289 lines
21 KiB
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289 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1145.96724" ID-Pensoft-Pub="1313-2970-1145-1" ID-Pensoft-UUID="ACBBD138B7375B0D9F63CE792A82F653" ID-ZooBank="830628C276CD4641BFC6144CD775ED6B" ModsDocID="1313-2970-1145-1" checkinTime="1675506069289" checkinUser="pensoft" docAuthor="Hedin, Marshal & Milne, Marc A." docDate="2023" docId="8D969825BFA85396B06E5A3E57D02852" docLanguage="en" docName="ZooKeys 1145: 1-130" docOrigin="ZooKeys 1145" docPubDate="2023-02-03" docSource="http://dx.doi.org/10.3897/zookeys.1145.96724" docTitle="Nesticus barrowsi Gertsch 1984" docType="treatment" docVersion="1" id="ACBBD138B7375B0D9F63CE792A82F653" lastPageNumber="1" masterDocId="ACBBD138B7375B0D9F63CE792A82F653" masterDocTitle="New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains" masterLastPageNumber="130" masterPageNumber="1" pageNumber="1" updateTime="1675506069289" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hedin, Marshal</mods:namePart>
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<mods:affiliation>Department of Biology, San Diego State University, San Diego, California 92182 - 4614, USA</mods:affiliation>
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<mods:nameIdentifier type="email">mhedin@sdsu.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Milne, Marc A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1943-0161</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, University of Indianapolis, Indianapolis, Indiana 46227, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-03</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1145</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>130</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1145.96724</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1145.96724</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1145-1</mods:identifier>
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<mods:identifier type="ZooBank">830628C276CD4641BFC6144CD775ED6B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">ACBBD138B7375B0D9F63CE792A82F653</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:8D969825BFA85396B06E5A3E57D02852" httpUri="http://treatment.plazi.org/id/8D969825BFA85396B06E5A3E57D02852" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName ID-CoL="478WM" ID-ENA="60722" LSID="8D969825-BFA8-5396-B06E-5A3E57D02852" authority="Gertsch, 1984" authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus barrowsi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="barrowsi">Nesticus barrowsi Gertsch, 1984</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Nesticus barrowsi ♂ palps. Tennessee, Blount Co., Great Smoky Mountains NP, Gregory Cave, MCH specimen # 1295, ventral (A), dorsal (B). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure44" httpUri="https://binary.pensoft.net/fig/804495" pageId="0" pageNumber="1">Figs 44A, B</figureCitation>
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<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Nesticus barrowsi epigynal variation. Tennessee, Blount Co., Great Smoky Mountains NP, Calf Cave, MCH 00 _ 147, ventral (A), dorsal (B). Tennessee, Blount Co., Great Smoky Mountains NP, White Oak Sinks, Rainbow Cave, MCH specimen # 1303, ventral (C), dorsal (D). Tennessee, Blount Co., Tuckaleechee Caverns, MCH specimen # 1568, ventral (E), dorsal (F). Tennessee, Blount Co., Great Smoky Mountains NP, Gregory Cave, MCH specimen # 1297, ventral (G), dorsal (H). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure45" httpUri="https://binary.pensoft.net/fig/804496" pageId="0" pageNumber="1">, 45A-H</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus barrowsi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="barrowsi">Nesticus barrowsi</taxonomicName>
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Gertsch, 1984: 35, figs 103-105, 118-120;
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<bibRefCitation author="Reeves, WK" journalOrPublisher="Journal of the Elisha Mitchell Scientific Society" pageId="0" pageNumber="1" pagination="334 - 343" publicationUrl="https://www.jstor.org/stable/24335885" refId="B50" refString="Reeves, WK, 2000. Invertebrate cavernicoles of the Great Smoky Mountains National Park, USA. Journal of the Elisha Mitchell Scientific Society 116 (4): 334 - 343, https://www.jstor.org/stable/24335885" title="Invertebrate cavernicoles of the Great Smoky Mountains National Park, USA." url="https://www.jstor.org/stable/24335885" volume="116" year="2000">Reeves 2000</bibRefCitation>
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: 338.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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Type material:
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<materialsCitation collectingDate="1938-11-01" collectingDateMax="2000-08-02" collectingDateMin="1938-11-01" collectorName="Tuckaleechee Caverns, Tuckaleechee Cove, W. B. Jones, New, M. Hedin, S. O'Kane, Non, Great Smoky Mountains NP, Cades Cove, Rich Mountain Blowhole, J. Cokendolpher, W. Reeves, White Oak Sinks" country="USA" county="Blount Co." location="Blount Co." municipality="Blount Co." specimenCount="9" specimenCount-female="9" stateProvince="Tennessee" typeStatus="Holotype">
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<emphasis italics="true" pageId="0" pageNumber="1">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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<collectingCountry name="United States of America">USA</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="Tennessee">Tennessee</collectingRegion>
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,
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<collectingCounty>Blount Co.</collectingCounty>
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</emphasis>
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•
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<specimenCount>♂</specimenCount>
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<typeStatus>holotype</typeStatus>
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;
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<collectorName>Tuckaleechee Caverns</collectorName>
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,
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<collectorName>Tuckaleechee Cove</collectorName>
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;
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<collectingDate value="1938-11-01">1 Nov. 1938</collectingDate>
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;
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<collectorName>W.B. Jones</collectorName>
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leg.; AMNH.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectorName>New</collectorName>
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collections from type locality
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</emphasis>
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: -
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingMunicipality>Blount Co.</collectingMunicipality>
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</emphasis>
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•
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<specimenCount>♀</specimenCount>
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;
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<collectorName>Tuckaleechee Caverns</collectorName>
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,
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<collectorName>Tuckaleechee Cove</collectorName>
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;
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<collectingDate value="1992-09-22">22 Sep. 1992</collectingDate>
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;
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<collectorName>M. Hedin</collectorName>
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,
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<collectorName>
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S.
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<normalizedToken originalValue="O’Kane">O'Kane</normalizedToken>
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</collectorName>
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leg.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectorName>Non</collectorName>
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type material
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</emphasis>
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: -
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<emphasis bold="true" pageId="0" pageNumber="1">
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<location LSID="urn:lsid:plazi:treatment:8D969825BFA85396B06E5A3E57D02852:0F5DF9D724B1AF29C89CF18FD19CC654" country="USA" county="Blount Co." municipality="Blount Co." name="Blount Co." stateProvince="Tennessee">Blount Co.</location>
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</emphasis>
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•
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<specimenCount>♂</specimenCount>
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,
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<specimenCount type="female">7♀</specimenCount>
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;
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<collectorName>
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<location LSID="urn:lsid:plazi:treatment:8D969825BFA85396B06E5A3E57D02852:94CDE405B1A84F1A88510A611CE9396F" country="USA" county="Blount Co." municipality="Blount Co." name="Great Smoky Mountains" stateProvince="Tennessee">Great Smoky Mountains</location>
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NP
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</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:8D969825BFA85396B06E5A3E57D02852:E12FD0B3B5644FE470588317C5F2E4B2" country="USA" county="Blount Co." municipality="Blount Co." name="Gregory Cave" stateProvince="Tennessee">Gregory Cave</location>
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,
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<collectorName>Cades Cove</collectorName>
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;
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<collectingDate value="1992-08-21">21 Aug. 1992</collectingDate>
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;
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<collectorName>M. Hedin</collectorName>
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leg.; •
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<specimenCount type="female">2♀</specimenCount>
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;
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<collectorName>
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<location LSID="urn:lsid:plazi:treatment:8D969825BFA85396B06E5A3E57D02852:63EB8B689A6AFA7AC46902E279E1774D" country="USA" county="Blount Co." municipality="Blount Co." name="Great Smoky Mountains" stateProvince="Tennessee">Great Smoky Mountains</location>
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NP
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</collectorName>
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,
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<collectorName>
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<location LSID="urn:lsid:plazi:treatment:8D969825BFA85396B06E5A3E57D02852:A955A2097F0B452FCF836CB82EB02B74" country="USA" county="Blount Co." municipality="Blount Co." name="Rich Mountain" stateProvince="Tennessee">
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Rich
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<location LSID="urn:lsid:plazi:treatment:8D969825BFA85396B06E5A3E57D02852:D0D457EC31CFD7D439F9B0DFD1648BD2" country="USA" county="Blount Co." municipality="Blount Co." name="Mountain Blowhole" stateProvince="Tennessee">Mountain Blowhole</location>
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</location>
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</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:8D969825BFA85396B06E5A3E57D02852:F191670A064BBEF6AB7BEF12DD44A2B0" country="USA" county="Blount Co." municipality="Blount Co." name="Calf Cave" stateProvince="Tennessee">Calf Cave</location>
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;
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<collectingDate value="2000-08-02">2 Aug. 2000</collectingDate>
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;
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<collectorName>M. Hedin</collectorName>
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,
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<collectorName>J. Cokendolpher</collectorName>
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,
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<collectorName>W. Reeves</collectorName>
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leg.; MCH 00_147; •
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<specimenCount>♀</specimenCount>
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;
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<collectorName>
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<location LSID="urn:lsid:plazi:treatment:8D969825BFA85396B06E5A3E57D02852:5A6410320832A89FF3B8D29912D9FC85" country="USA" county="Blount Co." municipality="Blount Co." name="Great Smoky Mountains" stateProvince="Tennessee">Great Smoky Mountains</location>
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NP
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</collectorName>
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,
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<collectorName>White Oak Sinks</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:8D969825BFA85396B06E5A3E57D02852:77F71C64BAE7A821767006BF71C8FEE9" country="USA" county="Blount Co." municipality="Blount Co." name="Rainbow Cave" stateProvince="Tennessee">Rainbow Cave</location>
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;
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<collectingDate value="1992-08-21">21 Aug. 1992</collectingDate>
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;
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<collectorName>M. Hedin</collectorName>
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leg.
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</materialsCitation>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The diagnosis of
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<bibRefCitation author="Gertsch, WJ" journalOrPublisher="Texas Memorial Museum Bulletin" pageId="0" pageNumber="1" pagination="1 - 91" publicationUrl="http://hdl.handle.net/2152/29952" refId="B12" refString="Gertsch, WJ, 1984. The spider family Nesticidae (Araneae) in North America, Central America, and the West Indies. Texas Memorial Museum Bulletin 31: 1 - 91, http://hdl.handle.net/2152/29952" title="The spider family Nesticidae (Araneae) in North America, Central America, and the West Indies." url="http://hdl.handle.net/2152/29952" volume="31" year="1984">Gertsch (1984)</bibRefCitation>
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is revised here to recognize the phylogenetic affinities within the
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<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus barrowsi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="barrowsi">
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<emphasis italics="true" pageId="0" pageNumber="1">Nesticus barrowsi</emphasis>
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</taxonomicName>
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group.
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<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus barrowsi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="barrowsi">
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<emphasis italics="true" pageId="0" pageNumber="1">Nesticus barrowsi</emphasis>
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</taxonomicName>
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is troglomorphic (long-legged, pale, approximately eyeless, relatively large-bodied), unlike other species in the species group. The male tegular apophysis curves to lie behind a quadrate median apophysis, is sharply tipped, without a basal keel. The paracymbial dorsal process is translucent, skinny and finger-like, while the distal process includes a well-sclerotized pointed tip and a small ventral keel (Fig.
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<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Nesticus barrowsi ♂ palps. Tennessee, Blount Co., Great Smoky Mountains NP, Gregory Cave, MCH specimen # 1295, ventral (A), dorsal (B). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure44" httpUri="https://binary.pensoft.net/fig/804495" pageId="0" pageNumber="1">44A, B</figureCitation>
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). Female
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<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. barrowsi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="barrowsi">
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<emphasis italics="true" pageId="0" pageNumber="1">N. barrowsi</emphasis>
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</taxonomicName>
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differ from other members of the species group in overall morphology of the epigynum, including the pear-shaped spermathecae (Fig.
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<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Nesticus barrowsi epigynal variation. Tennessee, Blount Co., Great Smoky Mountains NP, Calf Cave, MCH 00 _ 147, ventral (A), dorsal (B). Tennessee, Blount Co., Great Smoky Mountains NP, White Oak Sinks, Rainbow Cave, MCH specimen # 1303, ventral (C), dorsal (D). Tennessee, Blount Co., Tuckaleechee Caverns, MCH specimen # 1568, ventral (E), dorsal (F). Tennessee, Blount Co., Great Smoky Mountains NP, Gregory Cave, MCH specimen # 1297, ventral (G), dorsal (H). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure45" httpUri="https://binary.pensoft.net/fig/804496" pageId="0" pageNumber="1">45A-H</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1145.96724.figure44" httpUri="https://binary.pensoft.net/fig/804495" pageId="0" pageNumber="1" start="Figure 44" startId="F44">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 44.</emphasis>
|
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<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus barrowsi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="barrowsi">
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<emphasis italics="true" pageId="0" pageNumber="1">Nesticus barrowsi</emphasis>
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</taxonomicName>
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♂ palps. Tennessee, Blount Co., Great Smoky Mountains NP, Gregory Cave, MCH specimen #1295, ventral (
|
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), dorsal (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
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). Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.1145.96724.figure45" httpUri="https://binary.pensoft.net/fig/804496" pageId="0" pageNumber="1" start="Figure 45" startId="F45">
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<paragraph pageId="0" pageNumber="1">
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||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 45.</emphasis>
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<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus barrowsi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="barrowsi">
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<emphasis italics="true" pageId="0" pageNumber="1">Nesticus barrowsi</emphasis>
|
||
</taxonomicName>
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epigynal variation. Tennessee, Blount Co., Great Smoky Mountains NP, Calf Cave, MCH 00_147, ventral (
|
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), dorsal (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
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). Tennessee, Blount Co., Great Smoky Mountains NP, White Oak Sinks, Rainbow Cave, MCH specimen #1303, ventral (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), dorsal (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
). Tennessee, Blount Co., Tuckaleechee Caverns, MCH specimen #1568, ventral (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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), dorsal (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
). Tennessee, Blount Co., Great Smoky Mountains NP, Gregory Cave, MCH specimen #1297, ventral (
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<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
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), dorsal (
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<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
|
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). Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="variation">
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<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Minor variation is observed in the height and width of the paired epigynal plates across geographic locations (Fig.
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<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Nesticus barrowsi epigynal variation. Tennessee, Blount Co., Great Smoky Mountains NP, Calf Cave, MCH 00 _ 147, ventral (A), dorsal (B). Tennessee, Blount Co., Great Smoky Mountains NP, White Oak Sinks, Rainbow Cave, MCH specimen # 1303, ventral (C), dorsal (D). Tennessee, Blount Co., Tuckaleechee Caverns, MCH specimen # 1568, ventral (E), dorsal (F). Tennessee, Blount Co., Great Smoky Mountains NP, Gregory Cave, MCH specimen # 1297, ventral (G), dorsal (H). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1145.96724.figure45" httpUri="https://binary.pensoft.net/fig/804496" pageId="0" pageNumber="1">45A-H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution and natural history.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
This troglomorphic species is only known from caves in karst windows along the northwestern edge of Great Smoky Mountains National Park (Cades Cove, Tuckaleechee Cove; Fig.
|
||
<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Distribution of the Nesticus barrowsi group. Type localities designated with yellow circles. State boundaries and major cities shown for geographic context. Dashed lines circumscribe known species distributions." figureDoi="10.3897/zookeys.1145.96724.figure42" httpUri="https://binary.pensoft.net/fig/804493" pageId="0" pageNumber="1">42</figureCitation>
|
||
).
|
||
<bibRefCitation author="Reeves, WK" journalOrPublisher="Journal of the Elisha Mitchell Scientific Society" pageId="0" pageNumber="1" pagination="334 - 343" publicationUrl="https://www.jstor.org/stable/24335885" refId="B50" refString="Reeves, WK, 2000. Invertebrate cavernicoles of the Great Smoky Mountains National Park, USA. Journal of the Elisha Mitchell Scientific Society 116 (4): 334 - 343, https://www.jstor.org/stable/24335885" title="Invertebrate cavernicoles of the Great Smoky Mountains National Park, USA." url="https://www.jstor.org/stable/24335885" volume="116" year="2000">Reeves (2000)</bibRefCitation>
|
||
reported
|
||
<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus barrowsi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="barrowsi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nesticus barrowsi</emphasis>
|
||
</taxonomicName>
|
||
in sympatry with
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. stupkai" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stupkai">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. stupkai</emphasis>
|
||
</taxonomicName>
|
||
at two cave locations in Great Smoky Mountains National Park. We also collected these species in near syntopy at White Oak Sinks, with
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. barrowsi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="barrowsi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. barrowsi</emphasis>
|
||
</taxonomicName>
|
||
found in the dark zone of caves and
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="N. stupkai" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stupkai">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. stupkai</emphasis>
|
||
</taxonomicName>
|
||
found closer to cave entrances (twilight zone).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |