396 lines
48 KiB
XML
396 lines
48 KiB
XML
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<mods:title id="DDAA97CC55FFF3722AE595A6A824DAD5">Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar’s rainforest</mods:title>
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<mods:affiliation id="385932F66841CB62E006CEE3057BCEF6">Mention Zoologie et Biodiversité Animale, Université d’Antananarivo, BP 906, Antananarivo, 101 Madagascar</mods:affiliation>
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<emphasis id="35D6531A1301BF10FF28FC2DFE8E41F0" box="[151,285,939,963]" italics="true" pageId="43" pageNumber="44">Lygodactylus</emphasis>
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sp.
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21:
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<bibRefCitation id="6333F2F91301BF10FED1FC2DFDD041F0" author="Gippner, S. & Travers S. L. & Scherz M. D. & Colston T. J. & Lyra M. L. & Mohan A. V. & Multzsch M. & Nielsen S. V. & Rancilhac L. & Glaw F. & Bauer A. M. & Vences M." box="[366,579,939,963]" pageId="43" pageNumber="44" pagination="107311" refId="ref37826" refString="Gippner, S., Travers S. L., Scherz M. D., Colston T. J., Lyra M. L., Mohan A. V., Multzsch M., Nielsen S. V., Rancilhac L., Glaw F., Bauer A. M. & Vences M. (2021) A comprehensive phylogeny of dwarf geckos of the genus Lygodactylus, with insights into their systematics and morphological variation. Molecular Phylogenetics and Evolution, 165, 107311. https: // doi. org / 10.1016 / j. ympev. 2021.107311" type="journal article" year="2021">
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Gippner
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(2021)
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||
<geoCoordinate id="6296E9CF1301BF10FA94FB93FA0B461C" box="[1323,1432,1045,1071]" degrees="14.4377" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="5" value="-14.4377">S14.4377</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6296E9CF1301BF10FF28FBBFFE954667" box="[151,262,1081,1108]" degrees="49.7756" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5" value="49.7756">E49.7756</geoCoordinate>
|
||
,
|
||
<elevation id="8C8F683B1301BF10FEAEFBBFFE064660" box="[273,405,1081,1108]" metricMagnitude="2" metricUnit="m" metricValue="4.81" pageId="43" pageNumber="44" unit="m" value="481.0">
|
||
<quantity id="C05A22ED1301BF10FEAEFBBFFECB4667" box="[273,344,1081,1108]" metricMagnitude="2" metricUnit="m" metricValue="4.81" pageId="43" pageNumber="44" unit="m" value="481.0">481 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, on
|
||
<date id="731CA9C81301BF10FE7BFBBFFD184667" box="[452,651,1081,1108]" pageId="43" pageNumber="44" value="2005-02-14">
|
||
<collectingDate id="635850201301BF10FE7BFBBFFD184667" box="[452,651,1081,1108]" pageId="43" pageNumber="44" value="2005-02-14">14 February 2005</collectingDate>
|
||
</date>
|
||
(
|
||
<figureCitation id="9F99938D1301BF10FD24FBBFFD7D4667" box="[667,750,1081,1108]" captionStart="FIGURE 15" captionStartId="25.[151,250,1943,1967]" captionTargetBox="[244,1351,181,1906]" captionTargetId="figure-18@25.[236,1352,181,1919]" captionTargetPageId="25" captionText="FIGURE 15. Dorsal and ventral views of preserved holotypes of new species of the Lygodactylus madagascariensis group named in the current study." figureDoi="http://doi.org/10.5281/zenodo.7040784" httpUri="https://zenodo.org/record/7040784/files/figure.png" pageId="43" pageNumber="44">Fig. 15</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="071D8F081301BF10FF78FBDBFB7846A8" blockId="43.[151,1437,1009,2007]" pageId="43" pageNumber="44">
