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<document id="F2C6DF2F518291964B65D88DC14FDDB8" ID="10.11646/zootaxa.4273.4.3SLASH11267" ID-CLB-Dataset="32891" ID-DOI="10.11646/zootaxa.4273.4.3" ID-GBIF-Dataset="0935deaf-8f18-434e-b4a0-0468cefc9577" ID-ISSN="1175-5326" ID-Zenodo-Dep="803704" ID-ZooBank="4CD96097-1016-47CB-8DC3-AD6E9EDA330C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1496820946147" checkinUser="plazi" docAuthor="Enghoff, Henrik" docDate="2017" docId="8C70BE3B2441196CFF676AFEAAA4FC53" docLanguage="en" docName="zootaxa.4273.4.3SLASH11267.pdf" docOrigin="Zootaxa 4273 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tropostreptus austerus Attems 1950, comb. nov." docType="treatment" docVersion="7" lastPageNumber="522" masterDocId="7049C64324551979FFF06F72AF21FF9B" masterDocTitle="A new East African genus of spirostreptid millipedes (Diplopoda, Spirostreptida, Spirostreptidae), with notes on their fungal ectoparasite Rickia gigas" masterLastPageNumber="530" masterPageNumber="501" pageNumber="521" updateTime="1698440246528" updateUser="ExternalLinkService">
<mods:mods id="FF317DA15CE472C2B0E0B72EB7B43A6C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0B015F1E84213720B98B71975C14CC1A">
<mods:title id="B2F6B9502F0B2E39F62F0F93AE646603">A new East African genus of spirostreptid millipedes (Diplopoda, Spirostreptida, Spirostreptidae), with notes on their fungal ectoparasite Rickia gigas</mods:title>
</mods:titleInfo>
<mods:name id="0D390891FF48E2E0DB1BB7CB4C248F5A" type="personal">
<mods:role id="5F9CC5A2C65A1D4F29BD0E4AF70BE740">
<mods:roleTerm id="CF21E32EC30F6AFE92BBED35C398FE94">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5C9CAFD2F17DFC4AAFCE155CA6D29AB2">Enghoff, Henrik</mods:namePart>
</mods:name>
<mods:typeOfResource id="9DC5FC2B9759EB7809E5E4E02711D452">text</mods:typeOfResource>
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<mods:title id="8A109A66B37143583D54E0FEA84B2108">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1669929B1C02C616A3770361094AF97C">
<mods:date id="DBE7843D255880FE80A60D421DFD4DF1">2017</mods:date>
<mods:detail id="9154C73808B9BB255A5CF5A0D07F6DC9" type="volume">
<mods:number id="0DB79D2ECCBBA24CFD60D88775EEB5AC">4273</mods:number>
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<mods:number id="61EF667D5A78D45B8A9468405A4BAD66">4</mods:number>
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<mods:start id="FEF1C3F2267CC1127269A8A2909EE3CF">501</mods:start>
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<mods:classification id="C2CF99454680F39E1DE0FBDF063167EA">journal article</mods:classification>
<mods:identifier id="59733386AFED6634C41B41A1CE8BCA13" type="CLB-Dataset">32891</mods:identifier>
<mods:identifier id="B888C4AB0BE77F32765565CB11EA436F" type="DOI">10.11646/zootaxa.4273.4.3</mods:identifier>
<mods:identifier id="68D83996156BD32B8799E802E09892FB" type="GBIF-Dataset">0935deaf-8f18-434e-b4a0-0468cefc9577</mods:identifier>
<mods:identifier id="97C62BAAF9C32528F5CE39000C77B5C6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="776B71F0846367692FF5B4EE75BC4142" type="Zenodo-Dep">803704</mods:identifier>
<mods:identifier id="F827CF0E83FE776A9F078A719FEE5C8D" type="ZooBank">4CD96097-1016-47CB-8DC3-AD6E9EDA330C</mods:identifier>
</mods:mods>
<treatment id="8C70BE3B2441196CFF676AFEAAA4FC53" ID-DOI="http://doi.org/10.5281/zenodo.6015543" ID-GBIF-Taxon="131812502" ID-Zenodo-Dep="6015543" LSID="urn:lsid:plazi:treatment:8C70BE3B2441196CFF676AFEAAA4FC53" httpUri="http://treatment.plazi.org/id/8C70BE3B2441196CFF676AFEAAA4FC53" lastPageId="21" lastPageNumber="522" pageId="20" pageNumber="521">
<subSubSection id="4CC35CA62441196DFF676AFEAE8EFA5C" pageId="20" pageNumber="521" type="nomenclature">
<paragraph id="04660F2D2441196DFF676AFEADD7FA3D" blockId="20.[151,758,1420,1479]" box="[151,758,1420,1446]" pageId="20" pageNumber="521">
<heading id="5F2EB8412441196DFF676AFEADD7FA3D" bold="true" box="[151,758,1420,1446]" fontSize="11" level="1" pageId="20" pageNumber="521" reason="1">
<emphasis id="36ADD33F2441196DFF676AFEADD7FA3D" bold="true" box="[151,758,1420,1446]" pageId="20" pageNumber="521">
<taxonomicName id="C3D974AE2441196DFF676AFEAD48FA3D" ID-CoL="7CWQW" authority="Attems, 1950" authorityName="Attems" authorityYear="1950" box="[151,617,1420,1446]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="austerus" status="comb. nov.">
<emphasis id="36ADD33F2441196DFF676AFEAE8AFA3D" bold="true" box="[151,427,1420,1446]" italics="true" pageId="20" pageNumber="521">Tropostreptus austerus</emphasis>
(
<bibRefCitation id="604872DC2441196DFE4B6AFEAD40FA3D" author="Attems" box="[443,609,1420,1446]" pageId="20" pageNumber="521" refString="Attems, C. G. (1950) Uber Spirostreptiden (Diplopoda). Annalen des naturhistorischen Museums in Wien, 57, 179 - 257." type="journal article" year="1950">Attems, 1950</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="2D9E6E442441196DFD806AFEADD7FA3D" box="[624,758,1420,1446]" pageId="20" pageNumber="521">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="04660F2D2441196DFF676ADCAE8EFA5C" blockId="20.[151,758,1420,1479]" box="[151,431,1454,1479]" pageId="20" pageNumber="521">
<figureCitation id="9CE213A82441196DFF676ADCAFFFFA5C" box="[151,222,1454,1479]" captionStart="FIGURE 2" captionStartId="5.[151,250,1182,1204]" captionTargetBox="[151,1436,193,1096]" captionTargetId="figure@5.[151,1436,193,1161]" captionTargetPageId="5" captionText="FIGURE 2. Size of Tropostreptus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males. See Figs. 11 and 13 for details on T. hamatus and T. sigmatospinus sp. nov." httpUri="https://zenodo.org/record/803710/files/figure.png" pageId="20" pageNumber="521">Figs 2</figureCitation>
,
<figureCitation id="9CE213A82441196DFF1A6ADDAFD7FA5C" box="[234,246,1455,1479]" captionStart="FIGURE 8" captionStartId="14.[151,250,1827,1849]" captionTargetBox="[213,1375,193,1788]" captionTargetId="figure@14.[151,1436,153,1834]" captionTargetPageId="14" captionText="FIGURE 8. Tropostretus spp., left gonopods, proplica, anterior view. A: T. hamatus, East Usambara Mts, B: T. hamatus, Udzungwa Mts., C: T. sigmatospinus sp. nov., D: T. microcephalus sp. nov., E: T. droides sp. nov., F: T. kipunji sp. nov., G: T. austerus, H: T. severus sp. nov. — not to scale. The apicolateral proplical lobes (apl) are marked with yellow stars." httpUri="https://zenodo.org/record/803722/files/figure.png" pageId="20" pageNumber="521">8</figureCitation>
G, 10A, 14, 15A
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62441196DFF676A80AD06F99C" box="[151,551,1521,1543]" pageId="20" pageNumber="521" type="reference_group">
<paragraph id="04660F2D2441196DFF676A80AD06F99C" blockId="20.[151,1437,1521,2010]" box="[151,551,1521,1543]" pageId="20" pageNumber="521">
<treatmentCitationGroup id="24C928032441196DFF676A80AD06F99C" box="[151,551,1521,1543]" pageId="20" pageNumber="521">
<taxonomicName id="C3D974AE2441196DFF676A80AD06F99C" ID-CoL="6FVVJ" authority="Attems, 1950: 217" authorityName="Attems" authorityPageNumber="217" authorityYear="1950" box="[151,551,1521,1543]" class="Diplopoda" family="Spirostreptidae" genus="Epistreptus" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="austerus">
<emphasis id="36ADD33F2441196DFF676A80AE44F99C" box="[151,357,1522,1543]" italics="true" pageId="20" pageNumber="521">Epistreptus austerus</emphasis>
<treatmentCitation id="8578293C2441196DFE9C6A80AD06F99C" author="Attems" box="[364,551,1521,1543]" page="217" pageId="20" pageNumber="521" year="1950">
<bibRefCitation id="604872DC2441196DFE9C6A80AED6F99C" author="Attems" box="[364,503,1521,1543]" pageId="20" pageNumber="521" refString="Attems, C. G. (1950) Uber Spirostreptiden (Diplopoda). Annalen des naturhistorischen Museums in Wien, 57, 179 - 257." type="journal article" year="1950">Attems, 1950</bibRefCitation>
: 217
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62441196DFF67697DAE82F917" pageId="20" pageNumber="521" type="diagnosis">
<paragraph id="04660F2D2441196DFF67697DAE82F917" blockId="20.[151,1437,1521,2010]" pageId="20" pageNumber="521">
<emphasis id="36ADD33F2441196DFF67697DAE30F9B3" bold="true" box="[151,273,1551,1576]" pageId="20" pageNumber="521">Diagnosis.</emphasis>
A large species of
<taxonomicName id="C3D974AE2441196DFDFA6963AD85F9B3" box="[522,676,1553,1576]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="528" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="36ADD33F2441196DFDFA6963AD85F9B3" box="[522,676,1553,1576]" italics="true" pageId="20" pageNumber="521">Tropostreptus</emphasis>
</taxonomicName>
(
<figureCitation id="9CE213A82441196DFD49697DAC24F9B3" box="[697,773,1551,1576]" captionStart="FIGURE 2" captionStartId="5.[151,250,1182,1204]" captionTargetBox="[151,1436,193,1096]" captionTargetId="figure@5.[151,1436,193,1161]" captionTargetPageId="5" captionText="FIGURE 2. Size of Tropostreptus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males. See Figs. 11 and 13 for details on T. hamatus and T. sigmatospinus sp. nov." httpUri="https://zenodo.org/record/803710/files/figure.png" pageId="20" pageNumber="521">Fig. 2</figureCitation>
). Differs from other species, except
<taxonomicName id="C3D974AE2441196DFB356963AA1AF9B3" box="[1221,1339,1553,1576]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="severus">
<emphasis id="36ADD33F2441196DFB356963AA1AF9B3" box="[1221,1339,1553,1576]" italics="true" pageId="20" pageNumber="521">T. severus</emphasis>
</taxonomicName>
, by the combination of size and the presence of three apical lobes/processes on the gonopodal metaplica. Differs from
<taxonomicName id="C3D974AE2441196DFA776940AFCAF9F0" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="severus">
<emphasis id="36ADD33F2441196DFA776940AFCAF9F0" italics="true" pageId="20" pageNumber="521">T. severus</emphasis>
</taxonomicName>
by the smaller size of the lateral proplical lobe (
<emphasis id="36ADD33F2441196DFCD46920AC60F9F0" box="[804,833,1618,1643]" italics="true" pageId="20" pageNumber="521">lpl</emphasis>
) and the basad(-laterad) curvature of the telopodital antetorsal process (
<emphasis id="36ADD33F2441196DFE9F6907AEB2F917" box="[367,403,1653,1676]" italics="true" pageId="20" pageNumber="521">atp</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62441196DFF3769E7AC12F842" pageId="20" pageNumber="521" type="materials_examined">
<paragraph id="04660F2D2441196DFF3769E7ADB1F935" blockId="20.[151,1437,1521,2010]" box="[199,656,1685,1710]" pageId="20" pageNumber="521">
<emphasis id="36ADD33F2441196DFF3769E7AEACF935" bold="true" box="[199,397,1685,1710]" pageId="20" pageNumber="521">Material studied</emphasis>
(total: 18 ♂♂, 10 ♀♀)
</paragraph>
<paragraph id="04660F2D2441196DFF3769C4AAB1F954" blockId="20.[151,1437,1521,2010]" box="[199,1424,1718,1743]" pageId="20" pageNumber="521">
<emphasis id="36ADD33F2441196DFF3769C4AE1CF954" bold="true" box="[199,317,1718,1743]" pageId="20" pageNumber="521">Holotype:</emphasis>
♂ “
<taxonomicName id="C3D974AE2441196DFE9C69C5ADD0F954" authority="Attems, 1950" authorityName="Attems" authorityYear="1950" box="[364,753,1718,1743]" class="Diplopoda" family="Spirostreptidae" genus="Epistreptus" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="austerus">
<emphasis id="36ADD33F2441196DFE9C69C5AD70F955" box="[364,593,1719,1742]" italics="true" pageId="20" pageNumber="521">Epistreptus austerus</emphasis>
<bibRefCitation id="604872DC2441196DFDA769C5ADD0F954" author="Attems" box="[599,753,1718,1743]" pageId="20" pageNumber="521" refString="Attems, C. G. (1950) Uber Spirostreptiden (Diplopoda). Annalen des naturhistorischen Museums in Wien, 57, 179 - 257." type="journal article" year="1950">Attems, 1950</bibRefCitation>
</taxonomicName>
, E-Afrika Umgebung v. Amani, leg. Bogert” (NMW 2430).
</paragraph>
<paragraph id="04660F2D2441196DFF3769A5ADFDF822" blockId="20.[151,1437,1521,2010]" pageId="20" pageNumber="521">
<materialsCitation id="B4B105702441196DFF3769A5AB60F8A9" ID-GBIF-Occurrence="1563407758" collectingDate="1974" collectingDateMax="1995" collectingDateMin="1974" collectionCode="ZMUC" collectorName="P. Bjorn &amp; I. B. &amp; H. Enghoff &amp; C. Griswold &amp; O. Lomholdt &amp; O. Martin &amp; N. Scharff &amp; L. Sorensen &amp; D. Ubick" country="Tanzania" county="Amani" elevation="975" latitude="-5.095" location="New" longLatPrecision="926" longitude="38.633335" municipality="East Usambara Mts" pageId="20" pageNumber="521" specimenCount="18" specimenCount-female="5" specimenCount-male="13" stateProvince="Tanga Region">
<emphasis id="36ADD33F2441196DFF3769A5AE50F96B" bold="true" box="[199,369,1751,1776]" pageId="20" pageNumber="521">
<location id="010659F62441196DFF3769A5AFDDF96B" LSID="urn:lsid:plazi:treatment:8C70BE3B2441196CFF676AFEAAA4FC53:010659F62441196DFF3769A5AFDDF96B" box="[199,252,1751,1776]" country="Tanzania" county="Amani" latitude="-5.095" longLatPrecision="926" longitude="38.633335" municipality="East Usambara Mts" name="New" pageId="20" pageNumber="521" stateProvince="Tanga Region">New</location>
material:
</emphasis>
<location id="010659F62441196DFE8769A5AEBAF96B" LSID="urn:lsid:plazi:treatment:8C70BE3B2441196CFF676AFEAAA4FC53:010659F62441196DFE8769A5AEBAF96B" box="[375,411,1751,1776]" country="Tanzania" county="Amani" latitude="-5.095" longLatPrecision="926" longitude="38.633335" municipality="East Usambara Mts" name="All" pageId="20" pageNumber="521" stateProvince="Tanga Region">All</location>
from
<collectingCountry id="7CCE4FBD2441196DFE2E69AAAD51F96B" box="[478,624,1752,1776]" name="Tanzania" pageId="20" pageNumber="521">TANZANIA</collectingCountry>
,
<collectingRegion id="C61DC1CF2441196DFD8A69AAAC38F96B" box="[634,793,1751,1776]" country="Tanzania" name="Tanga" pageId="20" pageNumber="521">Tanga Region</collectingRegion>
,
<collectingMunicipality id="E40295572441196DFCD469AAAB21F974" box="[804,1024,1751,1776]" pageId="20" pageNumber="521">East Usambara Mts</collectingMunicipality>
, all in
<collectionCode id="62C897E82441196DFBBA69AAABBEF96B" box="[1098,1183,1752,1776]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="20" pageNumber="521">ZMUC</collectionCode>
:
<specimenCount id="12DFC4A42441196DFB5A69AAABD9F974" box="[1194,1272,1752,1776]" pageId="20" pageNumber="521" type="male">13 ♂♂</specimenCount>
,
<specimenCount id="12DFC4A42441196DFAF469A5AA63F974" box="[1284,1346,1751,1776]" pageId="20" pageNumber="521" type="female">5 ♀♀</specimenCount>
<collectingCounty id="ED0777A12441196DFAB969A5AAB9F96B" box="[1353,1432,1751,1776]" pageId="20" pageNumber="521">Amani</collectingCounty>
,
<geoCoordinate id="61ED69EA2441196DFF67698BAFCDF889" box="[151,236,1785,1810]" direction="south" orientation="latitude" pageId="20" pageNumber="521" precision="92" value="-5.095">5°5.7’S</geoCoordinate>
<geoCoordinate id="61ED69EA2441196DFF036988AE72F889" box="[243,339,1786,1810]" direction="east" orientation="longitude" pageId="20" pageNumber="521" precision="925" value="38.633335">38°38’E</geoCoordinate>
,
<elevation id="8FF4E81E2441196DFEAF698BAD34F889" box="[351,533,1785,1810]" metricMagnitude="2" metricUnit="m" metricValue="9.75" metricValueMax="10.0" metricValueMin="9.5" pageId="20" pageNumber="521" unit="m" value="975.0" valueMax="1000.0" valueMin="950.0">
<quantity id="C321A2C82441196DFEAF698BAECDF88A" box="[351,492,1785,1810]" metricMagnitude="2" metricUnit="m" metricValue="9.75" metricValueMax="10.0" metricValueMin="9.5" pageId="20" pageNumber="521" unit="m" value="975.0" valueMax="1000.0" valueMin="950.0">950–1000 m</quantity>
asl
</elevation>
,
<collectingDate id="6023D0052441196DFDD1698BAD80F889" box="[545,673,1785,1810]" pageId="20" pageNumber="521" value="1974" valueMax="1995">
<date id="706729ED2441196DFDD1698BAD80F889" box="[545,673,1785,1810]" pageId="20" pageNumber="521" value="1974" valueMax="1995">1974–1995</date>
</collectingDate>
,
<collectorName id="A92C6AFB2441196DFD5D6988AC2BF889" box="[685,778,1785,1810]" pageId="20" pageNumber="521">P. Bjørn</collectorName>
,
<collectorName id="A92C6AFB2441196DFCE66988ACC5F889" box="[790,996,1785,1810]" pageId="20" pageNumber="521">I.B. &amp; H. Enghoff</collectorName>
,
<collectorName id="A92C6AFB2441196DFC006988AB5DF889" box="[1008,1148,1785,1810]" pageId="20" pageNumber="521">C. Griswold</collectorName>
,
<collectorName id="A92C6AFB2441196DFB776988AA3AF889" box="[1159,1307,1785,1810]" pageId="20" pageNumber="521">O. Lomholdt</collectorName>
,
<collectorName id="A92C6AFB2441196DFAD66988AAB9F889" box="[1318,1432,1785,1810]" pageId="20" pageNumber="521">O. Martin</collectorName>
,
<collectorName id="A92C6AFB2441196DFF676869AE2EF8A8" box="[151,271,1818,1843]" pageId="20" pageNumber="521">N. Scharff</collectorName>
,
<collectorName id="A92C6AFB2441196DFEEA6869AE81F8A8" box="[282,416,1819,1843]" pageId="20" pageNumber="521">L. Sørensen</collectorName>
&amp;
<collectorName id="A92C6AFB2441196DFE356869AD0CF8A8" box="[453,557,1818,1843]" pageId="20" pageNumber="521">D. Ubick</collectorName>
leg., “on tree stump”, “in decaying wood” etc.
</materialsCitation>
;
<materialsCitation id="B4B105702441196DFBBD6869AE0AF8ED" ID-GBIF-Occurrence="1563407776" collectorName="Frontier Tanzania" country="Tanzania" county="Nilo Forest" elevation="940" latitude="-4.942389" location="Muheza" longLatPrecision="1" longitude="38.660694" municipality="Nilo Forest Reserve" pageId="20" pageNumber="521" specimenCount="1" specimenCount-male="1" stateProvince="Tanga Region">
<specimenCount id="12DFC4A42441196DFBBD6869AB59F8A9" box="[1101,1144,1819,1843]" pageId="20" pageNumber="521" type="male">1 ♂</specimenCount>
<collectingRegion id="C61DC1CF2441196DFB706869AA3EF8A8" box="[1152,1311,1818,1843]" country="Tanzania" name="Tanga" pageId="20" pageNumber="521">Tanga Region</collectingRegion>
,
<location id="010659F62441196DFADB6868AAA4F8A8" LSID="urn:lsid:plazi:treatment:8C70BE3B2441196CFF676AFEAAA4FC53:010659F62441196DFADB6868AAA4F8A8" box="[1323,1413,1818,1843]" country="Tanzania" county="Nilo Forest" latitude="-4.942389" longLatPrecision="1" longitude="38.660694" municipality="Nilo Forest Reserve" name="Muheza" pageId="20" pageNumber="521" stateProvince="Tanga Region">Muheza</location>
+
<location id="010659F62441196DFF67684FAE7CF8CE" LSID="urn:lsid:plazi:treatment:8C70BE3B2441196CFF676AFEAAA4FC53:010659F62441196DFF67684FAE7CF8CE" box="[151,349,1852,1877]" country="Tanzania" county="Nilo Forest" latitude="-4.942389" longLatPrecision="1" longitude="38.660694" municipality="Nilo Forest Reserve" name="Korogwe District" pageId="20" pageNumber="521" stateProvince="Tanga Region">Korogwe District</location>
, (
<collectingMunicipality id="E40295572441196DFE80684EAD72F8CF" box="[368,595,1852,1877]" pageId="20" pageNumber="521">
<collectingCounty id="ED0777A12441196DFE80684EAED1F8CF" box="[368,496,1852,1877]" pageId="20" pageNumber="521">Nilo Forest</collectingCounty>
Reserve
</collectingMunicipality>
)
<elevation id="8FF4E81E2441196DFD92684EADF3F8CE" box="[610,722,1852,1877]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="20" pageNumber="521" unit="m" value="940.0">
<quantity id="C321A2C82441196DFD92684EAD88F8CF" box="[610,681,1852,1876]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="20" pageNumber="521" unit="m" value="940.0">940 m</quantity>
asl
</elevation>
,
<geoCoordinate id="61ED69EA2441196DFD2D684EAC48F8CE" box="[733,873,1852,1877]" direction="south" orientation="latitude" pageId="20" pageNumber="521" precision="1" value="-4.942389">4°56’32.6”S</geoCoordinate>
<geoCoordinate id="61ED69EA2441196DFC80684EAB2CF8CE" box="[880,1037,1852,1877]" direction="east" orientation="longitude" pageId="20" pageNumber="521" precision="1" value="38.660694">38°39’38.5”E</geoCoordinate>
, trapsite 8 plot 59A, date?,
<collectorName id="A92C6AFB2441196DFAB2684EAFDDF8ED" pageId="20" pageNumber="521">Frontier Tanzania</collectorName>
leg.
</materialsCitation>
—
<materialsCitation id="B4B105702441196DFEB9682FAD01F80C" ID-GBIF-Occurrence="1563407766" collectingDate="2001-02-27" collectingDateMax="2001-06-21" collectingDateMin="2001-02-27" collectorName="Frontier Tanzania" country="Tanzania" county="Nilo Forest" elevation="1125" location="Muheza district" municipality="Nilo Forest Reserve" pageId="20" pageNumber="521" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Tanga Region">
<specimenCount id="12DFC4A42441196DFEB9682FAEADF8EE" box="[329,396,1885,1910]" pageId="20" pageNumber="521" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="12DFC4A42441196DFE69682CAEF8F8EE" box="[409,473,1886,1910]" pageId="20" pageNumber="521" type="female">4 ♀♀</specimenCount>
<location id="010659F62441196DFE12682FADB7F8ED" LSID="urn:lsid:plazi:treatment:8C70BE3B2441196CFF676AFEAAA4FC53:010659F62441196DFE12682FADB7F8ED" box="[482,662,1885,1910]" country="Tanzania" county="Nilo Forest" municipality="Nilo Forest Reserve" name="Muheza district" pageId="20" pageNumber="521" stateProvince="Tanga Region">Muheza district</location>
,
<collectingMunicipality id="E40295572441196DFD52682FACA9F8EE" box="[674,904,1885,1910]" pageId="20" pageNumber="521">
<collectingCounty id="ED0777A12441196DFD52682FAC03F8EE" box="[674,802,1885,1910]" pageId="20" pageNumber="521">Nilo Forest</collectingCounty>
Reserve
</collectingMunicipality>
,
<elevation id="8FF4E81E2441196DFC64682CAB7AF8ED" box="[916,1115,1885,1910]" metricMagnitude="3" metricUnit="m" metricValue="1.125" metricValueMax="1.24" metricValueMin="1.01" pageId="20" pageNumber="521" unit="m" value="1125.0" valueMax="1240.0" valueMin="1010.0">
<quantity id="C321A2C82441196DFC64682CAB10F8EE" box="[916,1073,1886,1910]" metricMagnitude="3" metricUnit="m" metricValue="1.125" metricValueMax="1.24" metricValueMin="1.01" pageId="20" pageNumber="521" unit="m" value="1125.0" valueMax="1240.0" valueMin="1010.0">1010–1240 m</quantity>
asl
</elevation>
, submontane forest,
<collectingDate id="6023D0052441196DFAB7682FAE30F80C" pageId="20" pageNumber="521" value="2001-02-27" valueMax="2001-06-21" valueMin="2001-02-27">
<date id="706729ED2441196DFAB7682FAE30F80C" pageId="20" pageNumber="521" value="2001-02-27" valueMax="2001-06-21" valueMin="2001-02-27">27.ii.— 21.vi.2001</date>
</collectingDate>
,
<collectorName id="A92C6AFB2441196DFED0680CAECBF80C" box="[288,490,1918,1943]" pageId="20" pageNumber="521">Frontier Tanzania</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="B4B105702441196DFDDF680DADFDF822" ID-GBIF-Occurrence="1563407775" collectingDate="2000-06-10" collectorName="Frontier Tanzania" country="Tanzania" county="Nilo Forest" elevation="820" latitude="-4.9044447" location="Muheza District" longLatPrecision="21" longitude="38.662502" municipality="Nilo Forest Reserve" pageId="20" pageNumber="521" specimenCount="1" specimenCount-female="1" stateProvince="Tanga Region">
<specimenCount id="12DFC4A42441196DFDDF680DAD7CF80D" box="[559,605,1919,1943]" pageId="20" pageNumber="521" type="female">1 ♀</specimenCount>
<location id="010659F62441196DFD97680CAC02F80C" LSID="urn:lsid:plazi:treatment:8C70BE3B2441196CFF676AFEAAA4FC53:010659F62441196DFD97680CAC02F80C" box="[615,803,1918,1943]" country="Tanzania" county="Nilo Forest" latitude="-4.9044447" longLatPrecision="21" longitude="38.662502" municipality="Nilo Forest Reserve" name="Muheza District" pageId="20" pageNumber="521" stateProvince="Tanga Region">Muheza District</location>
,
<collectingMunicipality id="E40295572441196DFCC1680CAB3AF80D" box="[817,1051,1918,1943]" pageId="20" pageNumber="521">
<collectingCounty id="ED0777A12441196DFCC1680CAC92F80D" box="[817,947,1918,1943]" pageId="20" pageNumber="521">Nilo Forest</collectingCounty>
Reserve
</collectingMunicipality>
,
<geoCoordinate id="61ED69EA2441196DFBD9680CAB81F80C" box="[1065,1184,1918,1943]" direction="south" orientation="latitude" pageId="20" pageNumber="521" precision="15" value="-4.9044447">4°54’16”S</geoCoordinate>
<geoCoordinate id="61ED69EA2441196DFB5A680CAA13F80C" box="[1194,1330,1918,1943]" direction="east" orientation="longitude" pageId="20" pageNumber="521" precision="15" value="38.662502">38°39’45”E</geoCoordinate>
, lowland forest,
<elevation id="8FF4E81E2441196DFF1468D2AE75F822" box="[228,340,1952,1977]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="20" pageNumber="521" unit="m" value="820.0">
<quantity id="C321A2C82441196DFF1468D2AE0AF823" box="[228,299,1952,1976]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="20" pageNumber="521" unit="m" value="820.0">820 m</quantity>
asl
</elevation>
,
<collectingDate id="6023D0052441196DFEAF68D2AEF8F822" box="[351,473,1952,1977]" pageId="20" pageNumber="521" value="2000-06-10">
<date id="706729ED2441196DFEAF68D2AEF8F822" box="[351,473,1952,1977]" pageId="20" pageNumber="521" value="2000-06-10">10.vi.2000</date>
</collectingDate>
,
<collectorName id="A92C6AFB2441196DFE1568D2AD8DF822" box="[485,684,1952,1977]" pageId="20" pageNumber="521">Frontier Tanzania</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="04660F2D2441196DFF3768B3AC12F842" blockId="20.[151,1437,1521,2010]" box="[199,819,1985,2010]" pageId="20" pageNumber="521">
<emphasis id="36ADD33F2441196DFF3768B3AE44F841" bold="true" box="[199,357,1985,2010]" pageId="20" pageNumber="521">Type locality.</emphasis>
Environs of Amani, East Usambara Mts.
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62440196CFF376FE5AE58FD6B" pageId="21" pageNumber="522" type="description">
<paragraph id="04660F2D2440196CFF376FE5AED2FF4E" blockId="21.[151,1437,151,968]" pageId="21" pageNumber="522">
<emphasis id="36ADD33F2440196CFF376FE5AE78FF2B" bold="true" box="[199,345,151,176]" pageId="21" pageNumber="522">Description.</emphasis>
SIZE (
<figureCitation id="9CE213A82440196CFE566FE5AECFFF2B" box="[422,494,151,176]" captionStart="FIGURE 2" captionStartId="5.[151,250,1182,1204]" captionTargetBox="[151,1436,193,1096]" captionTargetId="figure@5.[151,1436,193,1161]" captionTargetPageId="5" captionText="FIGURE 2. Size of Tropostreptus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males. See Figs. 11 and 13 for details on T. hamatus and T. sigmatospinus sp. nov." httpUri="https://zenodo.org/record/803710/files/figure.png" pageId="21" pageNumber="522">Fig. 2</figureCitation>
). Length
<quantity id="C321A2C82440196CFDAE6FEAADF5FF34" box="[606,724,152,176]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="21" pageNumber="522" unit="cm" value="13.5" valueMax="14.0" valueMin="13.0">13–14 cm</quantity>
; vertical body diameter
<quantity id="C321A2C82440196CFC1C6FE5ABA5FF34" box="[1004,1156,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" metricValueMax="10.6" metricValueMin="8.6" pageId="21" pageNumber="522" unit="mm" value="9.6" valueMax="10.6" valueMin="8.6">8.6–10.6 mm</quantity>
(males), up to
<quantity id="C321A2C82440196CFAC76FEAAABCFF34" box="[1335,1437,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="21" pageNumber="522" unit="mm" value="11.8">11.8 mm</quantity>
(females), 51–57 podous rings.
</paragraph>
<paragraph id="04660F2D2440196CFF376F92AB46FEDB" blockId="21.[151,1437,151,968]" pageId="21" pageNumber="522">COLOUR. Live colour dark brown (pers. obs.). After 16 years in alcohol head, antennae, legs and telson orange yellow or brownish, body rings marbled greyish back to suture, then brownish, posterior ca. 10% amber brown; alternatively each body ring dark orange in anterior half, almost black in posterior half.</paragraph>
<paragraph id="04660F2D2440196CFF376E3FAABCFEFF" blockId="21.[151,1437,151,968]" box="[199,1437,332,357]" pageId="21" pageNumber="522">HEAD. Eyes with more ommatidia than in congeners: up to 60 ommatidia in up to 17 vertical and 8 horizontal rows.</paragraph>
<paragraph id="04660F2D2440196CFF376E02AE58FD6B" blockId="21.[151,1437,151,968]" pageId="21" pageNumber="522">
GONOPODS (
<figureCitation id="9CE213A82440196CFEA56E1DAEBAFE13" box="[341,411,367,392]" captionStart="FIGURE 8" captionStartId="14.[151,250,1827,1849]" captionTargetBox="[213,1375,193,1788]" captionTargetId="figure@14.[151,1436,153,1834]" captionTargetPageId="14" captionText="FIGURE 8. Tropostretus spp., left gonopods, proplica, anterior view. A: T. hamatus, East Usambara Mts, B: T. hamatus, Udzungwa Mts., C: T. sigmatospinus sp. nov., D: T. microcephalus sp. nov., E: T. droides sp. nov., F: T. kipunji sp. nov., G: T. austerus, H: T. severus sp. nov. — not to scale. The apicolateral proplical lobes (apl) are marked with yellow stars." httpUri="https://zenodo.org/record/803722/files/figure.png" pageId="21" pageNumber="522">Figs 8</figureCitation>
G, 14, 15A). Sternum triangular, reaching almost as far distad as paracoxites, lateral edges slightly convex. Proplica ending in blunt-triangular process (
<emphasis id="36ADD33F2440196CFCB06EE7AC51FE37" box="[832,880,405,428]" italics="true" pageId="21" pageNumber="522">map</emphasis>
) separated by sinus from large apico-lateral, subsemicircular lobe (
<emphasis id="36ADD33F2440196CFE816EC5AEAFFE4B" box="[369,398,439,464]" italics="true" pageId="21" pageNumber="522">lpl</emphasis>
). Apico-lateral lobe (
<emphasis id="36ADD33F2440196CFD7C6EC5AD88FE4B" box="[652,681,439,464]" italics="true" pageId="21" pageNumber="522">lpl</emphasis>
) reaching further distad than triangular lobe (
<emphasis id="36ADD33F2440196CFB366EC5ABD1FE4B" box="[1222,1264,439,464]" italics="true" pageId="21" pageNumber="522">mpl</emphasis>
). Metaplica in anterior view basally with straight ridge continuing ca. to mid-length of coxite, apically with small mesal process (
<emphasis id="36ADD33F2440196CFF506D73AFF1FD83" box="[160,208,513,536]" italics="true" pageId="21" pageNumber="522">map</emphasis>
) separated by sinus from sub-semicircular mid-distal process which again is separated by deep sinus from lateral birdhead-shaped process (
<emphasis id="36ADD33F2440196CFDFB6D56AD06FDA6" box="[523,551,548,573]" italics="true" pageId="21" pageNumber="522">bh</emphasis>
), lateral “beak” of the latter long, slender, pointed, directed laterad. Birdheadshaped process reaching further distad than any other part of gonopod. Telopodite shortly after emergence from coxa with very long, slender, pointed, sinuous antetorsal process (
<emphasis id="36ADD33F2440196CFC6D6D1FACE0FD1F" box="[925,961,621,644]" italics="true" pageId="21" pageNumber="522">atp</emphasis>
); overall direction of process basad (the basad-mesad direction seen in
<figureCitation id="9CE213A82440196CFDFC6DFDAD42FD33" box="[524,611,655,680]" captionStart="FIGURE 14" captionStartId="21.[151,250,1888,1910]" captionTargetBox="[217,1374,1013,1848]" captionTargetId="figure@21.[155,1440,980,1898]" captionTargetPageId="21" captionText="FIGURE 14. Tropostreptus austerus, gonopods, anterior view. The birdhead-shaped process of the right gonopod is misshapen; the basad-mesad direction of the antetorsal process is due to drying of the gonopods for scanning electron microscopy. Scale 0.5 mm." httpUri="https://zenodo.org/record/803732/files/figure.png" pageId="21" pageNumber="522">Fig. 14</figureCitation>
is an artefact due to drying of the gonopods for scanning electron microscopy). Telopodite distal to antetorsal process with subrectangular flange, distal to flange slender, simple, without outgrowths.
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62440196CFF376D8EAA14FCC6" pageId="21" pageNumber="522" type="distribution">
<paragraph id="04660F2D2440196CFF376D8EACA4FCA3" blockId="21.[151,1437,151,968]" pageId="21" pageNumber="522">
<emphasis id="36ADD33F2440196CFF376D8EAE78FC8E" bold="true" box="[199,345,764,789]" pageId="21" pageNumber="522">Distribution</emphasis>
. Known only from lowland and submontane forest in the East Usambara Mts, 820–1010 (1240) m asl. Microhabitat notes include a tree stump and decaying wood.
</paragraph>
<paragraph id="04660F2D2440196CFF376C36AA14FCC6" blockId="21.[151,1437,151,968]" box="[199,1333,836,861]" pageId="21" pageNumber="522">
<emphasis id="36ADD33F2440196CFF376C36AE9CFCC6" bold="true" box="[199,445,836,861]" pageId="21" pageNumber="522">Coexisting congener.</emphasis>
Coexists with
<taxonomicName id="C3D974AE2440196CFD946C37ADFEFCC6" box="[612,735,836,861]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F2440196CFD946C37ADFEFCC6" box="[612,735,836,861]" italics="true" pageId="21" pageNumber="522">T. hamatus</emphasis>
</taxonomicName>
in the East Usambara Mts, especially around Amani.
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62440196CFF376C15AAA4FC53" pageId="21" pageNumber="522" type="description">
<paragraph id="04660F2D2440196CFF376C15AAA4FC53" blockId="21.[151,1437,151,968]" pageId="21" pageNumber="522">
<emphasis id="36ADD33F2440196CFF376C15AD21FC1B" bold="true" box="[199,512,871,896]" pageId="21" pageNumber="522">
Infection with
<taxonomicName id="C3D974AE2440196CFE846C15AD21FC1B" box="[372,512,871,896]" class="Laboulbeniomycetes" family="Laboulbeniaceae" genus="Rickia" kingdom="Fungi" order="Laboulbeniales" pageId="21" pageNumber="522" phylum="Ascomycota" rank="species" species="gigas">
<emphasis id="36ADD33F2440196CFE846C15AD21FC1B" box="[372,512,871,896]" italics="true" pageId="21" pageNumber="522">Rickia gigas</emphasis>
</taxonomicName>
</emphasis>
. The long-celled
<typeStatus id="DB62B18F2440196CFD366C1AADD4FC1B" box="[710,757,872,896]" pageId="21" pageNumber="522">type</typeStatus>
of
<taxonomicName id="C3D974AE2440196CFCEC6C15AC84FC1B" box="[796,933,871,896]" class="Laboulbeniomycetes" family="Laboulbeniaceae" genus="Rickia" kingdom="Fungi" order="Laboulbeniales" pageId="21" pageNumber="522" phylum="Ascomycota" rank="species" species="gigas">
<emphasis id="36ADD33F2440196CFCEC6C15AC84FC1B" box="[796,933,871,896]" italics="true" pageId="21" pageNumber="522">Rickia gigas</emphasis>
</taxonomicName>
was found on the mesal-ventral sides of legpairs
<quantity id="C321A2C82440196CFF236CFEAE39FC3E" box="[211,280,908,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.778" metricValueMin="0.508" pageId="21" pageNumber="522" unit="in" value="4.5" valueMax="7.0" valueMin="2.0">2–7 in</quantity>
males. Most specimens, including the
<typeStatus id="DB62B18F2440196CFD316CFEAC02FC3E" box="[705,803,908,933]" pageId="21" pageNumber="522" type="holotype">holotype</typeStatus>
, were found to be infested with the short-celled
<typeStatus id="DB62B18F2440196CFAC16CFFAA41FC3E" box="[1329,1376,909,933]" pageId="21" pageNumber="522">type</typeStatus>
of
<taxonomicName id="C3D974AE2440196CFA746CFFAFF3FC53" class="Laboulbeniomycetes" family="Laboulbeniaceae" genus="Rickia" kingdom="Fungi" order="Laboulbeniales" pageId="21" pageNumber="522" phylum="Ascomycota" rank="species" species="gigas">
<emphasis id="36ADD33F2440196CFA746CFFAFF3FC53" italics="true" pageId="21" pageNumber="522">R. gigas</emphasis>
</taxonomicName>
on the hind margin of the body rings of both sexes, in some cases on
<emphasis id="36ADD33F2440196CFC3C6CDDACC9FC53" box="[972,1000,943,968]" italics="true" pageId="21" pageNumber="522">all</emphasis>
body rings and even the preanal ring.
</paragraph>
</subSubSection>
</treatment>
</document>
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