335 lines
46 KiB
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335 lines
46 KiB
XML
<document ID-CLB-Dataset="9723" ID-DOI="http://dx.doi.org/10.3897/zookeys.976.53859" ID-GBIF-Dataset="3c8d8e79-b176-41e6-87c9-fb6f4b5ff9a6" ID-PMC="PMC7591553" ID-Pensoft-Pub="1313-2970-976-1" ID-Pensoft-UUID="1C1677B3CFE65ECEADF5CA56DACD0B9C" ID-PubMed="33173395" ID-ZooBank="B755A1D5D42D4CA589BE10C11EAB4580" ModsDocID="1313-2970-976-1" checkinTime="1603239875264" checkinUser="pensoft" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak, Tongkerd, Piyoros & Panha, Somsak" docDate="2020" docId="8C90A0D95A2A5543B265CA8E5F4CEC62" docLanguage="en" docName="ZooKeys 976: 1-100" docOrigin="ZooKeys 976" docSource="http://dx.doi.org/10.3897/zookeys.976.53859" docTitle="Sarika obesior" docType="treatment" docVersion="7" id="1C1677B3CFE65ECEADF5CA56DACD0B9C" lastPageNumber="1" masterDocId="1C1677B3CFE65ECEADF5CA56DACD0B9C" masterDocTitle="Integrative taxonomic revision of the land snail genus Sarika Godwin-Austen, 1907 in Thailand, with descriptions of nine new species (Eupulmonata, Ariophantidae)" masterLastPageNumber="100" masterPageNumber="1" pageNumber="1" updateTime="1732681712507" updateUser="ExternalLinkService">
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<mods:title id="40F919BB54F0A6091DD77FD055CB5CD7">Integrative taxonomic revision of the land snail genus Sarika Godwin-Austen, 1907 in Thailand, with descriptions of nine new species (Eupulmonata, Ariophantidae)</mods:title>
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<mods:namePart id="ABBC1DEC5A4A611BB87901081D84FABD">Pholyotha, Arthit</mods:namePart>
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<mods:affiliation id="2B1C9CE7B85562C30EF5EC5173F17710">Biological Sciences Program, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="89D307D1145583EBA07E682932DBBF07" type="orcid">https://orcid.org/0000-0001-6677-1164</mods:nameIdentifier>
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<mods:namePart id="19745746E1FA0C5EEC341CF1656B131B">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="63C7C479C1EF899555B5716889649FDF">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="F85A0D6ED10ADDEECB343B2A1AA17195" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="4A909B2DF4EF824A2CCAD5B5EFC07E94">Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation id="1A63E931549B17EBB5798E526AD08053">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="04A2D3CC5893A8161A8C6790F2A98527">Panha, Somsak</mods:namePart>
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<mods:affiliation id="34CC5D3B97BE3CDB18FE3A2F70741921">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:date id="98F4C30745C0B566D43142BA474BF4B8">2020</mods:date>
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<mods:number id="67479A122DE8DF3C7490D9D8C448038C">976</mods:number>
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<mods:classification id="D641D40C8C4BC68A549917C1FD68BAF4">journal article</mods:classification>
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<mods:identifier id="6236D10B4240BBBEE711C6A72C305405" type="DOI">http://dx.doi.org/10.3897/zookeys.976.53859</mods:identifier>
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<mods:identifier id="CC38DB8E8306010EE294487132BA409C" type="Pensoft-Pub">1313-2970-976-1</mods:identifier>
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<treatment id="8C90A0D95A2A5543B265CA8E5F4CEC62" ID-GBIF-Taxon="168504535" LSID="urn:lsid:plazi:treatment:8C90A0D95A2A5543B265CA8E5F4CEC62" httpUri="http://treatment.plazi.org/id/8C90A0D95A2A5543B265CA8E5F4CEC62" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Ariophantidae" scope_superOrder="Eupulmonata">
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<subSubSection id="B91E27B6F0B3290794D4EB49EFAC4757" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="ACF3FE3F597FD565193DB178A957B47A" pageId="0" pageNumber="1">
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<taxonomicName id="72E1F057A4CFA43ADEA2B8E31940CB6E" ID-CoL="6XNWL" LSID="8C90A0D9-5A2A-5543-B265-CA8E5F4CEC62" authority="(Martens, 1867)" baseAuthorityName="Martens" baseAuthorityYear="1867" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika obesior" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="obesior">Sarika obesior (Martens, 1867)</taxonomicName>
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<figureCitation id="D00BCCC5AB9747B3B561B771AD471568" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree showing the relationships among species of Sarika based on the mitochondrial COI gene sequences. Numbers by the nodes are the ML bootstrap values (left) and Bayesian posterior probabilities (right); shown only for the nodes supported by ML or BI (≥ 70 % and ≥ 0.95). Each clade colour refers to a genus." figureDoi="10.3897/zookeys.976.53859.figure1" httpUri="https://binary.pensoft.net/fig/464383" pageId="0" pageNumber="1">Figs 1</figureCitation>
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<figureCitation id="A7215CE52299D833A6D876A70CA58850" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic distribution of Sarika kawtaoensis, S. lactospira sp. nov., and S. obesior based on the specimens examined herein." figureDoi="10.3897/zookeys.976.53859.figure8" httpUri="https://binary.pensoft.net/fig/464390" pageId="0" pageNumber="1">, 8</figureCitation>
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<figureCitation id="CAAE0775CE84556178EF616C5C14750A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Living snails of group I: Sarika resplendens group. A Sarika resplendens specimen CUMZ 7843 showing left shell lobe lobes (lsl) B S. resplendens specimen CUMZ 7827 C mating pairs of S. resplendens CUMZ 7876 D S. dohrniana specimen CUMZ 7611 E S. obesior specimen CUMZ 7673 F S. limbata specimen CUMZ 7652. All not to scale." figureDoi="10.3897/zookeys.976.53859.figure9" httpUri="https://binary.pensoft.net/fig/464391" pageId="0" pageNumber="1">, 9E</figureCitation>
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<figureCitation id="57E68440C2036FD9470F0A95A9030747" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of group I: Sarika resplendens group. A, B Sarika resplendens A specimen NHMUK 1898.5.18.157 and B specimen CUMZ 7815 C, D S. dohrniana C syntype NHMUK 20160046 and D specimen CUMZ 7611. E, F S. obesior E specimen CUMZ 7675 and F specimen CUMZ 7673." figureDoi="10.3897/zookeys.976.53859.figure11" httpUri="https://binary.pensoft.net/fig/464393" pageId="0" pageNumber="1">, 11E, F</figureCitation>
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<figureCitation id="8D0FE9E3CE58378318CA056552611A81" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Genitalia. A, B Sarika dohrniana specimen CUMZ 7633 A general view of the genital system and B internal structure of the penis C, D Sarika obesior specimen CUMZ 7673 C general view of the genital system and D internal structure of penis. White arrowheads indicate the ends of the penes." figureDoi="10.3897/zookeys.976.53859.figure14" httpUri="https://binary.pensoft.net/fig/464396" pageId="0" pageNumber="1">, 14C, D</figureCitation>
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<figureCitation id="DAB5E889BAEC8079A1D9F27978F5A018" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Spermatophore of Sarika obesior specimen CUMZ 7678 A general view of the spermatophore B head filament C-E tail filament C three spines located close to the sperm sac D region with and without branching spines, and E branching spines on the tip region. Yellow arrowhead indicates the end of the spines from the tip." figureDoi="10.3897/zookeys.976.53859.figure16" httpUri="https://binary.pensoft.net/fig/464398" pageId="0" pageNumber="1">, 16</figureCitation>
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<figureCitation id="CC5140CF189B071C7179CD964439EE73" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Representative SEM images of the radula. A Sarika resplendens specimen CUMZ 7851 B S. dohrniana specimen CUMZ 7633 C S. obesior specimen CUMZ 7673 D S. limbata specimen CUMZ 7652 E S. heptagyra specimen CUMZ 7279. Central tooth indicated by ' C'; yellow colour indicates lateral teeth in the transition to marginal teeth." figureDoi="10.3897/zookeys.976.53859.figure30" httpUri="https://binary.pensoft.net/fig/464412" pageId="0" pageNumber="1">, 30C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="5B2472EAC9A5C42946A7F687883C652F" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="0D59603FD84369E5C1D5D17AAE349E78" pageId="0" pageNumber="1">
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<taxonomicName id="BAA8DFFA9F6053A75804455C7ECF7A0F" authorityName="Martens" authorityYear="1867" class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina (Orobia) resplendens subsp. var. var. obesior" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="resplendens" subGenus="Orobia" subSpecies="var." variety="obesior">Nanina (Orobia) resplendens var. obesior</taxonomicName>
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<bibRefCitation id="9D71F99DF79A2F6780F8DA3B4C7C207B" author="Martens von, E" journalOrPublisher="Unpublished Master's Thesis, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok" pageId="0" pageNumber="1" refId="B45" refString="Martens von, E, 1867. Die Preussische Expedition nach Ost-Asien. Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei, 477 pp." title="Die Preussische Expedition nach Ost-Asien. Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei, 477 pp." year="1867">Martens 1867</bibRefCitation>
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: 72, pl. 12, fig. 6. Type locality: "Um Petshaburi nicht selten; auch an der Ostseite des Golfs bei Bang-Pra beobachtet" [Petchaburi Province and Bang Phra, Si Racha District, Chonburi Province, Thailand].
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</paragraph>
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<paragraph id="EDB41BE7BD630B6698F41C48AA83CCCD" pageId="0" pageNumber="1">
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<taxonomicName id="D878AD89646C908CE2DB11CDD1C6199E" authorityName="Martens" authorityYear="1867" class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina (Macrochlamys) resplendens subsp. obesior" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="resplendens" subGenus="Macrochlamys" subSpecies="obesior">Nanina (Macrochlamys) resplendens obesior</taxonomicName>
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: Tryon 1886: 91, pl. 30, fig. 76.
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</paragraph>
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<paragraph id="80C689CC992D9FA3B23F4AB6849DA1A4" pageId="0" pageNumber="1">
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<taxonomicName id="860FE960EFC86AC243DE7ECBAE9BAF1B" baseAuthorityName="Martens" baseAuthorityYear="1867" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika obesior" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="obesior">Sarika obesior</taxonomicName>
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:
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<bibRefCitation id="06F5595CC2B51B705509BE1251688C72" author="Solem, A" journalOrPublisher="Spolia Zoologia Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B82" refString="Solem, A, 1966. Some non-marine mollusks from Thailand. Spolia Zoologia Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand." volume="24" year="1966">Solem 1966</bibRefCitation>
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: 36-38, fig. 5b, c;
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<bibRefCitation id="4FFF46EF893DAF39289023C3950C144C" author="Hemmen, J" journalOrPublisher="Schriften zur Malakozoologie aus dem Haus der Natur-Cismar" pageId="0" pageNumber="1" pagination="35 - 70" refId="B31" refString="Hemmen, J, Hemmen, C, 2001. Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie aus dem Haus der Natur-Cismar 18: 35 - 70" title="Aktualisierte Liste der terrestrischen Gastropoden Thailands." volume="18" year="2001">Hemmen and Hemmen 2001</bibRefCitation>
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: 45;
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<bibRefCitation id="C167A9E58D61FDAD96143A6834258EB6" author="Sutcharit, C" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="95 - 100" refId="B85" refString="Sutcharit, C, Panha, S, 2008. Taxonomic re-evaluation of Sarika diadema (Dall, 1897) and S. asamurai (Panha, 1997), two endemic land snails from Thailand (Pulmonata: Ariophantidae: Macrochlamydinae). The Raffles Bulletin of Zoology 56: 95 - 100" title="Taxonomic re-evaluation of Sarika diadema (Dall, 1897) and S. asamurai (Panha, 1997), two endemic land snails from Thailand (Pulmonata: Ariophantidae: Macrochlamydinae)." volume="56" year="2008">Sutcharit and Panha 2008</bibRefCitation>
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: 96;
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<bibRefCitation id="D385BCBF1321C797CD5B1DB7633B5C00" DOI="https://doi.org/10.1080/13235818.2020.1723041" author="Pholyotha, A" journalOrPublisher="Molluscan Research" pageId="0" pageNumber="1" pagination="183 - 204" refId="B69" refString="Pholyotha, A, Sutcharit, C, Tongkerd, P, Lin, A, Panha, S, 2020c. Taxonomic revision of the land snail genera Macrochlamys Gray, 1847 and Sarika Godwin-Austen 1907 (Eupulmonata: Ariophantidae) from Southeastern Myanmar, with descriptions of three new species. Molluscan Research 40: 183 - 204, DOI: https://doi.org/10.1080/13235818.2020.1723041" title="Taxonomic revision of the land snail genera Macrochlamys Gray, 1847 and Sarika Godwin-Austen 1907 (Eupulmonata: Ariophantidae) from Southeastern Myanmar, with descriptions of three new species." url="https://doi.org/10.1080/13235818.2020.1723041" volume="40" year="2020 c">Pholyotha et al. 2020c</bibRefCitation>
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: 18, 19, figs 2f, g, 7e, f, 10a, b, 13d-f.
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</paragraph>
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</subSubSection>
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<subSubSection id="63B7DFE2036F617A431D556E2E3A9F56" pageId="0" pageNumber="1" type="type material">
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<paragraph id="D3174A30A42337D1034A918C175774AF" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="41695546C80EC120BFC60A19491193A6" pageId="0" pageNumber="1">
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The type specimens could not be located and were probably missing from the Museum
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<normalizedToken id="364927D9939A32EE50652328AFCE49E8" originalValue="für">fuer</normalizedToken>
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Naturkunde (Berlin) collection (T. von Rintelen and C. Zorn, pers. comm., December 2018).
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</paragraph>
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</subSubSection>
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<subSubSection id="CB765BE838910D0D6D7C1F41985DF1A5" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="937DAF562B51F82CD4EA08897934121E" pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph id="E1633E928FD3AC3D36A69C5354F117B3" pageId="0" pageNumber="1">
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<emphasis id="D7CA28D6E2317CF6076C7B086748559A" bold="true" pageId="0" pageNumber="1">Myanmar.</emphasis>
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Forest on Kala Island, Myeik District, Tanintharyi Division,
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<geoCoordinate id="E7DACFBAD1A9AD6343306B8F10728C76" degrees="12" direction="north" minutes="25" orientation="latitude" precision="1" seconds="25.4" value="12.423722">12°25'25.4"N</geoCoordinate>
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,
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<geoCoordinate id="751E2505932041D49274F2153CC0CD9D" degrees="98" direction="east" minutes="29" orientation="longitude" precision="1" seconds="50.6" value="98.49738">98°29'50.6"E</geoCoordinate>
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: CUMZ 7140, 7141.
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<emphasis id="001303D9F75A446A85FD48ECAB177B1C" bold="true" pageId="0" pageNumber="1">Thailand-Western.</emphasis>
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Limestone outcrop in Kaeng Krachan, Kaeng Krachan, Phetchaburi,
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<geoCoordinate id="608D078836A0E72F30CCA5ADFB6ECFD9" degrees="12" direction="north" minutes="55" orientation="latitude" precision="1" seconds="01.0" value="12.9169445">12°55'01.0"N</geoCoordinate>
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,
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<geoCoordinate id="16807FCEDE0CA1331F1824E9F40A742D" degrees="99" direction="east" minutes="37" orientation="longitude" precision="1" seconds="48.0" value="99.630005">99°37'48.0"E</geoCoordinate>
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: CUMZ 7686. Wat Tham Rong, Ban Lat, Phetchaburi,
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<geoCoordinate id="13A349B80E437B20CADAA8D652E0AFFD" degrees="13" direction="north" minutes="01" orientation="latitude" precision="1" seconds="31.3" value="13.025361">13°01'31.3"N</geoCoordinate>
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,
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<geoCoordinate id="93876BF0AC60609DD3AA6AA7FE8A4A7E" degrees="99" direction="east" minutes="55" orientation="longitude" precision="1" seconds="07.9" value="99.91886">99°55'07.9"E</geoCoordinate>
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: CUMZ 7673 (Fig.
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<figureCitation id="0D1A2E6A53BFA2A19AB802D70EA5CC49" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of group I: Sarika resplendens group. A, B Sarika resplendens A specimen NHMUK 1898.5.18.157 and B specimen CUMZ 7815 C, D S. dohrniana C syntype NHMUK 20160046 and D specimen CUMZ 7611. E, F S. obesior E specimen CUMZ 7675 and F specimen CUMZ 7673." figureDoi="10.3897/zookeys.976.53859.figure11" httpUri="https://binary.pensoft.net/fig/464393" pageId="0" pageNumber="1">11F</figureCitation>
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), 7693. Wat Khao Krachiu, Tha Yang, Phetchaburi,
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<geoCoordinate id="A7E8977C655B3CBC547B6AE078C83FBC" degrees="12" direction="north" minutes="57" orientation="latitude" precision="1" seconds="42.9" value="12.961917">12°57'42.9"N</geoCoordinate>
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,
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|
<geoCoordinate id="63C41EB20B29F540C26CDB524D606E8E" degrees="99" direction="east" minutes="54" orientation="longitude" precision="1" seconds="48.8" value="99.91356">99°54'48.8"E</geoCoordinate>
|
|
: CUMZ 7670. Khao Tamo, Tha Yang, Phetchaburi,
|
|
<geoCoordinate id="5C80390CFBB5C86F3E09779DCEF4EAD1" degrees="12" direction="north" minutes="47" orientation="latitude" precision="1" seconds="49.0" value="12.796945">12°47'49.0"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="02ABB7A717454C0327212A032711499B" degrees="99" direction="east" minutes="44" orientation="longitude" precision="1" seconds="40.0" value="99.74444">99°44'40.0"E</geoCoordinate>
|
|
: CUMZ 7682. Wat Khiriwong, Khao Yoi, Phetchaburi,
|
|
<geoCoordinate id="3B26445E2532CC0ADEA874C56BED816F" degrees="13" direction="north" minutes="20" orientation="latitude" precision="1" seconds="02.2" value="13.333944">13°20'02.2"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="30C2C3D7C6FA10CAE53A3A5F8392848E" degrees="99" direction="east" minutes="45" orientation="longitude" precision="1" seconds="19.9" value="99.75553">99°45'19.9"E</geoCoordinate>
|
|
: CUMZ 7668. Wat Puang Malai, Khao Yoi, Phetchaburi,
|
|
<geoCoordinate id="8D9378F94321A2FFF046A940196B8FD9" degrees="13" direction="north" minutes="18" orientation="latitude" precision="1" seconds="40.4" value="13.311222">13°18'40.4"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="2A8D1687825C7DF422EE41F5D0714502" degrees="99" direction="east" minutes="47" orientation="longitude" precision="1" seconds="09.2" value="99.78589">99°47'09.2"E</geoCoordinate>
|
|
: CUMZ 7675 (Fig.
|
|
<figureCitation id="CF658865B3341761793BA674539A0253" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of group I: Sarika resplendens group. A, B Sarika resplendens A specimen NHMUK 1898.5.18.157 and B specimen CUMZ 7815 C, D S. dohrniana C syntype NHMUK 20160046 and D specimen CUMZ 7611. E, F S. obesior E specimen CUMZ 7675 and F specimen CUMZ 7673." figureDoi="10.3897/zookeys.976.53859.figure11" httpUri="https://binary.pensoft.net/fig/464393" pageId="0" pageNumber="1">11E</figureCitation>
|
|
). Khao Nang Panthurat, Cha-am, Phetchaburi,
|
|
<geoCoordinate id="CB2901F692F9BF65531628CE296E0AF7" degrees="12" direction="north" minutes="50" orientation="latitude" precision="1" seconds="22.2" value="12.839499">12°50'22.2"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="AA29A8A7E78E4D1E264F36784A12DBB6" degrees="99" direction="east" minutes="57" orientation="longitude" precision="1" seconds="10.0" value="99.952774">99°57'10.0"E</geoCoordinate>
|
|
: CUMZ 7680, 7681, 7694, 7695. Khao Na Kwang Cave Cha-am, Phetchaburi,
|
|
<geoCoordinate id="73C0EF1E4F5798BEDD78851AEEDB9AFF" degrees="12" direction="north" minutes="51" orientation="latitude" precision="1" seconds="31.3" value="12.858695">12°51'31.3"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="C89D6FE557CEA22CD95189E656F74D97" degrees="99" direction="east" minutes="56" orientation="longitude" precision="1" seconds="29.3" value="99.941475">99°56'29.3"E</geoCoordinate>
|
|
: CUMZ 7683, 7690. Wat Tham Jaeng, Cha-am, Phetchaburi,
|
|
<geoCoordinate id="8CB8762E606019813D5A2533E6B50D0F" degrees="12" direction="north" minutes="49" orientation="latitude" precision="1" seconds="39.3" value="12.827583">12°49'39.3"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="13DC5E632A5C099799BD758E9224603E" degrees="99" direction="east" minutes="56" orientation="longitude" precision="1" seconds="24.5" value="99.94014">99°56'24.5"E</geoCoordinate>
|
|
: CUMZ 7667. Khao Ma Rong Cave, Bang Saphan, Prachuap Khiri Khan,
|
|
<geoCoordinate id="B5354D3F186BC469300BA45FDEECB4DE" degrees="11" direction="north" minutes="12" orientation="latitude" precision="1" seconds="09.3" value="11.202583">11°12'09.3"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="6E7FFC11A235E2FD3A2454FB71961303" degrees="99" direction="east" minutes="29" orientation="longitude" precision="1" seconds="39.8" value="99.494385">99°29'39.8"E</geoCoordinate>
|
|
: CUMZ 7664, 7669, 7702. Wat Tham Siriwong, Bang Saphan, Prachuap Khiri Khan,
|
|
<geoCoordinate id="068CAF52BA080A889583D95CA7BAC75F" degrees="11" direction="north" minutes="23" orientation="latitude" precision="1" seconds="27.6" value="11.391">11°23'27.6"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="8E344AAA039DD0F658C0D1C3B5CEAD80" degrees="99" direction="east" minutes="34" orientation="longitude" precision="1" seconds="55.3" value="99.58202">99°34'55.3"E</geoCoordinate>
|
|
: CUMZ 7665, 7697, 7704. Wat Tham Khao Wong, Bang Saphan, Prachuap Khiri Khan,
|
|
<geoCoordinate id="35F4DC03C4538B1C241C67011C97F777" degrees="11" direction="north" minutes="17" orientation="latitude" precision="1" seconds="28.0" value="11.291112">11°17'28.0"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="2DD8E558667F072A7146E1380F69CCAA" degrees="99" direction="east" minutes="29" orientation="longitude" precision="1" seconds="45.4" value="99.49594">99°29'45.4"E</geoCoordinate>
|
|
: CUMZ 7666. Ko Thalu, Bang Saphan Noi, Prachuap Khiri Khan,
|
|
<geoCoordinate id="AC57235642A1362CC1C5F74091C0DDAD" degrees="11" direction="north" minutes="04" orientation="latitude" precision="1" seconds="34.4" value="11.076222">11°04'34.4"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="48518CF722A58FB14EB0D505A7D91765" degrees="99" direction="east" minutes="33" orientation="longitude" precision="1" seconds="37.4" value="99.560394">99°33'37.4"E</geoCoordinate>
|
|
: CUMZ 7691. Khao Ta Mong Lai, Mueang, Prachuap Khiri Khan,
|
|
<geoCoordinate id="8962CEB69C7E8AE5C7A18193F358F3E2" degrees="11" direction="north" minutes="50" orientation="latitude" precision="1" seconds="00.3" value="11.833416">11°50'00.3"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="F1149204F2BFE0B2C1FFFFA27A26839B" degrees="99" direction="east" minutes="49" orientation="longitude" precision="1" seconds="47.2" value="99.82977">99°49'47.2"E</geoCoordinate>
|
|
: CUMZ 7674. Khao Lom Muak, Mueang, Prachuap Khiri Khan,
|
|
<geoCoordinate id="C6CC7810A3016ABCB5233DC723102E0E" degrees="11" direction="north" minutes="47" orientation="latitude" precision="1" seconds="08.5" value="11.785695">11°47'08.5"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="9F62CD252B3E5B47C2E5ADB6A8CA54DD" degrees="99" direction="east" minutes="48" orientation="longitude" precision="1" seconds="57.9" value="99.816086">99°48'57.9"E</geoCoordinate>
|
|
: CUMZ 7676. Wat Ao Noi, Mueang, Prachuap Khiri Khan,
|
|
<geoCoordinate id="F0C8A00FD86F3A581795242B617078FE" degrees="11" direction="north" minutes="51" orientation="latitude" precision="1" seconds="44.0" value="11.862223">11°51'44.0"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="01361760A20A8CA75F70755E3D9D3D83" degrees="99" direction="east" minutes="49" orientation="longitude" precision="1" seconds="20.0" value="99.82222">99°49'20.0"E</geoCoordinate>
|
|
: CUMZ 7701, 7703. Tham Khao Rak Mai, Thap Sakae, Prachuap Khiri Khan,
|
|
<geoCoordinate id="28B194832F3D4936AB1A1C2BCFADA819" degrees="11" direction="north" minutes="25" orientation="latitude" precision="1" seconds="42.1" value="11.428362">11°25'42.1"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="AAED72EE24024A911FA21C4C00D84B4E" degrees="99" direction="east" minutes="36" orientation="longitude" precision="1" seconds="18.5" value="99.60514">99°36'18.5"E</geoCoordinate>
|
|
: CUMZ 7677. Khao Kalok, Sam Roi Yot, Prachuap Khiri Khan,
|
|
<geoCoordinate id="66E6F7EABD58B91FFA203A4257189B26" degrees="12" direction="north" minutes="20" orientation="latitude" precision="1" seconds="13.0" value="12.336945">12°20'13.0"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="389C55B9955503FEC1FB95D9542C889B" degrees="99" direction="east" minutes="59" orientation="longitude" precision="1" seconds="59.0" value="99.99972">99°59'59.0"E</geoCoordinate>
|
|
: CUMZ 7678, 7698. Phraya Nakhon Cave, Sam Roi Yot, Prachuap Khiri Khan,
|
|
<geoCoordinate id="30E07B1B20CA5E550049AE5D8318A519" degrees="12" direction="north" minutes="11" orientation="latitude" precision="1" seconds="45.3" value="12.195917">12°11'45.3"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="B61CC67419B3DF1475FFD9F13EEB7B06" degrees="100" direction="east" minutes="00" orientation="longitude" precision="1" seconds="41.8" value="100.01161">100°00'41.8"E</geoCoordinate>
|
|
: CUMZ 7685. Limestone outcrop near Hat Sam Phraya, Kui Buri, Prachuap Khiri Khan,
|
|
<geoCoordinate id="38759A1392B2518C95C543981AC27CE1" degrees="12" direction="north" minutes="09" orientation="latitude" precision="1" seconds="04.0" value="12.151111">12°09'04.0"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="62CB3D61E1487D60F9AA95E51842E8A1" degrees="99" direction="east" minutes="58" orientation="longitude" precision="1" seconds="58.0" value="99.98278">99°58'58.0"E</geoCoordinate>
|
|
: CUMZ 7705. Limestone outcrop in Kui Buri, Kui Buri, Prachuap Khiri Khan,
|
|
<geoCoordinate id="99576CF0E58E2CA6A81F92648700413F" degrees="12" direction="north" minutes="08" orientation="latitude" precision="1" seconds="06.1" value="12.135028">12°08'06.1"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="260279BA77987F523ED7F8267C928CAB" degrees="99" direction="east" minutes="38" orientation="longitude" precision="1" seconds="41.2" value="99.644775">99°38'41.2"E</geoCoordinate>
|
|
: CUMZ 7684, 7692, 7699. Wat Nong Phlap, Hua Hin, Prachuap Khiri Khan,
|
|
<geoCoordinate id="A9BADE03EBD92B2E85864613524C5897" degrees="12" direction="north" minutes="35" orientation="latitude" precision="1" seconds="29.9" value="12.591639">12°35'29.9"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="493BBCB2B11F1FFD7DA017DD7AD1A4C5" degrees="99" direction="east" minutes="43" orientation="longitude" precision="1" seconds="45.8" value="99.72939">99°43'45.8"E</geoCoordinate>
|
|
: CUMZ 7233. Pa La-U Waterfall, Hua Hin, Prachuap Khiri Khan,
|
|
<geoCoordinate id="933FD60C03FFA79BBF8016C04F0235D3" degrees="12" direction="north" minutes="32" orientation="latitude" precision="1" seconds="17.2" value="12.538112">12°32'17.2"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="4CEE1B2C94D5821ED8ACB96927027FB3" degrees="99" direction="east" minutes="27" orientation="longitude" precision="1" seconds="48.7" value="99.463524">99°27'48.7"E</geoCoordinate>
|
|
: CUMZ 7700. Lublae Cave, Hua Hin, Prachuap Khiri Khan,
|
|
<geoCoordinate id="29052FA0C6A4AE5C6A1458A631404B3E" degrees="12" direction="north" minutes="36" orientation="latitude" precision="1" seconds="15.1" value="12.604195">12°36'15.1"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="ED20D70ADE7C8760F87A70C6FEC9769B" degrees="99" direction="east" minutes="43" orientation="longitude" precision="1" seconds="20.9" value="99.72247">99°43'20.9"E</geoCoordinate>
|
|
: CUMZ 7679.
|
|
<emphasis id="82C100F0A927C28CE5E9A86C11BBAD59" bold="true" pageId="0" pageNumber="1">Thailand-Southern.</emphasis>
|
|
Wat Bonphot Phisai, Lang Suan, Chumphon,
|
|
<geoCoordinate id="AFF782A7143D9F9ABC7041CDA961B468" degrees="9" direction="north" minutes="56" orientation="latitude" precision="1" seconds="12.5" value="9.936806">9°56'12.5"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="AAAAF121F6C64AEC429462D1CD09F9CF" degrees="99" direction="east" minutes="08" orientation="longitude" precision="1" seconds="44.2" value="99.14561">99°08'44.2"E</geoCoordinate>
|
|
: CUMZ 7671. Limestone outcrop in Khlong Sok, Phanom, Surat Thani,
|
|
<geoCoordinate id="8E461907FF0118CF57F4472E4C08600F" degrees="8" direction="north" minutes="50" orientation="latitude" precision="1" seconds="53.0" value="8.848055">8°50'53.0"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="6E3064D57E4B983B91C12673F9999A11" degrees="98" direction="east" minutes="44" orientation="longitude" precision="1" seconds="35.3" value="98.74313">98°44'35.3"E</geoCoordinate>
|
|
: CUMZ 7688. Limestone outcrop near Anurak Community Lodge, Phanom, Surat Thani,
|
|
<geoCoordinate id="171F9D6E4E2A9F12BAC03224D4C1C30B" degrees="8" direction="north" minutes="53" orientation="latitude" precision="1" seconds="16.3" value="8.887861">8°53'16.3"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="19CDBE58C858F726942386B3720E3763" degrees="98" direction="east" minutes="40" orientation="longitude" precision="1" seconds="45.2" value="98.67922">98°40'45.2"E</geoCoordinate>
|
|
: CUMZ 7689. Wat Tham Wararam, Phanom, Surat Thani: CUMZ 7696. Khao Kloi, Don Sak, Surat Thani,
|
|
<geoCoordinate id="E7905544978DC330A2ACCF5790305D3B" degrees="8" direction="north" minutes="52" orientation="latitude" precision="1" seconds="43.9" value="8.878861">8°52'43.9"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="1E75FC8BE2922642345026A4989EBE5B" degrees="98" direction="east" minutes="39" orientation="longitude" precision="1" seconds="26.5" value="98.657364">98°39'26.5"E</geoCoordinate>
|
|
: CUMZ 7672.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="6E8A67AE2342866C6FBBD732A516A261" pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph id="3A9303FB66FCA91799D5C58DB6546EB7" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph id="786D6B5A1021BB860BE5FBAB25475402" pageId="0" pageNumber="1">Shell medium to large, depressed and well rounded body whorl. Animal with greyish to slightly dark grey body and five mantle lobes. Genitalia with straight epiphallic caecum and triangular prism penial pilasters. Spermatophore: head filament with irregularly obtuse-serrate longitudinal ridges; tail filament near sperm sac with three spines and terminal part more than ca. one-third of its length with series of several branching spines.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="7DA44E533B7A3AC66B0F9AB5BD5BA162" pageId="0" pageNumber="1" type="description">
|
|
<paragraph id="3BDAB3643705DEA9F0B3AFB3AC1368CA" pageId="0" pageNumber="1">Description.</paragraph>
|
|
<paragraph id="325C43F421E2DF98CD41BD28B2010C62" pageId="0" pageNumber="1">
|
|
The unique shell characters of
|
|
<taxonomicName id="CD381488DE8040FDA87DFFE70F46DAB5" lsidName="S. obesior" pageId="0" pageNumber="1" rank="species" species="obesior">
|
|
<emphasis id="97141BCC5E60C8C3DE4664A2E860A8F3" italics="true" pageId="0" pageNumber="1">S. obesior</emphasis>
|
|
</taxonomicName>
|
|
are depressed, medium to large size (shell width up to 22.1 mm; shell height up to 11.3 mm), pale brown, well-rounded body whorl, spire elevated, and impressed suture (Fig.
|
|
<figureCitation id="6BF96C4AA50B422001E5D341D5F2D5A9" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of group I: Sarika resplendens group. A, B Sarika resplendens A specimen NHMUK 1898.5.18.157 and B specimen CUMZ 7815 C, D S. dohrniana C syntype NHMUK 20160046 and D specimen CUMZ 7611. E, F S. obesior E specimen CUMZ 7675 and F specimen CUMZ 7673." figureDoi="10.3897/zookeys.976.53859.figure11" httpUri="https://binary.pensoft.net/fig/464393" pageId="0" pageNumber="1">11E, F</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="089D23E2BD7577D51EDC962DA8B92BAC" pageId="0" pageNumber="1">
|
|
The unique genitalia characters are straight epiphallic caecum; inner wall of penis with very fine longitudinal penial pilasters near atrium, changing to large rhomboid pilasters with acute angle on top (triangular prism shape) (Fig.
|
|
<figureCitation id="2160946634823F16032E38FBA310A944" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Genitalia. A, B Sarika dohrniana specimen CUMZ 7633 A general view of the genital system and B internal structure of the penis C, D Sarika obesior specimen CUMZ 7673 C general view of the genital system and D internal structure of penis. White arrowheads indicate the ends of the penes." figureDoi="10.3897/zookeys.976.53859.figure14" httpUri="https://binary.pensoft.net/fig/464396" pageId="0" pageNumber="1">14C, D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="EFCF40C152A017965BFB1A364ADEF1DE" pageId="0" pageNumber="1">
|
|
Spermatophore long and needle-shaped. Sperm sac (ss) enlarged and elliptical. Head filament gourd shape with irregularly obtuse-serrate longitudinal ridges. Tail filament very long tube; region near sperm sac with three spines. Spine I located near base of spine II, simple and short. Spine II broken. Spine III large with complicated branching into small and many spinules. Region furthest away smooth and without spine; terminal part (more than ca. one-third of its length) with series of short to long branching spines arranged in a row or encircling the tail filament tip (Fig.
|
|
<figureCitation id="2BCC3965F0A1FCF5BD23AF881DFF6415" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Spermatophore of Sarika obesior specimen CUMZ 7678 A general view of the spermatophore B head filament C-E tail filament C three spines located close to the sperm sac D region with and without branching spines, and E branching spines on the tip region. Yellow arrowhead indicates the end of the spines from the tip." figureDoi="10.3897/zookeys.976.53859.figure16" httpUri="https://binary.pensoft.net/fig/464398" pageId="0" pageNumber="1">16</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="882F9C1D2AA1468E72C0EC8C3A81A3FA" pageId="0" pageNumber="1">
|
|
<taxonomicName id="A404F6B560CFDE0F112C5B2A8F8F8015" class="Bivalvia" family="Limidae" genus="Radula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radula" order="Limida" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Radula</taxonomicName>
|
|
with half row formula: 1-(13-14)-60. The morphology of central tooth, lateral, and marginal teeth are similar to that described in
|
|
<bibRefCitation id="0FD23CCEA4B04F35278BA4CCF07A3FDF" DOI="https://doi.org/10.1080/13235818.2020.1723041" author="Pholyotha, A" journalOrPublisher="Molluscan Research" pageId="0" pageNumber="1" pagination="183 - 204" refId="B69" refString="Pholyotha, A, Sutcharit, C, Tongkerd, P, Lin, A, Panha, S, 2020c. Taxonomic revision of the land snail genera Macrochlamys Gray, 1847 and Sarika Godwin-Austen 1907 (Eupulmonata: Ariophantidae) from Southeastern Myanmar, with descriptions of three new species. Molluscan Research 40: 183 - 204, DOI: https://doi.org/10.1080/13235818.2020.1723041" title="Taxonomic revision of the land snail genera Macrochlamys Gray, 1847 and Sarika Godwin-Austen 1907 (Eupulmonata: Ariophantidae) from Southeastern Myanmar, with descriptions of three new species." url="https://doi.org/10.1080/13235818.2020.1723041" volume="40" year="2020 c">Pholyotha et al. (2020c</bibRefCitation>
|
|
: figs 13d-f). Marginal teeth starting at approximately row number 13 or 14 (Fig.
|
|
<figureCitation id="AFFC4F5DC23BE4BA682BE6AB27A4F40C" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Representative SEM images of the radula. A Sarika resplendens specimen CUMZ 7851 B S. dohrniana specimen CUMZ 7633 C S. obesior specimen CUMZ 7673 D S. limbata specimen CUMZ 7652 E S. heptagyra specimen CUMZ 7279. Central tooth indicated by ' C'; yellow colour indicates lateral teeth in the transition to marginal teeth." figureDoi="10.3897/zookeys.976.53859.figure30" httpUri="https://binary.pensoft.net/fig/464412" pageId="0" pageNumber="1">30C</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="745E38A86D7C7D652CE3A5162D7004D6" pageId="0" pageNumber="1">
|
|
Living snail with monochrome greyish to slightly dark grey body. Mantle edge well developed and same colour same body (Fig.
|
|
<figureCitation id="9BA61FF6561C6F44396688B95A1F7013" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Living snails of group I: Sarika resplendens group. A Sarika resplendens specimen CUMZ 7843 showing left shell lobe lobes (lsl) B S. resplendens specimen CUMZ 7827 C mating pairs of S. resplendens CUMZ 7876 D S. dohrniana specimen CUMZ 7611 E S. obesior specimen CUMZ 7673 F S. limbata specimen CUMZ 7652. All not to scale." figureDoi="10.3897/zookeys.976.53859.figure9" httpUri="https://binary.pensoft.net/fig/464391" pageId="0" pageNumber="1">9E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="EF37CDDD3F2F18C0CDC0A856DC48C2D7" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="A85050F37EFF7BD859B635B9099FD5BD" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="EB1594679687FDBED507CC52CF3AFA66" pageId="0" pageNumber="1">
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<taxonomicName id="454F0B0BD0D9DDB39262F9053335E268" baseAuthorityName="Martens" baseAuthorityYear="1867" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika obesior" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="obesior">
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<emphasis id="C836EA748775319FAFE4417E3BBCDC36" italics="true" pageId="0" pageNumber="1">Sarika obesior</emphasis>
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</taxonomicName>
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occurs in western and southern Thailand (Fig.
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<figureCitation id="C0DD5B63F7148240AECF5FC65F552AE5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic distribution of Sarika kawtaoensis, S. lactospira sp. nov., and S. obesior based on the specimens examined herein." figureDoi="10.3897/zookeys.976.53859.figure8" httpUri="https://binary.pensoft.net/fig/464390" pageId="0" pageNumber="1">8</figureCitation>
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) and Tanintharyi region in Myanmar (
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<bibRefCitation id="24E7EC7CAA8871BA4BFE97F5F2F0FE08" DOI="https://doi.org/10.1080/13235818.2020.1723041" author="Pholyotha, A" journalOrPublisher="Molluscan Research" pageId="0" pageNumber="1" pagination="183 - 204" refId="B69" refString="Pholyotha, A, Sutcharit, C, Tongkerd, P, Lin, A, Panha, S, 2020c. Taxonomic revision of the land snail genera Macrochlamys Gray, 1847 and Sarika Godwin-Austen 1907 (Eupulmonata: Ariophantidae) from Southeastern Myanmar, with descriptions of three new species. Molluscan Research 40: 183 - 204, DOI: https://doi.org/10.1080/13235818.2020.1723041" title="Taxonomic revision of the land snail genera Macrochlamys Gray, 1847 and Sarika Godwin-Austen 1907 (Eupulmonata: Ariophantidae) from Southeastern Myanmar, with descriptions of three new species." url="https://doi.org/10.1080/13235818.2020.1723041" volume="40" year="2020 c">Pholyotha et al. 2020c</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="AF2969CF7A4594D3E28A9F433EB71FAD" pageId="0" pageNumber="1" type="coi analysis">
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<paragraph id="80F3114A2B9A46BDAC7658055E0933E2" pageId="0" pageNumber="1">COI analysis.</paragraph>
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<paragraph id="5C2A47A182FC3ED943E8467B02DA1527" pageId="0" pageNumber="1">
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The ML and BI analyses showed that the individuals of
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<taxonomicName id="16507064E655FF56667ABDAA81216702" lsidName="S. obesior" pageId="0" pageNumber="1" rank="species" species="obesior">
|
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<emphasis id="988CBF7DBF8AB4669FF1BA1CB307CEAC" italics="true" pageId="0" pageNumber="1">S. obesior</emphasis>
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</taxonomicName>
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(n = 4) formed a monophyletic group with high support (Fig.
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<figureCitation id="EFD58B033D03DE7A9A59B2ED17DCF73B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree showing the relationships among species of Sarika based on the mitochondrial COI gene sequences. Numbers by the nodes are the ML bootstrap values (left) and Bayesian posterior probabilities (right); shown only for the nodes supported by ML or BI (≥ 70 % and ≥ 0.95). Each clade colour refers to a genus." figureDoi="10.3897/zookeys.976.53859.figure1" httpUri="https://binary.pensoft.net/fig/464383" pageId="0" pageNumber="1">1</figureCitation>
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; BS = 98%, PP = 1). The mean intraspecific genetic distance of
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<taxonomicName id="99E49D463899F579022F7DD9894B7D87" lsidName="S. obesior" pageId="0" pageNumber="1" rank="species" species="obesior">
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<emphasis id="4B560073CEE14E3B392B599EA2721686" italics="true" pageId="0" pageNumber="1">S. obesior</emphasis>
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</taxonomicName>
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|
was 3.0% (Table
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<tableCitation id="62CD0B855B558F1958A628271AD99CA0" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Mean intra-specific and inter-specific genetic divergences among species of Sarika from the mitochondrial COI gene sequences estimated by the K 2 P model. Taxa in bold are the new species described herein." httpUri="http://table.plazi.org/id/13BDBDF714F2FAA17ADB274D839659A1" pageId="0" pageNumber="1" tableUuid="13BDBDF714F2FAA17ADB274D839659A1">2</tableCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="488B2F4219AD0765C315812EF469BCCC" pageId="0" pageNumber="1" type="remarks">
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<paragraph id="8E517D0178630DF4BD4A11EFA7091FAF" pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph id="2BA281E5461150D56F29DD9EF65A3706" pageId="0" pageNumber="1">
|
|
This species has recently been re-described and illustrated based on the samples collected from Myeik, Myanmar by
|
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<bibRefCitation id="19C2D9DE06410FD9C5880E23BB52BA50" DOI="https://doi.org/10.1080/13235818.2020.1723041" author="Pholyotha, A" journalOrPublisher="Molluscan Research" pageId="0" pageNumber="1" pagination="183 - 204" refId="B69" refString="Pholyotha, A, Sutcharit, C, Tongkerd, P, Lin, A, Panha, S, 2020c. Taxonomic revision of the land snail genera Macrochlamys Gray, 1847 and Sarika Godwin-Austen 1907 (Eupulmonata: Ariophantidae) from Southeastern Myanmar, with descriptions of three new species. Molluscan Research 40: 183 - 204, DOI: https://doi.org/10.1080/13235818.2020.1723041" title="Taxonomic revision of the land snail genera Macrochlamys Gray, 1847 and Sarika Godwin-Austen 1907 (Eupulmonata: Ariophantidae) from Southeastern Myanmar, with descriptions of three new species." url="https://doi.org/10.1080/13235818.2020.1723041" volume="40" year="2020 c">Pholyotha et al. (2020c)</bibRefCitation>
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. The topotypic specimens from Petchaburi, Thailand agree well with the previous descriptions.
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<bibRefCitation id="9B22650793FDB23442121851C98F98E6" author="Solem, A" journalOrPublisher="Spolia Zoologia Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B82" refString="Solem, A, 1966. Some non-marine mollusks from Thailand. Spolia Zoologia Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand." volume="24" year="1966">Solem (1966)</bibRefCitation>
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noted that
|
|
<taxonomicName id="B622A21A852C6E8910B82C8B49C507C2" lsidName="S. obesior" pageId="0" pageNumber="1" rank="species" species="obesior">
|
|
<emphasis id="DB6A7413FE5E95F864556802B10408FE" italics="true" pageId="0" pageNumber="1">S. obesior</emphasis>
|
|
</taxonomicName>
|
|
differs from
|
|
<taxonomicName id="85D4CC8E357DB02141DFB0827E682B46" lsidName="S. hainesi" pageId="0" pageNumber="1" rank="species" species="hainesi">
|
|
<emphasis id="4B41EE77C3EC7A01AF33BE11BB857DE0" italics="true" pageId="0" pageNumber="1">S. hainesi</emphasis>
|
|
</taxonomicName>
|
|
in having a rounded body whorl and attributed several specimens from northern and eastern Thailand to
|
|
<taxonomicName id="31DA672DC2826734CE259FE9647C4FD9" lsidName="S. obesior" pageId="0" pageNumber="1" rank="species" species="obesior">
|
|
<emphasis id="18FAB81AC1AB867B88DBB0B3FD249FAC" italics="true" pageId="0" pageNumber="1">S. obesior</emphasis>
|
|
</taxonomicName>
|
|
. Yet, without the genital anatomy their identification could not be confirmed. Therefore, the specimens from northern and eastern Thailand as reported by
|
|
<bibRefCitation id="4D77032B29D56C86354B2D48F8760DA4" author="Solem, A" journalOrPublisher="Spolia Zoologia Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B82" refString="Solem, A, 1966. Some non-marine mollusks from Thailand. Spolia Zoologia Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand." volume="24" year="1966">Solem (1966)</bibRefCitation>
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are still doubtful, and to date, the distribution of
|
|
<taxonomicName id="B01432C4C98D2F748ACF70F8E72A06D7" lsidName="S. obesior" pageId="0" pageNumber="1" rank="species" species="obesior">
|
|
<emphasis id="D71B099FB6A9F376F84BBC7073F0A55A" italics="true" pageId="0" pageNumber="1">S. obesior</emphasis>
|
|
</taxonomicName>
|
|
is probably restricted to western and southern Thailand and south-eastern Myanmar.
|
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |