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<document id="106AF54AA327120E047928AA16CCFC62" ID-CLB-Dataset="55975" ID-DOI="10.1093/zoolinnean/zlab077" ID-GBIF-Dataset="5e350c66-bf9d-4f32-b68f-16a6b329a76f" ID-ISSN="0024-4082" ID-Zenodo-Dep="6461343" ID-ZooBank="FFC8945F-6A97-4DB7-99CD-E08162DD5819" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649833861871" checkinUser="plazi" docAuthor="Vilarino, Albane, Dias, Everton Santos &amp; Bispo, Pitágoras Da Conceição" docDate="2022" docId="8F15A94DFF9DFF8BB9F0F8420D1AFF26" docLanguage="en" docName="zlab077.pdf" docOrigin="Zoological Journal of the Linnean Society 194 (4)" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Caenocentron immaculatum FLINT 1991" docType="treatment" docVersion="7" lastPageNumber="1364" masterDocId="732CD135FF8BFF9CB937FD2E0956FFFD" masterDocTitle="Phylogeny indicates polyphyly in Cnodocentron (Trichoptera: Xiphocentronidae): biogeography and revision of New World species (Caenocentron)" masterLastPageNumber="1373" masterPageNumber="1341" pageNumber="1363" updateTime="1699328856895" updateUser="ExternalLinkService">
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<mods:title id="EE667169BBDF2B1C397C5B2C33098E04">Phylogeny indicates polyphyly in Cnodocentron (Trichoptera: Xiphocentronidae): biogeography and revision of New World species (Caenocentron)</mods:title>
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<mods:namePart id="912134501EDB9B19789F2FBC97990D10">Vilarino, Albane</mods:namePart>
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<mods:namePart id="5AA7E691319A0317E382D4C015C399A8">Dias, Everton Santos</mods:namePart>
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<mods:namePart id="A1BAEA3B727F89C92AF0C82DA63A1D6A">Bispo, Pitágoras Da Conceição</mods:namePart>
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<mods:title id="D395FFAFD069AFE88A5851C3FE02D6DD">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="4134EBA3BE89A33BA016F19A88CAAEDF">2022</mods:date>
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<mods:number id="EA97D202EEC0D23674F9569433FF6EB3">194</mods:number>
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<mods:identifier id="C982A398309BD8BB011F4603429DA990" type="DOI">10.1093/zoolinnean/zlab077</mods:identifier>
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<treatment id="8F15A94DFF9DFF8BB9F0F8420D1AFF26" ID-DOI="http://doi.org/10.5281/zenodo.6461401" ID-GBIF-Taxon="194598717" ID-Zenodo-Dep="6461401" LSID="urn:lsid:plazi:treatment:8F15A94DFF9DFF8BB9F0F8420D1AFF26" httpUri="http://treatment.plazi.org/id/8F15A94DFF9DFF8BB9F0F8420D1AFF26" lastPageId="23" lastPageNumber="1364" pageId="22" pageNumber="1363">
<subSubSection id="4FA64BD0FF9DFF8AB9F0F8420B94FA79" box="[199,706,1387,1412]" pageId="22" pageNumber="1363" type="nomenclature">
<paragraph id="0703185BFF9DFF8AB9F0F8420B94FA79" blockId="22.[199,706,1387,1412]" box="[199,706,1387,1412]" pageId="22" pageNumber="1363">
<heading id="5C4BAF37FF9DFF8AB9F0F8420B94FA79" box="[199,706,1387,1412]" centered="true" fontSize="9" level="2" pageId="22" pageNumber="1363" reason="2">
<taxonomicName id="C0BC63D8FF9DFF8AB9F0F8420B94FA79" ID-CoL="7cb1cf58-7066-4829-b8df-895eaf220919" authority="FLINT, 1991" authorityName="FLINT" authorityYear="1991" box="[199,706,1387,1412]" class="Insecta" family="Xiphocentronidae" genus="Caenocentron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="1363" phylum="Arthropoda" rank="species" species="immaculatum">
<emphasis id="35C8C449FF9DFF8AB9F0F8420B67FA7E" box="[199,561,1388,1411]" italics="true" pageId="22" pageNumber="1363">CAENOCENTRON IMMACULATUM</emphasis>
FLINT, 1991
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<caption id="53C348D3FF9DFF8ABA1DF8400D06FA5C" ID-DOI="http://doi.org/10.5281/zenodo.6461378" ID-Zenodo-Dep="6461378" httpUri="https://zenodo.org/record/6461378/files/figure.png" pageId="22" pageNumber="1363" startId="22.[810,888,1390,1412]" targetBox="[812,1421,196,1349]" targetPageId="22">
<paragraph id="0703185BFF9DFF8ABA1DF8400D06FA5C" blockId="22.[809,1425,1390,1441]" pageId="22" pageNumber="1363">
<emphasis id="35C8C449FF9DFF8ABA1DF8400AC9FA7E" bold="true" box="[810,927,1390,1412]" pageId="22" pageNumber="1363">Figure 17.</emphasis>
<taxonomicName id="C0BC63D8FF9DFF8ABA9FF8400D99FA79" authorityName="FLINT" authorityYear="1991" box="[936,1231,1390,1412]" class="Insecta" family="Xiphocentronidae" genus="Caenocentron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="1363" phylum="Arthropoda" rank="species" species="immaculatum">
<emphasis id="35C8C449FF9DFF8ABA9FF8400D99FA79" box="[936,1231,1390,1412]" italics="true" pageId="22" pageNumber="1363">Caenocentron immaculatum</emphasis>
</taxonomicName>
, male genitalia: A, lateral; B, dorsal; C, ventral.
</paragraph>
</caption>
<subSubSection id="4FA64BD0FF9DFF8AB84DF8BC0B59FA56" box="[378,527,1426,1451]" pageId="22" pageNumber="1363" type="description">
<paragraph id="0703185BFF9DFF8AB84DF8BC0B59FA56" blockId="22.[378,527,1426,1451]" box="[378,527,1426,1451]" pageId="22" pageNumber="1363">
(
<figureCitation id="9F8704DEFF9DFF8AB8B5F8BC0B51FA56" box="[386,519,1426,1451]" captionStart="Figure 17" captionStartId="22.[810,888,1390,1412]" captionTargetBox="[812,1421,196,1349]" captionTargetId="figure-422@22.[811,1422,195,1350]" captionTargetPageId="22" captionText="Figure 17. Caenocentron immaculatum, male genitalia: A, lateral; B, dorsal; C, ventral." figureDoi="http://doi.org/10.5281/zenodo.6461378" httpUri="https://zenodo.org/record/6461378/files/figure.png" pageId="22" pageNumber="1363">FIG. 17AC</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4FA64BD0FF9DFF8AB9A6F8940859FA13" pageId="22" pageNumber="1363" type="reference_group">
<paragraph id="0703185BFF9DFF8AB9A6F8940859FA13" blockId="22.[145,761,1466,1518]" pageId="22" pageNumber="1363">
<taxonomicName id="C0BC63D8FF9DFF8AB9A6F894085DFA13" ID-CoL="7SD3H" authority="Flint, 1991: 34" authorityName="Flint" authorityPageNumber="34" authorityYear="1991" class="Insecta" family="Xiphocentronidae" genus="Cnodocentron" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="1363" phylum="Arthropoda" rank="species" species="immaculatum" subGenus="Caenocentron">
<emphasis id="35C8C449FF9DFF8AB9A6F8940BFDFA32" box="[145,683,1466,1488]" italics="true" pageId="22" pageNumber="1363">Cnodocentron (Caenocentron) immaculatum</emphasis>
Flint, 1991: 34
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="4FA64BD0FF9DFF8BB9A6FB370D1AFF26" lastPageId="23" lastPageNumber="1364" pageId="22" pageNumber="1363" type="materials_examined">
<paragraph id="0703185BFF9DFF8AB9A6FB370B1DF996" blockId="22.[145,760,1560,1643]" pageId="22" pageNumber="1363">
<emphasis id="35C8C449FF9DFF8AB9A6FB370813F9D0" box="[145,325,1560,1582]" italics="true" pageId="22" pageNumber="1363">Type material:</emphasis>
<materialsCitation id="B7D41206FF9DFF8AB864FB360B1DF996" ID-GBIF-Occurrence="3751569318" collectingDate="1984-02-07" collectionCode="USNM" collectorName="U. Matthias" country="Colombia" location="Quebrada La Jimenez" pageId="22" pageNumber="1363" specimenCount="1" specimenCount-male="1" stateProvince="Antioquia" typeStatus="holotype">
<typeStatus id="D807A6F9FF9DFF8AB864FB360892F9D3" box="[339,452,1560,1582]" pageId="22" pageNumber="1363" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="7FAB58CBFF9DFF8AB8E5FB360B3DF9D0" box="[466,619,1560,1581]" name="Colombia" pageId="22" pageNumber="1363">COLOMBIA</collectingCountry>
:
<collectingRegion id="C578D6B9FF9DFF8ABB4FFB360BA2F9D3" box="[632,756,1560,1582]" country="Colombia" name="Antioquia" pageId="22" pageNumber="1363">Antioquia</collectingRegion>
:
<location id="02634E80FF9DFF8AB9A6FB1808FFF9B1" LSID="urn:lsid:plazi:treatment:8F15A94DFF9DFF8BB9F0F8420D1AFF26:02634E80FF9DFF8AB9A6FB1808FFF9B1" box="[145,425,1590,1612]" country="Colombia" name="Quebrada La Jimenez" pageId="22" pageNumber="1363" stateProvince="Antioquia">Quebrada La Jimenez</location>
, Mun. Sopetran.
<date id="73023E9BFF9DFF8ABBB1FB180BA5F9B1" box="[646,755,1590,1612]" pageId="22" pageNumber="1363" value="1984-02-07">
<collectingDate id="6346C773FF9DFF8ABBB1FB180BA5F9B1" box="[646,755,1590,1612]" pageId="22" pageNumber="1363" value="1984-02-07">7.ii.1984</collectingDate>
</date>
,
<collectorName id="AA497D8DFF9DFF8AB9A6FB780841F996" box="[145,279,1621,1643]" pageId="22" pageNumber="1363">U. Matthias</collectorName>
leg., (
<specimenCount id="11BAD3D2FF9DFF8AB86CFB7C083DF997" box="[347,363,1618,1642]" pageId="22" pageNumber="1363" type="male"></specimenCount>
in alcohol,
<collectionCode id="61AD809EFF9DFF8AB8DBFB7B0B69F996" box="[492,575,1621,1643]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="1363" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="0703185BFF9DFF8AB9A6FBBB0843F8F8" blockId="22.[145,761,1684,1797]" pageId="22" pageNumber="1363">
<emphasis id="35C8C449FF9DFF8AB9A6FBBB0878F954" box="[145,302,1684,1706]" italics="true" pageId="22" pageNumber="1363">
<typeStatus id="D807A6F9FF9DFF8AB9A6FBBB099EF957" box="[145,200,1685,1706]" pageId="22" pageNumber="1363">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="7FAB58CBFF9DFF8AB80CFBBA08FAF957" box="[315,428,1684,1706]" name="Colombia" pageId="22" pageNumber="1363">Colombia</collectingCountry>
, Dpto.
<collectingRegion id="C578D6B9FF9DFF8ABB37FBBA0B21F957" box="[512,631,1684,1706]" country="Colombia" name="Antioquia" pageId="22" pageNumber="1363">Antioquia</collectingRegion>
, Quebrada la Jimenez, Sopetran;
<collectionCode id="61AD809EFF9DFF8AB8AAFB9D08A6F935" box="[413,496,1715,1736]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="1363" type="Museum">USNM</collectionCode>
;
<specimenCount id="11BAD3D2FF9DFF8AB8CAFB9E0B5BF935" box="[509,525,1712,1736]" pageId="22" pageNumber="1363" type="male"></specimenCount>
.
<emphasis id="35C8C449FF9DFF8ABB2AFB9D0B7FF935" box="[541,553,1715,1736]" italics="true" pageId="22" pageNumber="1363"></emphasis>
<bibRefCitation id="632D65AAFF9DFF8ABB05FB9C09A9F91B" author="Munoz-Quesada F" pageId="22" pageNumber="1363" pagination="267 - 288" refId="ref20280" refString="Munoz-Quesada F. 2000. Especies del orden Trichoptera (Insecta) en Colombia. Biota Colombiana 1: 267 - 288." type="journal article" year="2000">Muñoz-Quesada, 2000: 281</bibRefCitation>
[checklist].
<emphasis id="35C8C449FF9DFF8AB8BDFBFC08C0F91A" box="[394,406,1746,1767]" italics="true" pageId="22" pageNumber="1363"></emphasis>
<bibRefCitation id="632D65AAFF9DFF8AB8A9FBFF0BAFF91A" author="Holzenthal RW &amp; Calor AR" box="[414,761,1745,1767]" pageId="22" pageNumber="1363" pagination="1 - 566" refId="ref19123" refString="Holzenthal RW, Calor AR. 2017. Catalog of the Neotropical Trichoptera (Caddisflies). ZooKeys 654: 1 - 566." type="journal article" year="2017">Holzenthal &amp; Calor, 2017: 454</bibRefCitation>
[catalogue].
</paragraph>
<paragraph id="0703185BFF9DFF8AB9A6FA190D14F9AE" blockId="22.[145,761,1846,1899]" lastBlockId="22.[809,1425,1536,1620]" pageId="22" pageNumber="1363">
<emphasis id="35C8C449FF9DFF8AB9A6FA190B30F8B6" box="[145,614,1846,1868]" italics="true" pageId="22" pageNumber="1363">
Diagnosis:
<taxonomicName id="C0BC63D8FF9DFF8AB829FA180B30F8B6" authorityName="Flint" authorityYear="1991" box="[286,614,1846,1867]" class="Insecta" family="Xiphocentronidae" genus="Cnodocentron" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="1363" phylum="Arthropoda" rank="species" species="immaculatum">Cnodocentron immaculatum</taxonomicName>
</emphasis>
is similar to
<taxonomicName id="C0BC63D8FF9DFF8AB9A6FA7B0852F897" authorityName="Schmid" authorityYear="1982" box="[145,260,1877,1898]" class="Insecta" family="Xiphocentronidae" genus="Cnodocentron" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="1363" phylum="Arthropoda" rank="species" species="galesus">
<emphasis id="35C8C449FF9DFF8AB9A6FA7B0852F897" box="[145,260,1877,1898]" italics="true" pageId="22" pageNumber="1363">C. galesus</emphasis>
</taxonomicName>
and
<taxonomicName id="C0BC63D8FF9DFF8AB877FA7B08F3F897" authorityName="Schmid" authorityYear="1982" box="[320,421,1877,1898]" class="Insecta" family="Xiphocentronidae" genus="Cnodocentron" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="1363" phylum="Arthropoda" rank="species" species="pallas">
<emphasis id="35C8C449FF9DFF8AB877FA7B08F3F897" box="[320,421,1877,1898]" italics="true" pageId="22" pageNumber="1363">C. pallas</emphasis>
</taxonomicName>
by presenting a pair of stout spines on the ventral projection of coxopodite. It can be differentiated by the greatly enlarged base of harpago bearing a brush of setae.
</paragraph>
<paragraph id="0703185BFF9DFF8BBA1EFB520806FF07" blockId="22.[809,1424,1657,1897]" lastBlockId="23.[163,778,197,250]" lastPageId="23" lastPageNumber="1364" pageId="22" pageNumber="1363">
<emphasis id="35C8C449FF9DFF8ABA1EFB520A21F96C" box="[809,887,1660,1681]" italics="true" pageId="22" pageNumber="1363">Adult:</emphasis>
<materialsCitation id="B7D41206FF9DFF8ABAB2FB520DE8F932" ID-GBIF-Occurrence="3751569310" collectionCode="IV" pageId="22" pageNumber="1363" specimenCount="1" specimenCount-male="1">
Forewing length 3.0 mm (
<emphasis id="35C8C449FF9DFF8ABDF8FB530DB5F96F" box="[1231,1251,1661,1682]" italics="true" pageId="22" pageNumber="1363">N</emphasis>
=
<specimenCount id="11BAD3D2FF9DFF8ABC34FB520C74F96C" box="[1283,1314,1657,1681]" pageId="22" pageNumber="1363" type="male">1♂</specimenCount>
). Colour (in alcohol) uniformly dark brown. Maxillary palp formula (I-II-III)-IV-V; segment
<collectionCode id="61AD809EFF9DFF8ABD95FB940DE8F932" box="[1186,1214,1722,1743]" pageId="22" pageNumber="1363">IV</collectionCode>
</materialsCitation>
shorter than sum of segments I-II-III; male tibial spurs 2-4-3, male hindleg apical spur unmodified. Venation: forewing fork
<collectionCode id="61AD809EFF9DFF8ABA52FA380A2CF8D6" box="[869,890,1814,1835]" pageId="22" pageNumber="1363">II</collectionCode>
and
<collectionCode id="61AD809EFF9DFF8ABA8AFA380A8CF8D6" box="[957,986,1814,1835]" pageId="22" pageNumber="1363">IV</collectionCode>
present, three anal veins present; hindwing fork
<collectionCode id="61AD809EFF9DFF8ABAD1FA1B0AADF8B7" box="[998,1019,1845,1866]" pageId="22" pageNumber="1363">II</collectionCode>
and
<collectionCode id="61AD809EFF9DFF8ABD08FA1B0D07F8B7" box="[1087,1105,1845,1866]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="1363" type="Museum">V</collectionCode>
<materialsCitation id="B7D41206FF9DFF8ABD6CFA1B0AE3F895" ID-GBIF-Occurrence="3751569317" collectionCode="R" pageId="22" pageNumber="1363" specimenCode="R1" specimenCount="1">
present, transverse vein between
<specimenCode id="571AB020FF9DFF8ABAA2FA7D0AE3F895" box="[917,949,1875,1896]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="1363">R1</specimenCode>
</materialsCitation>
and SR present. Abdominal sternum
<collectionCode id="61AD809EFF9DFF8ABC48FA7D0CC6F895" box="[1407,1424,1875,1896]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="1363" type="Museum">V</collectionCode>
with anterolateral oval region with cuticle modified and reticulate.
</paragraph>
<paragraph id="0703185BFF9CFF8BB994FC060827FBC1" blockId="23.[163,779,296,1084]" pageId="23" pageNumber="1364">
<emphasis id="35C8C449FF9CFF8BB994FC0608A0FEC0" box="[163,502,296,317]" italics="true" pageId="23" pageNumber="1364">
Male genitalia (
<figureCitation id="9F8704DEFF9CFF8BB855FC0608B3FEC0" box="[354,485,296,317]" captionStart="Figure 17" captionStartId="22.[810,888,1390,1412]" captionTargetBox="[812,1421,196,1349]" captionTargetId="figure-422@22.[811,1422,195,1350]" captionTargetPageId="22" captionText="Figure 17. Caenocentron immaculatum, male genitalia: A, lateral; B, dorsal; C, ventral." figureDoi="http://doi.org/10.5281/zenodo.6461378" httpUri="https://zenodo.org/record/6461378/files/figure.png" pageId="23" pageNumber="1364">Fig. 17AC</figureCitation>
):
</emphasis>
Tergum, IX in lateral view, narrow, height greater than length. Sternum IX, in lateral view, deltoid, about as high as long, anterior margin truncate, with narrow, elongate apodeme, apical margin rounded, dorsal margin prominent; in ventral view, apical margin concave. Segment X membranous, fused to paraproct. Paraproct, in lateral view, trapezoidal, dorsal margin with short process, ventroapically subacute; in dorsal view, mid-dorsal margins sclerotized, not fused. Pre-anal appendage, in lateral view, strongly sinuous, slender, slightly enlarged apically. Inferior appendage coxopodite and hapago distinct. Coxopodite, in lateral view, median region with short setae; apical margin produced, broad, apex truncate and bearing stout spine near ventral margin; basal surface with ventral projection. Ventral projection with stout spine and rows of setae mesally and along its margin, apical setae longer; in ventral view, ventral projection with long lateral setae, short mesal setae and two stout sublateral spines. Harpago, in lateral view, slender, basal third strongly enlarged, bearing short spine and basal setal brush. Basal plate, in lateral view, directed ventrad; in ventral view short. Phallus tubular, long and slender, base flared, reaching segment
<collectionCode id="61AD809EFF9CFF8BBB84F9260B90FBE0" box="[691,710,1032,1053]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="23" pageNumber="1364" type="Museum">V</collectionCode>
; apex slightly enlarged.
</paragraph>
<paragraph id="0703185BFF9CFF8BBA0CFDEB0D1AFF26" blockId="23.[827,1100,197,219]" box="[827,1100,197,219]" pageId="23" pageNumber="1364">
<emphasis id="35C8C449FF9CFF8BBA0CFDEB0A9AFF27" box="[827,972,197,218]" italics="true" pageId="23" pageNumber="1364">Distribution:</emphasis>
<collectingCountry id="7FAB58CBFF9CFF8BBAEEFDEB0D1EFF26" box="[985,1096,197,219]" name="Colombia" pageId="23" pageNumber="1364">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>