544 lines
46 KiB
XML
544 lines
46 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys..47983" ID-GBIF-Dataset="35113d62-0ccc-4e15-a673-8893bf800658" ID-PMC="PMC7316820" ID-Pensoft-Pub="1313-2970-942-105" ID-Pensoft-UUID="182B102E3B9357318B7B0A275F6D3560" ID-PubMed="32612444" ID-ZooBank="96C50618823C4800934897879B6B3B06" ModsDocID="1313-2970-942-105" checkinTime="1592583863628" checkinUser="pensoft" docAuthor="Lyu, Zhi-Tong, Li, Yuan-Qiu, Zeng, Zhao-Chi, Zhao, Jian, Liu, Zu-Yao, Guo, Guo-Xin & Wang, Ying-Yong" docDate="2020" docId="8EEA74E7F4775D9E9511762392E3F325" docLanguage="en" docName="ZooKeys 942: 105-140" docOrigin="ZooKeys 942" docSource="http://dx.doi.org/10.3897/zookeys..47983" docTitle="Megophrys (Panophrys) mirabilis Lyu, Wang & Zhao 2020, sp. nov." docType="treatment" docUuid="E624C3F8-5522-4A3C-B376-3519B7E5A377" docUuidSource="ZooBank" docVersion="5" id="182B102E3B9357318B7B0A275F6D3560" lastPageNumber="105" masterDocId="182B102E3B9357318B7B0A275F6D3560" masterDocTitle="Four new species of Asian horned toads (Anura, Megophryidae, Megophrys) from southern China" masterLastPageNumber="140" masterPageNumber="105" pageNumber="105" updateTime="1668168982812" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Four new species of Asian horned toads (Anura, Megophryidae, Megophrys) from southern China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lyu, Zhi-Tong</mods:namePart>
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<mods:affiliation>State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China & School of Ecology, Sun Yat-sen University, Guangzhou 510006, China</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4891-3859</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Yuan-Qiu</mods:namePart>
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<mods:affiliation>Guangdong Shimentai National Nature Reserve, Qingyuan 513000, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zeng, Zhao-Chi</mods:namePart>
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<mods:affiliation>State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China & Shenzhen Shuanghuamu Biological Technology Co., LTD, Shenzhen 51800, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhao, Jian</mods:namePart>
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<mods:affiliation>Shenzhen Shuanghuamu Biological Technology Co., LTD, Shenzhen 51800, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liu, Zu-Yao</mods:namePart>
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<mods:affiliation>State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China & Institute of Ecology and Evolution, University of Bern, Bern 3012, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Guo, Guo-Xin</mods:namePart>
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<mods:affiliation>Guangdong Shimentai National Nature Reserve, Qingyuan 513000, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wang, Ying-Yong</mods:namePart>
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<mods:affiliation>State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
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<mods:nameIdentifier type="email">wangyy@mail.sysu.edu.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>942</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>105</mods:start>
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<mods:end>140</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys..47983</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys..47983</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-942-105</mods:identifier>
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<mods:identifier type="ZooBank">96C50618823C4800934897879B6B3B06</mods:identifier>
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<mods:identifier type="Pensoft-UUID">182B102E3B9357318B7B0A275F6D3560</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="164674246" LSID="urn:lsid:zoobank.org:act:E624C3F8-5522-4A3C-B376-3519B7E5A377" httpUri="http://treatment.plazi.org/id/8EEA74E7F4775D9E9511762392E3F325" lastPageNumber="105" pageId="0" pageNumber="105">
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<subSubSection pageId="0" pageNumber="105" type="nomenclature">
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<paragraph pageId="0" pageNumber="105">
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<taxonomicName LSID="8EEA74E7-F477-5D9E-9511-762392E3F325" authority="Lyu, Wang & Zhao" authorityName="Lyu, Wang & Zhao" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophrys (Panophrys) mirabilis" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="species" species="mirabilis" status="sp. nov." subGenus="Panophrys">Megophrys (Panophrys) mirabilis Lyu, Wang & Zhao</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="105">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Adult male holotype SYS a 002917 of Megophrys (Panophrys) mirabilis sp. nov. in life A dorsolateral view B ventral view C hand D foot." figureDoi="10.3897/zookeys..47983.figure3" httpUri="https://binary.pensoft.net/fig/423057" pageId="0" pageNumber="105">Figures 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. The holotype specimen of each new species in preservative A Megophrys (Panophrys) mirabilis sp. nov. B M. (Pa.) shimentaina sp. nov. C M. (Pa.) xiangnanensis sp. nov. D M. (Pa.) yangmingensis sp. nov. 1 dorsal view 2 ventral view 3 lateral view." figureDoi="10.3897/zookeys..47983.figure4" httpUri="https://binary.pensoft.net/fig/423058" pageId="0" pageNumber="105">, 4A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="105" type="chresonymy">
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<paragraph pageId="0" pageNumber="105">Chresonymy.</paragraph>
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<paragraph pageId="0" pageNumber="105">
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<taxonomicName authorityName="Kuhl & van Hasselt" authorityYear="1822" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophrys" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="105">Megophrys</emphasis>
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</taxonomicName>
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sp25 (SYS a002192-93, 2289, 2917 in
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<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2018.06.020" author="Liu, ZY" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="105" pagination="723 - 731" refId="B15" refString="Liu, ZY, Chen, GL, Zhu, TQ, Zeng, ZC, Lyu, ZT, Wang, J, Messenger, K, Greenberg, AJ, Guo, ZX, Yang, ZH, Shi, SH, Wang, YY, 2018. Prevalence of cryptic species in morphologically uniform taxa - Fast speciation and evolutionary radiation in Asian frogs. Molecular Phylogenetics and Evolution 127: 723 - 731, DOI: https://doi.org/10.1016/j.ympev.2018.06.020" title="Prevalence of cryptic species in morphologically uniform taxa - Fast speciation and evolutionary radiation in Asian frogs." url="https://doi.org/10.1016/j.ympev.2018.06.020" volume="127" year="2018">Liu et al. 2018</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="105" type="type material">
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<paragraph pageId="0" pageNumber="105">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="105">
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<emphasis bold="true" pageId="0" pageNumber="105">
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<emphasis italics="true" pageId="0" pageNumber="105">Holotype</emphasis>
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.
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</emphasis>
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SYS a002917 (Figs
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Adult male holotype SYS a 002917 of Megophrys (Panophrys) mirabilis sp. nov. in life A dorsolateral view B ventral view C hand D foot." figureDoi="10.3897/zookeys..47983.figure3" httpUri="https://binary.pensoft.net/fig/423057" pageId="0" pageNumber="105">3</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. The holotype specimen of each new species in preservative A Megophrys (Panophrys) mirabilis sp. nov. B M. (Pa.) shimentaina sp. nov. C M. (Pa.) xiangnanensis sp. nov. D M. (Pa.) yangmingensis sp. nov. 1 dorsal view 2 ventral view 3 lateral view." figureDoi="10.3897/zookeys..47983.figure4" httpUri="https://binary.pensoft.net/fig/423058" pageId="0" pageNumber="105">4A</figureCitation>
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), adult male, collected on 16 June 2014 by Yu-Long Li and Ying-Yong Wang from Huaping Nature Reserve (
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<geoCoordinate degrees="25.5554" direction="north" orientation="latitude" precision="5" value="25.5554">25.5554N</geoCoordinate>
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,
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<geoCoordinate degrees="109.9490" direction="east" orientation="longitude" precision="5" value="109.949">109.9490E</geoCoordinate>
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; ca 1300 m a.s.l.), Lingui District, Guilin City, Guangxi Zhuang Autonomous Region, PR China.
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</paragraph>
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<paragraph pageId="0" pageNumber="105">
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<emphasis bold="true" pageId="0" pageNumber="105">
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<emphasis italics="true" pageId="0" pageNumber="105">Paratypes</emphasis>
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.
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</emphasis>
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Three adult specimens from the same locality as the holotype: male SYS a002192 and female SYS a002193 collected on 10 July 2013 by Jian Zhao and Yu-Long Li; female SYS a002289 collected on 9 September 2013 by Zu-Yao Liu.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="105" type="etymology">
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<paragraph pageId="0" pageNumber="105">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="105">
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The specific epithet
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<taxonomicName authorityName="Lyu, Wang & Zhao" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="mirabilis" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="species" species="mirabilis">
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<emphasis italics="true" pageId="0" pageNumber="105">mirabilis</emphasis>
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</taxonomicName>
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means marvelous, referring to its distinctive habitus and color pattern of this species within the subgenus
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<taxonomicName class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="(Panophrys)" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="subGenus" subGenus="Panophrys">Panophrys</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="105" type="common name">
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<paragraph pageId="0" pageNumber="105">Common name.</paragraph>
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<paragraph pageId="0" pageNumber="105">
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Huaping Horned Toad (in English) /
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<normalizedToken originalValue="Huā">Hua</normalizedToken>
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<normalizedToken originalValue="Píng">Ping</normalizedToken>
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<normalizedToken originalValue="Jiăo">Jiăo</normalizedToken>
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<normalizedToken originalValue="Chán">Chan</normalizedToken>
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(花坪角蟾 in Chinese)
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</paragraph>
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<caption doi="10.3897/zookeys..47983.figure3" httpUri="https://binary.pensoft.net/fig/423057" pageId="0" pageNumber="105" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="105">
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<emphasis bold="true" pageId="0" pageNumber="105">Figure 3.</emphasis>
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Adult male holotype SYS a002917 of
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<taxonomicName authorityName="Lyu, Wang & Zhao" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophrys (Panophrys) mirabilis" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="species" species="mirabilis" subGenus="Panophrys">Megophrys (Panophrys) mirabilis</taxonomicName>
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sp. nov. in life
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<emphasis bold="true" pageId="0" pageNumber="105">A</emphasis>
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dorsolateral view
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<emphasis bold="true" pageId="0" pageNumber="105">B</emphasis>
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ventral view
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<emphasis bold="true" pageId="0" pageNumber="105">C</emphasis>
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hand
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<emphasis bold="true" pageId="0" pageNumber="105">D</emphasis>
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foot.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="105" type="diagnosis">
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<paragraph pageId="0" pageNumber="105">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="105">
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(1) Body size relatively large, SVL 55.8-61.4 mm (
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<emphasis italics="true" pageId="0" pageNumber="105">N</emphasis>
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= 2) in adult males and SVL 68.5-74.8 (
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<emphasis italics="true" pageId="0" pageNumber="105">N</emphasis>
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= 2) mm in adult females; (2) snout rounded in dorsal view; (3) internasal distance smaller than interorbital distance; (4) tympanum clear, moderate size, TD/ED 0.49-0.63; (5) absence of vomerine ridge and vomerine teeth; (6) tongue small, majorly attached to the mandible, free margin small and rounded, not notched behind; (7) hindlimbs slender, heels overlapping and tibio-tarsal articulation reaching forward at the central eye; (8) fingers with distinct lateral fringes, presence of indistinct subarticular tubercles at the bases; (9) toes with distinct lateral fringes and rudiment of webs, presence of indistinct subarticular tubercles at the bases; (10) presence of slightly large horn-like tubercle at the edge of upper eyelid; (11) dorsal skin smooth with granules, (12) skin on flanks flabby, with spiny tubercles; (13) supratympanic fold distinct, with dense tubercles, forming an extremely swollen large shoulder gland above insertion of arm; (14) grayish brown above, tinged with blue in males, but dorsum of head and body reddish brown in females; (15) ventral surface of throat and chest with grayish blue latticed patches and black spots in males, but with orange latticed patches and black spots in females; (16) presence of underdeveloped nuptial pads on the dorsal surface of the first finger in adult males.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="105" type="comparisons">
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<paragraph pageId="0" pageNumber="105">Comparison.</paragraph>
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<paragraph pageId="0" pageNumber="105">
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<taxonomicName authorityName="Lyu, Wang & Zhao" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophrys (Panophrys) mirabilis" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="species" species="mirabilis" subGenus="Panophrys">Megophrys (Panophrys) mirabilis</taxonomicName>
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sp. nov. can be easily distinguished from all recognized congeners, by having a small tongue, majorly attached to the mandible, flank skin flabby with spiny tubercles, and supratympanic fold with dense tubercles forming an extremely swollen large shoulder gland above insertion of arm.
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</paragraph>
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<paragraph pageId="0" pageNumber="105">
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Further, detailed comparative data of
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<taxonomicName authorityName="Lyu, Wang & Zhao" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophrys (Panophrys) mirabilis" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="species" species="mirabilis" subGenus="Panophrys">Megophrys (Panophrys) mirabilis</taxonomicName>
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sp. nov. with 42 recognized congeners of
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<taxonomicName class="Amphibia" family="Megophryidae" genus="Panophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Panophrys" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="105">Panophrys</emphasis>
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</taxonomicName>
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are given in Table
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<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Diagnostic characters separating all 46 species of the Megophrys (Panophrys)." httpUri="http://table.plazi.org/id/3A3F83144332F1C21D91B504ADBE2530" pageId="0" pageNumber="105" tableUuid="3A3F83144332F1C21D91B504ADBE2530">4</tableCitation>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="105">
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Five
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<taxonomicName class="Amphibia" family="Megophryidae" genus="Panophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Panophrys" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="105">Panophrys</emphasis>
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</taxonomicName>
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species were previously recorded from the hilly areas among Guangdong, Guangxi, and Hunan, namely
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<taxonomicName authorityName="L.i.Wang & Jin" authorityYear="2014" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophrys (Panophrys) acuta" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="species" species="acuta" subGenus="Panophrys">Megophrys (Panophrys) acuta</taxonomicName>
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,
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<taxonomicName lsidName="M. (Pa.) brachykolos" pageId="0" pageNumber="105" rank="species" species="brachykolos" subGenus="Pa.">M. (Pa.) brachykolos</taxonomicName>
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,
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<taxonomicName lsidName="M. (Pa.) nanlingensis" pageId="0" pageNumber="105" rank="species" species="nanlingensis" subGenus="Pa.">M. (Pa.) nanlingensis</taxonomicName>
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,
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<taxonomicName lsidName="M. (Pa.) obesa" pageId="0" pageNumber="105" rank="species" species="obesa" subGenus="Pa.">M. (Pa.) obesa</taxonomicName>
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, and
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<taxonomicName lsidName="M. (Pa.) shunhuangensis" pageId="0" pageNumber="105" rank="species" species="shunhuangensis" subGenus="Pa.">M. (Pa.) shunhuangensis</taxonomicName>
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.
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<taxonomicName lsidName="M. (Pa.) mirabilis" pageId="0" pageNumber="105" rank="species" species="mirabilis" subGenus="Pa.">M. (Pa.) mirabilis</taxonomicName>
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sp. nov. differs from
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<taxonomicName lsidName="M. (Pa.) acuta" pageId="0" pageNumber="105" rank="species" species="acuta" subGenus="Pa.">M. (Pa.) acuta</taxonomicName>
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by the larger body size, SVL 55.8-61.4 mm in males and 68.5-74.8 mm in females (vs. 27.1-33.0 mm in males and 28.1-33.6 in females), snout rounded in dorsal view (vs. strongly remarkably pointed), fingers with distinct lateral fringes (vs. absent), and overlapping heels (vs. not meeting).
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<taxonomicName lsidName="M. (Pa.) mirabilis" pageId="0" pageNumber="105" rank="species" species="mirabilis" subGenus="Pa.">M. (Pa.) mirabilis</taxonomicName>
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sp. nov. differs from
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<taxonomicName lsidName="M. (Pa.) brachykolos" pageId="0" pageNumber="105" rank="species" species="brachykolos" subGenus="Pa.">M. (Pa.) brachykolos</taxonomicName>
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by the larger body size, SVL 55.8-61.4 mm in males and 68.5-74.8 mm in females (vs. 33.7-39.3 mm in males and 33.9-45.9 in females), slightly large horn-like tubercle at upper eyelid (vs. small), fingers and toes with distinct lateral fringes (vs. all absent), overlapping heels (vs. not meeting).
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<taxonomicName lsidName="M. (Pa.) mirabilis" pageId="0" pageNumber="105" rank="species" species="mirabilis" subGenus="Pa.">M. (Pa.) mirabilis</taxonomicName>
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sp. nov. differs from
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<taxonomicName lsidName="M. (Pa.) nanlingensis" pageId="0" pageNumber="105" rank="species" species="nanlingensis" subGenus="Pa.">M. (Pa.) nanlingensis</taxonomicName>
|
|
by the larger body size, SVL 55.8-61.4 mm in males (vs. 30.5-37.3 mm), slightly large horn-like tubercle at upper eyelid (vs. small), absence of vomerine ridge and vomerine teeth (vs. both present), tongue not notched behind (vs. notched), and fingers with distinct lateral fringes (vs. absent).
|
|
<taxonomicName lsidName="M. (Pa.) mirabilis" pageId="0" pageNumber="105" rank="species" species="mirabilis" subGenus="Pa.">M. (Pa.) mirabilis</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="M. (Pa.) obesa" pageId="0" pageNumber="105" rank="species" species="obesa" subGenus="Pa.">M. (Pa.) obesa</taxonomicName>
|
|
by larger body size, SVL 55.8-61.4 mm in males and 68.5-74.8 mm in females (vs. 35.6 mm in male and 37.5-41.2 in females), slightly large horn-like tubercle at upper eyelid (vs. small), absence of vomerine ridge (vs. present), fingers and toes with distinct lateral fringes (vs. all absent), and overlapping heels (vs. not meeting).
|
|
<taxonomicName lsidName="M. (Pa.) mirabilis" pageId="0" pageNumber="105" rank="species" species="mirabilis" subGenus="Pa.">M. (Pa.) mirabilis</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="M. (Pa.) shunhuangensis" pageId="0" pageNumber="105" rank="species" species="shunhuangensis" subGenus="Pa.">M. (Pa.) shunhuangensis</taxonomicName>
|
|
by larger body size, SVL 55.8-61.4 mm in males and 68.5-74.8 mm in females (vs. 30.3-33.7 mm in males and 37.6 in female), slightly large horn-like tubercle at upper eyelid (vs. small), and fingers and toes with lateral fringes (vs. all absent).
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys..47983.figure4" httpUri="https://binary.pensoft.net/fig/423058" pageId="0" pageNumber="105" start="Figure 4" startId="F4">
|
|
<paragraph pageId="0" pageNumber="105">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">Figure 4.</emphasis>
|
|
The holotype specimen of each new species in preservative
|
|
<emphasis bold="true" pageId="0" pageNumber="105">A</emphasis>
|
|
<taxonomicName authorityName="Lyu, Wang & Zhao" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophrys (Panophrys) mirabilis" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="species" species="mirabilis" subGenus="Panophrys">Megophrys (Panophrys) mirabilis</taxonomicName>
|
|
sp. nov.
|
|
<emphasis bold="true" pageId="0" pageNumber="105">B</emphasis>
|
|
<taxonomicName lsidName="M. (Pa.) shimentaina" pageId="0" pageNumber="105" rank="species" species="shimentaina" subGenus="Pa.">M. (Pa.) shimentaina</taxonomicName>
|
|
sp. nov.
|
|
<emphasis bold="true" pageId="0" pageNumber="105">C</emphasis>
|
|
<taxonomicName lsidName="M. (Pa.) xiangnanensis" pageId="0" pageNumber="105" rank="species" species="xiangnanensis" subGenus="Pa.">M. (Pa.) xiangnanensis</taxonomicName>
|
|
sp. nov.
|
|
<emphasis bold="true" pageId="0" pageNumber="105">D</emphasis>
|
|
<taxonomicName lsidName="M. (Pa.) yangmingensis" pageId="0" pageNumber="105" rank="species" species="yangmingensis" subGenus="Pa.">M. (Pa.) yangmingensis</taxonomicName>
|
|
sp. nov.
|
|
<emphasis bold="true" pageId="0" pageNumber="105">1</emphasis>
|
|
dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="105">2</emphasis>
|
|
ventral view
|
|
<emphasis bold="true" pageId="0" pageNumber="105">3</emphasis>
|
|
lateral view.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
With a large body size, SVL 55.8-61.4 mm in adult males and 68.5-74.8 mm in adult females,
|
|
<taxonomicName authorityName="Lyu, Wang & Zhao" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophrys (Panophrys) mirabilis" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="species" species="mirabilis" subGenus="Panophrys">Megophrys (Panophrys) mirabilis</taxonomicName>
|
|
sp. nov. is significantly different from 30 congeners whose SVL <50 mm in males or <60 mm in females, namely
|
|
<taxonomicName lsidName="M. (Pa.) baolongensis" pageId="0" pageNumber="105" rank="species" species="baolongensis" subGenus="Pa.">M. (Pa.) baolongensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) binchuanensis" pageId="0" pageNumber="105" rank="species" species="binchuanensis" subGenus="Pa.">M. (Pa.) binchuanensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) boettgeri" pageId="0" pageNumber="105" rank="species" species="boettgeri" subGenus="Pa.">M. (Pa.) boettgeri</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) cheni" pageId="0" pageNumber="105" rank="species" species="cheni" subGenus="Pa.">M. (Pa.) cheni</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) daweimontis" pageId="0" pageNumber="105" rank="species" species="daweimontis" subGenus="Pa.">M. (Pa.) daweimontis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) dongguanensis" pageId="0" pageNumber="105" rank="species" species="dongguanensis" subGenus="Pa.">M. (Pa.) dongguanensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) fansipanensis" pageId="0" pageNumber="105" rank="species" species="fansipanensis" subGenus="Pa.">M. (Pa.) fansipanensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) hoanglienensis" pageId="0" pageNumber="105" rank="species" species="hoanglienensis" subGenus="Pa.">M. (Pa.) hoanglienensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) huangshanensis" pageId="0" pageNumber="105" rank="species" species="huangshanensis" subGenus="Pa.">M. (Pa.) huangshanensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) insularis" pageId="0" pageNumber="105" rank="species" species="insularis" subGenus="Pa.">M. (Pa.) insularis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) jiangi" pageId="0" pageNumber="105" rank="species" species="jiangi" subGenus="Pa.">M. (Pa.) jiangi</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) jinggangensis" pageId="0" pageNumber="105" rank="species" species="jinggangensis" subGenus="Pa.">M. (Pa.) jinggangensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) jiulianensis" pageId="0" pageNumber="105" rank="species" species="jiulianensis" subGenus="Pa.">M. (Pa.) jiulianensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) kuatunensis" pageId="0" pageNumber="105" rank="species" species="kuatunensis" subGenus="Pa.">M. (Pa.) kuatunensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) latidactyla" pageId="0" pageNumber="105" rank="species" species="latidactyla" subGenus="Pa.">M. (Pa.) latidactyla</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) leishanensis" pageId="0" pageNumber="105" rank="species" species="leishanensis" subGenus="Pa.">M. (Pa.) leishanensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) lini" pageId="0" pageNumber="105" rank="species" species="lini" subGenus="Pa.">M. (Pa.) lini</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) lishuiensis" pageId="0" pageNumber="105" rank="species" species="lishuiensis" subGenus="Pa.">M. (Pa.) lishuiensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) minor" pageId="0" pageNumber="105" rank="species" species="minor" subGenus="Pa.">M. (Pa.) minor</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) mufumontana" pageId="0" pageNumber="105" rank="species" species="mufumontana" subGenus="Pa.">M. (Pa.) mufumontana</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) nankunensis" pageId="0" pageNumber="105" rank="species" species="nankunensis" subGenus="Pa.">M. (Pa.) nankunensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) ombrophila" pageId="0" pageNumber="105" rank="species" species="ombrophila" subGenus="Pa.">M. (Pa.) ombrophila</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) palpebralespinosa" pageId="0" pageNumber="105" rank="species" species="palpebralespinosa" subGenus="Pa.">M. (Pa.) palpebralespinosa</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) rubrimera" pageId="0" pageNumber="105" rank="species" species="rubrimera" subGenus="Pa.">M. (Pa.) rubrimera</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) spinata" pageId="0" pageNumber="105" rank="species" species="spinata" subGenus="Pa.">M. (Pa.) spinata</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) tuberogranulatus" pageId="0" pageNumber="105" rank="species" species="tuberogranulatus" subGenus="Pa.">M. (Pa.) tuberogranulatus</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) wugongensis" pageId="0" pageNumber="105" rank="species" species="wugongensis" subGenus="Pa.">M. (Pa.) wugongensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) wuliangshanensis" pageId="0" pageNumber="105" rank="species" species="wuliangshanensis" subGenus="Pa.">M. (Pa.) wuliangshanensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) wushanensis" pageId="0" pageNumber="105" rank="species" species="wushanensis" subGenus="Pa.">M. (Pa.) wushanensis</taxonomicName>
|
|
, and
|
|
<taxonomicName lsidName="M. (Pa.) xianjuensis" pageId="0" pageNumber="105" rank="species" species="xianjuensis" subGenus="Pa.">M. (Pa.) xianjuensis</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
<taxonomicName authorityName="Lyu, Wang & Zhao" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophrys (Panophrys) mirabilis" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="species" species="mirabilis" subGenus="Panophrys">Megophrys (Panophrys) mirabilis</taxonomicName>
|
|
sp. nov. can be further distinguished from the remaining seven congeners by the following characteristics: SVL 55.8-61.4 mm in adult males and 68.5-74.8 mm in adult females [vs. SVL 45.1-51.0 mm in adult males in
|
|
<taxonomicName lsidName="M. (Pa.) binlingensis" pageId="0" pageNumber="105" rank="species" species="binlingensis" subGenus="Pa.">M. (Pa.) binlingensis</taxonomicName>
|
|
; vs. SVL 81.3 mm in adult male in
|
|
<taxonomicName lsidName="M. (Pa.) caudoprocta" pageId="0" pageNumber="105" rank="species" species="caudoprocta" subGenus="Pa.">M. (Pa.) caudoprocta</taxonomicName>
|
|
; vs. SVL 63.5 mm in adult female in
|
|
<taxonomicName lsidName="M. (Pa.) jingdongensis" pageId="0" pageNumber="105" rank="species" species="jingdongensis" subGenus="Pa.">M. (Pa.) jingdongensis</taxonomicName>
|
|
; vs. SVL 102.0-118.3 mm in adult males and 99.8-115.6 mm in adult females in
|
|
<taxonomicName lsidName="M. (Pa.) shuichengensis" pageId="0" pageNumber="105" rank="species" species="shuichengensis" subGenus="Pa.">M. (Pa.) shuichengensis</taxonomicName>
|
|
]; slightly large horn-like tubercle at upper eyelid [vs. small in
|
|
<taxonomicName lsidName="M. (Pa.) binlingensis" pageId="0" pageNumber="105" rank="species" species="binlingensis" subGenus="Pa.">M. (Pa.) binlingensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) jingdongensis" pageId="0" pageNumber="105" rank="species" species="jingdongensis" subGenus="Pa.">M. (Pa.) jingdongensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) omeimontis" pageId="0" pageNumber="105" rank="species" species="omeimontis" subGenus="Pa.">M. (Pa.) omeimontis</taxonomicName>
|
|
, and
|
|
<taxonomicName lsidName="M. (Pa.) sangzhiensis" pageId="0" pageNumber="105" rank="species" species="sangzhiensis" subGenus="Pa.">M. (Pa.) sangzhiensis</taxonomicName>
|
|
]; vomerine teeth absent [vs. present in
|
|
<taxonomicName lsidName="M. (Pa.) caudoprocta" pageId="0" pageNumber="105" rank="species" species="caudoprocta" subGenus="Pa.">M. (Pa.) caudoprocta</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) jingdongensis" pageId="0" pageNumber="105" rank="species" species="jingdongensis" subGenus="Pa.">M. (Pa.) jingdongensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) liboensis" pageId="0" pageNumber="105" rank="species" species="liboensis" subGenus="Pa.">M. (Pa.) liboensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) omeimontis" pageId="0" pageNumber="105" rank="species" species="omeimontis" subGenus="Pa.">M. (Pa.) omeimontis</taxonomicName>
|
|
, and
|
|
<taxonomicName lsidName="M. (Pa.) sangzhiensis" pageId="0" pageNumber="105" rank="species" species="sangzhiensis" subGenus="Pa.">M. (Pa.) sangzhiensis</taxonomicName>
|
|
]; tongue not notched behind [vs. notched in
|
|
<taxonomicName lsidName="M. (Pa.) binlingensis" pageId="0" pageNumber="105" rank="species" species="binlingensis" subGenus="Pa.">M. (Pa.) binlingensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) jingdongensis" pageId="0" pageNumber="105" rank="species" species="jingdongensis" subGenus="Pa.">M. (Pa.) jingdongensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) liboensis" pageId="0" pageNumber="105" rank="species" species="liboensis" subGenus="Pa.">M. (Pa.) liboensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) omeimontis" pageId="0" pageNumber="105" rank="species" species="omeimontis" subGenus="Pa.">M. (Pa.) omeimontis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) sangzhiensis" pageId="0" pageNumber="105" rank="species" species="sangzhiensis" subGenus="Pa.">M. (Pa.) sangzhiensis</taxonomicName>
|
|
, and
|
|
<taxonomicName lsidName="M. (Pa.) shuichengensis" pageId="0" pageNumber="105" rank="species" species="shuichengensis" subGenus="Pa.">M. (Pa.) shuichengensis</taxonomicName>
|
|
]; lateral fringes on toes narrow [vs. wide in
|
|
<taxonomicName lsidName="M. (Pa.) jingdongensis" pageId="0" pageNumber="105" rank="species" species="jingdongensis" subGenus="Pa.">M. (Pa.) jingdongensis</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="M. (Pa.) liboensis" pageId="0" pageNumber="105" rank="species" species="liboensis" subGenus="Pa.">M. (Pa.) liboensis</taxonomicName>
|
|
, and
|
|
<taxonomicName lsidName="M. (Pa.) shuichengensis" pageId="0" pageNumber="105" rank="species" species="shuichengensis" subGenus="Pa.">M. (Pa.) shuichengensis</taxonomicName>
|
|
]; rudimentary webs on toes [vs. more than one-fourth webs in
|
|
<taxonomicName lsidName="M. (Pa.) jingdongensis" pageId="0" pageNumber="105" rank="species" species="jingdongensis" subGenus="Pa.">M. (Pa.) jingdongensis</taxonomicName>
|
|
and
|
|
<taxonomicName lsidName="M. (Pa.) shuichengensis" pageId="0" pageNumber="105" rank="species" species="shuichengensis" subGenus="Pa.">M. (Pa.) shuichengensis</taxonomicName>
|
|
].
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="105" type="description">
|
|
<paragraph pageId="0" pageNumber="105">Description of holotype.</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">Adult male. Body size large, SVL 61.4 mm; head width slightly larger than head length, HDW/HDL 1.02; snout rounded in dorsal view, projecting, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eyes large, ED 0.31 of HDL, pupil vertical; nostril oblique-ovoid; canthus rostralis well developed; loreal region slightly oblique; internasal distance smaller than interorbital distance; tympanum clear, TD/ED 0.49; large ovoid choanae at the base of the maxilla; absence of vomerine ridge and vomerine teeth; tongue small, majority attached at the mouth, margin rounded, not notched behind; absence of vocal sac.</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">Radio-ulna length 0.26 of SVL and hand 0.28 of SVL; hand without webs, fingers with distinct lateral fringes, relative finger length II <I <IV <III; tips of fingers slightly dilated, round; one indistinct subarticular tubercle at the bases of each finger; metacarpal tubercles indistinct, the inner one observably enlarged and the outer one smaller; presence of underdeveloped nuptial pad on the dorsal surface of the first finger, without nuptial spines. Hindlimbs slender, tibio-tarsal articulation reaching forward at the central eye when hindlimb is stretched along the side of the body; heels overlapping when the flexed hindlimbs are held at right angles to the body axis; tibia length 0.47 of SVL and foot length 0.71 of SVL; relative toe length I <II <V <III <IV; tips of toes round and slightly dilated; toes with narrow lateral fringes and rudiment of webs; one indistinct subarticular tubercle at the bases of each toe; inner metatarsal tubercle long ovoid and the outer one absent.</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">Dorsal skin smooth with sparse granules; flanks flabby with spiny tubercles; distinct supratympanic fold curving postero-ventrally from posterior corner of eye to a level above insertion of arm; small tubercles arranged from above the nostril, along the canthus rostralis, edge of upper eyelid and supratympanic fold, to the posterior margin of temporal region; a distinct horn-like prominent tubercle on the edge of upper eyelid; a discontinuous X-shaped ridge with several short ridges on two sides on the back; transverse skin ridges on the dorsal shank and thigh; ventral surface smooth; several tubercles on posterior hindlimbs; small pectoral gland closer to axilla; a single large femoral gland on rear of thigh.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="105" type="coloration">
|
|
<paragraph pageId="0" pageNumber="105">Coloration.</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">Grayish brown above in life; an dark interorbital triangle with light colored center and edge; a dark X-shaped making with light edge on the central of dorsum; dark brown transverse bands on forearms and hindlimbs; supratympanic fold light gray; dark vertical band below the eye; iris grayish brown; ventral surface grayish white; throat and chest with grayish blue latticed patches and black spots; ventral hands and feet grayish white, tips of digits creamy white, metacarpal tubercle and metatarsal tubercle grayish white; pectoral gland and femoral gland white.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="105" type="variation">
|
|
<paragraph pageId="0" pageNumber="105">Variations.</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
Measurement data of type series are listed in Table
|
|
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Measurements (in mm) of the type series of Megophrys (Panophrys) mirabilis sp. nov., * for the holotype." httpUri="http://table.plazi.org/id/E485D427D430B52C70EC0A29AEAB4143" pageId="0" pageNumber="105" tableUuid="E485D427D430B52C70EC0A29AEAB4143">5</tableCitation>
|
|
. All paratypes are similar to the holotype. Females (SVL 68.5-74.8 mm) are significantly larger than males (SVL 55.8-61.4 mm). Dorsal surfaces reddish brown and ventral surfaces with orange latticed patches and black spots in females SYS a002193, 2289.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="105" type="distribution">
|
|
<paragraph pageId="0" pageNumber="105">Distribution and ecology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
Currently,
|
|
<taxonomicName authorityName="Lyu, Wang & Zhao" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophrys (Panophrys) mirabilis" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="species" species="mirabilis" subGenus="Panophrys">Megophrys (Panophrys) mirabilis</taxonomicName>
|
|
sp. nov. is only known from Huaping Nature Reserve, northeastern Guangxi. The individuals were found on shrubbery branches near trail paths between elevations of 1300-1330 m a.s.l. from June to September. Males were not calling when found, but the collected female specimens bear mature yellowish oocytes. Tadpoles have not been found and ecological information remains unknown.
|
|
</paragraph>
|
|
<caption ID-Table-UUID="E485D427D430B52C70EC0A29AEAB4143" httpUri="http://table.plazi.org/id/E485D427D430B52C70EC0A29AEAB4143" pageId="0" pageNumber="105" start="Table 5" startId="T5">
|
|
<paragraph pageId="0" pageNumber="105">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">Table 5.</emphasis>
|
|
Measurements (in mm) of the type series of
|
|
<taxonomicName authorityName="Lyu, Wang & Zhao" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophrys (Panophrys) mirabilis" order="Anura" pageId="0" pageNumber="105" phylum="Chordata" rank="species" species="mirabilis" subGenus="Panophrys">Megophrys (Panophrys) mirabilis</taxonomicName>
|
|
sp. nov., * for the holotype.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="105">
|
|
<table pageId="0" pageNumber="105">
|
|
<tr pageId="0" pageNumber="105">
|
|
<th colspan="1" isEmpty="true" pageId="0" pageNumber="105" rowspan="1">-</th>
|
|
<th colspan="1" pageId="0" pageNumber="105" rowspan="1">SYS a002917 *</th>
|
|
<th colspan="1" pageId="0" pageNumber="105" rowspan="1">SYS a002192</th>
|
|
<th colspan="1" pageId="0" pageNumber="105" rowspan="1">SYS a002193</th>
|
|
<th colspan="1" pageId="0" pageNumber="105" rowspan="1">SYS a002289</th>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">Sex</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">Male</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">Male</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">Female</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">Female</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">SVL</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">61.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">55.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">74.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">68.5</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">HDL</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">21.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">18.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">23.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">22.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">HDW</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">21.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">18.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">23.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">22.4</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">SNT</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">7.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">7.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">9.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">8.8</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">IND</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">6.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">5.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">7.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">6.8</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">IOD</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">7.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">6.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">8.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">7.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">ED</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">6.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">5.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">8.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">6.8</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">TD</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">3.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">3.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">4.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">4.3</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">TED</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">3.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">3.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">4.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">3.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">HND</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">17.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">15.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">20.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">19.5</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">RAD</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">15.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">13.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">18.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">17.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">FTL</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">43.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">37.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">48.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">43.2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="105">
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="105">TIB</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">28.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">26.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">33.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="105" rowspan="1">30.5</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |