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<document id="F8A55E2D28EB1DA5DE4F6C192ED2F511" ID-CLB-Dataset="50294" ID-DOI="10.5281/zenodo.174866" ID-GBIF-Dataset="15363d41-dba5-4484-addb-9734f619bcf1" ID-ISSN="1175-5326" ID-Zenodo-Dep="174866" checkinTime="1459790442371" checkinUser="plazi" docAuthor="Short, Andrew Edward Z. &amp; Torres, Phillip J." docDate="2006" docId="8D7A87A6FFE5FFAB1F50B91EA9248770" docLanguage="en" docName="zt01369p017.pdf" docOrigin="Zootaxa 1369" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@632cf7d3" docStyleName="zootaxa.0000.journal_article" docTitle="Hemiosus selva Short, sp. n." docType="treatment" docVersion="7" lastPageNumber="7" masterDocId="7143FFDEFFE0FFAD1E58BC33AB088002" masterDocTitle="Review of the Hemiosus Sharp and Derallus Sharp of North America, Central America, and the Caribbean (Coleoptera: Hydrophilidae: Berosini)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="6" updateTime="1732741852905" updateUser="ExternalLinkService">
<mods:mods id="4266396A0A82604095CB663EF06F9727" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4C4BA892D39508060E50A673764E8463">
<mods:title id="A9D25DA9A9F695C7CD800BF4E97D062C">Review of the Hemiosus Sharp and Derallus Sharp of North America, Central America, and the Caribbean (Coleoptera: Hydrophilidae: Berosini)</mods:title>
</mods:titleInfo>
<mods:name id="5D8D4CB191FA06D28D267BB07343ABC9" type="personal">
<mods:role id="AB7BE69EB5E433E249497400C24B82F5">
<mods:roleTerm id="9FDA2EC085F5C8B1153ED433DFE555C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3EE3C63745187D4BC435FFDE915E46CB">Short, Andrew Edward Z.</mods:namePart>
</mods:name>
<mods:name id="2A9F37C01FB2ED779396C2DF791B53A3" type="personal">
<mods:role id="404C0D585854DBCFBDBB4E363D478AAC">
<mods:roleTerm id="AB0320850C5B076B5BDCC12714496EEF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2D8C255690AFFC25463CF54339581997">Torres, Phillip J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="F19CA75A6D346EE8B1EEAF52F5D3260C">text</mods:typeOfResource>
<mods:relatedItem id="FEE48F1A8BA4C89223BFEA341E3F5AF6" type="host">
<mods:titleInfo id="98A9CDE744D77B4BF019AC361C0D4F91">
<mods:title id="24DAFCFE95BF9F69E7F6BA79DF870217">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EBCD944BEB921E3C0324895D5F8C7132">
<mods:date id="F41DDC60A4ED625CD089FC0C9EF8CFBB">2006</mods:date>
<mods:detail id="7A56E6609BEB1E4D3518E0A989DBD731" type="volume">
<mods:number id="0864A3E5CB753BA08AA299A95EE312B3">1369</mods:number>
</mods:detail>
<mods:extent id="FCFAB03D684C6DE07FCD1066C395B96A" unit="page">
<mods:start id="67587B6F949135E632BBDFF91446D49D">1</mods:start>
<mods:end id="782B4EE82ACB5B73E042A2D2FADEFAF8">17</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6C7D12CCFBAC042729B770B8D9E1DFA8">journal article</mods:classification>
<mods:identifier id="C3DAD23C401B982F63AEE616234A5162" type="CLB-Dataset">50294</mods:identifier>
<mods:identifier id="C726BC17EF429646983480C645280A74" type="DOI">10.5281/zenodo.174866</mods:identifier>
<mods:identifier id="E1D2C5041EA151DAE75CB9BA701DE06A" type="GBIF-Dataset">15363d41-dba5-4484-addb-9734f619bcf1</mods:identifier>
<mods:identifier id="13641ECA906A5784B92F06351E187703" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CB8670461AABDD10C44DD681E9F89762" type="Zenodo-Dep">174866</mods:identifier>
</mods:mods>
<treatment id="8D7A87A6FFE5FFAB1F50B91EA9248770" ID-GBIF-Taxon="119350291" LSID="urn:lsid:plazi:treatment:8D7A87A6FFE5FFAB1F50B91EA9248770" httpUri="http://treatment.plazi.org/id/8D7A87A6FFE5FFAB1F50B91EA9248770" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" scope_family="Hydrophilidae" scope_order="Coleoptera" scope_tribe="Berosini">
<subSubSection id="4DC9653BFFE5FFA81F50B91EAA80856B" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="056C36B0FFE5FFA81F50B91EA96A8544" blockId="5.[264,610,1324,1385]" box="[264,610,1324,1351]" pageId="5" pageNumber="6">
<heading id="5E2481DCFFE5FFA81F50B91EA96A8544" bold="true" box="[264,610,1324,1351]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="37A7EAA2FFE5FFA81F50B91EA96A8544" bold="true" box="[264,610,1324,1351]" pageId="5" pageNumber="6">
<taxonomicName id="C2D34D33FFE5FFA81F50B91EA91C8544" ID-CoL="3KR7R" authority="Short" authorityName="Short" box="[264,532,1324,1351]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="selva" status="sp. n.">
<emphasis id="37A7EAA2FFE5FFA81F50B91EAACC8545" bold="true" box="[264,452,1325,1351]" italics="true" pageId="5" pageNumber="6">Hemiosus selva</emphasis>
Short
</taxonomicName>
,
<taxonomicNameLabel id="2C9457D9FFE5FFA81C78B91FA96A8544" box="[544,610,1324,1350]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="056C36B0FFE5FFA81F50B97CAA80856B" blockId="5.[264,610,1324,1385]" box="[264,392,1359,1385]" pageId="5" pageNumber="6">
(
<figureCitation id="9DE82A35FFE5FFA81F49B97CAA77856B" box="[273,383,1359,1385]" captionStart="FIGURES 3 – 5" captionStartId="7.[264,382,1004,1028]" captionTargetBox="[418,1160,325,975]" captionTargetId="figure@7.[409,1177,304,994]" captionTargetPageId="7" captionText="FIGURES 3 – 5. Details of Hemiosus selva Short, sp. n. (3, 5) and H. exilis (LeConte) (4). 3 – 4: pronotal coloration, scale bar 0.18 mm; 5: aedeagus, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.174868" httpUri="https://zenodo.org/record/174868/files/figure.png" pageId="5" pageNumber="6">Figs. 3, 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4DC9653BFFE5FFAB1F50B9ACAFE982C7" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="056C36B0FFE5FFA81F50B9ACAA4585E3" blockId="5.[264,1323,1439,1904]" pageId="5" pageNumber="6">
<emphasis id="37A7EAA2FFE5FFA81F50B9ACAAA685BB" bold="true" box="[264,430,1439,1465]" pageId="5" pageNumber="6">
<typeStatus id="DA688812FFE5FFA81F50B9ACAA4D85BB" box="[264,325,1439,1465]" pageId="5" pageNumber="6">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="7DC47620FFE5FFA81FE6B9ACA96785BB" box="[446,623,1439,1465]" name="Costa Rica" pageId="5" pageNumber="6">COSTA RICA</collectingCountry>
: Heredia Provice, La
<taxonomicName id="C2D34D33FFE5FFA81D2FB9ACA8B085BB" box="[887,952,1439,1465]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="selva">Selva</taxonomicName>
Biological Station, near Puerto Viejo.
</paragraph>
<paragraph id="056C36B0FFE5FFAB1F60B9DCAFDF8227" blockId="5.[264,1323,1439,1904]" lastBlockId="6.[264,1323,283,1906]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="37A7EAA2FFE5FFA81F60B9DCA976860B" bold="true" box="[312,638,1519,1545]" pageId="5" pageNumber="6">
<typeStatus id="DA688812FFE5FFA81F60B9DCAA7D860B" box="[312,373,1519,1545]" pageId="5" pageNumber="6">Type</typeStatus>
Material.
<typeStatus id="DA688812FFE5FFA81C54B9DCA976860B" box="[524,638,1519,1545]" pageId="5" pageNumber="6">Holotype</typeStatus>
</emphasis>
(male): “
<collectingCountry id="7DC47620FFE5FFA81CA7B9DCA8BE860B" box="[767,950,1519,1545]" name="Costa Rica" pageId="5" pageNumber="6">COSTA RICA</collectingCountry>
: Heredia Province/ nr. Puerto Viejo, La
<taxonomicName id="C2D34D33FFE5FFA81FD2BA24AAC38633" box="[394,459,1559,1585]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="selva">Selva</taxonomicName>
Bio. St./
<quantity id="C22B9B55FFE5FFA81C08BA24A9A98633" box="[592,673,1559,1585]" metricMagnitude="1" metricUnit="m" metricValue="5.45592" pageId="5" pageNumber="6" unit="ft" value="179.0">179 ft</quantity>
., Station Grounds/
<date id="716D1070FFE5FFA81DCDBA24AF2E8633" box="[917,1062,1559,1585]" pageId="5" pageNumber="6" value="2003-08-19">
<number id="B31B2497FFE5FFA81DCDBA24A8BA8633" box="[917,946,1559,1585]" pageId="5" pageNumber="6" value="19.0">19</number>
.viii.
<number id="B31B2497FFE5FFA81DBDBA24AF2E8633" box="[997,1062,1559,1585]" pageId="5" pageNumber="6" value="2003.0">2003</number>
</date>
, C.R. Bartlett, leg.”, “
<typeStatus id="DA688812FFE5FFA81F4ABA0CAAA7865B" box="[274,431,1599,1625]" pageId="5" pageNumber="6">HOLOTYPE</typeStatus>
/
<taxonomicName id="C2D34D33FFE5FFA81F99BA0CA93D865B" box="[449,565,1599,1625]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Hemiosus</taxonomicName>
/
<taxonomicName id="C2D34D33FFE5FFA81C1FBA0CA98A865B" box="[583,642,1599,1625]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="selva">selva</taxonomicName>
/ des. A.E.Z. Short” (
<collectionCode id="63C2AE75FFE5FFA81DD9BA0CA8D9865B" box="[897,977,1599,1625]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="5" pageNumber="6">INBio</collectionCode>
).
<emphasis id="37A7EAA2FFE5FFA81DB7BA0CAAE98683" bold="true" pageId="5" pageNumber="6">
<typeStatus id="DA688812FFE5FFA81DB7BA0CAF64865B" box="[1007,1132,1599,1625]" pageId="5" pageNumber="6">Paratypes</typeStatus>
(
<number id="B31B2497FFE5FFA81A26BA0CAFA7865B" box="[1150,1199,1599,1625]" pageId="5" pageNumber="6" value="129.0">129</number>
):
<collectingCountry id="7DC47620FFE5FFA81A90BA0CAA5F8683" name="Costa Rica" pageId="5" pageNumber="6">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
Turrialba,
<date id="716D1070FFE5FFA81C31BA54A81E8683" box="[617,790,1639,1665]" pageId="5" pageNumber="6" value="1965-03-13" valueMax="1965-03-17" valueMin="1965-03-13">
<number id="B31B2497FFE5FFA81C31BA54A9BC8683" box="[617,692,1639,1665]" pageId="5" pageNumber="6" value="15.0" valueMax="17.0" valueMin="13.0">13–17</number>
.iii.
<number id="B31B2497FFE5FFA81C8FBA54A81E8683" box="[727,790,1639,1665]" pageId="5" pageNumber="6" value="1965.0">1965</number>
</date>
, S.S. &amp; W.D. Duckworth leg. (
<number id="B31B2497FFE5FFA81ACCBA54AFBD8683" box="[1172,1205,1639,1665]" pageId="5" pageNumber="6" value="18.0">18</number>
:
<collectionCode id="63C2AE75FFE5FFA81A9CBA54AE168683" LSID="urn:lsid:biocol.org:col:34871" box="[1220,1310,1639,1665]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="6">USNM</collectionCode>
); same data but
<date id="716D1070FFE5FFA81FE4BABCA96886AB" box="[444,608,1679,1705]" pageId="5" pageNumber="6" value="1965-02-22" valueMax="1965-02-28" valueMin="1965-02-22">
<number id="B31B2497FFE5FFA81FE4BABCA90286AB" box="[444,522,1679,1705]" pageId="5" pageNumber="6" value="25.0" valueMax="28.0" valueMin="22.0">22–28</number>
.ii.
<number id="B31B2497FFE5FFA81C7BBABCA96886AB" box="[547,608,1679,1705]" pageId="5" pageNumber="6" value="1965.0">1965</number>
</date>
(
<number id="B31B2497FFE5FFA81C2CBABCA99E86AB" box="[628,662,1679,1705]" pageId="5" pageNumber="6" value="22.0">22</number>
:
<collectionCode id="63C2AE75FFE5FFA81CFDBABCA9F786AB" LSID="urn:lsid:biocol.org:col:34871" box="[677,767,1679,1705]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="6">USNM</collectionCode>
); Turrialba,
<date id="716D1070FFE5FFA81DCDBABCAF1586AB" box="[917,1053,1679,1705]" pageId="5" pageNumber="6" value="1965-07-15">
<number id="B31B2497FFE5FFA81DCDBABCA8BA86AB" box="[917,946,1679,1705]" pageId="5" pageNumber="6" value="15.0">15</number>
.vii.
<number id="B31B2497FFE5FFA81D85BABCAF1586AB" box="[989,1053,1679,1705]" pageId="5" pageNumber="6" value="1965.0">1965</number>
</date>
, P.J. Spangler leg. (
<number id="B31B2497FFE5FFA81B4DBABCAE2F86AB" box="[1301,1319,1679,1705]" pageId="5" pageNumber="6" value="9.0">9</number>
:
<collectionCode id="63C2AE75FFE5FFA81F50BA84AA6A86D3" LSID="urn:lsid:biocol.org:col:34871" box="[264,354,1719,1745]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="6">USNM</collectionCode>
); Turrialba,
<quantity id="C22B9B55FFE5FFA81FA8BA84A93586D3" box="[496,573,1719,1745]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="6" unit="m" value="650.0">650 m</quantity>
,
<date id="716D1070FFE5FFA81C10BA84A9FE86D3" box="[584,758,1719,1745]" pageId="5" pageNumber="6" value="1970-08-04" valueMax="1970-08-13" valueMin="1970-08-04">
<number id="B31B2497FFE5FFA81C10BA84A98D86D3" box="[584,645,1719,1745]" pageId="5" pageNumber="6" value="8.5" valueMax="13.0" valueMin="4.0">4–13</number>
.viii.
<number id="B31B2497FFE5FFA81CEFBA84A9FE86D3" box="[695,758,1719,1745]" pageId="5" pageNumber="6" value="1970.0">1970</number>
</date>
, HG-vapor light trap, J. &amp; M. Sedlacek leg. (
<number id="B31B2497FFE5FFA81B5CBA84AE2E86D3" box="[1284,1318,1719,1745]" pageId="5" pageNumber="6" value="29.0">29</number>
:
<collectionCode id="63C2AE75FFE5FFA81F50BAECAA5C86FB" box="[264,340,1759,1785]" pageId="5" pageNumber="6">AEZS</collectionCode>
,
<collectionCode id="63C2AE75FFE5FFA81F30BAECAAB486FB" LSID="urn:lsid:biocol.org:col:34697" box="[360,444,1759,1785]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="5" pageNumber="6">BPBM</collectionCode>
,
<collectionCode id="63C2AE75FFE5FFA81F89BAECA92986FB" box="[465,545,1759,1785]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="6">NMW</collectionCode>
);
<emphasis id="37A7EAA2FFE5FFA81C66BAEDA9D286FA" bold="true" box="[574,730,1758,1784]" pageId="5" pageNumber="6">Guanacaste:</emphasis>
Las Cañas,
<date id="716D1070FFE5FFA81DDBBAECAF0386FB" box="[899,1035,1759,1785]" pageId="5" pageNumber="6" value="1965-07-13">
<number id="B31B2497FFE5FFA81DDBBAECA8A886FB" box="[899,928,1759,1785]" pageId="5" pageNumber="6" value="13.0">13</number>
.vii.
<number id="B31B2497FFE5FFA81D93BAECAF0386FB" box="[971,1035,1759,1785]" pageId="5" pageNumber="6" value="1965.0">1965</number>
</date>
, P.J. Spangler leg. (
<number id="B31B2497FFE5FFA81B4FBAECAE2D86FB" box="[1303,1317,1759,1785]" pageId="5" pageNumber="6" value="1.0">1</number>
:
<collectionCode id="63C2AE75FFE5FFA81F50BB34AA6A8723" LSID="urn:lsid:biocol.org:col:34871" box="[264,354,1799,1825]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="6">USNM</collectionCode>
);
<emphasis id="37A7EAA2FFE5FFA81F24BB35AAE28722" bold="true" box="[380,490,1798,1824]" pageId="5" pageNumber="6">Heredia:</emphasis>
same data as
<typeStatus id="DA688812FFE5FFA81CFBBB34A8028723" box="[675,778,1799,1825]" pageId="5" pageNumber="6">holotype</typeStatus>
(
<number id="B31B2497FFE5FFA81D7BBB34A8398723" box="[803,817,1799,1825]" pageId="5" pageNumber="6" value="1.0">1</number>
:
<collectionCode id="63C2AE75FFE5FFA81D1CBB34A89B8723" box="[836,915,1799,1825]" pageId="5" pageNumber="6">AEZS</collectionCode>
); La
<taxonomicName id="C2D34D33FFE5FFA81D80BB34AF118723" box="[984,1049,1799,1825]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="selva">Selva</taxonomicName>
Biological Station nr. Puerto Viejo, HG Vapor light, (
<number id="B31B2497FFE5FFA81CDBBB1DA9D8874A" box="[643,720,1838,1864]" pageId="5" pageNumber="6" value="14.5" valueMax="15.0" valueMin="14.0">14–15</number>
).
<date id="716D1070FFE5FFA81C84BB1DA822874A" box="[732,810,1838,1864]" pageId="5" pageNumber="6" value="2005-01">
i.
<number id="B31B2497FFE5FFA81CB0BB1DA822874A" box="[744,810,1838,1864]" pageId="5" pageNumber="6" value="2005.0">2005</number>
</date>
, A.E.Z. Short leg., AS-05-
<number id="B31B2497FFE5FFA81A28BB1DAF93874A" box="[1136,1179,1838,1864]" pageId="5" pageNumber="6" value="153.0">153</number>
(
<number id="B31B2497FFE5FFA81AE9BB1DAFB7874A" box="[1201,1215,1838,1864]" pageId="5" pageNumber="6" value="1.0">1</number>
:
<collectionCode id="63C2AE75FFE5FFA81A97BB1DAE15874A" box="[1231,1309,1838,1864]" pageId="5" pageNumber="6">AEZS</collectionCode>
); same data but
<date id="716D1070FFE5FFA81F9DBB65A9768772" box="[453,638,1878,1904]" pageId="5" pageNumber="6" value="2003-08-18" valueMax="2003-08-19" valueMin="2003-08-18">
<number id="B31B2497FFE5FFA81F9DBB65A9058772" box="[453,525,1878,1904]" pageId="5" pageNumber="6" value="18.5" valueMax="19.0" valueMin="18.0">18–19</number>
.viii.
<number id="B31B2497FFE5FFA81C67BB65A9768772" box="[575,638,1878,1904]" pageId="5" pageNumber="6" value="2003.0">2003</number>
</date>
(
<number id="B31B2497FFE5FFA81CCFBB65A9A08772" box="[663,680,1878,1904]" pageId="5" pageNumber="6" value="5.0">5</number>
:
<collectionCode id="63C2AE75FFE5FFA81CE3BB65A8018772" box="[699,777,1878,1904]" pageId="5" pageNumber="6">AEZS</collectionCode>
); same data but (
<number id="B31B2497FFE5FFA81DB1BB65AF308772" box="[1001,1080,1878,1904]" pageId="5" pageNumber="6" value="23.0" valueMax="24.0" valueMin="22.0">22–24</number>
).
<date id="716D1070FFE5FFA81A1CBB65AF878772" box="[1092,1167,1878,1904]" pageId="5" pageNumber="6" value="2000-01">
i.
<number id="B31B2497FFE5FFA81A17BB65AF878772" box="[1103,1167,1878,1904]" pageId="5" pageNumber="6" value="2000.0">2000</number>
</date>
(
<number id="B31B2497FFE5FFA81AF0BB65AFCF8772" box="[1192,1223,1878,1904]" pageId="5" pageNumber="6" value="11.0">11</number>
:
<collectionCode id="63C2AE75FFE5FFA81A82BB65AE2E8772" box="[1242,1318,1878,1904]" pageId="5" pageNumber="6">AEZS</collectionCode>
,
<collectionCode id="63C2AE75FFE6FFAB1F50BD2FAA5C8134" box="[264,340,284,310]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="6" pageNumber="7">INBio</collectionCode>
,
<collectionCode id="63C2AE75FFE6FFAB1F39BD2FAAB38134" LSID="urn:lsid:biocol.org:col:34871" box="[353,443,284,310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="7">USNM</collectionCode>
);
<emphasis id="37A7EAA2FFE6FFAB1F88BD28A9248137" bold="true" box="[464,556,283,309]" pageId="6" pageNumber="7">Limón:</emphasis>
La Lola nr. Matina,
<date id="716D1070FFE6FFAB1D7ABD2FA8AB8134" box="[802,931,284,310]" pageId="6" pageNumber="7" value="1965-03-11">
<number id="B31B2497FFE6FFAB1D7ABD2FA8368134" box="[802,830,284,310]" pageId="6" pageNumber="7" value="11.0">11</number>
.iii.
<number id="B31B2497FFE6FFAB1D3ABD2FA8AB8134" box="[866,931,284,310]" pageId="6" pageNumber="7" value="1965.0">1965</number>
</date>
, S.S. &amp; W.D. Duckworth leg. (
<number id="B31B2497FFE6FFAB1B4DBD2FAE2E8134" box="[1301,1318,284,310]" pageId="6" pageNumber="7" value="8.0">8</number>
:
<collectionCode id="63C2AE75FFE6FFAB1F50BD77AA6A815C" LSID="urn:lsid:biocol.org:col:34871" box="[264,354,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="7">USNM</collectionCode>
); Hitoy Cerere Biological Reserve, Hitoy Cerere Station,
<quantity id="C22B9B55FFE6FFAB1A7FBD77AF70815C" box="[1063,1144,324,350]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="7" unit="m" value="160.0">160 m</quantity>
,
<date id="716D1070FFE6FFAB1AD1BD77AFF2815C" box="[1161,1274,324,350]" pageId="6" pageNumber="7" value="2000-01-17">
<number id="B31B2497FFE6FFAB1AD1BD77AFAE815C" box="[1161,1190,324,350]" pageId="6" pageNumber="7" value="17.0">17</number>
.i.
<number id="B31B2497FFE6FFAB1AEEBD77AFF2815C" box="[1206,1274,324,350]" pageId="6" pageNumber="7" value="2000.0">2000</number>
</date>
, W. Arana leg. (
<number id="B31B2497FFE6FFAB1FFABD5FAAB88184" box="[418,432,364,390]" pageId="6" pageNumber="7" value="1.0">1</number>
:
<collectionCode id="63C2AE75FFE6FFAB1F9BBD5FA9198184" box="[451,529,364,390]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="6" pageNumber="7">INBio</collectionCode>
), same locality but
<quantity id="C22B9B55FFE6FFAB1D55BD5FA8578184" box="[781,863,364,390]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="m" value="200.0">200 m</quantity>
,
<date id="716D1070FFE6FFAB1D29BD5FA8E08184" box="[881,1000,364,390]" pageId="6" pageNumber="7" value="2000-02-10">
<number id="B31B2497FFE6FFAB1D29BD5FA8868184" box="[881,910,364,390]" pageId="6" pageNumber="7" value="10.0">10</number>
.ii.
<number id="B31B2497FFE6FFAB1DFFBD5FA8E08184" box="[935,1000,364,390]" pageId="6" pageNumber="7" value="2000.0">2000</number>
</date>
(
<number id="B31B2497FFE6FFAB1A5ABD5FAF188184" box="[1026,1040,364,390]" pageId="6" pageNumber="7" value="1.0">1</number>
:
<collectionCode id="63C2AE75FFE6FFAB1A7CBD5FAF7A8184" box="[1060,1138,364,390]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="6" pageNumber="7">INBio</collectionCode>
); Hitoy Cerere Biological Reserve, “send. a catarata” [trail to waterfall],
<quantity id="C22B9B55FFE6FFAB1DE6BDA0AF4081AF" box="[958,1096,403,429]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="6" pageNumber="7" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100–200 m</quantity>
,
<date id="716D1070FFE6FFAB1A0EBDA0AFEE81AF" box="[1110,1254,403,429]" pageId="6" pageNumber="7" value="2002-08-26">
<number id="B31B2497FFE6FFAB1A0EBDA0AF7D81AF" box="[1110,1141,403,429]" pageId="6" pageNumber="7" value="26.0">26</number>
.viii.
<number id="B31B2497FFE6FFAB1AFEBDA0AFEE81AF" box="[1190,1254,403,429]" pageId="6" pageNumber="7" value="2002.0">2002</number>
</date>
, light trap, W. Arana leg. (
<number id="B31B2497FFE6FFAB1FAFBD88A90D81D7" box="[503,517,443,469]" pageId="6" pageNumber="7" value="1.0">1</number>
:
<collectionCode id="63C2AE75FFE6FFAB1C4ABD88A96881D7" box="[530,608,443,469]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="6" pageNumber="7">INBio</collectionCode>
); same data but
<date id="716D1070FFE6FFAB1D42BD88A89E81D7" box="[794,918,443,469]" pageId="6" pageNumber="7" value="1999-11-11">
<number id="B31B2497FFE6FFAB1D42BD88A83D81D7" box="[794,821,443,469]" pageId="6" pageNumber="7" value="11.0">11</number>
.xi.
<number id="B31B2497FFE6FFAB1D0EBD88A89E81D7" box="[854,918,443,469]" pageId="6" pageNumber="7" value="1999.0">1999</number>
</date>
(
<number id="B31B2497FFE6FFAB1DF1BD88A8BF81D7" box="[937,951,443,469]" pageId="6" pageNumber="7" value="1.0">1</number>
:
<collectionCode id="63C2AE75FFE6FFAB1D9CBD88AF1A81D7" box="[964,1042,443,469]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="6" pageNumber="7">INBio</collectionCode>
); Guapiles, Santa Clara,
<date id="716D1070FFE6FFAB1F50BDD0AA8F81FF" box="[264,391,483,509]" pageId="6" pageNumber="7" value="1935-03-24">
<number id="B31B2497FFE6FFAB1F50BDD0AA2081FF" box="[264,296,483,509]" pageId="6" pageNumber="7" value="24.0">24</number>
.iii.
<number id="B31B2497FFE6FFAB1F12BDD0AA8F81FF" box="[330,391,483,509]" pageId="6" pageNumber="7" value="1935.0">1935</number>
</date>
(
<number id="B31B2497FFE6FFAB1FC6BDD0AAA481FF" box="[414,428,483,509]" pageId="6" pageNumber="7" value="1.0">1</number>
:
<collectionCode id="63C2AE75FFE6FFAB1FE4BDD0A91E81FF" LSID="urn:lsid:biocol.org:col:34871" box="[444,534,483,509]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="7">USNM</collectionCode>
); Hamburg Farm, Reventezon,
<date id="716D1070FFE6FFAB1DCABDD0AF1A81FF" box="[914,1042,483,509]" pageId="6" pageNumber="7" value="1929-06-27">
<number id="B31B2497FFE6FFAB1DCABDD0A8BA81FF" box="[914,946,483,509]" pageId="6" pageNumber="7" value="27.0">27</number>
.vi.
<number id="B31B2497FFE6FFAB1D8ABDD0AF1A81FF" box="[978,1042,483,509]" pageId="6" pageNumber="7" value="1929.0">1929</number>
</date>
, F. Nevermann leg. (
<number id="B31B2497FFE6FFAB1B4CBDD0AE2E81FF" box="[1300,1318,483,509]" pageId="6" pageNumber="7" value="7.0">7</number>
:
<collectionCode id="63C2AE75FFE6FFAB1F50BE38AA6A8227" LSID="urn:lsid:biocol.org:col:34871" box="[264,354,523,549]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="7">USNM</collectionCode>
); same data but
<date id="716D1070FFE6FFAB1C46BE38A99D8227" box="[542,661,523,549]" pageId="6" pageNumber="7" value="1938-02-26">
<number id="B31B2497FFE6FFAB1C46BE38A9368227" box="[542,574,523,549]" pageId="6" pageNumber="7" value="26.0">26</number>
.ii.
<number id="B31B2497FFE6FFAB1C0FBE38A99D8227" box="[599,661,523,549]" pageId="6" pageNumber="7" value="1938.0">1938</number>
</date>
(
<number id="B31B2497FFE6FFAB1CF1BE38A9BF8227" box="[681,695,523,549]" pageId="6" pageNumber="7" value="1.0">1</number>
:
<collectionCode id="63C2AE75FFE6FFAB1C9CBE38A8168227" LSID="urn:lsid:biocol.org:col:34871" box="[708,798,523,549]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="7">USNM</collectionCode>
); same data but
<date id="716D1070FFE6FFAB1D82BE38AF358227" box="[986,1085,523,549]" pageId="6" pageNumber="7" value="1933-01-06">
<number id="B31B2497FFE6FFAB1D82BE38A8E28227" box="[986,1002,523,549]" pageId="6" pageNumber="7" value="6.0">6</number>
.i.
<number id="B31B2497FFE6FFAB1DA3BE38AF358227" box="[1019,1085,523,549]" pageId="6" pageNumber="7" value="1933.0">1933</number>
</date>
, (
<number id="B31B2497FFE6FFAB1A0EBE38AF6C8227" box="[1110,1124,523,549]" pageId="6" pageNumber="7" value="1.0">1</number>
:
<collectionCode id="63C2AE75FFE6FFAB1A29BE38AFC38227" LSID="urn:lsid:biocol.org:col:34871" box="[1137,1227,523,549]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="7">USNM</collectionCode>
).
</paragraph>
<paragraph id="056C36B0FFE6FFAB1F60BE00AFE982C7" blockId="6.[264,1323,283,1906]" pageId="6" pageNumber="7">
<emphasis id="37A7EAA2FFE6FFAB1F60BE00A958824F" bold="true" box="[312,592,563,589]" pageId="6" pageNumber="7">Differential Diagnosis.</emphasis>
Total length
<number id="B31B2497FFE6FFAB1CA0BE00A814824F" box="[760,796,563,589]" pageId="6" pageNumber="7" value="2.8">2.8</number>
–3.0 mm. This species can be distinguished from the remaining two mesoamerican species by the shape of the black iridescent spot on the pronotum (
<figureCitation id="9DE82A35FFE6FFAB1FE0BEB0A909829F" box="[440,513,643,669]" captionStart="FIGURES 3 – 5" captionStartId="7.[264,382,1004,1028]" captionTargetBox="[418,1160,325,975]" captionTargetId="figure@7.[409,1177,304,994]" captionTargetPageId="7" captionText="FIGURES 3 – 5. Details of Hemiosus selva Short, sp. n. (3, 5) and H. exilis (LeConte) (4). 3 – 4: pronotal coloration, scale bar 0.18 mm; 5: aedeagus, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.174868" httpUri="https://zenodo.org/record/174868/files/figure.png" pageId="6" pageNumber="7">
Fig.
<number id="B31B2497FFE6FFAB1FA9BEB0A909829F" box="[497,513,643,669]" pageId="6" pageNumber="7" value="3.0">3</number>
</figureCitation>
), which does not reach the lateral margins but narrowly reaches the medial anterior margin; and by the strongly acuminate median lobe of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="4DC9653BFFE6FFAB1F60BEE1AAA68680" pageId="6" pageNumber="7" type="description">
<paragraph id="056C36B0FFE6FFAB1F60BEE1AAA68680" blockId="6.[264,1323,283,1906]" pageId="6" pageNumber="7">
<emphasis id="37A7EAA2FFE6FFAB1F60BEE1A97C82EE" bold="true" box="[312,628,722,748]" pageId="6" pageNumber="7">Description. Coloration.</emphasis>
Head entirely iridescent black. Maxillary palps largely yellow with tip of apical segment dark brown. Pronotum largely iridescent black centrally and reaching posterior margin in medial half and anterior margin in medial fifth to sixth (
<figureCitation id="9DE82A35FFE6FFAB1F49BF79AA508366" box="[273,344,842,868]" captionStart="FIGURES 3 – 5" captionStartId="7.[264,382,1004,1028]" captionTargetBox="[418,1160,325,975]" captionTargetId="figure@7.[409,1177,304,994]" captionTargetPageId="7" captionText="FIGURES 3 – 5. Details of Hemiosus selva Short, sp. n. (3, 5) and H. exilis (LeConte) (4). 3 – 4: pronotal coloration, scale bar 0.18 mm; 5: aedeagus, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.174868" httpUri="https://zenodo.org/record/174868/files/figure.png" pageId="6" pageNumber="7">
Fig.
<number id="B31B2497FFE6FFAB1F10BF79AA508366" box="[328,344,842,868]" pageId="6" pageNumber="7" value="3.0">3</number>
</figureCitation>
); remainder of pronotal margins light brown to yellow. Elytra largely light brown to yellow with rows of serial punctures medium to dark brown. Dark brown maculae are present at posterior end of scutellary stria along elytral suture and medially on each elytron in interstriae
<number id="B31B2497FFE6FFAB1FFCBFF1AABB83DE" box="[420,435,962,988]" pageId="6" pageNumber="7" value="7.0">7</number>
and
<number id="B31B2497FFE6FFAB1FB6BFF1AAF783DE" box="[494,511,962,988]" pageId="6" pageNumber="7" value="8.0">8</number>
. Elytral color patterns exhibit moderate intraspecific variation, with some individuals lacking elytral maculae.
<emphasis id="37A7EAA2FFE6FFAB1D51BFD9A85B8406" bold="true" box="[777,851,1002,1028]" pageId="6" pageNumber="7">Head.</emphasis>
Ground punctation on head dense and moderately coarse, slightly less coarse than punctation on pronotum. Maxillary palps slightly longer than anterior margin of clypeus.
<emphasis id="37A7EAA2FFE6FFAB1D66B80AA8AA8451" bold="true" box="[830,930,1081,1107]" pageId="6" pageNumber="7">Thorax.</emphasis>
Ground punctation on pronotum very coarse, uneven. Lateral margins of pronotum finely to moderately crenulate, often bearing long setae. Scutellum with coarse punctures. Median process of mesoventrite strongly elevated in anterior two-fifths, with posterior half only moderately elevated. Elytra with interstrial punctures coarse and distinct medially, subequal to slightly smaller than serial punctures, and becoming finer and nearly obsolete in lateral interstriae. Protarsi of males with segments
<number id="B31B2497FFE6FFAB1CFDB91AA9DA8541" box="[677,722,1321,1347]" pageId="6" pageNumber="7" value="2.5" valueMax="3.0" valueMin="2.0">2–3</number>
slightly enlarged and with longer ventral setae.
<emphasis id="37A7EAA2FFE6FFAB1F50B962AA818569" bold="true" box="[264,393,1361,1387]" pageId="6" pageNumber="7">Abdomen.</emphasis>
First abdominal ventrite with median carina that is narrow and strongly raised anteriorly, gradually becoming wider and lower posteriorly. Abdominal ventrites
<number id="B31B2497FFE6FFAB1A9BB94AAFE78591" box="[1219,1263,1401,1427]" pageId="6" pageNumber="7" value="3.0" valueMax="4.0" valueMin="2.0">2–4</number>
with trace of broad, low median carina in anterior half. Emargination of fifth ventrite with single acute projection medially. All ventrites very finely crenulate on lateral margins. Aedeagus (
<figureCitation id="9DE82A35FFE6FFAB1FD6B9C2AADE8609" box="[398,470,1521,1547]" captionStart="FIGURES 3 – 5" captionStartId="7.[264,382,1004,1028]" captionTargetBox="[418,1160,325,975]" captionTargetId="figure@7.[409,1177,304,994]" captionTargetPageId="7" captionText="FIGURES 3 – 5. Details of Hemiosus selva Short, sp. n. (3, 5) and H. exilis (LeConte) (4). 3 – 4: pronotal coloration, scale bar 0.18 mm; 5: aedeagus, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.174868" httpUri="https://zenodo.org/record/174868/files/figure.png" pageId="6" pageNumber="7">
Fig.
<number id="B31B2497FFE6FFAB1F9FB9C2AADE8609" box="[455,470,1521,1547]" pageId="6" pageNumber="7" value="5.0">5</number>
</figureCitation>
) with median lobe strongly pointed apically, almost reaching the same level as apices of parameres. Parameres moderately narrowed apically to more or less acute apices, but not as acute as median lobe. Basal piece strongly asymmetrical in proximal half.
</paragraph>
</subSubSection>
<subSubSection id="4DC9653BFFE6FFAB1F60BAA3A8B086D0" pageId="6" pageNumber="7" type="etymology">
<paragraph id="056C36B0FFE6FFAB1F60BAA3A8B086D0" blockId="6.[264,1323,283,1906]" pageId="6" pageNumber="7">
<emphasis id="37A7EAA2FFE6FFAB1F60BAA3AACC86A8" bold="true" box="[312,452,1680,1706]" pageId="6" pageNumber="7">Etymology.</emphasis>
This species is named after “
<emphasis id="37A7EAA2FFE6FFAB1D63BAA3A89086AB" box="[827,920,1680,1705]" italics="true" pageId="6" pageNumber="7">
la
<taxonomicName id="C2D34D33FFE6FFAB1D04BAA3A89086AB" box="[860,920,1680,1705]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="selva">selva</taxonomicName>
</emphasis>
”, the Spanish word for “jungle”. The species name is to be considered a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="4DC9653BFFE6FFAB1F60BAECA9FC8720" pageId="6" pageNumber="7" type="distribution">
<paragraph id="056C36B0FFE6FFAB1F60BAECA9FC8720" blockId="6.[264,1323,283,1906]" pageId="6" pageNumber="7">
<emphasis id="37A7EAA2FFE6FFAB1F60BAECAAD086FB" bold="true" box="[312,472,1759,1785]" pageId="6" pageNumber="7">Distribution.</emphasis>
<collectingCountry id="7DC47620FFE6FFAB1FB1BAD3A96586F8" box="[489,621,1760,1786]" name="Costa Rica" pageId="6" pageNumber="7">Costa Rica</collectingCountry>
, with all but one locality (i.e. Las Cañas in Guanacaste) on the Atlantic side of the continental divide.
</paragraph>
</subSubSection>
<subSubSection id="4DC9653BFFE6FFAB1F60BB1CA9248770" pageId="6" pageNumber="7" type="discussion">
<paragraph id="056C36B0FFE6FFAB1F60BB1CA9248770" blockId="6.[264,1323,283,1906]" pageId="6" pageNumber="7">
<emphasis id="37A7EAA2FFE6FFAB1F60BB1CAAB9874B" bold="true" box="[312,433,1839,1865]" pageId="6" pageNumber="7">Remarks.</emphasis>
Most known specimens were collected at lights. At least one was collected in an “aquatic” situation.
</paragraph>
</subSubSection>
</treatment>
</document>
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