|
||
<emphasis id="35D6531A1301BF10FF78FBDBFE344644" bold="true" box="[199,423,1117,1143]" pageId="43" pageNumber="44">Referred material.</emphasis>
|
||
<materialsCitation id="B7CA85551301BF10FE10FBD8FB7B46A8" ID-GBIF-Occurrence="3897988302" collectingDate="2016-11-23" collectionCode="UADBA-R" collectorName="M. D. Scherz & Scherz, J & A. Razafimanantsoa" country="Madagascar" county="North-East" elevation="481" latitude="-14.4377" location="Marojejy National Park" longLatPrecision="7" longitude="49.7756" municipality="Camp" pageId="43" pageNumber="44" specimenCode="MSZC 0272" specimenCount="1">
|
||
<collectionCode id="61B317CD1301BF10FE10FBD8FDBC464B" box="[431,559,1118,1144]" pageId="43" pageNumber="44">UADBA-R</collectionCode>
|
||
uncatalogued (
|
||
<specimenCode id="570427731301BF10FD62FBDBFCE34644" box="[733,880,1117,1143]" collectionCode="MSZC" pageId="43" pageNumber="44">MSZC 0272</specimenCode>
|
||
), female, collected by
|
||
<collectorName id="AA57EADE1301BF10FBCCFBD8FA954644" box="[1139,1286,1117,1144]" pageId="43" pageNumber="44">M.D. Scherz</collectorName>
|
||
, J. Razafindraibe, and
|
||
<collectorName id="AA57EADE1301BF10FEB4FB04FE7D46A8" box="[267,494,1153,1180]" pageId="43" pageNumber="44">A. Razafimanantsoa</collectorName>
|
||
at the same locality as the holotype, at night on
|
||
<date id="731CA9C81301BF10FBB4FB07FB7B46A8" box="[1035,1256,1153,1179]" pageId="43" pageNumber="44" value="2016-11-23">
|
||
<collectingDate id="635850201301BF10FBB4FB07FB7B46A8" box="[1035,1256,1153,1179]" pageId="43" pageNumber="44" value="2016-11-23">23 November 2016</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FB8DC831301BF10FF78FB23FA0E4574" pageId="43" pageNumber="44" type="diagnosis">
|
||
<paragraph id="071D8F081301BF10FF78FB23FB2C4788" blockId="43.[151,1437,1009,2007]" pageId="43" pageNumber="44">
|
||
<emphasis id="35D6531A1301BF10FF78FB23FED1468C" bold="true" box="[199,322,1189,1215]" pageId="43" pageNumber="44">Diagnosis.</emphasis>
|
||
<taxonomicName id="C0A2F48B1301BF10FEF5FB23FD9F468C" authority="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz, 2022" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[330,524,1189,1215]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="ulli" status="sp. nov.">
|
||
<emphasis id="35D6531A1301BF10FEF5FB23FD9F468C" box="[330,524,1189,1215]" italics="true" pageId="43" pageNumber="44">Lygodactylus ulli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="35D6531A1301BF10FDAAFB23FDE2468C" bold="true" box="[533,625,1189,1215]" pageId="43" pageNumber="44">
|
||
<taxonomicNameLabel id="2EE5EE611301BF10FDAAFB23FDE2468C" box="[533,625,1189,1215]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
corresponds to a lineage forming part of subclade A5 of
|
||
<taxonomicName id="C0A2F48B1301BF10FABCFB23FA05468C" authorityName="Pasteur" authorityYear="1964" box="[1283,1430,1189,1215]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
|
||
