId="ref68211" refString="Lourenco, W. R. 2007. New considerations on the taxonomic status of the genus Physoctonus Mello-Leitao, 1934 (Scorpiones, Buthidae). Boletin de la Sociedad Entomologica Aragonesa 40: 359 - 365." title="New considerations on the taxonomic status of the genus Physoctonus Mello-Leitao, 1934 (Scorpiones, Buthidae)" type="journal article" year="2007">Lourenço, 2007: 359</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FFBEB12038AECF95B2416DA4" author="Lourenco" box="[206,285,953,977]" firstAuthor="Lourenco" journalOrPublisher="Euscorpius" pageId="116" pageNumber="116" pagination="1 - 14" part="73" refId="ref68250" refString="Lourenco, W. R. 2008. The geographic pattern of distribution of the genus Rhopalurus Thorell, 1876 in the Guayana-Amazon region (Scorpiones: Buthidae). Euscorpius 73: 1 - 14." title="The geographic pattern of distribution of the genus Rhopalurus Thorell, 1876 in the Guayana-Amazon region (Scorpiones: Buthidae)" type="journal article" year="2008">2008: 3</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FFBEB1203949CF95B2246DA4" author="Lourenco" box="[297,376,953,977]" firstAuthor="Lourenco" journalOrPublisher="Euscorpius" pageId="116" pageNumber="116" pagination="1 - 14" part="73" refId="ref68250" refString="Lourenco, W. R. 2008. The geographic pattern of distribution of the genus Rhopalurus Thorell, 1876 in the Guayana-Amazon region (Scorpiones: Buthidae). Euscorpius 73: 1 - 14." title="The geographic pattern of distribution of the genus Rhopalurus Thorell, 1876 in the Guayana-Amazon region (Scorpiones: Buthidae)" type="journal article" year="2008">2008: 3</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FFBEB12039E4CF95B3986D81" author="Prendini" etAl="et al." firstAuthor="Prendini" journalOrPublisher="Journal of Arachnology" pageId="116" pageNumber="116" pagination="206 - 224" part="37" refId="ref70522" refString="Prendini, L., L. A. Esposito, J. Huff, and E. S. Volschenk. 2009. Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola. Journal of Arachnology 37: 206 - 224." title="Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola" type="journal article" year="2009">Prendini et al., 2009: 222</bibRefCitation>
|
||
, 223;
|
||
<bibRefCitation id="635D21B0FFBEB1203962CFF0B1026D81" author="Outeda-Jorge" box="[258,606,988,1012]" etAl="et al." firstAuthor="Outeda-Jorge" journalOrPublisher="Zoologia" pageId="116" pageNumber="116" pagination="43 - 53" part="26" refId="ref69741" refString="Outeda-Jorge, S., T. Mello, and R. Pinto-da-Rocha. 2009. Litter size, effects of maternal body size, and date of birth in South American scorpions (Arachnida: Scorpiones). Zoologia 26 (1): 43 - 53." title="Litter size, effects of maternal body size, and date of birth in South American scorpions (Arachnida: Scorpiones)" type="journal article" year="2009">Outeda-Jorge et al., 2009: 44–46</bibRefCitation>
|
||
, 48, 49;
|
||
<bibRefCitation id="635D21B0FFBEB1203889CFD3B15E6A63" author="Brazil and Porto" bookContentInfo="84 pp." box="[233,514,1022,1047]" firstAuthor="Brazil" journalOrPublisher="Salvador, BA, Brazil: EDUFBA" pageId="116" pageNumber="116" refId="ref64569" refString="Brazil, T. K., and Porto, T. J. 2010. Os escorpioes. Salvador, BA, Brazil: EDUFBA. 84 pp." title="Os escorpioes" type="book" year="2010">Brazil and Porto, 2010: 50</bibRefCitation>
|
||
, 62;
|
||
<bibRefCitation id="635D21B0FFBEB1203A53CFD3B2246A4C" author="Porto" etAl="et al." firstAuthor="Porto" journalOrPublisher="Check List" pageId="116" pageNumber="116" pagination="292 - 297" part="6" refId="ref70138" refString="Porto, T. J., T. K. Brazil, and R. M. Lira-da-Silva. 2010. Scorpions, state of Bahia, northeastern Brazil. Check List 6 (2): 292 - 297." title="Scorpions, state of Bahia, northeastern Brazil" type="journal article" year="2010">Porto et al., 2010: 293–295</bibRefCitation>
|
||
,
|
||
<tableCitation id="4A4E69FAFFBEB12039E4C80DB2976A4C" box="[388,459,1057,1081]" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="116" pageNumber="116" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">table 1</tableCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FFBEB12039B8C80EB2426A29" author="Lourenco" firstAuthor="Lourenco" journalOrPublisher="Acta Biologica Paranaense" pageId="116" pageNumber="116" pagination="69 - 76" part="43" refId="ref68287" refString="Lourenco, W. R. 2014. The genus Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) in northeast Brazil; a possible case of a vicariant species. Acta Biologica Paranaense 43: 69 - 76." title="The genus Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) in northeast Brazil; a possible case of a vicariant species" type="journal article" year="2014">Lourenço, 2014: 69–75</bibRefCitation>
|
||
, figs. 1–12;
|
||
<bibRefCitation id="635D21B0FFBEB12039FDC868B2076A0A" author="Gallao and Bichuette" firstAuthor="Gallao" journalOrPublisher="Zoologia" pageId="116" pageNumber="116" pagination="1 - 13" part="33" refId="ref65887" refString="Gallao, J. E., and M. E. Bichuette. 2016. On the enigmatic troglobitic scorpion Troglorhopalurus translucidus: distribution, description of adult females, life history and comments on Rhopalurus lacrau (Scorpiones: Buthidae). Zoologia 33 (6): 1 - 13." title="On the enigmatic troglobitic scorpion Troglorhopalurus translucidus: distribution, description of adult females, life history and comments on Rhopalurus lacrau (Scorpiones: Buthidae)" type="journal article" year="2016">Gallão and Bichuette, 2016: 2</bibRefCitation>
|
||
, 3, 9–11, figs. 1, 18;
|
||
<bibRefCitation id="635D21B0FFBEB1203A55C84AB2366AD4" author="Ubinski" etAl="et al." firstAuthor="Ubinski" journalOrPublisher="Cytogenetic and Genome Research" pageId="116" pageNumber="116" pagination="83 - 155" part="148" refId="ref72292" refString="Ubinski, C. V., L. S. Carvalho, and M. C. Schneider. 2016. Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil. Cytogenetic and Genome Research 148: 83 - 155. [doi: 10.1159 / 000446523]" title="Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil" type="journal article" year="2016">Ubinski et al., 2016: 122</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFFBEB12038EBC891B06D6C99" pageId="116" pageNumber="116" type="diagnosis">
|
||
<paragraph id="07735C41FFBEB12038EBC891B06D6C99" blockId="116.[107,636,1213,1741]" lastBlockId="116.[684,1214,228,1740]" pageId="116" pageNumber="116">
|
||
DIAGNOSIS:
|
||
<taxonomicName id="C0CC27C2FFBEB120397DC891B2806AA0" authorityName="Lourenco et al., 2004:" authorityYear="1151" box="[285,476,1213,1237]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB120397DC891B2806AA0" box="[285,476,1213,1237]" italics="true" pageId="116" pageNumber="116">Troglorhopalurus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="C0CC27C2FFBEB120380BC8CCB23D6A82" authority="Jaguajir" authorityName=", Ischnotelson, Jaguajir" authorityYear="2017" box="[107,353,1247,1271]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="35B88053FFBEB120380BC8CCB3A56A82" box="[107,249,1248,1271]" italics="true" pageId="116" pageNumber="116">Heteroctenus</emphasis>
|
||
,
|
||
<emphasis id="35B88053FFBEB120396BC8CCB23D6A82" box="[267,353,1248,1271]" italics="true" pageId="116" pageNumber="116">Jaguajir</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="2E8B3D28FFBEB1203914C8F3B2836A83" box="[372,479,1247,1271]" pageId="116" pageNumber="116" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="C0CC27C2FFBEB1203990C8F3B12A6A82" authorityName="Mello-Leitao" authorityYear="1934" box="[496,630,1247,1271]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB1203990C8F3B12A6A82" box="[496,630,1247,1271]" italics="true" pageId="116" pageNumber="116">Physoctonus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="C0CC27C2FFBEB12038FAC92CB2496B6D" authorityName="Thorell" authorityYear="1876" box="[154,277,1280,1304]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB12038FAC92CB2496B6D" box="[154,277,1280,1304]" italics="true" pageId="116" pageNumber="116">Rhopalurus</emphasis>
|
||
</taxonomicName>
|
||
by the proximal dentate margins of the chela fixed and movable fingers of the adult male that are linear, with no gap evident between them, when closed; from
|
||
<taxonomicName id="C0CC27C2FFBEB120398BC94AB3E26BEB" authority=", Jaguajir" authorityName=", Ischnotelson, Jaguajir" authorityYear="2017" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB120398BC94AB1296B08" box="[491,629,1382,1405]" italics="true" pageId="116" pageNumber="116">Heteroctenus</emphasis>
|
||
,
|
||
<emphasis id="35B88053FFBEB120380BC9ABB3E26BEB" box="[107,190,1415,1438]" italics="true" pageId="116" pageNumber="116">Jaguajir</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="C0CC27C2FFBEB120389AC9ABB2296BEA" authorityName="Thorell" authorityYear="1876" box="[250,373,1415,1439]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB120389AC9ABB2296BEA" box="[250,373,1415,1439]" italics="true" pageId="116" pageNumber="116">Rhopalurus</emphasis>
|
||
</taxonomicName>
|
||
by the absence of a pecten-sternite stridulatory organ; from
|
||
<taxonomicName id="C0CC27C2FFBEB1203A74C984B3D36B97" authority="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" class="Arachnida" family="Buthidae" genus="Ischnotelson" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="35B88053FFBEB1203A74C984B3D36B97" italics="true" pageId="116" pageNumber="116">Ischnotelson</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="2E8B3D28FFBEB12038C0C9E6B2546B94" box="[160,264,1482,1506]" pageId="116" pageNumber="116" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="C0CC27C2FFBEB1203929C9E6B29A6B97" authorityName="Thorell" authorityYear="1876" box="[329,454,1482,1506]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB1203929C9E6B29A6B97" box="[329,454,1482,1506]" italics="true" pageId="116" pageNumber="116">Rhopalurus</emphasis>
|
||
</taxonomicName>
|
||
by the separate (unfused) central lateral and posterior central submedian carinae of the carapace; from
|
||
<taxonomicName id="C0CC27C2FFBEB1203A21CA22B39A6833" authorityName=", Ischnotelson, Jaguajir" authorityYear="2017" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB1203A21CA22B39A6833" italics="true" pageId="116" pageNumber="116">Heteroctenus</emphasis>
|
||
</taxonomicName>
|
||
by the presence of two lateral depressions in the male pectinal plate, and a subaculear tubercle on the telson; from
|
||
<taxonomicName id="C0CC27C2FFBEB12039CCCA5DB17168FC" authority="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[428,557,1649,1673]" class="Arachnida" family="Buthidae" genus="Ischnotelson" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="35B88053FFBEB12039CCCA5DB17168FC" box="[428,557,1649,1673]" italics="true" pageId="116" pageNumber="116">Ischnotelson</emphasis>
|
||
</taxonomicName>
|
||
by the separate (unfused) lateral ocular and central lateral carinae of the carapace and the telson vesicle not being laterally compressed; from
|
||
<taxonomicName id="C0CC27C2FFBEB1203C28CCC9B7C76E89" authority="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[1096,1179,229,252]" class="Arachnida" family="Buthidae" genus="Jaguajir" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="35B88053FFBEB1203C28CCC9B7C76E89" box="[1096,1179,229,252]" italics="true" pageId="116" pageNumber="116">Jaguajir</emphasis>
|
||
</taxonomicName>
|
||
by the separate (unfused) lateral ocular and anterior central submedian carinae of the carapace; from
|
||
<taxonomicName id="C0CC27C2FFBEB1203ACCCD6BB0736F2A" authorityName="Mello-Leitao" authorityYear="1934" box="[684,815,327,351]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB1203ACCCD6BB0736F2A" box="[684,815,327,351]" italics="true" pageId="116" pageNumber="116">Physoctonus</emphasis>
|
||
</taxonomicName>
|
||
by the larger size (35–40 mm), the more distinct carapacial carinae, the setose proximal dorsal fulcra of the pectines, the bifurcate prolateral pedal spur of leg I, and the oblique subrows of primary denticles on the pedipalp chela fingers flanked closely by pro- and retrolateral accessory (supernumerary) denticles; from
|
||
<taxonomicName id="C0CC27C2FFBEB1203ACCCE03B0726C32" authorityName="Thorell" authorityYear="1876" box="[684,814,559,583]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB1203ACCCE03B0726C32" box="[684,814,559,583]" italics="true" pageId="116" pageNumber="116">Rhopalurus</emphasis>
|
||
</taxonomicName>
|
||
by the slender metasoma, not increasing in width posteriorly; and from
|
||
<taxonomicName id="C0CC27C2FFBEB1203CE9CE7CB0576CFC" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & CLOSE Lorenzo Prendini Scorpion Systematics Research Group" authorityYear="2017" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB1203CE9CE7CB0576CFC" italics="true" pageId="116" pageNumber="116">Centruroides</emphasis>
|
||
</taxonomicName>
|
||
by the obsolete retromedian carina on the pedipalp chela manus, and the well-separated dorsointernal and proventral carinae of the pedipalp patella.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFFBEB1223AACCEDAB7426B6B" lastPageId="118" lastPageNumber="118" pageId="116" pageNumber="116" type="description">
|
||
<paragraph id="07735C41FFBEB1203AACCEDAB0DB6D3A" blockId="116.[684,1214,228,1740]" pageId="116" pageNumber="116">
|
||
DESCRIPTION: The following general description outlines characters common to both species of
|
||
<taxonomicName id="C0CC27C2FFBEB1203AA9CF1BB0DC6D3A" authorityName="Lourenco et al., 2004:" authorityYear="1151" box="[713,896,823,847]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBEB1203AA9CF1BB0DC6D3A" box="[713,896,823,847]" italics="true" pageId="116" pageNumber="116">Troglorhopalurus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBEB1203AACCF74B0ED6DC7" blockId="116.[684,1214,228,1740]" pageId="116" pageNumber="116">
|
||
<emphasis id="35B88053FFBEB1203AACCF74B0106D05" box="[716,844,856,880]" italics="true" pageId="116" pageNumber="116">Total length</emphasis>
|
||
: Medium-sized, gracile scorpions (total length, 35–40 mm) with soma slightly dorsoventrally compressed.
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBEB1203AACCF90B7746B4A" blockId="116.[684,1214,228,1740]" pageId="116" pageNumber="116">
|
||
<emphasis id="35B88053FFBEB1203AACCF90B0526DA1" box="[716,782,956,980]" italics="true" pageId="116" pageNumber="116">Color:</emphasis>
|
||
Base color predominantly pale brown with pigmentation somewhat reduced, especially in the immature stages (fig. 1G–H). Carapace and tergites similar to or slightly darker than chelicerae, pedipalps, legs, sternites, metasoma and telson vesicle; pedipalp chela fingers darker than chela manus, patella, and femur, carinae darker than intercarinal surfaces; pectines pale yellow; metasomal segments IV and V darker than preceding segments in
|
||
<taxonomicName id="C0CC27C2FFBEB1203BE9C8CAB0B46A88" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" baseAuthorityYear="2017" box="[905,1000,1253,1277]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="species" species="lacrau" status="comb. nov.">
|
||
<emphasis id="35B88053FFBEB1203BE9C8CAB0B46A88" box="[905,1000,1253,1277]" italics="true" pageId="116" pageNumber="116">T. lacrau</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="2E8B3D28FFBEB1203B98C8C9B72D6A88" box="[1016,1137,1253,1277]" pageId="116" pageNumber="116">comb. nov.</taxonomicNameLabel>
|
||
; metasomal carinae darker than intercarinal surfaces; telson aculeus dark brown to black.
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBEB1203AACC964B7D86BF4" blockId="116.[684,1214,228,1740]" pageId="116" pageNumber="116">
|
||
<emphasis id="35B88053FFBEB1203AACC964B0676B15" box="[716,827,1352,1376]" italics="true" pageId="116" pageNumber="116">Chelicerae</emphasis>
|
||
: Base, dorsal surface with medial transverse row of well-developed tubercles.
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBEB1203AACC9A7B74E68B9" blockId="116.[684,1214,228,1740]" pageId="116" pageNumber="116">
|
||
<emphasis id="35B88053FFBEB1203AACC9A7B06D6BD7" box="[716,817,1419,1442]" italics="true" pageId="116" pageNumber="116">Carapace</emphasis>
|
||
: Median ocular tubercle low (figs. 14F, 15E); two median ocelli reduced, more so in
|
||
<taxonomicName id="C0CC27C2FFBEB1203ACCC9E1B0146B90" box="[684,840,1485,1509]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="116" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBEB1203ACCC9E1B0146B90" box="[684,840,1485,1509]" italics="true" pageId="116" pageNumber="116">T. translucidus</emphasis>
|
||
</taxonomicName>
|
||
; three pairs of lateral macroocelli; one pair of lateral microocelli. Anteromedian, median ocular, and posteromedian sulci well developed, forming single, almost continuous, longitudinal sulcus. Lateral ocular, central lateral, anterior central submedian and posterior central submedian carinae somewhat indistinct, finely granular and separate (unfused).
|
||
</paragraph>
|
||
<caption id="53B30CC9FFBFB121380CCAA2B2B468AA" httpUri="https://zenodo.org/record/851508/files/figure.png" pageId="117" pageNumber="117" targetBox="[108,1212,224,1650]" targetPageId="117">
|
||
<paragraph id="07735C41FFBFB121380CCAA2B2B468AA" blockId="117.[108,1214,1676,1759]" pageId="117" pageNumber="117">
|
||
FIG. 63.
|
||
<taxonomicName id="C0CC27C2FFBFB12138A8CAA0B10468D0" authority="Lourenco et al., 2004" authorityName="Lourenco et al." authorityYear="2004" box="[200,600,1676,1701]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="117" pageNumber="117" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBFB12138A8CAA0B22F68D1" box="[200,371,1676,1700]" italics="true" pageId="117" pageNumber="117">Troglorhopalurus</emphasis>
|
||
<bibRefCitation id="635D21B0FFBFB121391BCAA1B10468D0" author="Lourenco" box="[379,600,1676,1701]" etAl="et al." firstAuthor="Lourenco" journalOrPublisher="Comptes Rendus Biologies" pageId="117" pageNumber="117" pagination="1151 - 1156" part="327" refId="ref68662" refString="Lourenco, W. R., R. L. C. Baptista, and A. P. L. Giupponi. 2004. Troglobitic scorpions: a new genus and species from Brazil. Comptes Rendus Biologies 327: 1151 - 1156." title="Troglobitic scorpions: a new genus and species from Brazil" type="journal article" year="2004">Lourenço et al., 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
, habitus, dorsal (
|
||
<emphasis id="35B88053FFBFB1213B6ACAA0B06768D1" bold="true" box="[778,827,1676,1700]" pageId="117" pageNumber="117">A, C</emphasis>
|
||
) and ventral (
|
||
<emphasis id="35B88053FFBFB1213BB2CAA1B75868D0" bold="true" box="[978,1028,1677,1701]" pageId="117" pageNumber="117">B, D</emphasis>
|
||
) aspects.
|
||
<emphasis id="35B88053FFBFB1213C0CCAA0B7FE68D0" bold="true" box="[1132,1186,1676,1701]" pageId="117" pageNumber="117">A, B.</emphasis>
|
||
<taxonomicName id="C0CC27C2FFBFB1213CCACAA1B3F668B7" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" baseAuthorityYear="2017" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="117" pageNumber="117" phylum="Arthropoda" rank="species" species="lacrau">
|
||
<emphasis id="35B88053FFBFB1213CCACAA1B3F668B7" italics="true" pageId="117" pageNumber="117">T. lacrau</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="635D21B0FFBFB12138D8CA86B16D68B7" author="Lourenco and Pinto-da-Rocha" box="[184,561,1706,1730]" firstAuthor="Lourenco" journalOrPublisher="Biogeographica (Paris) 73" pageId="117" pageNumber="117" pagination="181 - 191" part="4" refId="ref68476" refString="Lourenco, W. R., and R. Pinto-da-Rocha. 1997. A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species. Biogeographica (Paris) 73 (4): 181 - 191." title="A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species" type="journal article" year="1997">Lourenço and Pinto-da-Rocha, 1997</bibRefCitation>
|
||
), comb. nov., ♀ (AMNH).
|
||
<emphasis id="35B88053FFBFB1213B27CA86B02268B7" bold="true" box="[839,894,1706,1730]" pageId="117" pageNumber="117">C, D.</emphasis>
|
||
<taxonomicName id="C0CC27C2FFBFB1213BE5CA87B3C368AA" authority="Lourenco et al., 2004" authorityName="Lourenco et al." authorityYear="2004" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="117" pageNumber="117" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBFB1213BE5CA87B74468B7" box="[901,1048,1706,1730]" italics="true" pageId="117" pageNumber="117">T. translucidus</emphasis>
|
||
<bibRefCitation id="635D21B0FFBFB1213C7FCA86B3C368AA" author="Lourenco" etAl="et al." firstAuthor="Lourenco" journalOrPublisher="Comptes Rendus Biologies" pageId="117" pageNumber="117" pagination="1151 - 1156" part="327" refId="ref68662" refString="Lourenco, W. R., R. L. C. Baptista, and A. P. L. Giupponi. 2004. Troglobitic scorpions: a new genus and species from Brazil. Comptes Rendus Biologies 327: 1151 - 1156." title="Troglobitic scorpions: a new genus and species from Brazil" type="journal article" year="2004">Lourenço et al., 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
, ♀ (MZSP). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="07735C41FFBCB12238EBCCC8B2696DE4" blockId="118.[107,637,228,1740]" pageId="118" pageNumber="118">
|
||
<emphasis id="35B88053FFBCB12238EBCCC8B3AD6E89" box="[139,241,228,252]" italics="true" pageId="118" pageNumber="118">Pedipalps</emphasis>
|
||
: Pedipalp femur retrolateral accessory carinae absent. Pedipalp chela manus of males incrassate in
|
||
<taxonomicName id="C0CC27C2FFBCB1223920CD0BB2C36F4B" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" baseAuthorityYear="2017" box="[320,415,294,318]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="species" species="lacrau">
|
||
<emphasis id="35B88053FFBCB1223920CD0BB2C36F4B" box="[320,415,294,318]" italics="true" pageId="118" pageNumber="118">T. lacrau</emphasis>
|
||
</taxonomicName>
|
||
, slender in
|
||
<taxonomicName id="C0CC27C2FFBCB1223A41CD0BB3E46F2A" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBCB1223A41CD0BB3E46F2A" italics="true" pageId="118" pageNumber="118">T. translucidus</emphasis>
|
||
</taxonomicName>
|
||
(fig. 61), fixed and movable fingers not curved, such that proximal dentate margin linear, no gap present between fingers proximally, when closed; manus, proventral carina present, promedian carina absent; fixed and movable fingers, median denticle rows each comprising eight (
|
||
<taxonomicName id="C0CC27C2FFBCB1223A08CDC1B3F26C53" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" baseAuthorityYear="2017" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="species" species="lacrau">
|
||
<emphasis id="35B88053FFBCB1223A08CDC1B3F26C53" italics="true" pageId="118" pageNumber="118">T. lacrau</emphasis>
|
||
</taxonomicName>
|
||
) or nine (
|
||
<taxonomicName id="C0CC27C2FFBCB122397DCE22B2E66C53" box="[285,442,526,550]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBCB122397DCE22B2E66C53" box="[285,442,526,550]" italics="true" pageId="118" pageNumber="118">T. translucidus</emphasis>
|
||
</taxonomicName>
|
||
) oblique subrows of primary denticles flanked closely by pro- and retrolateral accessory (supernumerary) denticles, smaller and more sparse in
|
||
<taxonomicName id="C0CC27C2FFBCB12239F8CE5EB1696CFC" box="[408,565,625,649]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBCB12239F8CE5EB1696CFC" box="[408,565,625,649]" italics="true" pageId="118" pageNumber="118">T. translucidus</emphasis>
|
||
</taxonomicName>
|
||
; movable finger without proximal lobe (
|
||
<figureCitation id="9FF740C4FFBCB1223A70CEBEB1026CDF" box="[528,606,658,682]" captionStart="FIG. 17" captionStartId="36.[106,149,1153,1174]" captionTargetBox="[106,634,224,1125]" captionTargetId="figure@36.[106,634,224,1125]" captionTargetPageId="36" captionText="FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851416/files/figure.png" pageId="118" pageNumber="118">fig. 17</figureCitation>
|
||
F). Pedipalps orthobothriotaxic Type A, α configuration; femur with five dorsal trichobothria, trichobothrium
|
||
<emphasis id="35B88053FFBCB122397CCED9B2766D78" box="[284,298,757,781]" italics="true" pageId="118" pageNumber="118">d</emphasis>
|
||
<subScript id="9B485E04FFBCB122394ACF2CB26F6D64" attach="left" box="[298,307,768,785]" fontSize="7" pageId="118" pageNumber="118">2</subScript>
|
||
situated on prolateral surface; patella trichobothrium
|
||
<emphasis id="35B88053FFBCB122390BCF3AB2256D5B" box="[363,377,790,814]" italics="true" pageId="118" pageNumber="118">d</emphasis>
|
||
<subScript id="9B485E04FFBCB1223919CF0DB2DE6D47" attach="left" box="[377,386,801,818]" fontSize="7" pageId="118" pageNumber="118">3</subScript>
|
||
situated retrolateral to dorsomedian carina; chela fixed finger trichobothrium
|
||
<emphasis id="35B88053FFBCB122395ACF74B20A6D05" box="[314,342,856,880]" italics="true" pageId="118" pageNumber="118">db</emphasis>
|
||
situated proximal to trichobothrium
|
||
<emphasis id="35B88053FFBCB122397CCF56B2736DE4" box="[284,303,890,913]" italics="true" pageId="118" pageNumber="118">et</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBCB12238EBCFB7B11B6A60" blockId="118.[107,637,228,1740]" pageId="118" pageNumber="118">
|
||
<emphasis id="35B88053FFBCB12238EBCFB7B3E56DC7" box="[139,185,923,946]" italics="true" pageId="118" pageNumber="118">Legs</emphasis>
|
||
: Legs III and IV, tibial spurs absent; I–IV, basitarsi each with bifurcate prolateral pedal spur; telotarsi each with distinct pro- and retroventral rows of fine, acuminate macrosetae.
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBCB12238EBC80CB16B6B4A" blockId="118.[107,637,228,1740]" pageId="118" pageNumber="118">
|
||
<emphasis id="35B88053FFBCB12238EBC80CB3826A42" box="[139,222,1056,1079]" italics="true" pageId="118" pageNumber="118">Pectines</emphasis>
|
||
: Pectinal plate with two lateral depressions (male), anterior margin with or without sulcus (figs. 18F, 19E). Pectines not proximally expanded; proximal dorsal fulcra setose; pectinal teeth almost straight, slightly curved laterally, proximal teeth, dorsal surfaces without nodules and with irregular striations (figs. 11F, 12D), dorsobasal surfaces with or without macrosetae; pectinal sensillae elongate and acuminate.
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBCB12238EBC965B7656F4B" blockId="118.[107,637,228,1740]" lastBlockId="118.[684,1214,228,1740]" pageId="118" pageNumber="118">
|
||
<emphasis id="35B88053FFBCB12238EBC965B3A16B15" box="[139,253,1353,1376]" italics="true" pageId="118" pageNumber="118">Mesosoma</emphasis>
|
||
: Tergites IV and VI wider than than I–III and VII; dorsosubmedian carinae vestigial or absent, dorsomedian carinae finely granular, vestigial, restricted to posterior margins of segments I–V, complete on VI (fig. 63). Tergite VII pentacarinate, dorsomedian carina restricted to anterior half of segment. Sternites III–V smooth, carinae absent or obsolete, ventromedian carina present on VI and VII; sternite III, lateral margins not forming smooth, raised carina, ventromedian carina not elevated anteriorly, ventrosubmedian surfaces not forming paired depressions, finely and irregularly granular; respiratory spiracles (stigmata) small and short, width ca. 2× length (fig. 10F).
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBCB1223AACCD64B0576DC7" blockId="118.[684,1214,228,1740]" pageId="118" pageNumber="118">
|
||
<emphasis id="35B88053FFBCB1223AACCD64B0666F2A" box="[716,826,328,351]" italics="true" pageId="118" pageNumber="118">Metasoma</emphasis>
|
||
: Metasoma slender, not increasing in width posteriorly, I and V similar width in both sexes (fig. 62). Segments I and II each with 10 distinct, costate-granular carinae, III and IV each with eight distinct, costate-granular carinae, V with seven distinct but less pronounced, granular carinae; dorsosubmedian carinae absent or obsolete, reduced to rows of granules on dorsal surfaces of segments I–IV; dorsolateral carinae complete on segments I–IV, often terminating in prominent, spiniform granules posteriorly on III and IV, absent on V; lateral supramedian carinae complete on segments I–V; lateral inframedian carinae complete on segment I, partial on II, absent on III–V; ventrosubmedian carinae complete on segments I–IV, restricted to anterior third on V; ventromedian carina absent on segments I–IV, complete on V. Intercarinal surfaces finely granular.
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBCB1223AACCF90B0DC6AEF" blockId="118.[684,1214,228,1740]" pageId="118" pageNumber="118">
|
||
<emphasis id="35B88053FFBCB1223AACCF90B0516DA1" box="[716,781,956,980]" italics="true" pageId="118" pageNumber="118">Telson</emphasis>
|
||
: Vesicle slightly elongate, not laterally compressed, width similar to metasoma V width; anterodorsal lateral lobes reduced or absent; lateral and ventral surfaces granular (
|
||
<taxonomicName id="C0CC27C2FFBCB1223C59C80CB18F6A2D" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBCB1223C59C80CB18F6A2D" italics="true" pageId="118" pageNumber="118">T. translucidus</emphasis>
|
||
</taxonomicName>
|
||
) or smooth (
|
||
<taxonomicName id="C0CC27C2FFBCB1223BEEC86DB0A26A2D" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" baseAuthorityYear="2017" box="[910,1022,1088,1112]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="species" species="lacrau">
|
||
<emphasis id="35B88053FFBCB1223BEEC86DB0A26A2D" box="[910,1022,1088,1112]" italics="true" pageId="118" pageNumber="118">T. lacrau</emphasis>
|
||
</taxonomicName>
|
||
), with distinct ventromedian carina; subaculear tubercle well developed, spinoid.
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBCB1223AACC88FB77A6ACE" blockId="118.[684,1214,228,1740]" box="[716,1062,1187,1211]" pageId="118" pageNumber="118">
|
||
<emphasis id="35B88053FFBCB1223AACC88FB0F86ACE" box="[716,932,1187,1211]" italics="true" pageId="118" pageNumber="118">Hemispermatophore</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
<paragraph id="07735C41FFBCB1223AACC8E9B7426B6B" blockId="118.[684,1214,228,1740]" pageId="118" pageNumber="118">
|
||
<emphasis id="35B88053FFBCB1223AACC8E9B00C6AA9" box="[716,848,1221,1244]" italics="true" pageId="118" pageNumber="118">Cytogenetics</emphasis>
|
||
: The diploid chromosome number of
|
||
<taxonomicName id="C0CC27C2FFBCB1223A94C8CAB00E6A88" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" baseAuthorityYear="2017" box="[756,850,1253,1277]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="species" species="lacrau">
|
||
<emphasis id="35B88053FFBCB1223A94C8CAB00E6A88" box="[756,850,1253,1277]" italics="true" pageId="118" pageNumber="118">T. lacrau</emphasis>
|
||
</taxonomicName>
|
||
is 2n = 20 and of
|
||
<taxonomicName id="C0CC27C2FFBCB1223C7AC8CAB7EB6A88" box="[1050,1207,1253,1277]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBCB1223C7AC8CAB7EB6A88" box="[1050,1207,1253,1277]" italics="true" pageId="118" pageNumber="118">T. translucidus</emphasis>
|
||
</taxonomicName>
|
||
, 2n = 20–22 (
|
||
<bibRefCitation id="635D21B0FFBCB1223B58C92AB74E6B6B" author="Ubinski" box="[824,1042,1286,1310]" etAl="et al." firstAuthor="Ubinski" journalOrPublisher="Cytogenetic and Genome Research" pageId="118" pageNumber="118" pagination="83 - 155" part="148" refId="ref72292" refString="Ubinski, C. V., L. S. Carvalho, and M. C. Schneider. 2016. Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil. Cytogenetic and Genome Research 148: 83 - 155. [doi: 10.1159 / 000446523]" title="Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil" type="journal article" year="2016">Ubinski et al., 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFFBCB1223AACC904B07F6BD7" pageId="118" pageNumber="118" type="discussion">
|
||
<paragraph id="07735C41FFBCB1223AACC904B07F6BD7" blockId="118.[684,1214,228,1740]" pageId="118" pageNumber="118">
|
||
INCLUDED SPECIES:
|
||
<taxonomicName id="C0CC27C2FFBCB1223BC9C90BB0AC6B15" authority="Lourenco et al., 2004" authorityName="Lourenco et al." authorityYear="2004" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBCB1223BC9C90BB1B96B15" italics="true" pageId="118" pageNumber="118">Troglorhopalurus translucidus</emphasis>
|
||
<bibRefCitation id="635D21B0FFBCB1223A91C965B0AC6B15" author="Lourenco" box="[753,1008,1352,1376]" etAl="et al." firstAuthor="Lourenco" journalOrPublisher="Comptes Rendus Biologies" pageId="118" pageNumber="118" pagination="1151 - 1156" part="327" refId="ref68662" refString="Lourenco, W. R., R. L. C. Baptista, and A. P. L. Giupponi. 2004. Troglobitic scorpions: a new genus and species from Brazil. Comptes Rendus Biologies 327: 1151 - 1156." title="Troglobitic scorpions: a new genus and species from Brazil" type="journal article" year="2004">Lourenço et al., 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="C0CC27C2FFBCB1223C60C964B1AC6BF4" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" baseAuthorityYear="2017" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="species" species="lacrau">
|
||
<emphasis id="35B88053FFBCB1223C60C964B1AC6BF4" italics="true" pageId="118" pageNumber="118">Troglorhopalurus lacrau</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="635D21B0FFBCB1223B65C946B7ED6BF7" author="Lourenco and Pinto-da-Rocha" box="[773,1201,1385,1410]" firstAuthor="Lourenco" journalOrPublisher="Biogeographica (Paris) 73" pageId="118" pageNumber="118" pagination="181 - 191" part="4" refId="ref68476" refString="Lourenco, W. R., and R. Pinto-da-Rocha. 1997. A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species. Biogeographica (Paris) 73 (4): 181 - 191." title="A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species" type="journal article" year="1997">Lourenço and Pinto-da-Rocha, 1997</bibRefCitation>
|
||
), comb. nov.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFFBCB1223AACC980B7646873" pageId="118" pageNumber="118" type="distribution">
|
||
<paragraph id="07735C41FFBCB1223AACC980B7646873" blockId="118.[684,1214,228,1740]" pageId="118" pageNumber="118">
|
||
DISTRIBUTION:
|
||
<taxonomicName id="C0CC27C2FFBCB1223B14C980B77B6BB1" authorityName="Lourenco et al." authorityYear="2004" box="[884,1063,1452,1476]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBCB1223B14C980B77B6BB1" box="[884,1063,1452,1476]" italics="true" pageId="118" pageNumber="118">Troglorhopalurus</emphasis>
|
||
</taxonomicName>
|
||
is endemic to northeastern Brazil, where it has been recorded in the states of Bahía and Ceará (fig. 9).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFFBCB1223AACCA23B08C681C" pageId="118" pageNumber="118" type="biology_ecology">
|
||
<paragraph id="07735C41FFBCB1223AACCA23B08C681C" blockId="118.[684,1214,228,1740]" pageId="118" pageNumber="118">
|
||
ECOLOGY: Both species of
|
||
<taxonomicName id="C0CC27C2FFBCB1223C62CA23B7E16852" authorityName="Lourenco et al." authorityYear="2004" box="[1026,1213,1551,1575]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBCB1223C62CA23B7E16852" box="[1026,1213,1551,1575]" italics="true" pageId="118" pageNumber="118">Troglorhopalurus</emphasis>
|
||
</taxonomicName>
|
||
occur under stones either inside or in close proximity to caves (fig. 2G, H).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFFBCB1233AACCA5EB21C6ADF" lastPageId="119" lastPageNumber="119" pageId="118" pageNumber="118" type="discussion">
|
||
<paragraph id="07735C41FFBCB1233AACCA5EB21C6ADF" blockId="118.[684,1214,228,1740]" lastBlockId="119.[107,637,228,1194]" lastPageId="119" lastPageNumber="119" pageId="118" pageNumber="118">
|
||
REMARKS:
|
||
<taxonomicName id="C0CC27C2FFBCB1223B2ECA5EB75668FF" authorityName="Lourenco et al." authorityYear="2004" box="[846,1034,1650,1674]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBCB1223B2ECA5EB75668FF" box="[846,1034,1650,1674]" italics="true" pageId="118" pageNumber="118">Troglorhopalurus</emphasis>
|
||
</taxonomicName>
|
||
was originally monotypic, created to accommodate
|
||
<taxonomicName id="C0CC27C2FFBCB1223C5ECAB8B18C68B9" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBCB1223C5ECAB8B18C68B9" italics="true" pageId="118" pageNumber="118">T. translucidus</emphasis>
|
||
</taxonomicName>
|
||
. In comparing
|
||
<taxonomicName id="C0CC27C2FFBCB1223B18CA98B77768B9" authorityName="Lourenco et al." authorityYear="2004" box="[888,1067,1716,1740]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBCB1223B18CA98B77768B9" box="[888,1067,1716,1740]" italics="true" pageId="118" pageNumber="118">Troglorhopalurus</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="C0CC27C2FFBCB1233C0ACA98B2956E89" authority=", Lourenco et al." authorityName="Lourenco et al." class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lastPageId="119" lastPageNumber="119" order="Scorpiones" pageId="118" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBCB1233C0ACA98B3C66E89" italics="true" lastPageId="119" lastPageNumber="119" pageId="118" pageNumber="118">Rhopalurus</emphasis>
|
||
,
|
||
<bibRefCitation id="635D21B0FFBDB12338C6CCC9B2956E89" author="Lourenco" box="[166,457,228,252]" etAl="et al." firstAuthor="Lourenco" journalOrPublisher="Comptes Rendus Biologies" pageId="119" pageNumber="119" pagination="1151 - 1156" part="327" refId="ref68662" refString="Lourenco, W. R., R. L. C. Baptista, and A. P. L. Giupponi. 2004. Troglobitic scorpions: a new genus and species from Brazil. Comptes Rendus Biologies 327: 1151 - 1156." title="Troglobitic scorpions: a new genus and species from Brazil" type="journal article" year="2004">Lourenço et al. (2004: 1153</bibRefCitation>
|
||
</taxonomicName>
|
||
, 1156) noted “all modifications presented by the new troglobitic scorpion are the result of adaptation to a cave dwelling life,” prompting
|
||
<bibRefCitation id="635D21B0FFBDB123391ACD65B1026F14" author="Prendini" box="[378,606,329,353]" etAl="et al." firstAuthor="Prendini" journalOrPublisher="Journal of Arachnology" pageId="119" pageNumber="119" pagination="206 - 224" part="37" refId="ref70522" refString="Prendini, L., L. A. Esposito, J. Huff, and E. S. Volschenk. 2009. Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola. Journal of Arachnology 37: 206 - 224." title="Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola" type="journal article" year="2009">Prendini et al. (2009)</bibRefCitation>
|
||
to suggest that
|
||
<taxonomicName id="C0CC27C2FFBDB1233897CD47B2F16FF6" authorityName="Lourenco et al." authorityYear="2004" box="[247,429,363,387]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBDB1233897CD47B2F16FF6" box="[247,429,363,387]" italics="true" pageId="119" pageNumber="119">Troglorhopalurus</emphasis>
|
||
</taxonomicName>
|
||
might be a junior synonym of
|
||
<taxonomicName id="C0CC27C2FFBDB1233898CDA0B22E6FD1" box="[248,370,396,420]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBDB1233898CDA0B22E6FD1" box="[248,370,396,420]" italics="true" pageId="119" pageNumber="119">Rhopalurus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="C0CC27C2FFBDB12339E2CDA0B1156FD1" box="[386,585,396,420]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
|
||
<emphasis id="35B88053FFBDB12339E2CDA0B1156FD1" box="[386,585,396,420]" italics="true" pageId="119" pageNumber="119">Rhopalurus lacrau</emphasis>
|
||
</taxonomicName>
|
||
had earlier been described from caves belonging to the same subterranean formation in Brazil and, in the description of
|
||
<taxonomicName id="C0CC27C2FFBDB123396ECDDDB3ED6C5E" authority=", Lourenco et al. (2004)" authorityName="Lourenco et al." authorityYear="2004" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBDB123396ECDDDB2986C7C" box="[270,452,497,521]" italics="true" pageId="119" pageNumber="119">Troglorhopalurus</emphasis>
|
||
,
|
||
<bibRefCitation id="635D21B0FFBDB12339B4CDDEB3ED6C5E" author="Lourenco" etAl="et al." firstAuthor="Lourenco" journalOrPublisher="Comptes Rendus Biologies" pageId="119" pageNumber="119" pagination="1151 - 1156" part="327" refId="ref68662" refString="Lourenco, W. R., R. L. C. Baptista, and A. P. L. Giupponi. 2004. Troglobitic scorpions: a new genus and species from Brazil. Comptes Rendus Biologies 327: 1151 - 1156." title="Troglobitic scorpions: a new genus and species from Brazil" type="journal article" year="2004">Lourenço et al. (2004)</bibRefCitation>
|
||
</taxonomicName>
|
||
suggested the relationship between these taxa should be investigated using molecular data.
|
||
<bibRefCitation id="635D21B0FFBDB123380BCE7AB2C66C1B" author="Gallao and Bichuette" box="[107,410,598,622]" firstAuthor="Gallao" journalOrPublisher="Zoologia" pageId="119" pageNumber="119" pagination="1 - 13" part="33" refId="ref65887" refString="Gallao, J. E., and M. E. Bichuette. 2016. On the enigmatic troglobitic scorpion Troglorhopalurus translucidus: distribution, description of adult females, life history and comments on Rhopalurus lacrau (Scorpiones: Buthidae). Zoologia 33 (6): 1 - 13." title="On the enigmatic troglobitic scorpion Troglorhopalurus translucidus: distribution, description of adult females, life history and comments on Rhopalurus lacrau (Scorpiones: Buthidae)" type="journal article" year="2016">Gallão and Bichuette (2016)</bibRefCitation>
|
||
subsequently identified four morphological characters shared by
|
||
<taxonomicName id="C0CC27C2FFBDB1233A05CE55B3F06CC4" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
|
||
<emphasis id="35B88053FFBDB1233A05CE55B3F06CC4" italics="true" pageId="119" pageNumber="119">R. lacrau</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C0CC27C2FFBDB1233880CEB6B2246CC4" box="[224,376,665,689]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBDB1233880CEB6B2246CC4" box="[224,376,665,689]" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
|
||
</taxonomicName>
|
||
, in addition to their troglomorphic habitus (metasomal carination, pectinal tooth count, pectinal peg sensillar shape, and the absence of a pecten-sternite stridulatory apparatus). A close association between the two species was independently verified in the cytogenetic study of
|
||
<bibRefCitation id="635D21B0FFBDB12338ACCF4FB2F16D0E" author="Ubinski" box="[204,429,867,891]" etAl="et al." firstAuthor="Ubinski" journalOrPublisher="Cytogenetic and Genome Research" pageId="119" pageNumber="119" pagination="83 - 155" part="148" refId="ref72292" refString="Ubinski, C. V., L. S. Carvalho, and M. C. Schneider. 2016. Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil. Cytogenetic and Genome Research 148: 83 - 155. [doi: 10.1159 / 000446523]" title="Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil" type="journal article" year="2016">Ubinski et al. (2016)</bibRefCitation>
|
||
which identified a diploid chromosome number of 2n = 20 for
|
||
<taxonomicName id="C0CC27C2FFBDB1233A05CFAAB3F06DCA" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
|
||
<emphasis id="35B88053FFBDB1233A05CFAAB3F06DCA" italics="true" pageId="119" pageNumber="119">R. lacrau</emphasis>
|
||
</taxonomicName>
|
||
and 2n = 20–22 for
|
||
<taxonomicName id="C0CC27C2FFBDB12339E1CF8BB1456DCA" box="[385,537,935,959]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBDB12339E1CF8BB1456DCA" box="[385,537,935,959]" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="4A4E69FAFFBDB1233A45CF8AB1336DCA" box="[549,623,934,959]" captionStart="TABLE 2" captionStartId="7.[610,690,228,252]" captionTargetBox="[235,1058,354,747]" captionText="TABLE 2 Diploid chromosome numbers (2 n) for Brazilian species of Rhopalurusinae from Ubinski et al., (2016) with corresponding genera indicated." httpUri="http://table.plazi.org/id/53B30CC9FFCDB1533A02CCC8B0866F32" pageId="119" pageNumber="119" tableUuid="53B30CC9FFCDB1533A02CCC8B0866F32">table 2</tableCitation>
|
||
). Accordingly, the consistent placement of
|
||
<taxonomicName id="C0CC27C2FFBDB1233A7ECFE5B1276D95" box="[542,635,968,992]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
|
||
<emphasis id="35B88053FFBDB1233A7ECFE5B1276D95" box="[542,635,968,992]" italics="true" pageId="119" pageNumber="119">R. lacrau</emphasis>
|
||
</taxonomicName>
|
||
sister to
|
||
<taxonomicName id="C0CC27C2FFBDB12338A4CFC7B2016A77" box="[196,349,1002,1026]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
|
||
<emphasis id="35B88053FFBDB12338A4CFC7B2016A77" box="[196,349,1002,1026]" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
|
||
</taxonomicName>
|
||
in the analyses of Esposito et al. (in review) is rather predictable, and justifies the transfer of
|
||
<taxonomicName id="C0CC27C2FFBDB123396FC802B2336A30" box="[271,367,1069,1093]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
|
||
<emphasis id="35B88053FFBDB123396FC802B2336A30" box="[271,367,1069,1093]" italics="true" pageId="119" pageNumber="119">R. lacrau</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="C0CC27C2FFBDB12339F7C801B1166A30" box="[407,586,1069,1093]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FFBDB12339F7C801B1166A30" box="[407,586,1069,1093]" italics="true" pageId="119" pageNumber="119">Troglorhopalurus</emphasis>
|
||
</taxonomicName>
|
||
and the following new combination:
|
||
<taxonomicName id="C0CC27C2FFBDB12339A8C863B3F36AFD" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" baseAuthorityYear="2017" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
|
||
<emphasis id="35B88053FFBDB12339A8C863B3F36AFD" italics="true" pageId="119" pageNumber="119">Troglorhopalurus lacrau</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="635D21B0FFBDB12338A5C85CB12C6AFD" author="Lourenco and Pinto-da-Rocha" box="[197,624,1136,1160]" firstAuthor="Lourenco" journalOrPublisher="Biogeographica (Paris) 73" pageId="119" pageNumber="119" pagination="181 - 191" part="4" refId="ref68476" refString="Lourenco, W. R., and R. Pinto-da-Rocha. 1997. A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species. Biogeographica (Paris) 73 (4): 181 - 191." title="A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species" type="journal article" year="1997">Lourenço and Pinto-da-Rocha 1997</bibRefCitation>
|
||
), comb. nov. (fig. 13).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |