213 lines
25 KiB
XML
213 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.395.7110" ID-GBIF-Dataset="d9627d89-aad8-4b48-b024-c7a97d580231" ID-PMC="PMC3978236" ID-Pensoft-Pub="1313-2970-395-33" ID-PubMed="24715788" ID-ZBK="B4D4D101C9AE47F09B7B320735B69D1D" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-395-33" ModsDocOrigin="ZooKeys 395" ModsDocTitle="Six new species of Philiris Röber, 1891 (Lepidoptera, Lycaenidae) from Papua New Guinea" checkinTime="1451246046685" checkinUser="pensoft" docAuthor="Mueller, Chris J." docDate="2014" docId="8E3BD29FECCC130221416C6B1CDD977A" docLanguage="en" docName="ZooKeys 395: 33-55" docOrigin="ZooKeys 395" docSource="http://dx.doi.org/10.3897/zookeys.395.7110" docTitle="Philiris radicala Mueller, sp. n." docType="treatment" docUuid="EEF3D73A-0478-45A0-83B8-997BBECA6B9B" docUuidSource="ZooBank" docVersion="4" lastPageNumber="43" masterDocId="FFEBBE73FFD8D16FFFC64B5A751FFFB3" masterDocTitle="Six new species of Philiris Roeber, 1891 (Lepidoptera, Lycaenidae) from Papua New Guinea" masterLastPageNumber="55" masterPageNumber="33" pageNumber="42" updateTime="1668158131885" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Six new species of Philiris Roeber, 1891 (Lepidoptera, Lycaenidae) from Papua New Guinea</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Mueller, Chris J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>395</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>55</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.395.7110</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.395.7110</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-395-33</mods:identifier>
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<mods:identifier type="ZBK">B4D4D101C9AE47F09B7B320735B69D1D</mods:identifier>
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<mods:identifier type="ZooBank">B4D4D101C9AE47F09B7B320735B69D1D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152052285" LSID="urn:lsid:zoobank.org:act:EEF3D73A-0478-45A0-83B8-997BBECA6B9B" httpUri="http://treatment.plazi.org/id/8E3BD29FECCC130221416C6B1CDD977A" lastPageId="11" lastPageNumber="43" pageId="9" pageNumber="42">
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<subSubSection pageId="9" pageNumber="42" type="nomenclature">
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<paragraph pageId="9" pageNumber="42">
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<taxonomicName LSID="http://zoobank.org/EEF3D73A-0478-45A0-83B8-997BBECA6B9B" authority="Mueller" class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="radicala">
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Philiris radicala
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<normalizedToken originalValue="Müller">Mueller</normalizedToken>
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</taxonomicName>
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<taxonomicNameLabel pageId="9" pageNumber="42">sp. n.</taxonomicNameLabel>
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Figs 38-43, 67, 68
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="42" type="type material">
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<paragraph pageId="9" pageNumber="42">Type material.</paragraph>
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<paragraph pageId="9" pageNumber="42">
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Holotype ♂ (Figs 38-40): "Papua New Guinea, Hindenburg Range, Western Province, 1000 m,
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-5.2166667">5°13'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="141.23334">141°14'E</geoCoordinate>
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, 13-17 Feb, 2013, Chris J.
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<normalizedToken originalValue="Müller">Mueller</normalizedToken>
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, genitalia dissected and held in vial pinned to specimen, (ANIC), Registration: ANIC Database No. 31-023125. Paratypes (7 ♂♂, 2 ♀♀): labelled the same as the holotype (1 ♂ AM, 1 ♂ BMNH, 1 ♂ NARI, 4 ♂♂ CJMC, 1 ♀ ANIC, 1 ♀ AM).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="43" pageId="9" pageNumber="42" type="diagnosis">
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<paragraph pageId="9" pageNumber="42">Diagnosis.</paragraph>
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<paragraph pageId="9" pageNumber="42">
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="radicala">Philiris radicala</taxonomicName>
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is unique within
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="genus">Philiris</taxonomicName>
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, with its very unusual colouration in the male upperside and matt white underside ground colour and dark border to the underside termen in both sexes. The taxon shows some relationship to
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris intensa" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="intensa">Philiris intensa</taxonomicName>
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(Butler, 1876) (Holotype; Figs 44, 45 and female; Fig. 46), with which it flies in the Hindenburg Range, although the latter species is more commonly encountered below 500 m, while
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="radicala">Philiris radicala</taxonomicName>
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flies between 700-1100 m. The male upperside of
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris intensa" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="intensa">Philiris intensa</taxonomicName>
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is a deep, bright, shining sky blue, whereas that in
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="radicala">Philiris radicala</taxonomicName>
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is an iridescent turquoise (green-blue). The dark border to both wings on the upperside of the male
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="radicala">Philiris radicala</taxonomicName>
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is much broader than in
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris intensa" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="intensa">Philiris intensa</taxonomicName>
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and the border of the forewing is straight, forming a line between the postmedian section of the costa and the tornus, while this border is curved in
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris intensa" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="intensa">Philiris intensa</taxonomicName>
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. The female upperside of
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="radicala">Philiris radicala</taxonomicName>
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bears very large white median patches on both wings, unlike
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris intensa" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="intensa">Philiris intensa</taxonomicName>
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and the discocellulars of
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="radicala">Philiris radicala</taxonomicName>
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are distinctly dark brown. On the underside of both sexes the ground colour is an unusual powdery matt white, the termen of both wings is heavily blackened, particularly at the vein ends, and dark scaling occurs in the subterminal area. These features are not present in
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris intensa" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="intensa">Philiris intensa</taxonomicName>
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.