<emphasis id="35D6531A1301BF10FABCFB23FA05468C" box="[1283,1430,1189,1215]" italics="true" pageId="43" pageNumber="44">Domerguella</emphasis>
|
||
</taxonomicName>
|
||
, and is the second representative of this subclade reaching northern
|
||
<collectingCountry id="7FB5CF981301BF10FC35FB4FFB8046D0" box="[906,1043,1225,1251]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
|
||
. It is characterized by the presence of distinct lateral spine-like scales at the base of the tail in males, as is found in several representatives of A5 from eastern
|
||
<collectingCountry id="7FB5CF981301BF10FF50FA97FEE54718" box="[239,374,1297,1323]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
|
||
but not in other
|
||
<taxonomicName id="C0A2F48B1301BF10FD89FA97FD5A4718" authorityName="Pasteur" authorityYear="1964" box="[566,713,1297,1323]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
|
||
<emphasis id="35D6531A1301BF10FD89FA97FD5A4718" box="[566,713,1297,1323]" italics="true" pageId="43" pageNumber="44">Domerguella</emphasis>
|
||
</taxonomicName>
|
||
. The smallest genetic distances are 10.7% uncorrected 16S distance to specimens of
|
||
<taxonomicName id="C0A2F48B1301BF10FE2FFAB0FE67477C" box="[400,500,1333,1359]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="guibei">
|
||
<emphasis id="35D6531A1301BF10FE2FFAB0FE67477C" box="[400,500,1333,1359]" italics="true" pageId="43" pageNumber="44">L. guibei</emphasis>
|
||
</taxonomicName>
|
||
. It can also be assigned to the subgenus
|
||
<taxonomicName id="C0A2F48B1301BF10FC04FAB3FBDD477C" authorityName="Pasteur" authorityYear="1964" box="[955,1102,1333,1359]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
|
||
<emphasis id="35D6531A1301BF10FC04FAB3FBDD477C" box="[955,1102,1333,1359]" italics="true" pageId="43" pageNumber="44">Domerguella</emphasis>
|
||
</taxonomicName>
|
||
by an undivided mental scale with two postmentals, absence of a claw on the first finger, and 7 preanal pores in males. Within
|
||
<taxonomicName id="C0A2F48B1301BF10FB66FADFFAFF4740" authorityName="Pasteur" authorityYear="1964" box="[1241,1388,1369,1395]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
|
||
<emphasis id="35D6531A1301BF10FB66FADFFAFF4740" box="[1241,1388,1369,1395]" italics="true" pageId="43" pageNumber="44">Domerguella</emphasis>
|
||
</taxonomicName>
|
||
, the new species (together with
|
||
<taxonomicName id="C0A2F48B1301BF10FE73FAF8FDBB47A4" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[460,552,1405,1431]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="winki">
|
||
<emphasis id="35D6531A1301BF10FE73FAF8FDBB47A4" box="[460,552,1405,1431]" italics="true" pageId="43" pageNumber="44">L. winki</emphasis>
|
||
</taxonomicName>
|
||
) is one of only two species of subclade A5 known from northern
|
||
<collectingCountry id="7FB5CF981301BF10FAB0FAFBFA0B47A4" box="[1295,1432,1405,1431]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
|
||
, and the only species of
|
||
<taxonomicName id="C0A2F48B1301BF10FE20FA27FDA14788" authorityName="Pasteur" authorityYear="1964" box="[415,562,1441,1467]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
|
||
<emphasis id="35D6531A1301BF10FE20FA27FDA14788" box="[415,562,1441,1467]" italics="true" pageId="43" pageNumber="44">Domerguella</emphasis>
|
||
</taxonomicName>
|
||
so far known from the rainforests of the Marojejy Massif.
|
||
</paragraph>
|
||
<paragraph id="071D8F081301BF10FF78FA40FB584510" blockId="43.[151,1437,1009,2007]" pageId="43" pageNumber="44">
|
||
It differs from the nominal species of
|
||
<taxonomicName id="C0A2F48B1301BF10FDC9FA43FC9A47EC" authorityName="Pasteur" authorityYear="1964" box="[630,777,1477,1503]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
|
||
<emphasis id="35D6531A1301BF10FDC9FA43FC9A47EC" box="[630,777,1477,1503]" italics="true" pageId="43" pageNumber="44">Domerguella</emphasis>
|
||
</taxonomicName>
|
||
occurring in northern
|
||
<collectingCountry id="7FB5CF981301BF10FBB5FA43FB0247EC" box="[1034,1169,1477,1503]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
|
||
and belonging to subclades A1–A4 as follows: from
|
||
<taxonomicName id="C0A2F48B1301BF10FE54FA6CFDED4430" authorityName=": Rosler" authorityYear="2000" box="[491,638,1514,1539]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="expectatus">
|
||
<emphasis id="35D6531A1301BF10FE54FA6CFDED4430" box="[491,638,1514,1539]" italics="true" pageId="43" pageNumber="44">L. expectatus</emphasis>
|
||
</taxonomicName>
|
||
by non-enlarged dorsolateral scales (longitudinal count of dorsal scales>250 vs. <170); from
|
||
<taxonomicName id="C0A2F48B1301BF10FE2AF988FE624414" authorityName=": Rosler" authorityYear="2000" box="[405,497,1550,1575]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="rarus">
|
||
<emphasis id="35D6531A1301BF10FE2AF988FE624414" box="[405,497,1550,1575]" italics="true" pageId="43" pageNumber="44">L. rarus</emphasis>
|
||
</taxonomicName>
|
||
by lack of regular crossbands on tail (vs. presence) and different body shape with less elongated limbs (relative hindlimb length 0.47 vs.>0.55); from
|
||
<emphasis id="35D6531A1301BF10FC28F9B4FAFF4478" box="[919,1388,1585,1611]" italics="true" pageId="43" pageNumber="44">
|
||
<taxonomicName id="C0A2F48B1301BF10FC28F9B4FB934478" authorityName="Pasteur & Blanc" authorityYear="1967" box="[919,1024,1586,1611]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="petteri">L. petteri</taxonomicName>
|
||
,
|
||
<taxonomicName id="C0A2F48B1301BF10FBB3F9B4FB1E4478" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[1036,1165,1585,1611]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="tantsaha">L. tantsaha</taxonomicName>
|
||
,
|
||
<taxonomicName id="C0A2F48B1301BF10FB27F9B4FB7D4478" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[1176,1262,1585,1611]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="salvi">L. salvi</taxonomicName>
|
||
,
|
||
<taxonomicName id="C0A2F48B1301BF10FB46F9B4FAFB4478" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[1273,1384,1585,1611]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="roellae">L. roellae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="C0A2F48B1301BF10FF28F9D0FF67445C" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[151,244,1621,1647]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="hapei">
|
||
<emphasis id="35D6531A1301BF10FF28F9D0FF67445C" box="[151,244,1621,1647]" italics="true" pageId="43" pageNumber="44">L. hapei</emphasis>
|
||
</taxonomicName>
|
||
by a higher longitudinal count of dorsal scales (253 vs. <241); and from
|
||
<taxonomicName id="C0A2F48B1301BF10FB84F9D0FAB7445C" authorityName=": Pasteur" authorityYear="1965" box="[1083,1316,1621,1647]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="madagascariensis">
|
||
<emphasis id="35D6531A1301BF10FB84F9D0FAB7445C" box="[1083,1316,1621,1647]" italics="true" pageId="43" pageNumber="44">L. madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
as well as most of the previously mentioned species by a small but distinct spine-like tubercle at the base of the tail in males. From the other nominal species in subclade A5 (
|
||
<emphasis id="35D6531A1301BF10FD7CF918FCBF4484" box="[707,812,1694,1719]" italics="true" pageId="43" pageNumber="44">
|
||
<taxonomicName id="C0A2F48B1301BF10FD7CF918FCBA4484" authorityName="Gunther" authorityYear="1891" box="[707,809,1694,1719]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="miops">L. miops</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="C0A2F48B1301BF10FCD9F918FC5F4484" box="[870,972,1693,1719]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="guibei">
|
||
<emphasis id="35D6531A1301BF10FCD9F918FC5F4484" box="[870,972,1693,1719]" italics="true" pageId="43" pageNumber="44">L. guibei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C0A2F48B1301BF10FBBAF918FBF14484" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[1029,1122,1693,1719]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="winki">
|
||
<emphasis id="35D6531A1301BF10FBBAF918FBF14484" box="[1029,1122,1693,1719]" italics="true" pageId="43" pageNumber="44">L. winki</emphasis>
|
||
</taxonomicName>
|
||
) the new species differs by higher longitudinal counts of dorsal scales (253 vs. 170–242) and ventral scales (110 vs. 87–109). We did not detect haplotype sharing in RAG1 or CMOS between
|
||
<taxonomicName id="C0A2F48B1301BF10FD11F960FD6244CC" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[686,753,1765,1791]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="ulli">
|
||
<emphasis id="35D6531A1301BF10FD11F960FD6244CC" box="[686,753,1765,1791]" italics="true" pageId="43" pageNumber="44">L. ulli</emphasis>
|
||
</taxonomicName>
|
||
and the other nominal species in subclade A5 (
|
||
<emphasis id="35D6531A1301BF10FABAF960FAFF44CC" box="[1285,1388,1766,1791]" italics="true" pageId="43" pageNumber="44">
|
||
<taxonomicName id="C0A2F48B1301BF10FABAF960FAFA44CC" authorityName="Gunther" authorityYear="1891" box="[1285,1385,1766,1791]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="miops">L. miops</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="C0A2F48B1301BF10FF28F88CFF684510" box="[151,251,1801,1827]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="guibei">
|
||
<emphasis id="35D6531A1301BF10FF28F88CFF684510" box="[151,251,1801,1827]" italics="true" pageId="43" pageNumber="44">L. guibei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C0A2F48B1301BF10FE8EF88CFE1F4510" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[305,396,1801,1827]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="winki">
|
||
<emphasis id="35D6531A1301BF10FE8EF88CFE1F4510" box="[305,396,1801,1827]" italics="true" pageId="43" pageNumber="44">L. winki</emphasis>
|
||
</taxonomicName>
|
||
). Haplotpe sharing was detected only at the CMOS marker with
|
||
<emphasis id="35D6531A1301BF10FBDCF88CFBE94510" box="[1123,1146,1802,1827]" italics="true" pageId="43" pageNumber="44">L.</emphasis>
|
||
sp. 20.
|
||
</paragraph>
|
||
<paragraph id="071D8F081301BF10FF78F8A8FA0E4574" blockId="43.[151,1437,1009,2007]" box="[199,1437,1837,1864]" pageId="43" pageNumber="44">For a distinction from additional species newly named and described herein, see the respective diagnoses below.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FB8DC831301BF10FF78F8D7FE2B45E4" pageId="43" pageNumber="44" type="etymology">
|
||
<paragraph id="071D8F081301BF10FF78F8D7FE2B45E4" blockId="43.[151,1437,1009,2007]" pageId="43" pageNumber="44">
|
||
<emphasis id="35D6531A1301BF10FF78F8D7FEDD4558" bold="true" box="[199,334,1873,1899]" pageId="43" pageNumber="44">Etymology.</emphasis>
|
||
We are pleased to dedicate this species to Ulrich “Ulli” Joger, director emeritus of the Braunschweig Natural History Museum, in recognition of his contribution to the taxonomy of reptiles, especially geckos. The species epithet name is defined as a noun in apposition (not a noun in the genitive case) to avoid ending with a non-euphonious double-i.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="53DDDF801306BF17FF28F8CEFA9645B7" ID-DOI="http://doi.org/10.5281/zenodo.7040809" ID-Zenodo-Dep="7040809" httpUri="https://zenodo.org/record/7040809/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,250,1864,1888]" targetBox="[151,1436,181,1820]" targetPageId="44">
|
||
<paragraph id="071D8F081306BF17FF28F8CEFA9645B7" blockId="44.[151,1436,1864,1924]" pageId="44" pageNumber="45">
|
||
<emphasis id="35D6531A1306BF17FF28F8CEFEB24553" bold="true" box="[151,289,1864,1888]" pageId="44" pageNumber="45">FIGURE 21.</emphasis>
|
||
Photos of
|
||
<taxonomicName id="C0A2F48B1306BF17FE2DF8CEFDD14553" authority="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz, 2022" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[402,578,1864,1888]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="ulli" status="sp. nov.">
|
||
<emphasis id="35D6531A1306BF17FE2DF8CEFDD14553" box="[402,578,1864,1888]" italics="true" pageId="44" pageNumber="45">Lygodactylus ulli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="35D6531A1306BF17FDF6F8CEFD084553" bold="true" box="[585,667,1864,1888]" pageId="44" pageNumber="45">
|
||
<taxonomicNameLabel id="2EE5EE611306BF17FDF6F8CEFD084553" box="[585,667,1864,1888]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, specimen from Marojejy in life. A, male holotype ZSM 154/2005 (FGZC 2811). B, C, D, female specimen UADBA-R-MSZC 272 in dorsolateral, dorsal, and ventral views. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="4FB8DC831307BF16FF78FF10FC624196" pageId="45" pageNumber="46" type="description">
|
||
<paragraph id="071D8F081307BF16FF78FF10FB544052" blockId="45.[151,1437,150,1041]" pageId="45" pageNumber="46">
|
||
<emphasis id="35D6531A1307BF16FF78FF10FD804282" bold="true" box="[199,531,150,177]" pageId="45" pageNumber="46">
|
||
Description of the
|
||
<typeStatus id="D81931AA1307BF16FE1AFF11FD9D4282" box="[421,526,151,177]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male, hemipenes everted, in moderate state of preservation, tail is broken and missing, right forelimb is removed as source of tissue for molecular analysis (
|
||
<figureCitation id="9F99938D1307BF16FBF5FF3DFB3342E6" box="[1098,1184,186,213]" captionStart="FIGURE 15" captionStartId="25.[151,250,1943,1967]" captionTargetBox="[244,1351,181,1906]" captionTargetId="figure-18@25.[236,1352,181,1919]" captionTargetPageId="25" captionText="FIGURE 15. Dorsal and ventral views of preserved holotypes of new species of the Lygodactylus madagascariensis group named in the current study." figureDoi="http://doi.org/10.5281/zenodo.7040784" httpUri="https://zenodo.org/record/7040784/files/figure.png" pageId="45" pageNumber="46">Fig. 15</figureCitation>
|
||
). SVL
|
||
<quantity id="C05A22ED1307BF16FB4BFF3DFACF42E6" box="[1268,1372,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.88" pageId="45" pageNumber="46" unit="mm" value="28.8">28.8 mm</quantity>
|
||
, TAL
|
||
<quantity id="C05A22ED1307BF16FF28FF59FF6342CA" box="[151,240,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="45" pageNumber="46" unit="mm" value="2.6">2.6 mm</quantity>
|
||
; for other measurements see
|
||
<tableCitation id="4A20BAB31307BF16FD89FF59FD1942CA" box="[566,650,223,249]" captionStart="TABLE 1" captionStartId="31.[151,239,459,485]" captionTargetPageId="31" captionText="TABLE 1. Selected morphometric and meristic data for voucher specimens of the Lygodactylus madagascariensis group examined for this study. All morphometric measurements (SVL, TAL, HIL, HW, ED, SED) in mm. All specimens in the table were genotyped by 16S sequences, except for historical types from BMNH, MNHN, and SMF, and two further individuals of L. expectatus. Abbreviations are those described in Materials and Methods section; additional abbreviations and symbols as follows: NA, not assessed or not applicable; * counts and measurements partly or fully taken by M. Puente (Puente et al. 2009); § specimen not genotyped; # tail broken or incomplete; ## tail regenerated. HT (sep) and HT (spi) mark the holotypes of Lygodactylus septemtuberculatus Angel, 1942 and Microscalabotes spinulifer Boettger, 1913, here considered to be junior synonyms of L. miops. First LCDS value for L. expectatus refers to the enlarged dorsolateral scales, the second value to the non-enlarged vertebral scales. See species account of L. fritzi sp. nov for the identity of the historical locality “Senbendrana”." httpUri="http://table.plazi.org/id/53DDDF801335BF24FF28FE4DFD0E408E" pageId="45" pageNumber="46" tableUuid="53DDDF801335BF24FF28FE4DFD0E408E">Table 1</tableCitation>
|
||
. Head broader than body. The distance from the tip of the snout to the anterior border of the eye (
|
||
<quantity id="C05A22ED1307BF16FE7BFE84FD8F432E" box="[452,540,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="45" pageNumber="46" unit="mm" value="3.5">3.5 mm</quantity>
|
||
) is lesser than the interorbital distance anteriorly (
|
||
<quantity id="C05A22ED1307BF16FBF0FE85FB38432E" box="[1103,1195,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="45" pageNumber="46" unit="mm" value="3.9">3.9 mm</quantity>
|
||
), and greater than the distance between the eye and ear opening. Snout covered with granular scales larger than those on the rest of the dorsum. Nostril surrounded by five scales: rostral, first supralabial, and three supranasals. Mental scale undivided; no contact between posterior projection of mental scale and first infralabial; three asymmetrical postmental scales with five postpostmental scales; six infralabial scales; seven supralabial scales; two internasal scales; granular dorsal scales; dorsum with small, homogeneous, granular, and unkeeled scales of similar size to those on trunk, no distinct size difference to scales on limbs; 253 dorsal scales longitudinally along the body; 110 ventral scales between mental and cloaca; venter with large homogeneous smooth scales; first finger present but very small, not bearing a claw; three pairs of subdigital lamellae on the fourth toe; two not very distinct dorsolateral tubercles, each consisting of one scale; seven preanal pores; tail without whorls; small lateral spines at the base of the tail.
|
||
</paragraph>
|
||
<paragraph id="071D8F081307BF16FF78FDEDFC664126" blockId="45.[151,1437,150,1041]" pageId="45" pageNumber="46">
|
||
Based on available photographs (
|
||
<figureCitation id="9F99938D1307BF16FD81FDEDFD0240B6" box="[574,657,619,645]" captionStart="FIGURE 21" captionStartId="44.[151,250,1864,1888]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-18@44.[151,1436,181,1840]" captionTargetPageId="44" captionText="FIGURE 21. Photos of Lygodactylus ulli sp. nov., specimen from Marojejy in life. A, male holotype ZSM 154/2005 (FGZC 2811). B, C, D, female specimen UADBA-R-MSZC 272 in dorsolateral, dorsal, and ventral views. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7040809" httpUri="https://zenodo.org/record/7040809/files/figure.png" pageId="45" pageNumber="46">Fig. 21</figureCitation>
|
||
), the live
|
||
<typeStatus id="D81931AA1307BF16FCBCFDEDFCF640B6" box="[771,869,619,645]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
displays a brownish to grayish pattern on the dorsum and the limbs. From the snout, a narrow black stripe runs irregularly to two elongated spots on the shoulder. At the forelimb level two symmetrical black spots are present on the spine (
|
||
<figureCitation id="9F99938D1307BF16FC6DFD35FBAF40FE" box="[978,1084,691,717]" captionStart="FIGURE 21" captionStartId="44.[151,250,1864,1888]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-18@44.[151,1436,181,1840]" captionTargetPageId="44" captionText="FIGURE 21. Photos of Lygodactylus ulli sp. nov., specimen from Marojejy in life. A, male holotype ZSM 154/2005 (FGZC 2811). B, C, D, female specimen UADBA-R-MSZC 272 in dorsolateral, dorsal, and ventral views. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7040809" httpUri="https://zenodo.org/record/7040809/files/figure.png" pageId="45" pageNumber="46">Fig. 21A</figureCitation>
|
||
). After 16 years in ethanol, the preserved specimen is more grayish. The ventral side is whitish. While there are only a few small brown spots on the venter, multiple larger brown spots are irregularly scattered on the throat.
|
||
</paragraph>
|
||
<paragraph id="071D8F081307BF16FF78FC98FC624196" blockId="45.[151,1437,150,1041]" pageId="45" pageNumber="46">
|
||
<emphasis id="35D6531A1307BF16FF78FC98FEAC410B" bold="true" box="[199,319,798,824]" pageId="45" pageNumber="46">Variation.</emphasis>
|
||
Comparing the two photographed specimens, dorsal coloration and patterns are less pronounced on the male (
|
||
<figureCitation id="9F99938D1307BF16FEB8FCC5FEFC416E" box="[263,367,835,861]" captionStart="FIGURE 21" captionStartId="44.[151,250,1864,1888]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-18@44.[151,1436,181,1840]" captionTargetPageId="44" captionText="FIGURE 21. Photos of Lygodactylus ulli sp. nov., specimen from Marojejy in life. A, male holotype ZSM 154/2005 (FGZC 2811). B, C, D, female specimen UADBA-R-MSZC 272 in dorsolateral, dorsal, and ventral views. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7040809" httpUri="https://zenodo.org/record/7040809/files/figure.png" pageId="45" pageNumber="46">Fig. 21A</figureCitation>
|
||
), maybe because it appears to be close to skin shedding. The female has a blackish and grayish alternating dorsal pattern, surrounded by a yellowish base layer (
|
||
<figureCitation id="9F99938D1307BF16FCC8FCE1FC4F41B2" box="[887,988,871,897]" captionStart="FIGURE 21" captionStartId="44.[151,250,1864,1888]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-18@44.[151,1436,181,1840]" captionTargetPageId="44" captionText="FIGURE 21. Photos of Lygodactylus ulli sp. nov., specimen from Marojejy in life. A, male holotype ZSM 154/2005 (FGZC 2811). B, C, D, female specimen UADBA-R-MSZC 272 in dorsolateral, dorsal, and ventral views. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7040809" httpUri="https://zenodo.org/record/7040809/files/figure.png" pageId="45" pageNumber="46">Fig. 21B</figureCitation>
|
||
+C). The female is laterally darker than the male (
|
||
<figureCitation id="9F99938D1307BF16FEB9FC0DFEFD4196" box="[262,366,907,933]" captionStart="FIGURE 21" captionStartId="44.[151,250,1864,1888]" captionTargetBox="[151,1436,181,1820]" captionTargetId="figure-18@44.[151,1436,181,1840]" captionTargetPageId="44" captionText="FIGURE 21. Photos of Lygodactylus ulli sp. nov., specimen from Marojejy in life. A, male holotype ZSM 154/2005 (FGZC 2811). B, C, D, female specimen UADBA-R-MSZC 272 in dorsolateral, dorsal, and ventral views. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7040809" httpUri="https://zenodo.org/record/7040809/files/figure.png" pageId="45" pageNumber="46">Fig. 21A</figureCitation>
|
||
+B), has a white throat and yellow venter and ventral tail.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FB8DC831307BF16FF78FC29FC3641FA" box="[199,933,942,969]" pageId="45" pageNumber="46" type="biology_ecology">
|
||
<paragraph id="071D8F081307BF16FF78FC29FC3641FA" blockId="45.[151,1437,150,1041]" box="[199,933,942,969]" pageId="45" pageNumber="46">
|
||
<emphasis id="35D6531A1307BF16FF78FC29FE1641FB" bold="true" box="[199,389,942,969]" pageId="45" pageNumber="46">Natural history.</emphasis>
|
||
Specimens were collected in primary rainforest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FB8DC831307BF16FF78FC54FEB64622" pageId="45" pageNumber="46" type="distribution">
|
||
<paragraph id="071D8F081307BF16FF78FC54FEB64622" blockId="45.[151,1437,150,1041]" pageId="45" pageNumber="46">
|
||
<emphasis id="35D6531A1307BF16FF78FC54FEF241DF" bold="true" box="[199,353,978,1004]" pageId="45" pageNumber="46">Distribution.</emphasis>
|
||
<taxonomicName id="C0A2F48B1307BF16FED4FC55FE2341DE" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[363,432,979,1005]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="ulli">
|
||
<emphasis id="35D6531A1307BF16FED4FC55FE2341DE" box="[363,432,979,1005]" italics="true" pageId="45" pageNumber="46">L. ulli</emphasis>
|
||
</taxonomicName>
|
||
is only known from its
|
||
<typeStatus id="D81931AA1307BF16FD70FC55FD6C41DE" box="[719,767,979,1005]" pageId="45" pageNumber="46">type</typeStatus>
|
||
locality, the Marojejy Massif in the North East region of
|
||
<collectingCountry id="7FB5CF981307BF16FF28FC71FEB34622" box="[151,288,1015,1041]" name="Madagascar" pageId="45" pageNumber="46">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |