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<document id="7676543D99257A69186F2CD0C77D5AFE" ID-DOI="10.5281/zenodo.5458372" ID-ISSN="2345-7600" ID-Zenodo-Dep="5458372" ID-ZooBank="08BE1873-6F6F-4255-9520-9A89F48E4F16" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630211029465" checkinUser="tatiana" docAuthor="Lemaitre, Rafael" docDate="2014" docId="8C6A8943FFB27824FF1DFA04FC02FEA6" docLanguage="en" docName="62rbz210-301.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Oncopagurus spiniartus Lemaitre 2014, new species" docType="treatment" docVersion="3" lastPageNumber="293" masterDocId="7053F13BFFFB7877FFE4FFD9FFDDFFB3" masterDocTitle="A worldwide taxonomic and distributional synthesis of the genus Oncopagurus Lemaitre, 1996 (Crustacea: Decapoda: Anomura: Parapaguridae), with descriptions of nine new species" masterLastPageNumber="301" masterPageNumber="210" pageNumber="283" updateTime="1699187482961" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C1A00A0E6817346EE49D70E1B0308731">A worldwide taxonomic and distributional synthesis of the genus Oncopagurus Lemaitre, 1996 (Crustacea: Decapoda: Anomura: Parapaguridae), with descriptions of nine new species</mods:title>
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<mods:namePart id="B7F0B87B491F388C957DFC38C51C4681">Lemaitre, Rafael</mods:namePart>
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<mods:date id="D9F3CD0DB3E4937265249B342F9072EC">2014</mods:date>
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<paragraph id="047C3855FFB2783EFF1DFA04FD4BFA46" blockId="73.[249,662,1501,1557]" box="[249,662,1501,1525]" pageId="73" pageNumber="283">
<heading id="5F348F39FFB2783EFF1DFA04FD4BFA46" box="[249,662,1501,1525]" centered="true" fontSize="10" level="2" pageId="73" pageNumber="283" reason="3">
<taxonomicName id="C3C343D6FFB2783EFF1DFA04FDD8FA46" authority="Lemaitre, 2014" authorityName="Lemaitre" authorityYear="2014" box="[249,517,1501,1525]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="73" pageNumber="283" phylum="Arthropoda" rank="species" species="spiniartus" status="sp. nov.">
<emphasis id="36B7E447FFB2783EFF1DFA04FDD8FA46" bold="true" box="[249,517,1501,1525]" italics="true" pageId="73" pageNumber="283">Oncopagurus spiniartus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2D84593CFFB2783EFDF0FA07FD4BFA46" box="[532,662,1501,1525]" pageId="73" pageNumber="283" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="4CD96BDEFFB2783EFE96FA24FDC0F9A6" box="[370,541,1533,1557]" pageId="73" pageNumber="283" type="description">
<paragraph id="047C3855FFB2783EFE96FA24FDC0F9A6" blockId="73.[249,662,1501,1557]" box="[370,541,1533,1557]" pageId="73" pageNumber="283">
<figureCitation id="9CF824D0FFB2783EFE96FA24FE28F9A6" box="[370,501,1533,1557]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="74.[113,148,1975,1997]" captionStartId-1="75.[136,171,2035,2057]" captionStartId-2="76.[113,148,2035,2057]" captionStartId-3="77.[136,171,2035,2057]" captionTargetBox-0="[116,1445,306,1862]" captionTargetBox-1="[140,1460,932,1997]" captionTargetBox-2="[119,1448,599,2005]" captionTargetBox-3="[135,1459,222,1997]" captionTargetId-0="figure-9@74.[101,1472,291,1881]" captionTargetId-1="figure-7@75.[130,1475,922,2007]" captionTargetId-2="figure-9@76.[91,1467,554,2009]" captionTargetId-3="figure-7@77.[125,1474,209,2005]" captionTargetPageId-0="74" captionTargetPageId-1="75" captionTargetPageId-2="76" captionTargetPageId-3="77" captionText-0="Fig. 42. Oncopagurus spiniartus, new species, southern Indian Ocean, Île Amsterdam and Île St.-Paul: AE, holotype, male 2.3 mm, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610); F, G, paratype, male 2.8 mm, CENTOB, MD 50, DC 114, sta 24, (MNHN-IU-5495, ex MNHN Pg 7609): A, shield and cephalic appendages; B, right antennal peduncle, lateral; C, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; D, exopod of left uropod, dorsal; E, telson, dorsal; F, left first gonopod, mesial; G, left second gonopod, anterior. Scale bars = 0.5 mm (A), 0.25 mm (BD), 0.2 mm (F, G)." captionText-1="Fig. 43. Oncopagurus spiniartus, new species, paratype, male 2.8 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 114, sta 24 (MNHN-IU-5495, ex MNHN Pg 7609). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.2 mm (AE); 0.5 mm (F)." captionText-2="Fig. 44. Oncopagurus spiniartus, new species, holotype, male 2.3 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610): A, right cheliped, dorsal; B, chela of same, mesial; C, left cheliped, dorsal; D, same, mesial. Scale bar = 1 mm." captionText-3="Fig. 45. Oncopagurus spiniartus, new species, holotype, male 2.3 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610): A, right first ambulatory leg, mesial; B, same, lateral; C, right second ambulatory leg, lateral, D, dactyl of same, mesial. Scale bars = 1 mm (AD); 0. 5 mm (E, F)." figureDoi-0="http://doi.org/10.5281/zenodo.5458460" figureDoi-1="http://doi.org/10.5281/zenodo.5458462" figureDoi-2="http://doi.org/10.5281/zenodo.5458464" figureDoi-3="http://doi.org/10.5281/zenodo.5458466" httpUri-0="https://zenodo.org/record/5458460/files/figure.png" httpUri-1="https://zenodo.org/record/5458462/files/figure.png" httpUri-2="https://zenodo.org/record/5458464/files/figure.png" httpUri-3="https://zenodo.org/record/5458466/files/figure.png" pageId="73" pageNumber="283">Figs. 4245</figureCitation>
,
<figureCitation id="9CF824D0FFB2783EFDE6FA24FDC0F9A6" box="[514,541,1533,1557]" captionStart="Fig" captionStartId="85.[136,171,2092,2114]" captionTargetBox="[139,1474,1084,2054]" captionTargetId="figure-224@85.[139,1476,1083,2054]" captionTargetPageId="85" captionText="Fig. 51. Bathymetric distribution of Oncopagurus species from the world, with maximum and minimum depths (meters) indicated for each." figureDoi="http://doi.org/10.5281/zenodo.5458478" httpUri="https://zenodo.org/record/5458478/files/figure.png" pageId="73" pageNumber="283">51</figureCitation>
</paragraph>
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<subSubSection id="4CD96BDEFFB2783EFF6CF9E5FDA9F8FA" pageId="73" pageNumber="283" type="materials_examined">
<paragraph id="047C3855FFB2783EFF6CF9E5FE60F91B" blockId="73.[136,775,1595,1704]" pageId="73" pageNumber="283">
<emphasis id="36B7E447FFB2783EFF6CF9E5FEC3F9E1" bold="true" box="[136,286,1596,1618]" pageId="73" pageNumber="283">Type material.</emphasis>
<materialsCitation id="B4AB3208FFB2783EFEC1F9E5FE60F91B" collectingDate="1986-07-17" collectionCode="MNHN" county="Ile St." elevation="165" location="Southern Indian Ocean" municipality="Ile Amsterdam" pageId="73" pageNumber="283" specimenCount="1" typeStatus="holotype">
<typeStatus id="DB7886F7FFB2783EFEC1F9E5FE5CF9E1" box="[293,385,1596,1618]" pageId="73" pageNumber="283" type="holotype">Holotype</typeStatus>
,
<location id="011C6E8EFFB2783EFE6FF9E5FDB0F9E2" LSID="urn:lsid:plazi:treatment:8C6A8943FFB27824FF1DFA04FC02FEA6:011C6E8EFFB2783EFE6FF9E5FDB0F9E2" box="[395,621,1596,1618]" county="Ile St." municipality="Ile Amsterdam" name="Southern Indian Ocean" pageId="73" pageNumber="283">Southern Indian Ocean</location>
,
<collectingMunicipality id="E418A22FFFB2783EFD9CF9E2FCDAF9E1" box="[632,775,1595,1618]" pageId="73" pageNumber="283">Île Amsterdam</collectingMunicipality>
and
<collectingCounty id="ED1D40D9FFB2783EFF51F981FF2EF9DC" box="[181,243,1624,1647]" pageId="73" pageNumber="283">Île St.</collectingCounty>
-Paul, CENTOB, MD 50, DC 82, sta 19, 38°428'S, 77°283'E,
<quantity id="C33B95B0FFB2783EFF0EF9AFFEF5F938" box="[234,296,1654,1676]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="73" pageNumber="283" unit="m" value="165.0">
<elevation id="8FEEDF66FFB2783EFF0EF9AFFEF5F938" box="[234,296,1654,1676]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="73" pageNumber="283" unit="m" value="165.0">165 m</elevation>
</quantity>
,
<date id="707D1E95FFB2783EFED4F9AFFE76F93F" box="[304,427,1654,1676]" pageId="73" pageNumber="283" value="1986-07-17">
<collectingDate id="6039E77DFFB2783EFED4F9AFFE76F93F" box="[304,427,1654,1676]" pageId="73" pageNumber="283" value="1986-07-17">17 July 1986</collectingDate>
</date>
: M
<quantity id="C33B95B0FFB2783EFE29F9AFFDC8F938" box="[461,533,1654,1675]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="73" pageNumber="283" unit="mm" value="2.3">2.3 mm</quantity>
[dismembered] (MNHN- IU-5494, ex
<collectionCode id="62D2A090FFB2783EFEE1F94AFE93F91B" box="[261,334,1683,1704]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="73" pageNumber="283">MNHN</collectionCode>
Pg. 7610).
</materialsCitation>
</paragraph>
<paragraph id="047C3855FFB2783EFF6CF908FDA9F8FA" blockId="73.[136,775,1744,1865]" pageId="73" pageNumber="283">
<materialsCitation id="B4AB3208FFB2783EFF6CF908FDADF8FA" collectingDate="1986-07-19" collectionCode="MNHN" county="Ile St." elevation="160" latitude="-39.0" location="Southern Indian Ocean" municipality="Ile Amsterdam" pageId="73" pageNumber="283" specimenCount="1" typeStatus="paratype">
<typeStatus id="DB7886F7FFB2783EFF6CF908FF2EF95A" box="[136,243,1745,1769]" pageId="73" pageNumber="283" type="paratype">Paratypes</typeStatus>
,
<location id="011C6E8EFFB2783EFEE5F908FDDDF95A" LSID="urn:lsid:plazi:treatment:8C6A8943FFB27824FF1DFA04FC02FEA6:011C6E8EFFB2783EFEE5F908FDDDF95A" box="[257,512,1745,1769]" county="Ile St." latitude="-39.0" municipality="Ile Amsterdam" name="Southern Indian Ocean" pageId="73" pageNumber="283">Southern Indian Ocean</location>
,
<collectingMunicipality id="E418A22FFFB2783EFDEAF909FD6DF95A" box="[526,688,1744,1769]" pageId="73" pageNumber="283">Île Amsterdam</collectingMunicipality>
and
<collectingCounty id="ED1D40D9FFB2783EFD0FF909FF78F8BA" pageId="73" pageNumber="283">Île St.</collectingCounty>
-Paul: CENTOB, MD 50, DC 114, sta 24,
<geoCoordinate id="61F75E92FFB2783EFD7FF928FCDFF8BA" box="[667,770,1777,1801]" degrees="39" direction="south" minutes="000" orientation="latitude" pageId="73" pageNumber="283" precision="925" value="-39.0">39°000'S</geoCoordinate>
, 77°464'E,
<quantity id="C33B95B0FFB2783EFF1EF8C8FE9FF89B" box="[250,322,1809,1833]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="73" pageNumber="283" unit="m" value="160.0">
<elevation id="8FEEDF66FFB2783EFF1EF8C8FE9FF89B" box="[250,322,1809,1833]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="73" pageNumber="283" unit="m" value="160.0">160 m</elevation>
</quantity>
,
<date id="707D1E95FFB2783EFEABF8C8FE3CF89A" box="[335,481,1809,1833]" pageId="73" pageNumber="283" value="1986-07-19">
<collectingDate id="6039E77DFFB2783EFEABF8C8FE3CF89A" box="[335,481,1809,1833]" pageId="73" pageNumber="283" value="1986-07-19">19 July 1986</collectingDate>
</date>
: 2 M 2.8,
<quantity id="C33B95B0FFB2783EFDB1F8C8FD76F89B" box="[597,683,1809,1833]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="73" pageNumber="283" unit="mm" value="2.9">2.9 mm</quantity>
, 1 F
<quantity id="C33B95B0FFB2783EFD01F8C8FF6CF8FB" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="73" pageNumber="283" unit="mm" value="2.1">2.1 mm</quantity>
(MNHN-IU-5495, ex
<collectionCode id="62D2A090FFB2783EFE4DF8E8FE27F8FB" box="[425,506,1841,1864]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="73" pageNumber="283">MNHN</collectionCode>
Pg. 7609)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4CD96BDEFFB27839FF6CF8A9FDC9FAE4" lastPageId="78" lastPageNumber="288" pageId="73" pageNumber="283" type="description">
<paragraph id="047C3855FFB2783EFF6CF8A9FAD6FE86" blockId="73.[136,775,1904,2089]" lastBlockId="73.[835,1474,189,309]" pageId="73" pageNumber="283">
<emphasis id="36B7E447FFB2783EFF6CF8A9FECEF83B" bold="true" box="[136,275,1904,1928]" pageId="73" pageNumber="283">Description.</emphasis>
Gills biserial. Shield (
<figureCitation id="9CF824D0FFB2783EFDE2F8A8FDB5F83A" box="[518,616,1905,1929]" captionStart="Fig" captionStartId="74.[113,148,1975,1997]" captionTargetBox="[116,1445,306,1862]" captionTargetId="figure-9@74.[101,1472,291,1881]" captionTargetPageId="74" captionText="Fig. 42. Oncopagurus spiniartus, new species, southern Indian Ocean, Île Amsterdam and Île St.-Paul: AE, holotype, male 2.3 mm, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610); F, G, paratype, male 2.8 mm, CENTOB, MD 50, DC 114, sta 24, (MNHN-IU-5495, ex MNHN Pg 7609): A, shield and cephalic appendages; B, right antennal peduncle, lateral; C, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; D, exopod of left uropod, dorsal; E, telson, dorsal; F, left first gonopod, mesial; G, left second gonopod, anterior. Scale bars = 0.5 mm (A), 0.25 mm (BD), 0.2 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.5458460" httpUri="https://zenodo.org/record/5458460/files/figure.png" pageId="73" pageNumber="283">Fig. 42A</figureCitation>
) about as long as broad; dorsal surface weakly calcified medially (weak calcification often reaching to anterior margin), with scattered short setae; rostrum broadly rounded, weakly produced, with short mid-dorsal ridge; anterior margins weakly concave; lateral projections subtriangular, terminating in small blunt or sharp spine; anterolateral margins sloping; posterior margin broadly rounded; ventrolateral margins of shield usually with slender spine on one or both sides. Anterodistal margin of branchiostegite rounded, unarmed, setose.
</paragraph>
<paragraph id="047C3855FFB2783EFCA7FE84FAB6FE46" blockId="73.[835,1475,349,502]" pageId="73" pageNumber="283">
Ocular peduncles (
<figureCitation id="9CF824D0FFB2783EFBEEFE84FBB7FEC5" box="[1034,1130,349,374]" captionStart="Fig" captionStartId="74.[113,148,1975,1997]" captionTargetBox="[116,1445,306,1862]" captionTargetId="figure-9@74.[101,1472,291,1881]" captionTargetPageId="74" captionText="Fig. 42. Oncopagurus spiniartus, new species, southern Indian Ocean, Île Amsterdam and Île St.-Paul: AE, holotype, male 2.3 mm, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610); F, G, paratype, male 2.8 mm, CENTOB, MD 50, DC 114, sta 24, (MNHN-IU-5495, ex MNHN Pg 7609): A, shield and cephalic appendages; B, right antennal peduncle, lateral; C, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; D, exopod of left uropod, dorsal; E, telson, dorsal; F, left first gonopod, mesial; G, left second gonopod, anterior. Scale bars = 0.5 mm (A), 0.25 mm (BD), 0.2 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.5458460" httpUri="https://zenodo.org/record/5458460/files/figure.png" pageId="73" pageNumber="283">Fig. 42A</figureCitation>
) more than half length of shield; with longitudinal row of long setae dorsally; lateral and ventral faces usually weakly calcified. Cornea weakly dilated. Ocular acicles subtriangular, each terminating in strong spine; separated basally by about basal width of 1 acicle.
</paragraph>
<paragraph id="047C3855FFB2783EFCA7FDC4FA1EFD46" blockId="73.[835,1475,541,758]" pageId="73" pageNumber="283">
Antennular peduncle (
<figureCitation id="9CF824D0FFB2783EFBDAFDC4FB79FD85" box="[1086,1188,541,566]" captionStart="Fig" captionStartId="74.[113,148,1975,1997]" captionTargetBox="[116,1445,306,1862]" captionTargetId="figure-9@74.[101,1472,291,1881]" captionTargetPageId="74" captionText="Fig. 42. Oncopagurus spiniartus, new species, southern Indian Ocean, Île Amsterdam and Île St.-Paul: AE, holotype, male 2.3 mm, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610); F, G, paratype, male 2.8 mm, CENTOB, MD 50, DC 114, sta 24, (MNHN-IU-5495, ex MNHN Pg 7609): A, shield and cephalic appendages; B, right antennal peduncle, lateral; C, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; D, exopod of left uropod, dorsal; E, telson, dorsal; F, left first gonopod, mesial; G, left second gonopod, anterior. Scale bars = 0.5 mm (A), 0.25 mm (BD), 0.2 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.5458460" httpUri="https://zenodo.org/record/5458460/files/figure.png" pageId="73" pageNumber="283">Fig. 42A</figureCitation>
) exceeding distal margin of cornea by 0.7 to nearly full length or ultimate segment. Ultimate segment slightly less than twice as long as penultimate segment, naked or with scattered setae dorsally. Basal segment with strong ventromesial spine; lateral face with distal subrectangular lobe armed with small spine, and strong spine proximally. Ventral flagellum with 5 or 6 articles.
</paragraph>
<paragraph id="047C3855FFB2783EFCA7FCC4FBDDFAA6" blockId="73.[835,1474,797,1301]" pageId="73" pageNumber="283">
Antennal peduncle (
<figureCitation id="9CF824D0FFB2783EFBD5FCC4FB1CFC85" box="[1073,1217,797,822]" captionStart="Fig" captionStartId="74.[113,148,1975,1997]" captionTargetBox="[116,1445,306,1862]" captionTargetId="figure-9@74.[101,1472,291,1881]" captionTargetPageId="74" captionText="Fig. 42. Oncopagurus spiniartus, new species, southern Indian Ocean, Île Amsterdam and Île St.-Paul: AE, holotype, male 2.3 mm, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610); F, G, paratype, male 2.8 mm, CENTOB, MD 50, DC 114, sta 24, (MNHN-IU-5495, ex MNHN Pg 7609): A, shield and cephalic appendages; B, right antennal peduncle, lateral; C, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; D, exopod of left uropod, dorsal; E, telson, dorsal; F, left first gonopod, mesial; G, left second gonopod, anterior. Scale bars = 0.5 mm (A), 0.25 mm (BD), 0.2 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.5458460" httpUri="https://zenodo.org/record/5458460/files/figure.png" pageId="73" pageNumber="283">Fig. 42A, B</figureCitation>
) not exceeding distal margin of cornea, reaching at most to about midportion of cornea. Fifth segment unarmed except for scattered setae and laterodistal tuft of setae. Fourth segment armed with strong dorsodistal spine. Third segment with strong ventromesial distal spine. Second segment with dorsolateral distal angle produced, terminating in strong, simple spine often with smaller dorsal subterminal spine; mesial margin with spine on dorsodistal angle. First segment with lateral face usually armed with small spine; ventromesial angle produced, with 2 or 3 small spines laterally. Acicle relatively short, reaching to about proximal margin of cornea, nearly straight (in dorsal view), terminating in strong spine; mesial margin armed with 610 spines, sparsely setose. Flagellum slightly exceeding extended right cheliped, articles with setae 1 or 2 flagellar articles in length.
</paragraph>
<paragraph id="047C3855FFB2783EFCA7FAE4FAFBF8A6" blockId="73.[835,1475,1341,1813]" pageId="73" pageNumber="283">
Mandible (
<figureCitation id="9CF824D0FFB2783EFC5FFAE4FBC2FAE5" box="[955,1055,1341,1366]" captionStart="Fig" captionStartId="75.[136,171,2035,2057]" captionTargetBox="[140,1460,932,1997]" captionTargetId="figure-7@75.[130,1475,922,2007]" captionTargetPageId="75" captionText="Fig. 43. Oncopagurus spiniartus, new species, paratype, male 2.8 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 114, sta 24 (MNHN-IU-5495, ex MNHN Pg 7609). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.2 mm (AE); 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458462" httpUri="https://zenodo.org/record/5458462/files/figure.png" pageId="73" pageNumber="283">Fig. 43A</figureCitation>
) with 3-segmented palp; cutting edge calcified, with small tooth medially; molar process with small tooth medially. Maxillule (
<figureCitation id="9CF824D0FFB2783EFB7EFAA4FB2AFA25" box="[1178,1271,1405,1430]" captionStart="Fig" captionStartId="75.[136,171,2035,2057]" captionTargetBox="[140,1460,932,1997]" captionTargetId="figure-7@75.[130,1475,922,2007]" captionTargetPageId="75" captionText="Fig. 43. Oncopagurus spiniartus, new species, paratype, male 2.8 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 114, sta 24 (MNHN-IU-5495, ex MNHN Pg 7609). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.2 mm (AE); 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458462" httpUri="https://zenodo.org/record/5458462/files/figure.png" pageId="73" pageNumber="283">Fig. 43B</figureCitation>
) with external lobe of endopod slender, moderately developed, not recurved, internal lobe with long, terminal seta. Maxilla (
<figureCitation id="9CF824D0FFB2783EFACEFA64FA5BFA65" box="[1322,1414,1469,1494]" captionStart="Fig" captionStartId="75.[136,171,2035,2057]" captionTargetBox="[140,1460,932,1997]" captionTargetId="figure-7@75.[130,1475,922,2007]" captionTargetPageId="75" captionText="Fig. 43. Oncopagurus spiniartus, new species, paratype, male 2.8 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 114, sta 24 (MNHN-IU-5495, ex MNHN Pg 7609). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.2 mm (AE); 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458462" httpUri="https://zenodo.org/record/5458462/files/figure.png" pageId="73" pageNumber="283">Fig. 43C</figureCitation>
) with endopod not exceeding distal margin of scaphognathite. First maxilliped (
<figureCitation id="9CF824D0FFB2783EFC22FA24FBF4F9A5" box="[966,1065,1533,1558]" captionStart="Fig" captionStartId="75.[136,171,2035,2057]" captionTargetBox="[140,1460,932,1997]" captionTargetId="figure-7@75.[130,1475,922,2007]" captionTargetPageId="75" captionText="Fig. 43. Oncopagurus spiniartus, new species, paratype, male 2.8 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 114, sta 24 (MNHN-IU-5495, ex MNHN Pg 7609). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.2 mm (AE); 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458462" httpUri="https://zenodo.org/record/5458462/files/figure.png" pageId="73" pageNumber="283">Fig. 43D</figureCitation>
) with endopod not exceeding exopod in distal extension. Second maxilliped (
<figureCitation id="9CF824D0FFB2783EFB18F9C4FA83F985" box="[1276,1374,1565,1590]" captionStart="Fig" captionStartId="75.[136,171,2035,2057]" captionTargetBox="[140,1460,932,1997]" captionTargetId="figure-7@75.[130,1475,922,2007]" captionTargetPageId="75" captionText="Fig. 43. Oncopagurus spiniartus, new species, paratype, male 2.8 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 114, sta 24 (MNHN-IU-5495, ex MNHN Pg 7609). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.2 mm (AE); 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458462" httpUri="https://zenodo.org/record/5458462/files/figure.png" pageId="73" pageNumber="283">Fig. 43E</figureCitation>
) without distinguishing characters. Third maxilliped (
<figureCitation id="9CF824D0FFB2783EFAC0F9E4FA5EF9E5" box="[1316,1411,1597,1622]" captionStart="Fig" captionStartId="75.[136,171,2035,2057]" captionTargetBox="[140,1460,932,1997]" captionTargetId="figure-7@75.[130,1475,922,2007]" captionTargetPageId="75" captionText="Fig. 43. Oncopagurus spiniartus, new species, paratype, male 2.8 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 114, sta 24 (MNHN-IU-5495, ex MNHN Pg 7609). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.2 mm (AE); 0.5 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458462" httpUri="https://zenodo.org/record/5458462/files/figure.png" pageId="73" pageNumber="283">Fig. 43F</figureCitation>
) with merus to dactyl each distinctly longer than broad, ischium about twice as long as broad; crista dentata with about 7 sharp teeth diminishing in strength distally, proximal tooth distinctly larger than others; basis with 1 small mesial spine; coxa lacking spine. Sternite of third maxillipeds unarmed or with small spine on each side of midline.
</paragraph>
<paragraph id="047C3855FFB2783CFCA7F8E4FAF3FEE6" blockId="73.[835,1474,1853,2101]" lastBlockId="75.[834,1474,189,342]" lastPageId="75" lastPageNumber="285" pageId="73" pageNumber="283">
Chelipeds markedly dissimilar. Right cheliped (
<figureCitation id="9CF824D0FFB2783EFAB3F8E4FC88F8C5" captionStart="Fig" captionStartId="76.[113,148,2035,2057]" captionTargetBox="[119,1448,599,2005]" captionTargetId="figure-9@76.[91,1467,554,2009]" captionTargetPageId="76" captionText="Fig. 44. Oncopagurus spiniartus, new species, holotype, male 2.3 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610): A, right cheliped, dorsal; B, chela of same, mesial; C, left cheliped, dorsal; D, same, mesial. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5458464" httpUri="https://zenodo.org/record/5458464/files/figure.png" pageId="73" pageNumber="283">Fig. 44A, B</figureCitation>
) massive, chela operculate; carpus and chela with sparse to moderately dense setae on dorsal surfaces, setae more dense on dorsal surfaces of fingers and mesial face of carpus. Fingers curved ventromesially, each terminating in small, usually blunt corneous claw; cutting edges with 1 or 2 large calcareous teeth and several irregularly sized smaller calcareous teeth, with short row closely set minute corneous teeth distally. Dactyl about 1.3 times as long as mesial margin of palm, set at strongly oblique angle to longitudinal axis of palm; mesial margin broadly curved, well delimited by row of spines diminishing in size distally; dorsal face with moderately dense small spines or tubercles decreasing in density distally; ventromesial face concave. Fixed finger broad at base, dorsal face with scattered small spines or tubercles, lateral margin well delimited by row of spines; ventrolateral face concave. Palm about 1.51.7 times as broad as long; dorsal surface with numerous small spines or tubercles; dorsolateral margin broadly rounded, well delimited by row of spines, dorsomesial margin delimited by row of spines or tubercles; dorsomesial face rounded, with small spines or tubercles and usually well delimited, blunt ventromesial margin; ventral surface smooth, at most with scattered short setae. Carpus longer than broad; dorsolateral margin usually well delimited by row of spines; dorsodistal margin with row of small spines; dorsal face with numerous small spines or tubercles; ventral face nearly flat, with scattered small tubercles or spines. Merus with scattered setae dorsally, and moderately dense setae ventrodistally; with scattered small tubercles on dorsal face and small dorsodistal spine; ventromesial margin with row of blunt to sharp spines; ventrolateral margin with row of small tubercles. Ischium with ventromesial row of small, blunt spines. Coxa with ventromesial row of setae.
</paragraph>
<caption id="50BC68DDFFB1783DFF95F86EFB71F7F3" ID-DOI="http://doi.org/10.5281/zenodo.5458460" ID-Zenodo-Dep="5458460" httpUri="https://zenodo.org/record/5458460/files/figure.png" pageId="74" pageNumber="284" startId="74.[113,148,1975,1997]" targetBox="[116,1445,306,1862]" targetPageId="74">
<paragraph id="047C3855FFB1783DFF95F86EFB71F7F3" blockId="74.[113,1452,1974,2112]" pageId="74" pageNumber="284">
Fig. 42.
<taxonomicName id="C3C343D6FFB1783DFF2CF860FE6BF87E" authority="Lemaitre, 2014" authorityName="Lemaitre" authorityYear="2014" box="[200,438,1977,1997]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="74" pageNumber="284" phylum="Arthropoda" rank="species" species="spiniartus" status="sp. nov.">
<emphasis id="36B7E447FFB1783DFF2CF860FE6BF87E" box="[200,438,1977,1997]" italics="true" pageId="74" pageNumber="284">Oncopagurus spiniartus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2D84593CFFB1783DFE21F86EFDE3F87E" box="[453,574,1975,1997]" pageId="74" pageNumber="284" rank="species">new species</taxonomicNameLabel>
, southern Indian Ocean, Île Amsterdam and Île St.-Paul: AE, holotype, male 2.3 mm, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610); F, G, paratype, male 2.8 mm, CENTOB, MD 50, DC 114, sta 24, (MNHN-IU-5495, ex MNHN Pg 7609): A, shield and cephalic appendages; B, right antennal peduncle, lateral; C, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; D, exopod of left uropod, dorsal; E, telson, dorsal; F, left first gonopod, mesial; G, left second gonopod, anterior. Scale bars = 0.5 mm (A), 0.25 mm (BD), 0.2 mm (F, G).
</paragraph>
</caption>
<paragraph id="047C3855FFB0783CFCA6FEA4FAEFFCE6" blockId="75.[834,1475,381,853]" pageId="75" pageNumber="285">
Left cheliped (
<figureCitation id="9CF824D0FFB0783CFC08FEA4FBAAFE25" box="[1004,1143,381,406]" captionStart="Fig" captionStartId="76.[113,148,2035,2057]" captionTargetBox="[119,1448,599,2005]" captionTargetId="figure-9@76.[91,1467,554,2009]" captionTargetPageId="76" captionText="Fig. 44. Oncopagurus spiniartus, new species, holotype, male 2.3 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610): A, right cheliped, dorsal; B, chela of same, mesial; C, left cheliped, dorsal; D, same, mesial. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5458464" httpUri="https://zenodo.org/record/5458464/files/figure.png" pageId="75" pageNumber="285">Fig. 44C, D</figureCitation>
) usually weakly calcified on dorsolateral face of carpus. Fingers each terminating in small corneous claw; dorsal and ventral surfaces unarmed except for scattered tufts of setae; cutting edge of dactyl with row of minute, fused corneous teeth; cutting edge of fixed finger with row of well-spaced small calcareous teeth. Dactyl longer than length of mesial margin of palm. Palm unarmed except for dorsomesial row of small sharp or blunt spines and scattered tufts of setae; ventral face smooth. Carpus with irregular dorsal row of about 9 small spines in addition to 1 dorsodistal and 1 dorsolateral spine on distal margin; dorsal surface with long setae; ventral face smooth. Merus with long setae on dorsal margin; with ventrolateral row of small spines; ventral face smooth. Ischium and coxa unarmed, but with ventromesial row of setae.
</paragraph>
<caption id="50BC68DDFFB0783CFF6CF82AFB09F7F0" ID-DOI="http://doi.org/10.5281/zenodo.5458462" ID-Zenodo-Dep="5458462" httpUri="https://zenodo.org/record/5458462/files/figure.png" pageId="75" pageNumber="285" startId="75.[136,171,2035,2057]" targetBox="[140,1460,932,1997]" targetPageId="75">
<paragraph id="047C3855FFB0783CFF6CF82AFB09F7F0" blockId="75.[136,1475,2034,2115]" pageId="75" pageNumber="285">
Fig. 43.
<taxonomicName id="C3C343D6FFB0783CFF3EF82CFE1EF7BA" authority="Lemaitre, 2014" authorityName="Lemaitre" authorityYear="2014" box="[218,451,2037,2057]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="75" pageNumber="285" phylum="Arthropoda" rank="species" species="spiniartus" status="sp. nov.">
<emphasis id="36B7E447FFB0783CFF3EF82CFE1EF7BA" box="[218,451,2037,2057]" italics="true" pageId="75" pageNumber="285">Oncopagurus spiniartus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2D84593CFFB0783CFE34F82DFD9AF7BA" box="[464,583,2035,2057]" pageId="75" pageNumber="285" rank="species">new species</taxonomicNameLabel>
, paratype, male 2.8 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 114, sta 24 (MNHN-IU-5495, ex MNHN Pg 7609). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.2 mm (AE); 0.5 mm (F).
</paragraph>
</caption>
<paragraph id="047C3855FFB7783BFF95FF66FBD6FEE4" blockId="76.[113,752,191,535]" lastBlockId="76.[812,1452,191,343]" pageId="76" pageNumber="286">
Ambulatory legs (
<figureCitation id="9CF824D0FFB7783BFED5FF66FE6CFF64" box="[305,433,191,215]" captionStart="Fig" captionStartId="77.[136,171,2035,2057]" captionTargetBox="[135,1459,222,1997]" captionTargetId="figure-7@77.[125,1474,209,2005]" captionTargetPageId="77" captionText="Fig. 45. Oncopagurus spiniartus, new species, holotype, male 2.3 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610): A, right first ambulatory leg, mesial; B, same, lateral; C, right second ambulatory leg, lateral, D, dactyl of same, mesial. Scale bars = 1 mm (AD); 0. 5 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.5458466" httpUri="https://zenodo.org/record/5458466/files/figure.png" pageId="76" pageNumber="286">Fig. 45AD</figureCitation>
) similar right from left except for longer segments and stronger spination on right, extending to about tip of dactyls of right cheliped or slightly exceeding them. Dactyls broadly curved, about 1.5 times as long as propodi, and terminating in sharp corneous claws; each with dorsal and dorsomesial distal rows of long setae, and 1013 small spinules on ventromesial margin. Propodi each with row of about 8 small spines (reduced or obscure on right side) and sparse setae on dorsal margin; ventral margin with few setae. Carpi each with row of 9 or 10 small spines and few setae on dorsal margin, and small dorsodistal spine. Meri each with row of 9 or 10 small spines and long setae on dorsal margin. Ischia with setae dorsally and ventrally. Coxae with ventromesial row of setae. Anterior lobe of sternite XII (between third pereopods,
<figureCitation id="9CF824D0FFB7783BFB00FEC6FA96FE84" box="[1252,1355,287,311]" captionStart="Fig" captionStartId="74.[113,148,1975,1997]" captionTargetBox="[116,1445,306,1862]" captionTargetId="figure-9@74.[101,1472,291,1881]" captionTargetPageId="74" captionText="Fig. 42. Oncopagurus spiniartus, new species, southern Indian Ocean, Île Amsterdam and Île St.-Paul: AE, holotype, male 2.3 mm, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610); F, G, paratype, male 2.8 mm, CENTOB, MD 50, DC 114, sta 24, (MNHN-IU-5495, ex MNHN Pg 7609): A, shield and cephalic appendages; B, right antennal peduncle, lateral; C, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; D, exopod of left uropod, dorsal; E, telson, dorsal; F, left first gonopod, mesial; G, left second gonopod, anterior. Scale bars = 0.5 mm (A), 0.25 mm (BD), 0.2 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.5458460" httpUri="https://zenodo.org/record/5458460/files/figure.png" pageId="76" pageNumber="286">Fig. 42C</figureCitation>
), setose, with subdistal spine.
</paragraph>
<paragraph id="047C3855FFB77839FCC8FEA6FEEAFEA5" blockId="76.[812,1451,383,535]" lastBlockId="78.[113,753,191,279]" lastPageId="78" lastPageNumber="288" pageId="76" pageNumber="286">
Fourth pereopod (
<figureCitation id="9CF824D0FFB7783BFC0FFEA6FB94FE24" box="[1003,1097,383,407]" captionStart="Fig" captionStartId="77.[136,171,2035,2057]" captionTargetBox="[135,1459,222,1997]" captionTargetId="figure-7@77.[125,1474,209,2005]" captionTargetPageId="77" captionText="Fig. 45. Oncopagurus spiniartus, new species, holotype, male 2.3 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610): A, right first ambulatory leg, mesial; B, same, lateral; C, right second ambulatory leg, lateral, D, dactyl of same, mesial. Scale bars = 1 mm (AD); 0. 5 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.5458466" httpUri="https://zenodo.org/record/5458466/files/figure.png" pageId="76" pageNumber="286">Fig. 45E</figureCitation>
) semichelate. Dactyl terminating in sharp corneous claw; with ventrolateral row of small corneous spinules. Propodus longer than broad, rasp with 1 row of rounded scales at least distally. Carpus with long setae on dorsal margin. Merus with rows of long setae on dorsal and ventral margins. Coxa with anteroventral row of setae. Anterior lobe of sternite XIII (between fourth pereopods) with row of setae.
</paragraph>
<caption id="50BC68DDFFB7783BFF95F82AFE09F7F1" ID-DOI="http://doi.org/10.5281/zenodo.5458464" ID-Zenodo-Dep="5458464" httpUri="https://zenodo.org/record/5458464/files/figure.png" pageId="76" pageNumber="286" startId="76.[113,148,2035,2057]" targetBox="[119,1448,599,2005]" targetPageId="76">
<paragraph id="047C3855FFB7783BFF95F82AFE09F7F1" blockId="76.[113,1452,2034,2115]" pageId="76" pageNumber="286">
Fig. 44.
<taxonomicName id="C3C343D6FFB7783BFF27F82CFE71F7BA" authority="Lemaitre, 2014" authorityName="Lemaitre" authorityYear="2014" box="[195,428,2037,2057]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="76" pageNumber="286" phylum="Arthropoda" rank="species" species="spiniartus" status="sp. nov.">
<emphasis id="36B7E447FFB7783BFF27F82CFE71F7BA" box="[195,428,2037,2057]" italics="true" pageId="76" pageNumber="286">Oncopagurus spiniartus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2D84593CFFB7783BFE5DF82DFDF2F7BA" box="[441,559,2035,2057]" pageId="76" pageNumber="286" rank="species">new species</taxonomicNameLabel>
, holotype, male 2.3 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610): A, right cheliped, dorsal; B, chela of same, mesial; C, left cheliped, dorsal; D, same, mesial. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="50BC68DDFFB6783AFF6CF82AFBF8F7F1" ID-DOI="http://doi.org/10.5281/zenodo.5458466" ID-Zenodo-Dep="5458466" httpUri="https://zenodo.org/record/5458466/files/figure.png" pageId="77" pageNumber="287" startId="77.[136,171,2035,2057]" targetBox="[135,1459,222,1997]" targetPageId="77">
<paragraph id="047C3855FFB6783AFF6CF82AFBF8F7F1" blockId="77.[136,1475,2034,2115]" pageId="77" pageNumber="287">
Fig. 45.
<taxonomicName id="C3C343D6FFB6783AFF3DF82CFE1EF7BA" authority="Lemaitre, 2014" authorityName="Lemaitre" authorityYear="2014" box="[217,451,2037,2057]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="77" pageNumber="287" phylum="Arthropoda" rank="species" species="spiniartus" status="sp. nov.">
<emphasis id="36B7E447FFB6783AFF3DF82CFE1EF7BA" box="[217,451,2037,2057]" italics="true" pageId="77" pageNumber="287">Oncopagurus spiniartus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2D84593CFFB6783AFE2BF82DFD9BF7BA" box="[463,582,2035,2057]" pageId="77" pageNumber="287" rank="species">new species</taxonomicNameLabel>
, holotype, male 2.3 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610): A, right first ambulatory leg, mesial; B, same, lateral; C, right second ambulatory leg, lateral, D, dactyl of same, mesial. Scale bars = 1 mm (AD); 0. 5 mm (E, F).
</paragraph>
</caption>
<paragraph id="047C3855FFB57839FF95FEE6FD3EFE05" blockId="78.[113,752,319,439]" pageId="78" pageNumber="288">
Fifth pereopod (
<figureCitation id="9CF824D0FFB57839FEC3FEE6FE55FEE4" box="[295,392,319,343]" captionStart="Fig" captionStartId="77.[136,171,2035,2057]" captionTargetBox="[135,1459,222,1997]" captionTargetId="figure-7@77.[125,1474,209,2005]" captionTargetPageId="77" captionText="Fig. 45. Oncopagurus spiniartus, new species, holotype, male 2.3 mm, southern Indian Ocean, Île Amsterdam and Île St.-Paul, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610): A, right first ambulatory leg, mesial; B, same, lateral; C, right second ambulatory leg, lateral, D, dactyl of same, mesial. Scale bars = 1 mm (AD); 0. 5 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.5458466" httpUri="https://zenodo.org/record/5458466/files/figure.png" pageId="78" pageNumber="288">Fig. 45F</figureCitation>
) semichelate. Propodal rasp not reaching midlength of segment. Coxa with row of long setae on anteroventral and ventrodistal margins. Anterior lobe of sternite XIII (between fifth pereopods) with row of setae.
</paragraph>
<paragraph id="047C3855FFB57839FF95FE06FD80FD44" blockId="78.[113,753,479,759]" pageId="78" pageNumber="288">
Uropods markedly asymmetrical; left exopod (
<figureCitation id="9CF824D0FFB57839FD66FE06FD34FE44" box="[642,745,479,503]" captionStart="Fig" captionStartId="74.[113,148,1975,1997]" captionTargetBox="[116,1445,306,1862]" captionTargetId="figure-9@74.[101,1472,291,1881]" captionTargetPageId="74" captionText="Fig. 42. Oncopagurus spiniartus, new species, southern Indian Ocean, Île Amsterdam and Île St.-Paul: AE, holotype, male 2.3 mm, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610); F, G, paratype, male 2.8 mm, CENTOB, MD 50, DC 114, sta 24, (MNHN-IU-5495, ex MNHN Pg 7609): A, shield and cephalic appendages; B, right antennal peduncle, lateral; C, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; D, exopod of left uropod, dorsal; E, telson, dorsal; F, left first gonopod, mesial; G, left second gonopod, anterior. Scale bars = 0.5 mm (A), 0.25 mm (BD), 0.2 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.5458460" httpUri="https://zenodo.org/record/5458460/files/figure.png" pageId="78" pageNumber="288">Fig. 42D</figureCitation>
) slender, slightly more than 3 times as long as broad, rasp narrow, consisting of 14 rows of small scales. Telson (
<figureCitation id="9CF824D0FFB57839FD21FDC6FF43FDE4" captionStart="Fig" captionStartId="74.[113,148,1975,1997]" captionTargetBox="[116,1445,306,1862]" captionTargetId="figure-9@74.[101,1472,291,1881]" captionTargetPageId="74" captionText="Fig. 42. Oncopagurus spiniartus, new species, southern Indian Ocean, Île Amsterdam and Île St.-Paul: AE, holotype, male 2.3 mm, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610); F, G, paratype, male 2.8 mm, CENTOB, MD 50, DC 114, sta 24, (MNHN-IU-5495, ex MNHN Pg 7609): A, shield and cephalic appendages; B, right antennal peduncle, lateral; C, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; D, exopod of left uropod, dorsal; E, telson, dorsal; F, left first gonopod, mesial; G, left second gonopod, anterior. Scale bars = 0.5 mm (A), 0.25 mm (BD), 0.2 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.5458460" httpUri="https://zenodo.org/record/5458460/files/figure.png" pageId="78" pageNumber="288">Fig. 42E</figureCitation>
) moderately asymmetrical, lacking transverse suture; dorsal surface with scattered setae; lateral margins with long setae medially; posterior lobes separated by shallow unarmed, U-shaped cleft; terminal margin of left lobe armed with about 10, often curved corneous spines, terminal margin of right lobe armed with 4 or 5 small spines.
</paragraph>
<paragraph id="047C3855FFB57839FF95FCC6FD2CFC64" blockId="78.[113,753,799,983]" pageId="78" pageNumber="288">
<collectingRegion id="C607F6B7FFB57839FF95FCC6FF75FC84" box="[113,168,799,823]" country="Maldives" name="Maale" pageId="78" pageNumber="288">Male</collectingRegion>
with paired first and second gonopods (
<figureCitation id="9CF824D0FFB57839FD83FCC6FD35FC84" box="[615,744,799,823]" captionStart="Fig" captionStartId="74.[113,148,1975,1997]" captionTargetBox="[116,1445,306,1862]" captionTargetId="figure-9@74.[101,1472,291,1881]" captionTargetPageId="74" captionText="Fig. 42. Oncopagurus spiniartus, new species, southern Indian Ocean, Île Amsterdam and Île St.-Paul: AE, holotype, male 2.3 mm, CENTOB, MD 50, DC 82, sta 19 (MNHN-IU-5494, ex MNHN Pg 7610); F, G, paratype, male 2.8 mm, CENTOB, MD 50, DC 114, sta 24, (MNHN-IU-5495, ex MNHN Pg 7609): A, shield and cephalic appendages; B, right antennal peduncle, lateral; C, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; D, exopod of left uropod, dorsal; E, telson, dorsal; F, left first gonopod, mesial; G, left second gonopod, anterior. Scale bars = 0.5 mm (A), 0.25 mm (BD), 0.2 mm (F, G)." figureDoi="http://doi.org/10.5281/zenodo.5458460" httpUri="https://zenodo.org/record/5458460/files/figure.png" pageId="78" pageNumber="288">Fig. 42F, G</figureCitation>
) occasionally with first pair missing and poorly developed second pair; first gonopod with distal portion subovate, with setae distally; second gonopod with distal segment flat and with long setae distally, proximal segment with long setae distomesially. Females with vestigial second right pleopod.
</paragraph>
<paragraph id="047C3855FFB57839FF95FC26FF3BFB64" blockId="78.[113,753,1023,1239]" pageId="78" pageNumber="288">
<emphasis id="36B7E447FFB57839FF95FC26FF2CFBA4" bold="true" box="[113,241,1023,1047]" pageId="78" pageNumber="288">Variations.</emphasis>
With only four known specimens, variations cannot be properly ascertained. However, it appears that in males, gonopods can vary in development irrespective of whether the individuals are mature or not. One of the males (
<typeStatus id="DB7886F7FFB57839FF9DFBA6FF01FB24" box="[121,220,1151,1175]" pageId="78" pageNumber="288" type="holotype">holotype</typeStatus>
), despite being an adult, lacks first gonopods, and the second gonopods are incompletely and asymmetrically developed.
</paragraph>
<paragraph id="047C3855FFB57839FF95FB26FEA6FAA4" blockId="78.[113,379,1279,1303]" box="[113,379,1279,1303]" pageId="78" pageNumber="288">
<emphasis id="36B7E447FFB57839FF95FB26FEDFFAA4" bold="true" box="[113,258,1279,1303]" pageId="78" pageNumber="288">Colouration.</emphasis>
Unknown.
</paragraph>
<paragraph id="047C3855FFB57839FF95FAE6FDC9FAE4" blockId="78.[113,532,1343,1367]" box="[113,532,1343,1367]" pageId="78" pageNumber="288">
<emphasis id="36B7E447FFB57839FF95FAE6FF0DFAE4" bold="true" box="[113,208,1343,1367]" pageId="78" pageNumber="288">Habitat.</emphasis>
Presumably gastropod shells.
</paragraph>
</subSubSection>
<subSubSection id="4CD96BDEFFB57839FF95FAA6FD39FA05" pageId="78" pageNumber="288" type="distribution">
<paragraph id="047C3855FFB57839FF95FAA6FD39FA05" blockId="78.[113,753,1406,1463]" pageId="78" pageNumber="288">
<emphasis id="36B7E447FFB57839FF95FAA6FED9FA24" bold="true" box="[113,260,1407,1431]" pageId="78" pageNumber="288">Distribution.</emphasis>
So far known only from Île Amsterdam and Île St.-Paul, in the southern Indian Ocean. Depth:
<specimenCount id="12C5F3DCFFB57839FD73FA46FD39FA05" box="[663,740,1439,1463]" pageId="78" pageNumber="288" type="generic">
<quantity id="C33B95B0FFB57839FD73FA46FD3DFA05" box="[663,736,1439,1463]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="78" pageNumber="288" unit="m" value="165.0">165 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="4CD96BDEFFB57839FF95FA06FF31F804" pageId="78" pageNumber="288" type="discussion">
<paragraph id="047C3855FFB57839FF95FA06FE80F964" blockId="78.[113,753,1503,1751]" pageId="78" pageNumber="288">
<emphasis id="36B7E447FFB57839FF95FA06FF3DFA44" bold="true" box="[113,224,1503,1527]" pageId="78" pageNumber="288">Remarks.</emphasis>
This new species is characterised by the spinose condition of the left cheliped and ambulatory legs. The carpus of the left cheliped has a row of nine small spines on the dorsal margin; the meri, carpi and propodi of the ambulatory legs (both sides), each have a row of nine small spines on the dorsal margin. Also, the ambulatory legs are not as slender as in other
<taxonomicName id="C3C343D6FFB57839FF0EF978FE0EF904" box="[234,467,1695,1719]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" kingdom="Animalia" order="Decapoda" pageId="78" pageNumber="288" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="36B7E447FFB57839FF0EF978FEA6F904" box="[234,379,1697,1719]" italics="true" pageId="78" pageNumber="288">Oncopagurus</emphasis>
species
</taxonomicName>
; this is particularly visible in the meri and carpi.
</paragraph>
<paragraph id="047C3855FFB57839FF95F926FF31F804" blockId="78.[113,752,1791,1975]" pageId="78" pageNumber="288">Not many samples of parapagurids are known from the isolated islands at higher latitudes of the southern Indian Ocean. The discovery of this new species in the only known samples from Île Amsterdam and Île St.-Paul, suggests that additional sampling could yield even more species form this region.</paragraph>
</subSubSection>
<subSubSection id="4CD96BDEFFB57839FF95F806FB1CFEE4" pageId="78" pageNumber="288" type="etymology">
<paragraph id="047C3855FFB57839FF95F806FD2DF7A4" blockId="78.[113,752,2015,2071]" pageId="78" pageNumber="288">
<emphasis id="36B7E447FFB57839FF95F806FF2EF844" bold="true" box="[113,243,2015,2039]" pageId="78" pageNumber="288">Etymology.</emphasis>
The specific name is derived from the Latin,
<emphasis id="36B7E447FFB57839FF95F7D8FF70F7A4" box="[113,173,2049,2071]" italics="true" pageId="78" pageNumber="288">spina</emphasis>
, spine, and
<emphasis id="36B7E447FFB57839FEDAF7D8FEAAF7A4" box="[318,375,2049,2071]" italics="true" pageId="78" pageNumber="288">artus</emphasis>
, limb, and is in reference to the
</paragraph>
<paragraph id="047C3855FFB57839FCC9FF66FBEBFF44" blockId="78.[813,1451,191,247]" pageId="78" pageNumber="288">spinose condition of the carpi and meri of the ambulatory legs in this new species.</paragraph>
<paragraph id="047C3855FFB57839FC72FEC6FB1CFEE4" blockId="78.[918,1346,287,343]" pageId="78" pageNumber="288">
<taxonomicName id="C3C343D6FFB57839FC72FEC6FA9FFE84" authority="Zhadan, 1997" authorityName="Zhadan" authorityYear="1997" box="[918,1346,287,311]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="78" pageNumber="288" phylum="Arthropoda" rank="species" species="stockmani">
<emphasis id="36B7E447FFB57839FC72FEC6FB78FE84" bold="true" box="[918,1189,287,311]" italics="true" pageId="78" pageNumber="288">Oncopagurus stockmani</emphasis>
<bibRefCitation id="605245A4FFB57839FB49FEC6FA9FFE84" author="Zhadan DG" box="[1197,1346,287,311]" pageId="78" pageNumber="288" pagination="55 - 79" refId="ref73241" refString="Zhadan DG (1997) Deep-sea hermit crabs from the submerged ridges Nazka and Sala-y-Gomez, southeastern Pacific (Decapoda Anomura Parapaguridae). Arthropoda Selecta, 6 (1 / 2): 55 - 79." type="journal article" year="1997">Zhadan, 1997</bibRefCitation>
</taxonomicName>
<figureCitation id="9CF824D0FFB57839FBF3FEE6FB44FEE4" box="[1047,1177,319,343]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="79.[136,171,1976,1998]" captionStartId-1="80.[113,148,2032,2054]" captionStartId-2="81.[138,173,2033,2055]" captionStartId-3="82.[113,148,2005,2027]" captionTargetBox-0="[142,1460,715,1943]" captionTargetBox-1="[120,1447,559,1991]" captionTargetBox-2="[140,1466,951,1993]" captionTargetBox-3="[105,1456,494,1965]" captionTargetId-0="figure-7@79.[130,1468,691,1950]" captionTargetId-1="figure-9@80.[110,1460,539,2011]" captionTargetId-2="figure-7@81.[130,1488,941,2010]" captionTargetId-3="figure-9@82.[95,1471,484,1974]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionTargetPageId-2="81" captionTargetPageId-3="82" captionText-0="Fig. 46. Oncopagurus stockmani Zhadan, 1997: AD, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233); E, F, holotype, male 6.1 mm, southeastern Pacific, Nazca and Sala-y-Gómez ridges, “Professor Stockman”, 18th cruise, sta 1873 (ZMUM Ma 4758). A, shield and cephalic appendages, dorsal; B, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; C, exopod of left uropod, dorsal; D, telson, dorsal; E, left first gonopod, mesial; F, left second gonopod, anterior. Scale bars = 2 mm (A); 1 mm (BE); 2 mm (F)." captionText-1="Fig. 47. Oncopagurus stockmani Zhadan, 1997: ov female 4.0 mm, eastern Pacific, Robinsoe Crusoe Island, Juan Fernández Islands, CIMAR 6, sta 38 (USNM 1211234). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.5 mm." captionText-2="Fig. 48. Oncopagurus stockmani Zhadan, 1997, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233): A, right cheliped, dorsal; BD, chela of same, ventral (B), lateral (C), mesial (D); E, left cheliped, dorsal; F, carpus of same, dorsolateral. Scale bars = 5 mm." captionText-3="Fig. 49. Oncopagurus stockmani Zhadan, 1997, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233): A, right first ambulatory leg, lateral; B, dactyl of same, mesial; C, right second ambulatory leg, lateral; D, dactyl of same, mesial; E, propodus and dactyl of left fourth pereopod, lateral; F, propodus and dactyl of left fifth pereopod, lateral. Scale bars = 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5458468" figureDoi-1="http://doi.org/10.5281/zenodo.5458470" figureDoi-2="http://doi.org/10.5281/zenodo.5458472" figureDoi-3="http://doi.org/10.5281/zenodo.5458474" httpUri-0="https://zenodo.org/record/5458468/files/figure.png" httpUri-1="https://zenodo.org/record/5458470/files/figure.png" httpUri-2="https://zenodo.org/record/5458472/files/figure.png" httpUri-3="https://zenodo.org/record/5458474/files/figure.png" pageId="78" pageNumber="288">Figs. 4649</figureCitation>
,
<figureCitation id="9CF824D0FFB57839FB42FEE6FB1CFEE4" box="[1190,1217,319,343]" captionStart="Fig" captionStartId="85.[136,171,2092,2114]" captionTargetBox="[139,1474,1084,2054]" captionTargetId="figure-224@85.[139,1476,1083,2054]" captionTargetPageId="85" captionText="Fig. 51. Bathymetric distribution of Oncopagurus species from the world, with maximum and minimum depths (meters) indicated for each." figureDoi="http://doi.org/10.5281/zenodo.5458478" httpUri="https://zenodo.org/record/5458478/files/figure.png" pageId="78" pageNumber="288">51</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4CD96BDEFFB57839FCCAFE59FAF2FD90" pageId="78" pageNumber="288" type="reference_group">
<paragraph id="047C3855FFB57839FCCAFE59FADAFE20" blockId="78.[813,1452,382,548]" box="[814,1287,382,404]" pageId="78" pageNumber="288">
<treatmentCitationGroup id="24D31F7BFFB57839FCCAFE59FADAFE20" box="[814,1287,382,404]" pageId="78" pageNumber="288">
<taxonomicName id="C3C343D6FFB57839FCCAFE59FBC1FE27" ID-CoL="40653cf0-7dba-4d39-ab5e-20ebbade44dc" baseAuthorityName="Studer" baseAuthorityYear="1883" box="[814,1052,384,404]" class="Malacostraca" family="Parapaguridae" genus="Parapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="78" pageNumber="288" phylum="Arthropoda" rank="species" species="dimorphus">
<emphasis id="36B7E447FFB57839FCCAFE59FBC1FE27" box="[814,1052,384,404]" italics="true" pageId="78" pageNumber="288">Parapagurus dimorphus</emphasis>
</taxonomicName>
<treatmentCitation id="85621E44FFB57839FBD1FEA7FADEFE20" author="Zarenkov NA" box="[1077,1283,382,404]" page="238" pageId="78" pageNumber="288" year="1990">
<bibRefCitation id="605245A4FFB57839FBD1FEA7FADEFE20" author="Zarenkov NA" box="[1077,1283,382,404]" pageId="78" pageNumber="288" pagination="218 - 244" refId="ref73200" refString="Zarenkov NA (1990) Decapods (Stenopodidea, Brachyura, Anomura) of the Naska and Sala-I-Comes underwater Ridges. Trudy Institute of Oceanology, 124: 218 - 244. [In Russian, with English summary]." type="journal article" year="1990">Zarenkov, 1990: 238</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="047C3855FFB57839FCC9FE45FAACFE03" blockId="78.[813,1452,382,548]" box="[813,1393,410,432]" pageId="78" pageNumber="288">
<treatmentCitationGroup id="24D31F7BFFB57839FCC9FE45FAACFE03" box="[813,1393,410,432]" pageId="78" pageNumber="288">
<taxonomicName id="C3C343D6FFB57839FCC9FE45FBDBFE03" authority="Lemaitre, 2014" authorityName="Lemaitre" authorityYear="2014" box="[813,1030,412,432]" class="Malacostraca" family="Parapaguridae" genus="Sympagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="78" pageNumber="288" phylum="Arthropoda" rank="species" species="africanus" status="subsp. nov.">
<emphasis id="36B7E447FFB57839FCC9FE45FBDBFE03" box="[813,1030,412,432]" italics="true" pageId="78" pageNumber="288">Sympagurus africanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2D84593CFFB57839FBE9FE43FBA6FE03" box="[1037,1147,410,432]" pageId="78" pageNumber="288" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
<treatmentCitation id="85621E44FFB57839FB72FE43FAACFE03" author="Parin NV &amp; Mironov AN &amp; Nesis KN" box="[1174,1393,410,432]" page="163" pageId="78" pageNumber="288" year="1997">
<bibRefCitation id="605245A4FFB57839FB72FE43FAACFE03" author="Parin NV &amp; Mironov AN &amp; Nesis KN" box="[1174,1393,410,432]" pageId="78" pageNumber="288" pagination="145 - 242" refId="ref71686" refString="Parin NV, Mironov AN &amp; Nesis KN (1997) Biology of the Nazca and Sala y Gomez submarine ridges, and outpost of the Indo- West Pacific fauna in the eastern Pacific Ocean: composition and distribution of the fauna, its communities and history. Advances in Marine Biology, 32: 145 - 242." type="journal article" year="1997">Parin et al., 1997: 163</bibRefCitation>
</treatmentCitation>
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</paragraph>
<paragraph id="047C3855FFB57839FCC9FE60FAF2FD90" blockId="78.[813,1452,382,548]" pageId="78" pageNumber="288">
<treatmentCitationGroup id="24D31F7BFFB57839FCC9FE60FAF2FD90" pageId="78" pageNumber="288">
<treatmentCitation id="85621E44FFB57839FCC9FE60FB15FE7E" author="Zhadan DG" box="[813,1224,439,461]" page="65" pageId="78" pageNumber="288" year="1997">
<taxonomicName id="C3C343D6FFB57839FCC9FE60FB15FE7E" ID-CoL="b98a575f-d5d1-48e3-b786-671ff4040cff" authority="Zhadan, 1997: 65" authorityName="Zhadan" authorityPageNumber="65" authorityYear="1997" box="[813,1224,439,461]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="78" pageNumber="288" phylum="Arthropoda" rank="species" species="stockmani">
<emphasis id="36B7E447FFB57839FCC9FE60FBCBFE7E" box="[813,1046,441,461]" italics="true" pageId="78" pageNumber="288">Oncopagurus stockmani</emphasis>
<bibRefCitation id="605245A4FFB57839FBF9FE6EFB15FE7E" author="Zhadan DG" box="[1053,1224,439,461]" pageId="78" pageNumber="288" pagination="55 - 79" refId="ref73241" refString="Zhadan DG (1997) Deep-sea hermit crabs from the submerged ridges Nazka and Sala-y-Gomez, southeastern Pacific (Decapoda Anomura Parapaguridae). Arthropoda Selecta, 6 (1 / 2): 55 - 79." type="journal article" year="1997">Zhadan, 1997: 65</bibRefCitation>
</taxonomicName>
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, figs 68 (
<typeStatus id="DB7886F7FFB57839FAC8FE6EFA8AFE7F" box="[1324,1367,439,460]" pageId="78" pageNumber="288">type</typeStatus>
locality: Nazca and Sala-y-Gómez ridges, southeastern Pacific, R/V Professor Stockman, sta 1873,
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,
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);
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<bibRefCitation id="605245A4FFB57839FAD1FE28FC37FD90" author="McLaughlin PA &amp; Komai T &amp; Lemaitre R &amp; Rahayu DL" pageId="78" pageNumber="288" pagination="5 - 107" refId="ref71034" refString="McLaughlin PA, Komai T, Lemaitre R &amp; Rahayu DL (2010) Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galatheidae of the Galatheoidea). Part I - Lithodoidea, Lomisoidea and Paguroidea. The Raffles Bulletin of Zoology, Supplement 23: 5 - 107." type="journal article" year="2010">McLaughlin et al., 2010: 39</bibRefCitation>
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;
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<bibRefCitation id="605245A4FFB57839FC12FDD7FAF6FD90" author="Retamal M &amp; Moyano HI" box="[1014,1323,526,548]" pageId="78" pageNumber="288" pagination="302 - 328" refId="ref72031" refString="Retamal M &amp; Moyano HI (2010). Zoogeografia de los crustaceos decapodos chilenos marinos y dulceacuicolas. Latin American Journal of Aquatic Research, 38 (3): 302 - 328." type="journal article" year="2010">Retamal &amp; Moyano, 2010: 313</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="4CD96BDEFFB57824FCC9FD9EFEAFFC66" lastPageId="83" lastPageNumber="293" pageId="78" pageNumber="288" type="materials_examined">
<paragraph id="047C3855FFB57839FCC9FD9EFB18FD00" blockId="78.[813,1452,583,692]" pageId="78" pageNumber="288">
<emphasis id="36B7E447FFB57839FCC9FD9EFC19FDEE" bold="true" box="[813,964,583,605]" pageId="78" pageNumber="288">Type material.</emphasis>
<materialsCitation id="B4AB3208FFB57839FC29FD9EFB18FD00" collectingDate="1987-04-22" collectionCode="ZMUM" collectorName="Sala-y-Gomez Ridges, R &amp; V Professor Stockman &amp; A. N. Mironov" elevation="235" latitude="-22.116667" location="southeastern Pacific" longLatPrecision="1261" longitude="-81.3" municipality="Nazca" pageId="78" pageNumber="288" specimenCount="1" typeStatus="holotype">
<typeStatus id="DB7886F7FFB57839FC29FD9EFBFBFDEE" box="[973,1062,583,605]" pageId="78" pageNumber="288" type="holotype">Holotype</typeStatus>
(not seen),
<location id="011C6E8EFFB57839FB7BFD9EFABFFDEE" LSID="urn:lsid:plazi:treatment:8C6A8943FFB27824FF1DFA04FC02FEA6:011C6E8EFFB57839FB7BFD9EFABFFDEE" box="[1183,1378,583,605]" latitude="-22.116667" longLatPrecision="1261" longitude="-81.3" municipality="Nazca" name="southeastern Pacific" pageId="78" pageNumber="288">southeastern Pacific</location>
,
<collectingMunicipality id="E418A22FFFB57839FA8BFD91FA76FDEE" box="[1391,1451,584,605]" pageId="78" pageNumber="288">Nazca</collectingMunicipality>
and
<collectorName id="A9365D83FFB57839FCBCFDBDFB90FDCA" box="[856,1101,612,634]" pageId="78" pageNumber="288">Sala-y-Gómez Ridges, R</collectorName>
/
<collectorName id="A9365D83FFB57839FBB0FDBDFAE9FDC9" box="[1108,1332,612,634]" pageId="78" pageNumber="288">V Professor Stockman</collectorName>
, 18
<superScript id="F3B6951DFFB57839FABCFDBAFABEFDDC" attach="left" box="[1368,1379,611,623]" fontSize="5" pageId="78" pageNumber="288">th</superScript>
cruise, sta 1873,
<geoCoordinate id="61F75E92FFB57839FC76FD58FC3DFD25" box="[914,992,641,662]" degrees="22" direction="south" minutes="07" orientation="latitude" pageId="78" pageNumber="288" precision="925" value="-22.116667">22°07'S</geoCoordinate>
,
<geoCoordinate id="61F75E92FFB57839FC0AFD58FB99FD25" box="[1006,1092,641,662]" degrees="81" direction="west" minutes="18" orientation="longitude" pageId="78" pageNumber="288" precision="925" value="-81.3">81°18'W</geoCoordinate>
,
<quantity id="C33B95B0FFB57839FBB5FD58FB49FD25" box="[1105,1172,641,663]" metricMagnitude="2" metricUnit="m" metricValue="2.35" pageId="78" pageNumber="288" unit="m" value="235.0">
<elevation id="8FEEDF66FFB57839FBB5FD58FB49FD25" box="[1105,1172,641,663]" metricMagnitude="2" metricUnit="m" metricValue="2.35" pageId="78" pageNumber="288" unit="m" value="235.0">235 m</elevation>
</quantity>
,
<date id="707D1E95FFB57839FB46FD58FAEFFD25" box="[1186,1330,641,663]" pageId="78" pageNumber="288" value="1987-04-22">
<collectingDate id="6039E77DFFB57839FB46FD58FAEFFD25" box="[1186,1330,641,663]" pageId="78" pageNumber="288" value="1987-04-22">22 April 1987</collectingDate>
</date>
, coll.
<collectorName id="A9365D83FFB57839FA97FD58FC58FD07" pageId="78" pageNumber="288">A. N. Mironov</collectorName>
: M
<quantity id="C33B95B0FFB57839FC48FD47FC2AFD00" box="[940,1015,670,692]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="78" pageNumber="288" unit="mm" value="6.1">6.1 mm</quantity>
(
<collectionCode id="62D2A090FFB57839FBE1FD47FB8CFD00" box="[1029,1105,670,691]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="78" pageNumber="288">ZMUM</collectionCode>
Ma4758).
</materialsCitation>
</paragraph>
<paragraph id="047C3855FFB57839FCC9FD01FC18FCB9" blockId="78.[813,1452,727,778]" pageId="78" pageNumber="288">
<typeStatus id="DB7886F7FFB57839FCC9FD01FC53FD5E" box="[813,910,728,749]" pageId="78" pageNumber="288" type="paratype">Paratypes</typeStatus>
: 2 M 4.5,
<quantity id="C33B95B0FFB57839FC0CFD0EFBEEFD5E" box="[1000,1075,727,749]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="78" pageNumber="288" unit="mm" value="6.4">6.4 mm</quantity>
, same station data as holotype (MNHN- IU-2013-6871).
</paragraph>
<paragraph id="047C3855FFB57839FCC9FCF4FA48FC67" blockId="78.[813,1452,813,980]" pageId="78" pageNumber="288">
<emphasis id="36B7E447FFB57839FCC9FCF4FB1BFCF0" bold="true" box="[813,1222,813,835]" pageId="78" pageNumber="288">Additional material. Eastern Pacific:</emphasis>
<materialsCitation id="B4AB3208FFB57839FB35FCF7FA4CFC67" collectingDate="1965-12-13" collectingDateMax="2000-10-23" collectingDateMin="1965-12-13" collectionCode="USNM" collectorName="V Anton Bruun &amp; Mas" county="Juan Fernandez Islands" elevation="150" latitude="-33.64139" location="Tierra Island" longLatPrecision="19" longitude="-78.84111" municipality="Robinsoe Crusoe Island" pageId="78" pageNumber="288" specimenCode="USNM 1211233, USNM 1211234" specimenCount="1">
SEPBOP, R/
<collectorName id="A9365D83FFB57839FAB6FCF7FCB6FCD3" pageId="78" pageNumber="288">V Anton Bruun</collectorName>
, cruise 12: sta 65MVIV54, off
<collectorName id="A9365D83FFB57839FB5EFC93FB3EFCD3" box="[1210,1251,842,864]" pageId="78" pageNumber="288">Más</collectorName>
a
<location id="011C6E8EFFB57839FB1DFC93FAABFCD3" LSID="urn:lsid:plazi:treatment:8C6A8943FFB27824FF1DFA04FC02FEA6:011C6E8EFFB57839FB1DFC93FAABFCD3" box="[1273,1398,842,864]" county="Juan Fernandez Islands" latitude="-33.64139" longLatPrecision="19" longitude="-78.84111" municipality="Robinsoe Crusoe Island" name="Tierra Island" pageId="78" pageNumber="288">Tierra Island</location>
,
<location id="011C6E8EFFB57839FA9BFC92FC02FCCE" LSID="urn:lsid:plazi:treatment:8C6A8943FFB27824FF1DFA04FC02FEA6:011C6E8EFFB57839FA9BFC92FC02FCCE" county="Juan Fernandez Islands" latitude="-33.64139" longLatPrecision="19" longitude="-78.84111" municipality="Robinsoe Crusoe Island" name="Juan Fernandez Islands" pageId="78" pageNumber="288">Juan Fernández Islands</location>
, [approx.
<geoCoordinate id="61F75E92FFB57839FBA0FCBEFB6CFCCF" box="[1092,1201,871,892]" degrees="33" direction="south" minutes="38" orientation="latitude" pageId="78" pageNumber="288" precision="15" seconds="29" value="-33.64139">33°38'29&quot;S</geoCoordinate>
<geoCoordinate id="61F75E92FFB57839FB5DFCBFFAE9FCCF" box="[1209,1332,870,892]" degrees="78" direction="west" minutes="50" orientation="longitude" pageId="78" pageNumber="288" precision="15" seconds="28" value="-78.84111">78°50'28&quot;W</geoCoordinate>
],
<quantity id="C33B95B0FFB57839FAA2FCBEFA5AFCCE" box="[1350,1415,871,893]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="78" pageNumber="288" unit="m" value="150.0">
<elevation id="8FEEDF66FFB57839FAA2FCBEFA5AFCCE" box="[1350,1415,871,893]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="78" pageNumber="288" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="707D1E95FFB57839FA77FCBEFC17FC29" pageId="78" pageNumber="288" value="1965-12-13">
<collectingDate id="6039E77DFFB57839FA77FCBEFC17FC29" pageId="78" pageNumber="288" value="1965-12-13">13 December 1965</collectingDate>
</date>
: ov F 7.0 mm (
<specimenCode id="5465902EFFB57839FBBBFC5DFAD9FC29" box="[1119,1284,900,922]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="78" pageNumber="288" type="Museum">USNM 1211233</specimenCode>
); CIMAR 6, stan 38,
<collectingMunicipality id="E418A22FFFB57839FCB0FC78FB9FFC04" box="[852,1090,929,951]" pageId="78" pageNumber="288">Robinsoe Crusoe Island</collectingMunicipality>
,
<collectingCounty id="ED1D40D9FFB57839FBABFC78FAE5FC04" box="[1103,1336,929,951]" pageId="78" pageNumber="288">Juan Fernández Islands</collectingCounty>
,
<quantity id="C33B95B0FFB57839FAA0FC78FA5AFC05" box="[1348,1415,929,951]" metricMagnitude="2" metricUnit="m" metricValue="1.46" pageId="78" pageNumber="288" unit="m" value="146.0">
<elevation id="8FEEDF66FFB57839FAA0FC78FA5AFC05" box="[1348,1415,929,951]" metricMagnitude="2" metricUnit="m" metricValue="1.46" pageId="78" pageNumber="288" unit="m" value="146.0">146 m</elevation>
</quantity>
,
<date id="707D1E95FFB57839FA77FC78FC6BFC60" pageId="78" pageNumber="288" value="2000-10-23">
<collectingDate id="6039E77DFFB57839FA77FC78FC6BFC60" pageId="78" pageNumber="288" value="2000-10-23">23 October 2000</collectingDate>
</date>
: 1 F
<quantity id="C33B95B0FFB57839FC0DFC67FBEAFC60" box="[1001,1079,958,979]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="78" pageNumber="288" unit="mm" value="4.9">4.9 mm</quantity>
, 1 ov F 4.0 mm (
<specimenCode id="5465902EFFB57839FB00FC67FA56FC67" box="[1252,1419,958,980]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="78" pageNumber="288" type="Museum">USNM 1211234</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="047C3855FFB57839FCC9FC25FCA8FA80" blockId="78.[813,1458,1020,1331]" pageId="78" pageNumber="288">
<emphasis id="36B7E447FFB57839FCC9FC25FC01FBA7" bold="true" box="[813,988,1020,1044]" pageId="78" pageNumber="288">Re-description.</emphasis>
Gills biserial. Shield (
<figureCitation id="9CF824D0FFB57839FB3DFC25FAE3FBA7" box="[1241,1342,1020,1044]" captionStart="Fig" captionStartId="79.[136,171,1976,1998]" captionTargetBox="[142,1460,715,1943]" captionTargetId="figure-7@79.[130,1468,691,1950]" captionTargetPageId="79" captionText="Fig. 46. Oncopagurus stockmani Zhadan, 1997: AD, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233); E, F, holotype, male 6.1 mm, southeastern Pacific, Nazca and Sala-y-Gómez ridges, “Professor Stockman”, 18th cruise, sta 1873 (ZMUM Ma 4758). A, shield and cephalic appendages, dorsal; B, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; C, exopod of left uropod, dorsal; D, telson, dorsal; E, left first gonopod, mesial; F, left second gonopod, anterior. Scale bars = 2 mm (A); 1 mm (BE); 2 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458468" httpUri="https://zenodo.org/record/5458468/files/figure.png" pageId="78" pageNumber="288">Fig. 46A</figureCitation>
) about as long as broad; dorsal surface weakly calcified medially, with scattered short setae; rostrum broadly rounded, weakly produced, with short mid-dorsal ridge; anterior margins weakly concave; lateral projections subtriangular, terminating in small blunt or sharp spine; anterolateral margins sloping; posterior margin broadly rounded; ventrolateral margins of shield usually with slender spine on one or both sides. Anterodistal margin of branchiostegite rounded, unarmed, setose.
</paragraph>
<paragraph id="047C3855FFB57839FCC9FA85FA88FA47" blockId="78.[813,1452,1372,1524]" pageId="78" pageNumber="288">
Ocular peduncles (
<figureCitation id="9CF824D0FFB57839FBE4FA85FBBAFAC7" box="[1024,1127,1372,1396]" captionStart="Fig" captionStartId="79.[136,171,1976,1998]" captionTargetBox="[142,1460,715,1943]" captionTargetId="figure-7@79.[130,1468,691,1950]" captionTargetPageId="79" captionText="Fig. 46. Oncopagurus stockmani Zhadan, 1997: AD, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233); E, F, holotype, male 6.1 mm, southeastern Pacific, Nazca and Sala-y-Gómez ridges, “Professor Stockman”, 18th cruise, sta 1873 (ZMUM Ma 4758). A, shield and cephalic appendages, dorsal; B, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; C, exopod of left uropod, dorsal; D, telson, dorsal; E, left first gonopod, mesial; F, left second gonopod, anterior. Scale bars = 2 mm (A); 1 mm (BE); 2 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458468" httpUri="https://zenodo.org/record/5458468/files/figure.png" pageId="78" pageNumber="288">Fig. 46A</figureCitation>
) about half length of shield, with longitudinal row of short setae dorsally; lateral and ventral faces usually weakly calcified. Cornea weakly dilated. Ocular acicles subtriangular, each terminating in strong spine; separated basally by about basal width of 1 acicle.
</paragraph>
<paragraph id="047C3855FFB57839FCC9F9C5FA71F947" blockId="78.[813,1452,1564,1780]" pageId="78" pageNumber="288">
Antennular peduncle (
<figureCitation id="9CF824D0FFB57839FBC3F9C5FB53F987" box="[1063,1166,1564,1588]" captionStart="Fig" captionStartId="79.[136,171,1976,1998]" captionTargetBox="[142,1460,715,1943]" captionTargetId="figure-7@79.[130,1468,691,1950]" captionTargetPageId="79" captionText="Fig. 46. Oncopagurus stockmani Zhadan, 1997: AD, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233); E, F, holotype, male 6.1 mm, southeastern Pacific, Nazca and Sala-y-Gómez ridges, “Professor Stockman”, 18th cruise, sta 1873 (ZMUM Ma 4758). A, shield and cephalic appendages, dorsal; B, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; C, exopod of left uropod, dorsal; D, telson, dorsal; E, left first gonopod, mesial; F, left second gonopod, anterior. Scale bars = 2 mm (A); 1 mm (BE); 2 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458468" httpUri="https://zenodo.org/record/5458468/files/figure.png" pageId="78" pageNumber="288">Fig. 46A</figureCitation>
) exceeding distal margin of cornea by about full length or slightly more, of ultimate segment. Ultimate segment about twice as long as penultimate segment, naked or with scattered setae dorsally. Basal segment with strong ventromesial spine; lateral face with distal subrectangular lobe armed with small spine distally, and strong spine proximally. Ventral flagellum with 8 articles.
</paragraph>
<paragraph id="047C3855FFB57838FCC9F8C5FD48FE26" blockId="78.[813,1452,1820,2068]" lastBlockId="79.[138,776,189,406]" lastPageId="79" lastPageNumber="289" pageId="78" pageNumber="288">
Antennal peduncle (
<figureCitation id="9CF824D0FFB57839FBEAF8C5FBA9F887" box="[1038,1140,1820,1844]" captionStart="Fig" captionStartId="79.[136,171,1976,1998]" captionTargetBox="[142,1460,715,1943]" captionTargetId="figure-7@79.[130,1468,691,1950]" captionTargetPageId="79" captionText="Fig. 46. Oncopagurus stockmani Zhadan, 1997: AD, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233); E, F, holotype, male 6.1 mm, southeastern Pacific, Nazca and Sala-y-Gómez ridges, “Professor Stockman”, 18th cruise, sta 1873 (ZMUM Ma 4758). A, shield and cephalic appendages, dorsal; B, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; C, exopod of left uropod, dorsal; D, telson, dorsal; E, left first gonopod, mesial; F, left second gonopod, anterior. Scale bars = 2 mm (A); 1 mm (BE); 2 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458468" httpUri="https://zenodo.org/record/5458468/files/figure.png" pageId="78" pageNumber="288">Fig. 46A</figureCitation>
) at most slightly exceeding distal margin of cornea. Fifth segment unarmed except for scattered setae and laterodistal tuft of setae. Fourth segment armed with strong dorsodistal spine. Third segment with strong ventromesial distal spine. Second segment with dorsolateral distal angle produced, terminating in strong simple spine (often with small spine dorsally); mesial margin with spine on dorsodistal angle. First segment with lateral face unarmed or with small blunt spine; ventromesial angle produced, with 2 or 3 small spines laterally. Acicle reaching or slightly exceeding distal margin of cornea, weakly curved outward (in dorsal view), terminating in strong spine; mesial margin armed with 69 spines, sparsely setose. Flagellum slightly exceeding extended right cheliped, articles with few setae less than 1 to 2 flagellar articles in length.
</paragraph>
<paragraph id="047C3855FFB47838FF6EFE64FA1EFE66" blockId="79.[138,776,445,629]" lastBlockId="79.[836,1475,189,469]" pageId="79" pageNumber="289">
Mandible (
<figureCitation id="9CF824D0FFB47838FEE5FE64FEBBFE65" box="[257,358,445,470]" captionStart="Fig" captionStartId="80.[113,148,2032,2054]" captionTargetBox="[120,1447,559,1991]" captionTargetId="figure-9@80.[110,1460,539,2011]" captionTargetPageId="80" captionText="Fig. 47. Oncopagurus stockmani Zhadan, 1997: ov female 4.0 mm, eastern Pacific, Robinsoe Crusoe Island, Juan Fernández Islands, CIMAR 6, sta 38 (USNM 1211234). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5458470" httpUri="https://zenodo.org/record/5458470/files/figure.png" pageId="79" pageNumber="289">Fig. 47A</figureCitation>
) with 3-segmented palp; cutting edge calcified, with small tooth medially; molar process with small tooth medially. Maxillule (
<figureCitation id="9CF824D0FFB47838FE04FE24FDE0FDA5" box="[480,573,509,534]" captionStart="Fig" captionStartId="80.[113,148,2032,2054]" captionTargetBox="[120,1447,559,1991]" captionTargetId="figure-9@80.[110,1460,539,2011]" captionTargetPageId="80" captionText="Fig. 47. Oncopagurus stockmani Zhadan, 1997: ov female 4.0 mm, eastern Pacific, Robinsoe Crusoe Island, Juan Fernández Islands, CIMAR 6, sta 38 (USNM 1211234). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5458470" httpUri="https://zenodo.org/record/5458470/files/figure.png" pageId="79" pageNumber="289">Fig. 47B</figureCitation>
) with external lobe of endopod slender, moderately developed, not recurved, internal lobe with long, terminal seta. Maxilla (
<figureCitation id="9CF824D0FFB47838FD78FDE4FCDDFDE5" box="[668,768,573,598]" captionStart="Fig" captionStartId="80.[113,148,2032,2054]" captionTargetBox="[120,1447,559,1991]" captionTargetId="figure-9@80.[110,1460,539,2011]" captionTargetPageId="80" captionText="Fig. 47. Oncopagurus stockmani Zhadan, 1997: ov female 4.0 mm, eastern Pacific, Robinsoe Crusoe Island, Juan Fernández Islands, CIMAR 6, sta 38 (USNM 1211234). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5458470" httpUri="https://zenodo.org/record/5458470/files/figure.png" pageId="79" pageNumber="289">Fig. 47C</figureCitation>
) with endopod not exceeding distal margin of scaphognathite. First maxilliped (
<figureCitation id="9CF824D0FFB47838FC1FFF64FB81FF65" box="[1019,1116,189,214]" captionStart="Fig" captionStartId="80.[113,148,2032,2054]" captionTargetBox="[120,1447,559,1991]" captionTargetId="figure-9@80.[110,1460,539,2011]" captionTargetPageId="80" captionText="Fig. 47. Oncopagurus stockmani Zhadan, 1997: ov female 4.0 mm, eastern Pacific, Robinsoe Crusoe Island, Juan Fernández Islands, CIMAR 6, sta 38 (USNM 1211234). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5458470" httpUri="https://zenodo.org/record/5458470/files/figure.png" pageId="79" pageNumber="289">Fig. 47D</figureCitation>
) with endopod slightly exceeding exopod in distal extension. Second maxilliped (
<figureCitation id="9CF824D0FFB47838FABCFF04FA67FF45" box="[1368,1466,221,246]" captionStart="Fig" captionStartId="80.[113,148,2032,2054]" captionTargetBox="[120,1447,559,1991]" captionTargetId="figure-9@80.[110,1460,539,2011]" captionTargetPageId="80" captionText="Fig. 47. Oncopagurus stockmani Zhadan, 1997: ov female 4.0 mm, eastern Pacific, Robinsoe Crusoe Island, Juan Fernández Islands, CIMAR 6, sta 38 (USNM 1211234). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5458470" httpUri="https://zenodo.org/record/5458470/files/figure.png" pageId="79" pageNumber="289">Fig. 47E</figureCitation>
) without distinguishing characters. Third maxilliped (
<figureCitation id="9CF824D0FFB47838FA72FF24FCB2FE85" captionStart="Fig" captionStartId="80.[113,148,2032,2054]" captionTargetBox="[120,1447,559,1991]" captionTargetId="figure-9@80.[110,1460,539,2011]" captionTargetPageId="80" captionText="Fig. 47. Oncopagurus stockmani Zhadan, 1997: ov female 4.0 mm, eastern Pacific, Robinsoe Crusoe Island, Juan Fernández Islands, CIMAR 6, sta 38 (USNM 1211234). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5458470" httpUri="https://zenodo.org/record/5458470/files/figure.png" pageId="79" pageNumber="289">Fig. 47F</figureCitation>
) with merus to dactyl each distinctly longer than broad, ischium about twice as long as broad; crista dentata with about 1218 blunt or sharp teeth gradually diminishing in strength distally, proximal tooth distinctly larger than others; basis with 1 small mesial spine; coxa with spine. Sternite of third maxillipeds with small spine on each side of midline.
</paragraph>
<paragraph id="047C3855FFB47826FCA0FE24FDEEFDA7" blockId="79.[836,1474,509,630]" lastBlockId="81.[138,774,189,533]" lastPageId="81" lastPageNumber="291" pageId="79" pageNumber="289">
Chelipeds markedly dissimilar. Right cheliped (
<figureCitation id="9CF824D0FFB47838FADEFE24FA67FDA5" box="[1338,1466,509,534]" captionStart="Fig" captionStartId="81.[138,173,2033,2055]" captionTargetBox="[140,1466,951,1993]" captionTargetId="figure-7@81.[130,1488,941,2010]" captionTargetPageId="81" captionText="Fig. 48. Oncopagurus stockmani Zhadan, 1997, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233): A, right cheliped, dorsal; BD, chela of same, ventral (B), lateral (C), mesial (D); E, left cheliped, dorsal; F, carpus of same, dorsolateral. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5458472" httpUri="https://zenodo.org/record/5458472/files/figure.png" pageId="79" pageNumber="289">Fig. 48AD</figureCitation>
) massive, chela operculate; carpus and chela with sparse to moderately dense setae on dorsal surfaces, setae more dense on dorsaldistal surfaces of fingers and mesial face of carpus. Fingers curved ventromesially, each terminating in small, usually blunt corneous claw; cutting edges with 1 or 2 large calcareous, mostly serrated teeth and, with short row of closely set minute corneous teeth distally on dactyl. Dactyl about as long as mesial margin of palm, set at strongly oblique angle to longitudinal axis of palm; mesial margin broadly curved, delimited by row of spines diminishing in size distally; dorsal face with numerous small spines or tubercles; ventromesial face concave and elevated longitudinally on midline. Fixed finger broad at base, dorsal face covered with numerous small spines or tubercles, lateral margin delimited by row of spines; ventral face concave mesially, elevated longitudinally on midline. Palm about 1.3 times broader than long; dorsal surface covered with numerous well-spaced small spines or turbercles; dorsolateral margin broadly rounded, or well delimited by irregular row of calcareous spines, dorsomesial margin delimited by row of small spines or tubercles; dorsomesial face nearly flat, sloping mesially, with many small tubercles; ventral surface with numerous small spines or tubercles, and scattered short setae. Carpus longer than broad, dorsal surface covered with numerous small spines or turbercles; dorsodistal margin with row of small spines; dorsolateral face rounded; mesial face strongly sloping, expanded distomesially and with row of strong ventromesial spines distally; ventral face nearly flat, with well spaced small tubercles. Merus with scattered setae dorsally, and moderately dense setae ventrodistally; dorsal surface with scattered small tubercles and transverse, distal row of setae near dorsodistal margin; ventromesial margin with row of blunt to sharp spines; ventrolateral margin with row of small tubercles. Ischium with ventromesial row of small, blunt spines. Coxa row of small spines on ventrodistal margin, and ventromesial row of setae.
</paragraph>
<caption id="50BC68DDFFB47838FF6CF861FB10F7F1" ID-DOI="http://doi.org/10.5281/zenodo.5458468" ID-Zenodo-Dep="5458468" httpUri="https://zenodo.org/record/5458468/files/figure.png" pageId="79" pageNumber="289" startId="79.[136,171,1976,1998]" targetBox="[142,1460,715,1943]" targetPageId="79">
<paragraph id="047C3855FFB47838FF6CF861FB10F7F1" blockId="79.[136,1474,1976,2114]" pageId="79" pageNumber="289">
Fig. 46.
<taxonomicName id="C3C343D6FFB47838FF3EF863FD89F87D" authority="Zhadan, 1997" authorityName="Zhadan" authorityYear="1997" box="[218,596,1976,1998]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="79" pageNumber="289" phylum="Arthropoda" rank="species" species="stockmani">
<emphasis id="36B7E447FFB47838FF3EF863FE19F87D" box="[218,452,1978,1998]" italics="true" pageId="79" pageNumber="289">Oncopagurus stockmani</emphasis>
<bibRefCitation id="605245A4FFB47838FE2FF861FD89F87D" author="Zhadan DG" box="[459,596,1976,1998]" pageId="79" pageNumber="289" pagination="55 - 79" refId="ref73241" refString="Zhadan DG (1997) Deep-sea hermit crabs from the submerged ridges Nazka and Sala-y-Gomez, southeastern Pacific (Decapoda Anomura Parapaguridae). Arthropoda Selecta, 6 (1 / 2): 55 - 79." type="journal article" year="1997">Zhadan, 1997</bibRefCitation>
</taxonomicName>
: AD, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233); E, F, holotype, male 6.1 mm, southeastern Pacific, Nazca and Sala-y-Gómez ridges, “Professor Stockman”, 18
<superScript id="F3B6951DFFB47838FD6EF828FD48F84E" attach="left" box="[650,661,2033,2045]" fontSize="5" pageId="79" pageNumber="289">th</superScript>
cruise, sta 1873 (ZMUM Ma 4758). A, shield and cephalic appendages, dorsal; B, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; C, exopod of left uropod, dorsal; D, telson, dorsal; E, left first gonopod, mesial; F, left second gonopod, anterior. Scale bars = 2 mm (A); 1 mm (BE); 2 mm (F).
</paragraph>
</caption>
<caption id="50BC68DDFFAB7827FF95F829FD3EF78C" ID-DOI="http://doi.org/10.5281/zenodo.5458470" ID-Zenodo-Dep="5458470" httpUri="https://zenodo.org/record/5458470/files/figure.png" pageId="80" pageNumber="290" startId="80.[113,148,2032,2054]" targetBox="[120,1447,559,1991]" targetPageId="80">
<paragraph id="047C3855FFAB7827FF95F829FD3EF78C" blockId="80.[113,1452,2032,2111]" pageId="80" pageNumber="290">
Fig. 47.
<taxonomicName id="C3C343D6FFAB7827FF2CF82BFD91F7B6" authority="Zhadan, 1997" authorityName="Zhadan" authorityYear="1997" box="[200,588,2032,2054]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="290" phylum="Arthropoda" rank="species" species="stockmani">
<emphasis id="36B7E447FFAB7827FF2CF82BFE6AF7B5" box="[200,439,2034,2054]" italics="true" pageId="80" pageNumber="290">Oncopagurus stockmani</emphasis>
<bibRefCitation id="605245A4FFAB7827FE25F829FD91F7B6" author="Zhadan DG" box="[449,588,2032,2054]" pageId="80" pageNumber="290" pagination="55 - 79" refId="ref73241" refString="Zhadan DG (1997) Deep-sea hermit crabs from the submerged ridges Nazka and Sala-y-Gomez, southeastern Pacific (Decapoda Anomura Parapaguridae). Arthropoda Selecta, 6 (1 / 2): 55 - 79." type="journal article" year="1997">Zhadan, 1997</bibRefCitation>
</taxonomicName>
: ov female 4.0 mm, eastern Pacific, Robinsoe Crusoe Island, Juan Fernández Islands, CIMAR 6, sta 38 (USNM 1211234). Left mouthparts, internal: A, distal half of mandible; B, maxillule; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="047C3855FFAA7826FF6EFDE4FC0FFE67" blockId="81.[138,774,573,885]" lastBlockId="81.[836,1474,189,469]" pageId="81" pageNumber="291">
Left cheliped (
<figureCitation id="9CF824D0FFAA7826FEC5FDE4FE4AFDE6" box="[289,407,573,597]" captionStart="Fig" captionStartId="81.[138,173,2033,2055]" captionTargetBox="[140,1466,951,1993]" captionTargetId="figure-7@81.[130,1488,941,2010]" captionTargetPageId="81" captionText="Fig. 48. Oncopagurus stockmani Zhadan, 1997, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233): A, right cheliped, dorsal; BD, chela of same, ventral (B), lateral (C), mesial (D); E, left cheliped, dorsal; F, carpus of same, dorsolateral. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5458472" httpUri="https://zenodo.org/record/5458472/files/figure.png" pageId="81" pageNumber="291">Fig. 48E, F</figureCitation>
) usually weakly calcified on nearly entire dorsolateral face of carpus. Fingers each terminating in small corneous claw; dorsal and ventral surfaces with scattered tufts of setae; cutting edge of dactyl with row of minute, fused corneous teeth; cutting edge of fixed finger with row of well-spaced small calcareous teeth. Dactyl about as long as length of mesial margin of palm; with small proximal spine on dorsal face. Fixed finger with 1 or 2 small spines or tubercles proximally on dorsal face. Palm with scattered setae on dorsal surface; with dorsomesial row of small spines or tubercles and scattered small tubercles or spines on rest of dorsal face; ventral face smooth except for scattered setae. Carpus with dorsal irregular row of small spines in addition to dorsodistal and dorsolateral spine on distal margin; dorsal margin with long setae; ventral face smooth except for scattered setae. Merus with setae on dorsal margin; with ventrolateral row of small spines; ventral face smooth. Ischium and coxa unarmed, but with ventromesial row of setae.
</paragraph>
<paragraph id="047C3855FFAA7825FCA0FE24FE3BFEC6" blockId="81.[836,1474,509,885]" lastBlockId="82.[113,752,189,373]" lastPageId="82" lastPageNumber="292" pageId="81" pageNumber="291">
Ambulatory legs (
<figureCitation id="9CF824D0FFAA7826FBE7FE24FB59FDA6" box="[1027,1156,509,533]" captionStart="Fig" captionStartId="82.[113,148,2005,2027]" captionTargetBox="[105,1456,494,1965]" captionTargetId="figure-9@82.[95,1471,484,1974]" captionTargetPageId="82" captionText="Fig. 49. Oncopagurus stockmani Zhadan, 1997, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233): A, right first ambulatory leg, lateral; B, dactyl of same, mesial; C, right second ambulatory leg, lateral; D, dactyl of same, mesial; E, propodus and dactyl of left fourth pereopod, lateral; F, propodus and dactyl of left fifth pereopod, lateral. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5458474" httpUri="https://zenodo.org/record/5458474/files/figure.png" pageId="81" pageNumber="291">Fig. 49AD</figureCitation>
) similar right from left except for longer segments and stronger spination on right, extending to about tip of dactyls of right cheliped or slightly exceeding them. Dactyls (
<figureCitation id="9CF824D0FFAA7826FC02FD84FBB9FDC6" box="[998,1124,605,629]" captionStart="Fig" captionStartId="82.[113,148,2005,2027]" captionTargetBox="[105,1456,494,1965]" captionTargetId="figure-9@82.[95,1471,484,1974]" captionTargetPageId="82" captionText="Fig. 49. Oncopagurus stockmani Zhadan, 1997, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233): A, right first ambulatory leg, lateral; B, dactyl of same, mesial; C, right second ambulatory leg, lateral; D, dactyl of same, mesial; E, propodus and dactyl of left fourth pereopod, lateral; F, propodus and dactyl of left fifth pereopod, lateral. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5458474" httpUri="https://zenodo.org/record/5458474/files/figure.png" pageId="81" pageNumber="291">Fig. 49B, D</figureCitation>
) broadly curved, about 1.5 times as long as propodi, terminating in sharp corneous claws; each with dorsal and dorsomesial distal rows of long setae, and 1535 small spinules on ventromesial margin (in large specimens such as
<typeStatus id="DB7886F7FFAA7826FBF3FD04FBAAFD46" box="[1047,1143,733,757]" pageId="81" pageNumber="291" type="holotype">holotype</typeStatus>
, sl 7.0 mm, spinules arranged irregularly on proximal 0.6 of ventromesial margin and in regular row distally). Propodi with setae on dorsal margin; ventral margin with few setae. Carpi each with setae on dorsal margin, and small dorsodistal spine; carpi of right with row of small spines on dorsal margins. Meri each with setae on dorsal margin, and 2 or 3 small spines distally on ventrolateral margin. Ischia with setae on dorsal and ventroproximal margins. Coxae with ventromesial row of setae. Anterior lobe of sternite XII (between second ambulatory legs,
<figureCitation id="9CF824D0FFA97825FD21FEE4FF7CFEC6" captionStart="Fig" captionStartId="79.[136,171,1976,1998]" captionTargetBox="[142,1460,715,1943]" captionTargetId="figure-7@79.[130,1468,691,1950]" captionTargetPageId="79" captionText="Fig. 46. Oncopagurus stockmani Zhadan, 1997: AD, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233); E, F, holotype, male 6.1 mm, southeastern Pacific, Nazca and Sala-y-Gómez ridges, “Professor Stockman”, 18th cruise, sta 1873 (ZMUM Ma 4758). A, shield and cephalic appendages, dorsal; B, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; C, exopod of left uropod, dorsal; D, telson, dorsal; E, left first gonopod, mesial; F, left second gonopod, anterior. Scale bars = 2 mm (A); 1 mm (BE); 2 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458468" httpUri="https://zenodo.org/record/5458468/files/figure.png" pageId="82" pageNumber="292">Fig. 46B</figureCitation>
), setose, with subdistal spine.
</paragraph>
<caption id="50BC68DDFFAA7826FF6EF828FB84F7F3" ID-DOI="http://doi.org/10.5281/zenodo.5458472" ID-Zenodo-Dep="5458472" httpUri="https://zenodo.org/record/5458472/files/figure.png" pageId="81" pageNumber="291" startId="81.[138,173,2033,2055]" targetBox="[140,1466,951,1993]" targetPageId="81">
<paragraph id="047C3855FFAA7826FF6EF828FB84F7F3" blockId="81.[138,1474,2033,2113]" pageId="81" pageNumber="291">
Fig. 48.
<taxonomicName id="C3C343D6FFAA7826FF05F82AFDB9F7B5" authority="Zhadan, 1997" authorityName="Zhadan" authorityYear="1997" box="[225,612,2033,2055]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="291" phylum="Arthropoda" rank="species" species="stockmani">
<emphasis id="36B7E447FFAA7826FF05F82AFE0DF7B4" box="[225,464,2035,2055]" italics="true" pageId="81" pageNumber="291">Oncopagurus stockmani</emphasis>
<bibRefCitation id="605245A4FFAA7826FE3DF828FDB9F7B5" author="Zhadan DG" box="[473,612,2033,2055]" pageId="81" pageNumber="291" pagination="55 - 79" refId="ref73241" refString="Zhadan DG (1997) Deep-sea hermit crabs from the submerged ridges Nazka and Sala-y-Gomez, southeastern Pacific (Decapoda Anomura Parapaguridae). Arthropoda Selecta, 6 (1 / 2): 55 - 79." type="journal article" year="1997">Zhadan, 1997</bibRefCitation>
</taxonomicName>
, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233): A, right cheliped, dorsal; BD, chela of same, ventral (B), lateral (C), mesial (D); E, left cheliped, dorsal; F, carpus of same, dorsolateral. Scale bars = 5 mm.
</paragraph>
</caption>
<paragraph id="047C3855FFA97825FCC8FF64FBACFE06" blockId="82.[812,1452,189,437]" pageId="82" pageNumber="292">
Fourth pereopod (
<figureCitation id="9CF824D0FFA97825FC0FFF64FB97FF65" box="[1003,1098,189,214]" captionStart="Fig" captionStartId="82.[113,148,2005,2027]" captionTargetBox="[105,1456,494,1965]" captionTargetId="figure-9@82.[95,1471,484,1974]" captionTargetPageId="82" captionText="Fig. 49. Oncopagurus stockmani Zhadan, 1997, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233): A, right first ambulatory leg, lateral; B, dactyl of same, mesial; C, right second ambulatory leg, lateral; D, dactyl of same, mesial; E, propodus and dactyl of left fourth pereopod, lateral; F, propodus and dactyl of left fifth pereopod, lateral. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5458474" httpUri="https://zenodo.org/record/5458474/files/figure.png" pageId="82" pageNumber="292">Fig. 49E</figureCitation>
) semichelate. Dactyl terminating in sharp corneous claw; with ventrolateral row of small corneous spinules. Propodus longer than broad, rasp with 2 or 3 rows of ovate scales at least distally. Carpus with long setae on dorsal margin. Merus with rows of long setae on dorsal and ventral margins. Coxa with anteroventral row of setae. Anterior lobe of sternite XIII (between fourth pereopods) with row of setae.
</paragraph>
<caption id="50BC68DDFFA97825FF95F80CFD44F7F2" ID-DOI="http://doi.org/10.5281/zenodo.5458474" ID-Zenodo-Dep="5458474" httpUri="https://zenodo.org/record/5458474/files/figure.png" pageId="82" pageNumber="292" startId="82.[113,148,2005,2027]" targetBox="[105,1456,494,1965]" targetPageId="82">
<paragraph id="047C3855FFA97825FF95F80CFD44F7F2" blockId="82.[113,1452,2005,2113]" pageId="82" pageNumber="292">
Fig. 49.
<taxonomicName id="C3C343D6FFA97825FF5BF80EFDEFF859" authority="Zhadan, 1997" authorityName="Zhadan" authorityYear="1997" box="[191,562,2005,2027]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="82" pageNumber="292" phylum="Arthropoda" rank="species" species="stockmani">
<emphasis id="36B7E447FFA97825FF5BF80EFE75F858" box="[191,424,2007,2027]" italics="true" pageId="82" pageNumber="292">Oncopagurus stockmani</emphasis>
<bibRefCitation id="605245A4FFA97825FE49F80CFDEFF859" author="Zhadan DG" box="[429,562,2005,2027]" pageId="82" pageNumber="292" pagination="55 - 79" refId="ref73241" refString="Zhadan DG (1997) Deep-sea hermit crabs from the submerged ridges Nazka and Sala-y-Gomez, southeastern Pacific (Decapoda Anomura Parapaguridae). Arthropoda Selecta, 6 (1 / 2): 55 - 79." type="journal article" year="1997">Zhadan, 1997</bibRefCitation>
</taxonomicName>
, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233): A, right first ambulatory leg, lateral; B, dactyl of same, mesial; C, right second ambulatory leg, lateral; D, dactyl of same, mesial; E, propodus and dactyl of left fourth pereopod, lateral; F, propodus and dactyl of left fifth pereopod, lateral. Scale bars = 2 mm.
</paragraph>
</caption>
<paragraph id="047C3855FFA87824FF6CFF64FECBFEE6" blockId="83.[136,776,189,341]" pageId="83" pageNumber="293">
Fifth pereopod (
<figureCitation id="9CF824D0FFA87824FEC9FF64FE5AFF65" box="[301,391,189,214]" captionStart="Fig" captionStartId="82.[113,148,2005,2027]" captionTargetBox="[105,1456,494,1965]" captionTargetId="figure-9@82.[95,1471,484,1974]" captionTargetPageId="82" captionText="Fig. 49. Oncopagurus stockmani Zhadan, 1997, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233): A, right first ambulatory leg, lateral; B, dactyl of same, mesial; C, right second ambulatory leg, lateral; D, dactyl of same, mesial; E, propodus and dactyl of left fourth pereopod, lateral; F, propodus and dactyl of left fifth pereopod, lateral. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5458474" httpUri="https://zenodo.org/record/5458474/files/figure.png" pageId="83" pageNumber="293">Fig. 49F</figureCitation>
) semichelate. Propodal rasp reaching to or slightly exceeding mid-length of segment. Coxa with row of long setae on anteroventral and ventrodistal margins. Anterior lobe of sternite XIII (between fifth pereopods) with row of setae.
</paragraph>
<paragraph id="047C3855FFA87824FF6CFEA4FD79FDC6" blockId="83.[136,775,381,629]" pageId="83" pageNumber="293">
Uropods markedly asymmetrical; left exopod (
<figureCitation id="9CF824D0FFA87824FD81FEA4FD1DFE26" box="[613,704,381,405]" captionStart="Fig" captionStartId="79.[136,171,1976,1998]" captionTargetBox="[142,1460,715,1943]" captionTargetId="figure-7@79.[130,1468,691,1950]" captionTargetPageId="79" captionText="Fig. 46. Oncopagurus stockmani Zhadan, 1997: AD, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233); E, F, holotype, male 6.1 mm, southeastern Pacific, Nazca and Sala-y-Gómez ridges, “Professor Stockman”, 18th cruise, sta 1873 (ZMUM Ma 4758). A, shield and cephalic appendages, dorsal; B, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; C, exopod of left uropod, dorsal; D, telson, dorsal; E, left first gonopod, mesial; F, left second gonopod, anterior. Scale bars = 2 mm (A); 1 mm (BE); 2 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458468" httpUri="https://zenodo.org/record/5458468/files/figure.png" pageId="83" pageNumber="293">Fig. 46C</figureCitation>
) about 2.7 times as long as broad, rasp consisting of 25 rows of small scales. Telson (
<figureCitation id="9CF824D0FFA87824FE92FE64FE06FE66" box="[374,475,445,469]" captionStart="Fig" captionStartId="79.[136,171,1976,1998]" captionTargetBox="[142,1460,715,1943]" captionTargetId="figure-7@79.[130,1468,691,1950]" captionTargetPageId="79" captionText="Fig. 46. Oncopagurus stockmani Zhadan, 1997: AD, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233); E, F, holotype, male 6.1 mm, southeastern Pacific, Nazca and Sala-y-Gómez ridges, “Professor Stockman”, 18th cruise, sta 1873 (ZMUM Ma 4758). A, shield and cephalic appendages, dorsal; B, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; C, exopod of left uropod, dorsal; D, telson, dorsal; E, left first gonopod, mesial; F, left second gonopod, anterior. Scale bars = 2 mm (A); 1 mm (BE); 2 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458468" httpUri="https://zenodo.org/record/5458468/files/figure.png" pageId="83" pageNumber="293">Fig. 46D</figureCitation>
) moderately asymmetrical, lacking transverse suture; dorsal surface with scattered setae; lateral margins with long setae medially; posterior lobes separated by shallow, armed, U-shaped cleft; terminal margin of left and right lobes armed with about 2 or 3 irregular rows of often weakly curved corneous, unequal spines.
</paragraph>
<paragraph id="047C3855FFA87824FF6CFD44FF38FC86" blockId="83.[136,775,669,821]" pageId="83" pageNumber="293">
<collectingRegion id="C607F6B7FFA87824FF6CFD44FF62FD06" box="[136,191,669,693]" country="Maldives" name="Maale" pageId="83" pageNumber="293">Male</collectingRegion>
with paired first and second gonopods well developed; first gonopod (
<figureCitation id="9CF824D0FFA87824FEFBFD64FEA7FD66" box="[287,378,701,725]" captionStart="Fig" captionStartId="79.[136,171,1976,1998]" captionTargetBox="[142,1460,715,1943]" captionTargetId="figure-7@79.[130,1468,691,1950]" captionTargetPageId="79" captionText="Fig. 46. Oncopagurus stockmani Zhadan, 1997: AD, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233); E, F, holotype, male 6.1 mm, southeastern Pacific, Nazca and Sala-y-Gómez ridges, “Professor Stockman”, 18th cruise, sta 1873 (ZMUM Ma 4758). A, shield and cephalic appendages, dorsal; B, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; C, exopod of left uropod, dorsal; D, telson, dorsal; E, left first gonopod, mesial; F, left second gonopod, anterior. Scale bars = 2 mm (A); 1 mm (BE); 2 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458468" httpUri="https://zenodo.org/record/5458468/files/figure.png" pageId="83" pageNumber="293">Fig. 46E</figureCitation>
) with ovate or subtriagular distal lobe; second gonopods (
<figureCitation id="9CF824D0FFA87824FEB1FD04FE72FD46" box="[341,431,733,757]" captionStart="Fig" captionStartId="79.[136,171,1976,1998]" captionTargetBox="[142,1460,715,1943]" captionTargetId="figure-7@79.[130,1468,691,1950]" captionTargetPageId="79" captionText="Fig. 46. Oncopagurus stockmani Zhadan, 1997: AD, ov female 7.0 mm, eastern Pacific, off Más a Tierra Island, Juan Fernández Islands, SEPBOP, R/V Anton Bruun, cruise 12: sta 65MVIV54 (USNM 1211233); E, F, holotype, male 6.1 mm, southeastern Pacific, Nazca and Sala-y-Gómez ridges, “Professor Stockman”, 18th cruise, sta 1873 (ZMUM Ma 4758). A, shield and cephalic appendages, dorsal; B, anterior and posterior lobes of sternite XII (between second ambulatory legs), ventral; C, exopod of left uropod, dorsal; D, telson, dorsal; E, left first gonopod, mesial; F, left second gonopod, anterior. Scale bars = 2 mm (A); 1 mm (BE); 2 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5458468" httpUri="https://zenodo.org/record/5458468/files/figure.png" pageId="83" pageNumber="293">Fig 46F</figureCitation>
) with spatulate distal segment, densely setose distally. Female with vestigial second right pleopod.
</paragraph>
<paragraph id="047C3855FFA87824FF6CFC84FE4FFCC6" blockId="83.[136,402,861,885]" box="[136,402,861,885]" pageId="83" pageNumber="293">
<emphasis id="36B7E447FFA87824FF6CFC84FEC4FCC6" bold="true" box="[136,281,861,885]" pageId="83" pageNumber="293">Colouration.</emphasis>
Unknown.
</paragraph>
<paragraph id="047C3855FFA87824FF6CFC44FEAFFC66" blockId="83.[136,774,925,982]" pageId="83" pageNumber="293">
<emphasis id="36B7E447FFA87824FF6CFC44FF36FC06" bold="true" box="[136,235,925,949]" pageId="83" pageNumber="293">Habitat.</emphasis>
Gastropod shells with or without zoanthid or
<taxonomicName id="C3C343D6FFA87824FF6CFC19FED4FC65" authorityName="Van Beneden" authorityYear="1844" box="[136,265,960,982]" class="Hydrozoa" family="Hydractiniidae" genus="Hydractinia" kingdom="Animalia" order="Anthoathecata" pageId="83" pageNumber="293" phylum="Cnidaria" rank="genus">
<emphasis id="36B7E447FFA87824FF6CFC19FED4FC65" box="[136,265,960,982]" italics="true" pageId="83" pageNumber="293">Hydractinia</emphasis>
</taxonomicName>
colonies.
</paragraph>
</subSubSection>
<subSubSection id="4CD96BDEFFA87824FF6CFC24FE86FBE6" pageId="83" pageNumber="293" type="distribution">
<paragraph id="047C3855FFA87824FF6CFC24FE86FBE6" blockId="83.[136,775,1021,1109]" pageId="83" pageNumber="293">
<emphasis id="36B7E447FFA87824FF6CFC24FEC2FBA6" bold="true" box="[136,287,1021,1045]" pageId="83" pageNumber="293">Distribution.</emphasis>
So far known only from the
<typeStatus id="DB7886F7FFA87824FD91FC27FD79FBA5" box="[629,676,1022,1046]" pageId="83" pageNumber="293">type</typeStatus>
locality, Nazca and Sala-y-Gómez Ridges, in the southeastern Pacific. Depth:
<quantity id="C33B95B0FFA87824FF3DFBE4FE8AFBE6" box="[217,343,1085,1109]" metricMagnitude="2" metricUnit="m" metricValue="2.38" metricValueMax="3.3" metricValueMin="1.46" pageId="83" pageNumber="293" unit="m" value="238.0" valueMax="330.0" valueMin="146.0">
146
<specimenCount id="12C5F3DCFFA87824FEEAFBE4FE8AFBE6" box="[270,343,1085,1109]" pageId="83" pageNumber="293" type="generic">330 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="4CD96BDEFFA87824FF6CFBA7FC02FEA6" pageId="83" pageNumber="293" type="discussion">
<paragraph id="047C3855FFA87824FF6CFBA7FDF1FA86" blockId="83.[136,775,1149,1334]" pageId="83" pageNumber="293">
<emphasis id="36B7E447FFA87824FF6CFBA7FF2AFB25" bold="true" box="[136,247,1150,1174]" pageId="83" pageNumber="293">Remarks.</emphasis>
<taxonomicName id="C3C343D6FFA87824FF18FB59FE22FB25" authorityName="Zhadan" authorityYear="1997" box="[252,511,1152,1174]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="species" species="stockmani">
<emphasis id="36B7E447FFA87824FF18FB59FE22FB25" box="[252,511,1152,1174]" italics="true" pageId="83" pageNumber="293">Oncopagurus stockmani</emphasis>
</taxonomicName>
is unique among species of
<taxonomicName id="C3C343D6FFA87824FF4CFB79FEE7FB05" authorityName="Lemaitre" authorityYear="2014" box="[168,314,1184,1206]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="36B7E447FFA87824FF4CFB79FEE7FB05" box="[168,314,1184,1206]" italics="true" pageId="83" pageNumber="293">Oncopagurus</emphasis>
</taxonomicName>
in having two rows of scales throughout the propodal rasp of the fourth pereopods. Individuals of this species are among the largest known in species of
<taxonomicName id="C3C343D6FFA87824FF6CFAD9FEC4FAA5" authorityName="Lemaitre" authorityYear="2014" box="[136,281,1280,1302]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="36B7E447FFA87824FF6CFAD9FEC4FAA5" box="[136,281,1280,1302]" italics="true" pageId="83" pageNumber="293">Oncopagurus</emphasis>
</taxonomicName>
, reaching up to 8.0 mm (a
<typeStatus id="DB7886F7FFA87824FDADFB27FD78FAA5" box="[585,677,1278,1302]" pageId="83" pageNumber="293" type="paratype">paratype</typeStatus>
reported by
<bibRefCitation id="605245A4FFA87824FF4FFAC4FEB1FA86" author="Zhadan DG" box="[171,364,1309,1334]" pageId="83" pageNumber="293" pagination="55 - 79" refId="ref73241" refString="Zhadan DG (1997) Deep-sea hermit crabs from the submerged ridges Nazka and Sala-y-Gomez, southeastern Pacific (Decapoda Anomura Parapaguridae). Arthropoda Selecta, 6 (1 / 2): 55 - 79." type="journal article" year="1997">Zhadan, 1997: 65</bibRefCitation>
) in shield length.
</paragraph>
<paragraph id="047C3855FFA87824FF6CFA87FCD5F806" blockId="83.[136,776,1373,1973]" pageId="83" pageNumber="293">
As noted by
<bibRefCitation id="605245A4FFA87824FEF4FA84FE6DFAC5" author="Zhadan DG" box="[272,432,1373,1398]" pageId="83" pageNumber="293" pagination="55 - 79" refId="ref73241" refString="Zhadan DG (1997) Deep-sea hermit crabs from the submerged ridges Nazka and Sala-y-Gomez, southeastern Pacific (Decapoda Anomura Parapaguridae). Arthropoda Selecta, 6 (1 / 2): 55 - 79." type="journal article" year="1997">Zhadan (1997)</bibRefCitation>
, the presence of several rows of scales on the propodal rasp of the fourth pereopod, misled
<bibRefCitation id="605245A4FFA87824FF6CFA44FEE7FA05" author="Zarenkov NA" box="[136,314,1437,1462]" pageId="83" pageNumber="293" pagination="218 - 244" refId="ref73200" refString="Zarenkov NA (1990) Decapods (Stenopodidea, Brachyura, Anomura) of the Naska and Sala-I-Comes underwater Ridges. Trudy Institute of Oceanology, 124: 218 - 244. [In Russian, with English summary]." type="journal article" year="1990">Zarenkov (1990)</bibRefCitation>
to report earlier specimens of
<taxonomicName id="C3C343D6FFA87824FD92FA79FF29FA65" authorityName="Zhadan" authorityYear="1997" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="species" species="stockmani">
<emphasis id="36B7E447FFA87824FD92FA79FF29FA65" italics="true" pageId="83" pageNumber="293">Oncopagurus stockmani</emphasis>
</taxonomicName>
collected at R/V Professor Stockman, sta 1873, as
<taxonomicName id="C3C343D6FFA87824FF6CFA39FE50FA45" baseAuthorityName="Studer" baseAuthorityYear="1883" box="[136,397,1504,1526]" class="Malacostraca" family="Parapaguridae" genus="Parapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="species" species="dimorphus">
<emphasis id="36B7E447FFA87824FF6CFA39FE50FA45" box="[136,397,1504,1526]" italics="true" pageId="83" pageNumber="293">Parapagurus dimorphus</emphasis>
</taxonomicName>
[=
<taxonomicName id="C3C343D6FFA87824FE55FA39FF17F9A5" authority="(Studer, 1883)" baseAuthorityName="Studer" baseAuthorityYear="1883" class="Malacostraca" family="Parapaguridae" genus="Sympagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="species" species="dimorphus">
<emphasis id="36B7E447FFA87824FE55FA39FD70FA45" box="[433,685,1504,1526]" italics="true" pageId="83" pageNumber="293">Sympagurus dimorphus</emphasis>
(
<bibRefCitation id="605245A4FFA87824FD5EFA04FF1FF9A5" author="Studer T" pageId="83" pageNumber="293" pagination="1 - 32" refId="ref72863" refString="Studer T (1883) Verzeichniss der Crustaceen, Welche Wahrend der Reise S. M. S. Gazelle an der Weskuste von Africa, Ascension und dem Cap der guten Hoffnung gesammelt wurden. Abandlungen der Preussischen Akademie der Wissenschaften, 2 (1882 - 1883): 1 - 32." type="journal article" year="1883">Studer, 1883</bibRefCitation>
)
</taxonomicName>
], a species which has two or more rows of scales on the propodal rasp. The ressemblance of these two species is only superficial as they differ in fundamental generic characters such as
<typeStatus id="DB7886F7FFA87824FEABF987FEA0F9C5" box="[335,381,1630,1654]" pageId="83" pageNumber="293">type</typeStatus>
and number of gills, and presence or shape of the epistomial spine.
<bibRefCitation id="605245A4FFA87824FE34F9A4FDB2F925" author="Zhadan DG" box="[464,623,1661,1686]" pageId="83" pageNumber="293" pagination="55 - 79" refId="ref73241" refString="Zhadan DG (1997) Deep-sea hermit crabs from the submerged ridges Nazka and Sala-y-Gomez, southeastern Pacific (Decapoda Anomura Parapaguridae). Arthropoda Selecta, 6 (1 / 2): 55 - 79." type="journal article" year="1997">Zhadan (1997)</bibRefCitation>
compared his
<taxonomicName id="C3C343D6FFA87824FF6CF979FE50F905" authorityName="Zhadan" authorityYear="1997" box="[136,397,1696,1718]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="species" species="stockmani">
<emphasis id="36B7E447FFA87824FF6CF979FE50F905" box="[136,397,1696,1718]" italics="true" pageId="83" pageNumber="293">Oncopagurus stockmani</emphasis>
</taxonomicName>
with other congenerics, and stated that two other species,
<taxonomicName id="C3C343D6FFA87824FE9AF919FDD9F965" authorityName="Lemaitre" authorityYear="2014" box="[382,516,1728,1750]" class="Malacostraca" family="Parapaguridae" genus="Sympagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="36B7E447FFA87824FE9AF919FDD9F965" box="[382,516,1728,1750]" italics="true" pageId="83" pageNumber="293">O. africanus</emphasis>
</taxonomicName>
and
<taxonomicName id="C3C343D6FFA87824FDDCF919FD7CF965" baseAuthorityName="de Saint Laurent" baseAuthorityYear="1972" box="[568,673,1728,1750]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="species" species="haigae">
<emphasis id="36B7E447FFA87824FDDCF919FD7CF965" box="[568,673,1728,1750]" italics="true" pageId="83" pageNumber="293">O. haigae</emphasis>
</taxonomicName>
, also had more than one row of scales on the propodal rasp of the fourth pereopod. However, in actuality the propodal rasp in
<taxonomicName id="C3C343D6FFA87824FF6CF8F9FED3F885" authorityName="Lemaitre" authorityYear="2014" box="[136,270,1824,1846]" class="Malacostraca" family="Parapaguridae" genus="Sympagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="36B7E447FFA87824FF6CF8F9FED3F885" box="[136,270,1824,1846]" italics="true" pageId="83" pageNumber="293">O. africanus</emphasis>
</taxonomicName>
and
<taxonomicName id="C3C343D6FFA87824FEA6F8F9FE71F885" baseAuthorityName="de Saint Laurent" baseAuthorityYear="1972" box="[322,428,1824,1846]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="species" species="haigae">
<emphasis id="36B7E447FFA87824FEA6F8F9FE71F885" box="[322,428,1824,1846]" italics="true" pageId="83" pageNumber="293">O. haigae</emphasis>
</taxonomicName>
may have more than one row of scales only on the proximal portion of the segment where the scales can often be arranged in a small cluster. Except for
<taxonomicName id="C3C343D6FFA87824FF6CF859FEC5F825" authorityName="Zhadan" authorityYear="1997" box="[136,280,1920,1942]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="species" species="stockmani">
<emphasis id="36B7E447FFA87824FF6CF859FEC5F825" box="[136,280,1920,1942]" italics="true" pageId="83" pageNumber="293">O. stockmani</emphasis>
</taxonomicName>
, all species of
<taxonomicName id="C3C343D6FFA87824FE27F859FD89F825" authorityName="Lemaitre" authorityYear="2014" box="[451,596,1920,1942]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="36B7E447FFA87824FE27F859FD89F825" box="[451,596,1920,1942]" italics="true" pageId="83" pageNumber="293">Oncopagurus</emphasis>
</taxonomicName>
invariably have only one row of scales on the propodal rasp, at least distally.
</paragraph>
<paragraph id="047C3855FFA87824FF6CF804FC02FEA6" blockId="83.[136,775,2013,2102]" lastBlockId="83.[835,1474,189,277]" pageId="83" pageNumber="293">
<materialsCitation id="B4AB3208FFA87824FF6CF804FC06FEA6" country="Chile" county="Juan Fernandez Islands" location="Nazca" municipality="southeastern Pacific. The" pageId="83" pageNumber="293" specimenCount="1">
Previous to this study,
<taxonomicName id="C3C343D6FFA87824FE9BF839FD59F845" authorityName="Zhadan" authorityYear="1997" box="[383,644,2016,2038]" class="Malacostraca" family="Parapaguridae" genus="Oncopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="83" pageNumber="293" phylum="Arthropoda" rank="species" species="stockmani">
<emphasis id="36B7E447FFA87824FE9BF839FD59F845" box="[383,644,2016,2038]" italics="true" pageId="83" pageNumber="293">Oncopagurus stockmani</emphasis>
</taxonomicName>
was known exclusively based on the
<typeStatus id="DB7886F7FFA87824FE7FF827FE14F7A5" box="[411,457,2046,2070]" pageId="83" pageNumber="293">type</typeStatus>
material from the
<location id="011C6E8EFFA87824FD71F827FD05F7A6" LSID="urn:lsid:plazi:treatment:8C6A8943FFB27824FF1DFA04FC02FEA6:011C6E8EFFA87824FD71F827FD05F7A6" box="[661,728,2046,2069]" country="Chile" county="Juan Fernandez Islands" municipality="southeastern Pacific. The" name="Nazca" pageId="83" pageNumber="293">Nazca</location>
and
<location id="011C6E8EFFA87824FF6CF7C4FEFFF786" LSID="urn:lsid:plazi:treatment:8C6A8943FFB27824FF1DFA04FC02FEA6:011C6E8EFFA87824FF6CF7C4FEFFF786" box="[136,290,2077,2101]" country="Chile" county="Juan Fernandez Islands" municipality="southeastern Pacific. The" name="Sala-y-Gomez" pageId="83" pageNumber="293">Sala-y-Gómez</location>
ridges, in the
<collectingMunicipality id="E418A22FFFA87824FE5EF7C4FD19F786" box="[442,708,2077,2101]" pageId="83" pageNumber="293">southeastern Pacific. The</collectingMunicipality>
range of this species is herein extended about
<quantity id="C33B95B0FFA87824FAF6FF67FAABFF66" box="[1298,1398,189,214]" metricMagnitude="6" metricUnit="m" metricValue="2.0" pageId="83" pageNumber="293" unit="km" value="2000.0">2000 km</quantity>
to the southeast, where it has been found in the
<collectingCounty id="ED1D40D9FFA87824FAFCFF07FC4CFEA6" pageId="83" pageNumber="293">Juan Fernández Islands</collectingCounty>
,
<collectingCountry id="7CD478C5FFA87824FC7AFF24FC06FEA6" box="[926,987,253,277]" name="Chile" pageId="83" pageNumber="293">Chile</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>