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</paragraph>
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<caption pageId="9" pageNumber="42">
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<paragraph pageId="9" pageNumber="42">
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Figures 46-60.
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="genus">Philiris</taxonomicName>
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adults (left side upperside and right side underside, where halved) and label data. 46
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris intensa" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="intensa">Philiris intensa</taxonomicName>
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♀ (halved) (Hindenburg Range, Western Province), 47
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris hindenburgensis" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="hindenburgensis">Philiris hindenburgensis</taxonomicName>
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holotype ♂ upperside 48
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris hindenburgensis" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="hindenburgensis">Philiris hindenburgensis</taxonomicName>
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holotype ♂ underside 49
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris hindenburgensis" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="hindenburgensis">Philiris hindenburgensis</taxonomicName>
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holotype ♂ label data 50
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris satis" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="satis">Philiris satis</taxonomicName>
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holotype ♂ (halved) 51
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris satis" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="satis">Philiris satis</taxonomicName>
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holotype ♂ label data 52
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris oreas" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="oreas">Philiris oreas</taxonomicName>
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holotype ♂ (halved) 53
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris oreas" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="oreas">Philiris oreas</taxonomicName>
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holotype ♂ label data 54
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris albihumerata" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="albihumerata">Philiris albihumerata</taxonomicName>
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holotype ♂ (halved) 55
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris albihumerata" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="albihumerata">Philiris albihumerata</taxonomicName>
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holotype ♂ label data 56
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris parsonsi" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="parsonsi">Philiris parsonsi</taxonomicName>
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holotype ♂ upperside 57
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris parsonsi" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="parsonsi">Philiris parsonsi</taxonomicName>
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holotype ♂ underside 58
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris parsonsi" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="parsonsi">Philiris parsonsi</taxonomicName>
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holotype ♂ label data 59
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris angabunga" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="angabunga">Philiris angabunga</taxonomicName>
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holotype ♂ (halved) 60
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris angabunga" order="Lepidoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="angabunga">Philiris angabunga</taxonomicName>
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holotype ♂ label data. Scale bar = 10 mm.
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</paragraph>
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</caption>
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<paragraph lastPageId="10" lastPageNumber="43" pageId="9" pageNumber="42">
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The male genitalia of
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="radicala">
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<pageBreakToken pageId="10" pageNumber="43" start="start">Philiris</pageBreakToken>
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radicala
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</taxonomicName>
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corroborates a relationship with
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris intensa" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="intensa">Philiris intensa</taxonomicName>
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, with both possessing laterally pointed socii (more so in the latter species than in
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="radicala">Philiris radicala</taxonomicName>
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, see Figs 69, 70) and similar aedeagi. Both taxa also bear triangular-shaped valvae, which are approximately equilateral in
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="radicala">Philiris radicala</taxonomicName>
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but much longer and apically tapered in
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris intensa" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="intensa">Philiris intensa</taxonomicName>
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. The valvae are slightly asymmetrical in
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="radicala">Philiris radicala</taxonomicName>
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but symmetrical in the latter species.
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</paragraph>
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<caption pageId="10" pageNumber="43">
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<paragraph pageId="10" pageNumber="43">
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Figures 61-70.
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="genus">Philiris</taxonomicName>
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male genitalia (a genitalia in ventral view with aedeagus removed, b genitalia in lateral view, c aedeagus in lateral view). 61
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris petriei" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="petriei">Philiris petriei</taxonomicName>
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(Whiteman Range, West New Britain Province) 62
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris harterti" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="harterti">Philiris harterti</taxonomicName>
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(Nr. Wabo, Gulf Province) 63
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris bubalisatina" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="bubalisatina">Philiris bubalisatina</taxonomicName>
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holotype ♂ 64
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris baiteta" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="baiteta">Philiris baiteta</taxonomicName>
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(Hindenburg Range, Western Province) 65
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris hemileuca" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="hemileuca">Philiris hemileuca</taxonomicName>
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(Telefomin, West Sepik Province) 66
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris hypoxantha" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="hypoxantha">Philiris hypoxantha</taxonomicName>
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(Nr. Oetakwa River, Papua) 67
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="radicala">Philiris radicala</taxonomicName>
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holotype ♂ 68
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="radicala">Philiris radicala</taxonomicName>
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(Hindenburg Range, Western Province) 69
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris intensa" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="intensa">Philiris intensa</taxonomicName>
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(Bulolo, Morobe Province), 70
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<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris intensa" order="Lepidoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="intensa">Philiris intensa</taxonomicName>
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(Nr. Wabo, Gulf Province). Scale bar = approx. 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="43" type="description">
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<paragraph pageId="10" pageNumber="43">Description.</paragraph>
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<paragraph pageId="10" pageNumber="43">♂ (Figs 38-40): Forewing length 17 mm, antenna 9 mm (holotype). Head, palpus and thorax dark grey dorsally, white ventrally, abdomen dark grey dorsally, white ventrally, frons dark grey with white eye ring; legs white with black areas on tibiae; antenna shaft black, ringed conspicuously with white between segments, apex of club tipped with brown.</paragraph>
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<paragraph pageId="10" pageNumber="43">Fore wing termen slightly convex, inner margin very slightly bowed near base, apex slightly rounded; upperside bright shining turquoise, costa and termen broadly black, 2 mm wide at tornus but up to 5 mm wide at apex, the boundary between the dark border and shining turquoise area is straight between veins 2 and 8, cilia white but broadly black at vein ends, giving chequered appearance; underside matt white with apical area suffused with black scales, termen and vein ends distinctly black, cilia as in upperside.</paragraph>
|
||
<paragraph pageId="10" pageNumber="43">Hind wing rounded; upperside bright shining turquoise, termen broadly black (2 mm wide), between inner margin and vein 1a dark brown-black, grading to light grey-brown basally, apex broadly black, costa broadly light grey, basally grey-brown, cilia white but broadly black at vein ends, giving chequered appearance; underside matt white, subterminal area narrowly suffused with black scales, termen and vein ends distinctly black, cilia as in upperside.</paragraph>
|
||
<paragraph pageId="10" pageNumber="43">
|
||
Male genitalia (Figs 67, 68): Vinculum and tegumen ring oval, enlarged towards sociuncus, sociuncus broad, socii with lateral margin pointed apically, dorsally sub-rounded, socii separated by slightly protruding sinus, saccus tapered posteriorly, brachium tapered dorsally and slightly hooked at apex; valva slightly asymmetrical, with left valva longer than left valva, valva equilateral triangle-shaped; phallus slender, with zone of intricate cornuti in post-zonal section, vesica with central
|
||
<normalizedToken originalValue="‘tongue’">'tongue'</normalizedToken>
|
||
apically.
|
||
</paragraph>
|
||
<paragraph pageId="10" pageNumber="43">♀ (Figs 41, 42): Forewing length (17 mm), antenna length (9 mm), antenna, head, palpus, thorax, legs and abdomen similar to male.</paragraph>
|
||
<paragraph pageId="10" pageNumber="43">Fore wing termen slightly convex, inner margin very slightly bowed near base, apex slightly rounded; upperside dark brown-black, large area of white extending from base along inner margin to postmedian area and occupying about two-thirds of cell, margins of this pale patch suffused with powder blue, costal half of discocellulars heavily brown-black, intruding into pale area, cilia as in male; underside as in male.</paragraph>
|
||
<paragraph pageId="10" pageNumber="43">Hind wing rounded; upperside dark brown-black, apical area broadly grey-white, cell and basal area along vein 1b grey-white with powder blue suffusion, inner margin and basal part of costa light grey-brown, discocellulars heavily brown-black, intruding into pale area, cilia as in male; underside as in male.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="43" type="etymology">
|
||
<paragraph pageId="10" pageNumber="43">Etymology.</paragraph>
|
||
<paragraph pageId="10" pageNumber="43">
|
||
The name
|
||
<normalizedToken originalValue="‘radicala’">'radicala'</normalizedToken>
|
||
reflects the extraordinary colouration of the male upperside and overall divergent morphology of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="43" type="distribution">
|
||
<paragraph pageId="10" pageNumber="43">Distribution.</paragraph>
|
||
<paragraph pageId="10" pageNumber="43">Western Province, Papua New Guinea.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="11" lastPageNumber="44" pageId="10" pageNumber="43" type="ecology">
|
||
<paragraph pageId="10" pageNumber="43">Ecology.</paragraph>
|
||
<paragraph lastPageId="11" lastPageNumber="44" pageId="10" pageNumber="43">
|
||
Males of
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris radicala" order="Lepidoptera" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="radicala">
|
||
<pageBreakToken pageId="11" pageNumber="44" start="start">Philiris</pageBreakToken>
|
||
radicala
|
||
</taxonomicName>
|
||
were taken around midday and early afternoon as they settled at the tops of tall saplings, between 8-10 metres above the ground, in small clearings created by tree falls in very steep terrain. Females were recorded in similar areas, where they fed at the small white flowers of an unidentified tree. Conversely, males of
|
||
<taxonomicName class="Insecta" family="Lycaenidae" genus="Philiris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philiris intensa" order="Lepidoptera" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="intensa">Philiris intensa</taxonomicName>
|
||
, flying at the same localities but generally at a lower altitude, were always seen to fly within 2-3 metres above the ground in areas of regrowth proximal to streams.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |