213 lines
26 KiB
XML
213 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.43.80969" ID-GBIF-Dataset="6b8ee741-b4b9-4636-9df2-94adf46aee53" ID-Pensoft-Pub="1314-2615-43-61" ID-Pensoft-UUID="94B591976E9051C7AA220F5AB7F30AF7" ID-ZooBank="9B9E6A5AA0784AC0A1DF6B1D32FF4DB3" ModsDocID="1314-2615-43-61" checkinTime="1654082233325" checkinUser="pensoft" docAuthor="Sasakawa, Koji & Ito, Hirotaro" docDate="2022" docId="8CD5F85F6B595FA08552C1864653A65E" docLanguage="en" docName="SubterBiol 43: 61-71" docOrigin="Subterranean Biology 43" docPubDate="2022-05-31" docSource="http://dx.doi.org/10.3897/subtbiol.43.80969" docTitle="Pterostichus (Nialoe) asahinus Habu & Baba 1960" docType="treatment" docVersion="4" id="94B591976E9051C7AA220F5AB7F30AF7" lastPageNumber="61" masterDocId="94B591976E9051C7AA220F5AB7F30AF7" masterDocTitle="The identities of two species in the Pterostichus macrogenys species group of subterranean carabid beetles (Coleoptera, Carabidae) revealed by external morphometric analysis and comparative genital morphology" masterLastPageNumber="71" masterPageNumber="61" pageNumber="61" updateTime="1732567018414" updateUser="ExternalLinkService">
|
|
<mods:mods id="6876CA050C8DA795351A3C000A679FA8" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="7B6EF1808174EEF7645AC2A010F8D7AE">
|
|
<mods:title id="6CF04D5644FE6BFB0CACA376C9AA46C1">The identities of two species in the Pterostichus macrogenys species group of subterranean carabid beetles (Coleoptera, Carabidae) revealed by external morphometric analysis and comparative genital morphology</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="F0F6353BBFC56F78789A2C711C31FFD7" type="personal">
|
|
<mods:role id="9A3C8E5795BEB8F431B37C546CB949B4">
|
|
<mods:roleTerm id="D5A31D6D8BE49B2AD96A1E92057DDE43">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="4B42463A86C139C963CC7B38F36015C9">Sasakawa, Koji</mods:namePart>
|
|
<mods:nameIdentifier id="8F86769B3EB0EB1B939B7822BC6FD841" type="ORCID">https://orcid.org/0000-0001-9246-5777</mods:nameIdentifier>
|
|
<mods:affiliation id="2B2103475CC7C14B37A663D691657C4E">Laboratory of Zoology, Department of Science Education, Faculty of Education, Chiba University, 1 - 33 Yayoi-cho, Inage-ku, Chiba-shi, Chiba 263 - 8522, Japan</mods:affiliation>
|
|
<mods:nameIdentifier id="64AB6B1851B3208C116925B4DD9F7079" type="email">ksasa@chiba-u.jp</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name id="E333CD3C5E03105A7D48BA6B6316A8C3" type="personal">
|
|
<mods:role id="3531DDD5F1B0A783ED0436D1624C4FB8">
|
|
<mods:roleTerm id="8385CBCDBED48BC5051EE695A8432789">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="3C7659E999DEFF2CB6619F6EBEE75CD5">Ito, Hirotaro</mods:namePart>
|
|
<mods:affiliation id="4EE7E46C87B95E72FCC40CAAE6904A33">1 - 14 - 16 Awayama, Niigata-shi, Niigata 950 - 0843, Japan</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource id="A2362BC628BE451505478A5DF175D0A3">text</mods:typeOfResource>
|
|
<mods:relatedItem id="D36D84320554C4E45C9C227E94387667" type="host">
|
|
<mods:titleInfo id="E4BF79261F9C403870A461B040EA7300">
|
|
<mods:title id="78830B0D939D7F141F23049E2C2D2C4C">Subterranean Biology</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="651A6CACFDB088E14426F209AE536DEF">
|
|
<mods:date id="EB798D17D538A5963B0B9101CC9AD197">2022</mods:date>
|
|
<mods:detail id="9E41CA36E7E3AA5D0D184DE758EF63C8" type="pubDate">
|
|
<mods:number id="139DDF23A7EC5F1E991B0C2AC0CF8E25">2022-05-31</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="D05780EB0E3A5B84578255E3562FB584" type="volume">
|
|
<mods:number id="76D22383673D334F7ED255BF978DC72B">43</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="28E30FF17FE62B1A348C0ADFBE8D2611" unit="page">
|
|
<mods:start id="37B04BFA4E2EF7E326CAB195A6B805FA">61</mods:start>
|
|
<mods:end id="B111D2DF5EF23CFA632B0F2FF05647C4">71</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location id="162245EA092595257FE52A311D6A5755">
|
|
<mods:url id="8AF28AE5A49CF51F3408FBB0494D60D8">http://dx.doi.org/10.3897/subtbiol.43.80969</mods:url>
|
|
</mods:location>
|
|
<mods:classification id="1B8DC5B1C8F21EC24C3FD1B6056F705F">journal article</mods:classification>
|
|
<mods:identifier id="46C7B99EE74EAD7C81B56DD5D839B404" type="DOI">http://dx.doi.org/10.3897/subtbiol.43.80969</mods:identifier>
|
|
<mods:identifier id="DB7EE63967FEFDE918D96654D520261B" type="Pensoft-Pub">1314-2615-43-61</mods:identifier>
|
|
<mods:identifier id="594BA2A4D33DD3A75620F3A8C7515A26" type="ZooBank">9B9E6A5AA0784AC0A1DF6B1D32FF4DB3</mods:identifier>
|
|
<mods:identifier id="71CF11C3729629C4C7161F1B72A2047F" type="Pensoft-UUID">94B591976E9051C7AA220F5AB7F30AF7</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="8CD5F85F6B595FA08552C1864653A65E" ID-GBIF-Taxon="196160410" LSID="urn:lsid:plazi:treatment:8CD5F85F6B595FA08552C1864653A65E" httpUri="http://treatment.plazi.org/id/8CD5F85F6B595FA08552C1864653A65E" lastPageNumber="61" pageId="0" pageNumber="61" scope_family="Carabidae" scope_order="Coleoptera">
|
|
<subSubSection id="861B1EB040BBD1D29BB399C9A489EBC5" pageId="0" pageNumber="61" type="nomenclature">
|
|
<paragraph id="4EA693A9BE074BB2F38AFEBD1B337C56" pageId="0" pageNumber="61">
|
|
<taxonomicName id="AD0AF95B5EC689D2C380903A6D6662D1" ID-CoL="8MHQC" LSID="8CD5F85F-6B59-5FA0-8552-C1864653A65E" authority="Habu & Baba, 1960" authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus" subGenus="Nialoe">Pterostichus (Nialoe) asahinus Habu & Baba, 1960</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="8957434F722AAEB2C6A61C264493DE76" pageId="0" pageNumber="61" type="description">
|
|
<paragraph id="C30A92733759E17FAB5BA1B928C747A7" pageId="0" pageNumber="61">
|
|
<figureCitation id="393B0757F055E7F6957DD689FE23CBA1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pterostichus asahinus male from Miomote, on the right bank of the Miomotegawa River A habitus dorsal view B-E Endophallus left lateral (B) ventral (C) right lateral (D) and dorsal (E) views F-H right paramere left lateral (F) apical (G) and dorsal (H) views. go, gonopore; lal, left apical lobe; lpl, left preapical lobe; rpl, right preapical lobe." figureDoi="10.3897/subtbiol.43.80969.figure2" httpUri="https://binary.pensoft.net/fig/690341" pageId="0" pageNumber="61">Fig. 2</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="47ABBA715AF85FDA7FA0FDE1DE373F85" pageId="0" pageNumber="61" type="reference_group">
|
|
<paragraph id="0FEF55F9230AE96CC971BED422D1EB63" pageId="0" pageNumber="61">
|
|
<taxonomicName id="95AAF06B8A7248265C60325104AB4362" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Paralianoe) macrogenys subsp. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="macrogenys" subGenus="Paralianoe" subSpecies="asahinus">Pterostichus (Paralianoe) macrogenys asahinus</taxonomicName>
|
|
:
|
|
<bibRefCitation id="7EC78C595846E55EDEA53305F77E44FC" author="Habu, A" journalOrPublisher="Entomological Review Japan" pageId="0" pageNumber="61" refId="B5" refString="Habu, A, Baba, K, 1960. A new subspecies of Pterostichus macrogenys Bates from Niigata Prefecture, Japan. Akitu 9: 62." title="A new subspecies of Pterostichus macrogenys Bates from Niigata Prefecture, Japan. Akitu 9: 62." year="1960">Habu and Baba (1960)</bibRefCitation>
|
|
: 62 (original description), holotype ♀: "Mt. Dorokujin, Mts. Asahi, Niigata Pref." [Miomote, Mt.
|
|
<normalizedToken id="C2A7EEC58DD77660FF701C54444D5D91" originalValue="Dôrokujinpô">Dorokujinpo</normalizedToken>
|
|
, Murakami-shi, Niigata Prefecture, Japan];
|
|
<bibRefCitation id="C705C02C2E02BF7D3472F26AD822877E" author="Habu, A" journalOrPublisher="Transactions of Essa Entomological Society" pageId="0" pageNumber="61" pagination="1 - 58" refId="B6" refString="Habu, A, Baba, K, 1972. List of Carabidae (excluding Carabini) from Niigata Prefecture, Japan. Transactions of Essa Entomological Society 41: 1 - 58" title="List of Carabidae (excluding Carabini) from Niigata Prefecture, Japan." volume="41" year="1972">Habu and Baba (1972)</bibRefCitation>
|
|
: 19.
|
|
</paragraph>
|
|
<paragraph id="1F059D92ED1D788F750A8D21F62F6123" pageId="0" pageNumber="61">
|
|
<taxonomicName id="2D8F9DCC603CABF1B8F01EA5950E3094" authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Paralianoe) asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus" subGenus="Paralianoe">Pterostichus (Paralianoe) asahinus</taxonomicName>
|
|
:
|
|
<bibRefCitation id="9F8F5EFBF96CAF2535C3C8E8D8900D84" author="Habu, A" journalOrPublisher="Entomological Review Japan" pageId="0" pageNumber="61" pagination="13 - 17" refId="B4" refString="Habu, A, 1977. On two Pterostichus species found in caves in Niigata Prefecture, Japan (Coleoptera, Carabidae). Entomological Review Japan 30: 13 - 17" title="On two Pterostichus species found in caves in Niigata Prefecture, Japan (Coleoptera, Carabidae)." volume="30" year="1977">Habu (1977)</bibRefCitation>
|
|
: 14 (part).
|
|
</paragraph>
|
|
<paragraph id="BB4EF9BE8F6D30B281F8033CB2CB180D" pageId="0" pageNumber="61">
|
|
<taxonomicName id="408FC0438DD280BF43B3DB7C2252BE2C" authorityName="Sasakawa & Itô" authorityYear="2022" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus macrogenys" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="macrogenys">Pterostichus macrogenys</taxonomicName>
|
|
:
|
|
<bibRefCitation id="56F391B676CEEF807776ED94C0AE077B" author="Tanaka, K" editor="Ueno, S" journalOrPublisher="Vol. II. Hoikusha, Osaka" pageId="0" pageNumber="61" pagination="105 - 138" refId="B21" refString="Tanaka, K, 1985. Carabidae (Pterostichinae, Zabrinae). In: Ueno, S, Kurosawa, Y, Sato, M, Eds., The Coleoptera of Japan in Color. Vol. II. Hoikusha, Osaka: 105 - 138" title="Carabidae (Pterostichinae, Zabrinae)." volumeTitle="The Coleoptera of Japan in Color." year="1985">Tanaka (1985)</bibRefCitation>
|
|
: 114 (part?).
|
|
</paragraph>
|
|
<paragraph id="2EBEA10A4406AD46F7CC8500716DA05F" pageId="0" pageNumber="61">
|
|
<taxonomicName id="680A06BFA7C707493255C9A76BF73096" authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus" subGenus="Nialoe">Pterostichus (Nialoe) asahinus</taxonomicName>
|
|
:
|
|
<bibRefCitation id="0FDC3CA1797737440E0AD8080A6973F0" author="Bousquet, Y" editor="Loebl, I" journalOrPublisher="1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden" pageId="0" pageNumber="61" pagination="675 - 755" refId="B2" refString="Bousquet, Y, 2017. Tribe Pterostichini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Vol. 1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden: 675 - 755" title="Tribe Pterostichini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol." year="2017">Bousquet (2017)</bibRefCitation>
|
|
: 724.
|
|
</paragraph>
|
|
<paragraph id="FA4CC34011D4666FE20EEA8E83A259F1" pageId="0" pageNumber="61">
|
|
<taxonomicName id="3DB29D4FBB63206EF2841C6F0E10960C" authorityName="Sasakawa" authorityYear="2005" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus" subGenus="Nialoe">Pterostichus (Nialoe) falcispinus</taxonomicName>
|
|
:
|
|
<bibRefCitation id="B092E54EEB450F612D3570608EF2E9FC" author="Sasakawa, K" journalOrPublisher="Biogeography" pageId="0" pageNumber="61" pagination="69 - 78" refId="B12" refString="Sasakawa, K, 2005. Pterostichus macrogenys Bates, 1883 (Coleoptera, Carabidae) and its allied species of northern Japan. Biogeography 7: 69 - 78" title="Pterostichus macrogenys Bates, 1883 (Coleoptera, Carabidae) and its allied species of northern Japan." volume="7" year="2005">Sasakawa (2005)</bibRefCitation>
|
|
: 75 (original description), holotype ♂: "Cave Ishikiri, Nakajo-Machi, N-Echigo" [Ishikiri Cave, Mt. Ishikiriyama, Haguro, Tainai-shi, Niigata Prefecture, Japan];
|
|
<bibRefCitation id="3CB245FA7C1FE7E401BB593BF9ADF34D" author="Bousquet, Y" editor="Loebl, I" journalOrPublisher="1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden" pageId="0" pageNumber="61" pagination="675 - 755" refId="B2" refString="Bousquet, Y, 2017. Tribe Pterostichini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Vol. 1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden: 675 - 755" title="Tribe Pterostichini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol." year="2017">Bousquet (2017)</bibRefCitation>
|
|
: 724;
|
|
<bibRefCitation id="D3FE0FAC5D74A198E0132972CDDA1BDA" DOI="https://doi.org/10.3390/insects11110803" author="Sasakawa, K" journalOrPublisher="Applied Entomology and Zoology" pageId="0" pageNumber="61" refId="B17" refString="Sasakawa, K, Mitsuduka, Y, Ito, H, 2020. Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803. https://doi.org/10.3390/insects11110803" title="Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803." url="https://doi.org/10.3390/insects11110803" year="2020">Sasakawa et al. (2020)</bibRefCitation>
|
|
: 7. Syn. nov.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="72096F1ED0BFA2B0D5B7AA786549A1FB" pageId="0" pageNumber="61" type="specimen examined">
|
|
<paragraph id="C4BB171E575DD5696609B2CFD4583485" pageId="0" pageNumber="61">Specimen examined.</paragraph>
|
|
<paragraph id="4AEBD80FC77B29C48B3D90869D728EEF" pageId="0" pageNumber="61">
|
|
1♂, Miomote, alt. 276 m, on the right bank of Miomotegawa River, Murakami-shi, Niigata Prefecture, Japan (
|
|
<geoCoordinate id="BCE1BEB97536BA98466CF9B540BE310C" degrees="38.273211" direction="north" orientation="latitude" precision="1" value="38.273212">38.273211°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="CAEF9F21A139E7E54FE23D1AFFE1E075" degrees="139.779922" direction="east" orientation="longitude" precision="1" value="139.77992">139.779922°E</geoCoordinate>
|
|
), 12.vi.-17.vii.2021,
|
|
<normalizedToken id="9C56742F357865B79F9E586B499B7B66" originalValue="Hirotarô">Hirotaro</normalizedToken>
|
|
<normalizedToken id="788573952A565B45C8A30F5FFB20CF21" originalValue="Itô">Ito</normalizedToken>
|
|
leg., in the collection of HI.
|
|
</paragraph>
|
|
<caption id="C530FDC105DF6C651F5F9D95AAE28BD4" doi="10.3897/subtbiol.43.80969.figure2" httpUri="https://binary.pensoft.net/fig/690341" pageId="0" pageNumber="61" start="Figure 2" startId="F2">
|
|
<paragraph id="63180431A8A196F322249253BF9C0E2F" pageId="0" pageNumber="61">
|
|
<emphasis id="77CC9E034167B20AFF189C63FE09F64D" bold="true" pageId="0" pageNumber="61">Figure 2.</emphasis>
|
|
<taxonomicName id="5E3F006BFC7C346B7841E94012D485D9" authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis id="CDCDF320E6B4F04B80CA42BDD6A4F8F5" italics="true" pageId="0" pageNumber="61">Pterostichus asahinus</emphasis>
|
|
</taxonomicName>
|
|
male from Miomote, on the right bank of the Miomotegawa River
|
|
<emphasis id="5A52D1363C0A9DE67567F41DF343BD4D" bold="true" pageId="0" pageNumber="61">A</emphasis>
|
|
habitus dorsal view
|
|
<emphasis id="5E31E372E1D28CB3DA3B334CBABA9446" bold="true" pageId="0" pageNumber="61">B-E</emphasis>
|
|
Endophallus left lateral (
|
|
<emphasis id="270EAFA9DE01BB2C46B12392296C65D7" bold="true" pageId="0" pageNumber="61">B</emphasis>
|
|
) ventral (
|
|
<emphasis id="0C9D21BCD388C6D44A8B0588BB8F199A" bold="true" pageId="0" pageNumber="61">C</emphasis>
|
|
) right lateral (
|
|
<emphasis id="CFC343AA6851C5F1FB3B34BCDA11917E" bold="true" pageId="0" pageNumber="61">D</emphasis>
|
|
) and dorsal (
|
|
<emphasis id="75ADEDA4467C6A6EE63BA27175E2FC5E" bold="true" pageId="0" pageNumber="61">E</emphasis>
|
|
) views
|
|
<emphasis id="8F6FE4635CC00D7544DDBB327FB09FEB" bold="true" pageId="0" pageNumber="61">F-H</emphasis>
|
|
right paramere left lateral (
|
|
<emphasis id="F039D909F18071805910AFDB3E5EC1C1" bold="true" pageId="0" pageNumber="61">F</emphasis>
|
|
) apical (
|
|
<emphasis id="594EFA5D434E3ED57403DAE2E17EB1C6" bold="true" pageId="0" pageNumber="61">G</emphasis>
|
|
) and dorsal (
|
|
<emphasis id="90384B4940F0FCCAA8C382EF30915A42" bold="true" pageId="0" pageNumber="61">H</emphasis>
|
|
) views.
|
|
<emphasis id="A8FFBEC35A379035961E8C21E2F42449" bold="true" pageId="0" pageNumber="61">go</emphasis>
|
|
, gonopore;
|
|
<emphasis id="0B4A2DA98A2215E40478C17FFF937155" bold="true" pageId="0" pageNumber="61">lal</emphasis>
|
|
, left apical lobe;
|
|
<emphasis id="DEB7972B8224460D4D62407D9EE06541" bold="true" pageId="0" pageNumber="61">lpl</emphasis>
|
|
, left preapical lobe;
|
|
<emphasis id="17D8C0C24D946BFEA40838978CAD3DF5" bold="true" pageId="0" pageNumber="61">rpl</emphasis>
|
|
, right preapical lobe.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="548F36E7DB8E63985EE1A15792D6006F" pageId="0" pageNumber="61" type="notes">
|
|
<paragraph id="13CBE3F09CD0FC405424C15CA8E9D513" pageId="0" pageNumber="61">Notes.</paragraph>
|
|
<paragraph id="99DAEED5CA60976833F46137E027074B" pageId="0" pageNumber="61">
|
|
<bibRefCitation id="FFCE57C2F8FED6D18C494D21A6A13EF5" DOI="https://doi.org/10.3390/insects11110803" author="Sasakawa, K" journalOrPublisher="Applied Entomology and Zoology" pageId="0" pageNumber="61" refId="B17" refString="Sasakawa, K, Mitsuduka, Y, Ito, H, 2020. Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803. https://doi.org/10.3390/insects11110803" title="Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803." url="https://doi.org/10.3390/insects11110803" year="2020">Sasakawa et al. (2020)</bibRefCitation>
|
|
suggested that
|
|
<taxonomicName id="1C2FC4793F2A71812B8971CE9FF7D068" class="Insecta" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis id="81A1D4B254BBB521CBFB58F1E73F342E" italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="E59ABAD79ACF52444610E86ED69B7692" class="Insecta" kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
|
<emphasis id="9B61B23B1F0D346B1546DBC0658B7086" italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
|
</taxonomicName>
|
|
might be conspecific. However, this hypothesis remained untested because males from the type locality of
|
|
<taxonomicName id="D71D3548982F3ED827F1F6EB18653CC6" class="Insecta" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis id="FAA19F3FFCB610ECD53CC5DBB2B6EF8E" italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
|
</taxonomicName>
|
|
have not been examined. The male specimen examined here was obtained from a valley southwest of Mt.
|
|
<normalizedToken id="E451D16D69926DB16ABA81ED288F9A37" originalValue="Dôrokujinpô">Dorokujinpo</normalizedToken>
|
|
, which can virtually be regarded as the type locality (Fig.
|
|
<figureCitation id="DCD2E592295600B4358C8033E282E551" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the Pterostichus macrogenys species group in northern Chubu, northern Kanto, and southern Tohoku Districts, Honshu, based on specimens with unambiguous identities, modified from Sasakawa et al. (2020) and Sato (2021) 1 P. chokaisanus Sasakawa, 2009 2 P. kurikomasanus Sasakawa, 2005 3 P. asahinus Habu and Baba, 1960 4 P. takadateyamanus Sasakawa, 2009 5 P. adatarasanus Sasakawa, 2005 6 P. gassanus Sasakawa, 2009 7 P. shinbodakensis Sasakawa & Ito, 2017 8 P. awashimaensis sp. nov. 9 P. iwakiensis Sasakawa, 2009 10 P. eboshiyamanus Sasakawa, 2009 11 P. tateishiyamanus Sasakawa & Ito, 2017 12 P. monolineatus Sasakawa, Mitsuduka, & Ito, 2020 13 P. yamizosanus Sasakawa, 2005 14 P. yahikosanus Sasakawa, 2009 15 P. ohsawacavus Sasakawa, 2005 16 P. sumondakensis Sasakawa, 2005 17 P. isolatus Sasakawa, 2005 18 P. nakamiyorinus Morita, Ohkawa & Kurihara, 2013 19 P. momuranus Morita, Ohkawa & Kurihara, 2013 20 P. macrogenys Bates, 1883 21 P. nagasawai Ito & Ogai, 2015. Red letters denote the type localities of each species. The blue letter indicates where the male specimen of P. asahinus newly examined here was collected." figureDoi="10.3897/subtbiol.43.80969.figure1" httpUri="https://binary.pensoft.net/fig/690340" pageId="0" pageNumber="61">1</figureCitation>
|
|
). The structures of the endophallus and right paramere of this specimen are identical to those of the eastern type of
|
|
<taxonomicName id="B41238298508BE887CDCA0CC00620A31" class="Insecta" kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
|
<emphasis id="19EB4568B64EF843840664D754FEE54E" italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
|
</taxonomicName>
|
|
in
|
|
<bibRefCitation id="5A9901DF8F95663E96AD7A7D6795A3AE" DOI="https://doi.org/10.3390/insects11110803" author="Sasakawa, K" journalOrPublisher="Applied Entomology and Zoology" pageId="0" pageNumber="61" refId="B17" refString="Sasakawa, K, Mitsuduka, Y, Ito, H, 2020. Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803. https://doi.org/10.3390/insects11110803" title="Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803." url="https://doi.org/10.3390/insects11110803" year="2020">Sasakawa et al. (2020)</bibRefCitation>
|
|
. Its body length is intermediate between that of the eastern and western types of
|
|
<taxonomicName id="15D598B8812832DC7A5E727EA11AAAAD" class="Insecta" kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
|
<emphasis id="EDAC239DA8C5136EAE651B4B2CF6D2C6" italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
|
</taxonomicName>
|
|
as follows: body length from mandible apices to elytral end (BLm) 16.32 mm, that from anterior margin of labrum (BLl) 14.72 mm, and that from clypeal apex (BLc) 14.26 mm. Based on these results, we regard
|
|
<taxonomicName id="EC582803158B1A838C851F1A47D732AB" class="Insecta" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis id="1C6CE692801B7ABFA746AD06387132A9" italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="2B5A579630DD1D4E3A2EB8B081008196" class="Insecta" kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
|
<emphasis id="B60EA8A39C94A3D547E21BAF4F6E4DA8" italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
|
</taxonomicName>
|
|
as conspecific. The results of the discriminant analysis support this conclusion. The
|
|
<taxonomicName id="EA142E96EBC1ECB51C9DF3B28626E5B2" class="Insecta" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis id="E2346FE6EC75BA6F3C67A344B86B2ADF" italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
|
</taxonomicName>
|
|
holotype was classified in this newly defined
|
|
<taxonomicName id="6A87BD3C14AA0F52EF4AA856309EAA78" class="Insecta" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis id="7CDBCADA82C88EF46CEA0D1CD279DF1B" italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
|
</taxonomicName>
|
|
in the discriminant function analysis, and was within the area of
|
|
<taxonomicName id="963053FF9C22785DDE3F0AEFA0656630" class="Insecta" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis id="DDDA5B5D2C6207B18282B7FFB0DE903C" italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
|
</taxonomicName>
|
|
on the scatterplot of the first two canonical variates (Fig.
|
|
<figureCitation id="E55AC29C51F45B2201611131D3C8D199" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scatterplot of the first two canonical variates obtained from the discriminant analysis of four measurements of the female pronotum." figureDoi="10.3897/subtbiol.43.80969.figure3" httpUri="https://binary.pensoft.net/fig/690342" pageId="0" pageNumber="61">3</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="72CF9B3270DD4E0DAE1C6A9E9F0EB1B6" doi="10.3897/subtbiol.43.80969.figure3" httpUri="https://binary.pensoft.net/fig/690342" pageId="0" pageNumber="61" start="Figure 3" startId="F3">
|
|
<paragraph id="5FC33BCDEE140833096E753CA012C09D" pageId="0" pageNumber="61">
|
|
<emphasis id="8B6983CF2932C65FECA19B9D7CDAF4C6" bold="true" pageId="0" pageNumber="61">Figure 3.</emphasis>
|
|
Scatterplot of the first two canonical variates obtained from the discriminant analysis of four measurements of the female pronotum.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |