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(
<figureCitation id="9CF72A44FFABFFEC174C7288FEED5DDE" box="[262,337,1417,1442]" captionStart="Figure 2" captionStartId="6.[163,244,1709,1731]" captionTargetBox="[241,1358,197,1670]" captionTargetId="figure-27@6.[240,1360,195,1672]" captionTargetPageId="6" captionText="Figure 2. Phylogenetic relationships of the family Velutinidae (Bayesian tree based on the combined dataset), with clades (AL) collapsed by genera. Numbers at nodes indicate branch support values [posterior probability (PP) and ultrafast bootstrap (UFb) values, respectively]; support values are shown only when at least one of them is ≥ 80%; white circles at nodes indicate maximum support (PP = 1, Ufb = 100). On the right side of the tree, for each genus is reported a representative photograph of the shell, a schematic representation of the radula, the depth range (in metres) and the geographical distribution." figureDoi="http://doi.org/10.5281/zenodo.7814312" httpUri="https://zenodo.org/record/7814312/files/figure.png" pageId="31" pageNumber="955">FIGS 2</figureCitation>
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<figureCitation id="9CF72A44FFABFFEC17147288FED05DDE" box="[350,364,1417,1442]" captionStart="Figure 3" captionStartId="7.[145,225,1585,1607]" captionTargetBox="[271,1296,200,1554]" captionTargetId="figure-25@7.[269,1299,195,1557]" captionTargetPageId="7" captionText="Figure 3. Phylogenetic relationships of the family Velutinidae (Bayesian tree based on the combined dataset) showing clades AL expanded. Numbers at nodes indicate branch support values [posterior probability (PP) and ultrafast bootstrap (UFb) values, respectively]; support values are shown only when at least one of them is ≥ 80%; white circles at nodes indicate maximum support (PP = 1, Ufb = 100). Boxes indicate hypothetical species; dashed lines indicate possible alternative species partitions. Coloured boxes on the right indicate genera, and white boxes indicate subfamilies. Letters at the end of sample codes broadly indicate the area of origin:AK, Alaska; ANT, Antarctica; AR, Argentina; BC, British Columbia; BRE, Bretagne; CA, California; CH, China; COR, Corse; CR, Costa Rica; FL, Florida; GF, French Guiana; GP, Guadeloupe; HI, Hawaii; IT, Italy; MG, Madagascar; MQ, Martinique; MX, Mexico; MZ, Mozambique; NC, New Caledonia; NL, Newfoundland; NO, Norway; OR, Oregon; PA, Panama; PF, French Polynesia; PG, Papua New Guinea; PH, Philippines; QLS, Queensland; RE, Reunion; SA, Saudi Arabia; SP, Spain; TAS, Tasmania; TW, Taiwan; VU, Vanuatu; WA, Washington State; ZNZ, Zanzibar." figureDoi="http://doi.org/10.5281/zenodo.7814314" httpUri="https://zenodo.org/record/7814314/files/figure.png" pageId="31" pageNumber="955">3</figureCitation>
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<figureCitation id="9CF72A44FFABFFEC17337288FE055DDE" box="[377,441,1417,1442]" captionStart="Figure 5" captionStartId="12.[163,242,1701,1723]" captionTargetBox="[214,1391,198,1668]" captionTargetId="figure-27@12.[211,1394,195,1671]" captionTargetPageId="12" captionText="Figure 5. A, Pacifica lentiginosa sp. nov., MNHN-IM-2009-16140, holotype. B, Pacifica sp. (L18), MNHN-IM-2007-33407. C, Djiboutia sp. (L22), MNHN-IM-2019-9053. D, Coriocella sp., UF-IZ-523481. E, Coriocella sp., MNHN-IM-2019-26097. F, Lamellaria mopsicolor, MNHN-IM-2013-73086. G, Lamellaria latens complex (L41), MNHN-IM-2019-6391. H, Marsenia herberti, MNHN-IM-2019-8219. I, Marsenia perspicua complex (L55), MNHN-IM-2019-13522. Photograph credits: AC, F, MNHN, Philippe Maestrati; D, Florida Museum of Natural History—Invertebrate Zoology, Gustav Paulay; E, MNHN, Giulia Fassio; G, I, MNHN, Gilles Devauchelle; H, MNHN, Laurent Charles. For the sequenced specimens, the corresponding clade code (see Fig. 3) is reported in parentheses." figureDoi="http://doi.org/10.5281/zenodo.7814322" httpUri="https://zenodo.org/record/7814322/files/figure.png" pageId="31" pageNumber="955">5D, E</figureCitation>
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<figureCitation id="9CF72A44FFABFFEC178D7288FE5A5DDE" box="[455,486,1417,1442]" captionStart="Figure 8" captionStartId="15.[145,223,1631,1653]" captionTargetBox="[224,1345,198,1599]" captionTargetId="figure-107@15.[221,1348,195,1602]" captionTargetPageId="15" captionText="Figure 8. Velutinidae shells. A, Djiboutia sp. (L26), MNHN-IM-2019-7668. B, Coriocella sp. (L27), MNHN-IM-2013-84230. C, Lamellaria latens complex (L41), MNHN-IM-2019-14179. D, Marsenia perspicua complex (L55), MNHN-IM-2019-14574. Scale bars: A, C, D, 1 mm; B, 5 mm. For the sequenced specimens, the corresponding species code (see Fig. 3) is reported in parentheses." figureDoi="http://doi.org/10.5281/zenodo.7814330" httpUri="https://zenodo.org/record/7814330/files/figure.png" pageId="31" pageNumber="955">8B</figureCitation>
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<figureCitation id="9CF72A44FFABFFEC17B97288FDAA5DDE" box="[499,534,1417,1442]" captionStart="Figure 9" captionStartId="16.[163,246,1664,1686]" captionTargetBox="[230,1376,198,1631]" captionTargetId="figure-26@16.[227,1379,195,1634]" captionTargetPageId="16" captionText="Figure 9. Velutinidae protoconchs. A, Marseniopsis conica, BAS 03-764. B, Marsenina rhombica, BAU 4229. C, Calyptoconcha pellucida, BAU 3758. D, Variolipallium cerebroides, NIWA 74086.A. E, Pacifica lentiginosa sp. nov., MNHN-IM-2009-16140, holotype.F, Pacifica sp. (L16), MNHN-IM-2013-58543. G, Djiboutia sp. (L22), MNHN-IM-2019-9053. H, Coriocella sp. (L27), MNHN-IM-2013-84230. I, Lamellaria latens complex (L41), MNHN-IM-2019-14179. J, Lamellaria mopsicolor, MNHN-IM-2013-73086. K, Marsenia perspicua complex (L55), MNHN-IM-2019-14574. L, Marsenia sp. (L42), MNHN-IM-2019-7641. Scale bars: AL, 500 µm. For the sequenced specimens, the corresponding species code (see Fig. 3) is reported in parentheses. Arrows indicate protoconch I protoconch II boundary." figureDoi="http://doi.org/10.5281/zenodo.7814334" httpUri="https://zenodo.org/record/7814334/files/figure.png" pageId="31" pageNumber="955">9H</figureCitation>
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<figureCitation id="9CF72A44FFABFFEC146E7288FDF05DDE" box="[548,588,1417,1442]" captionStart="Figure 10" captionStartId="17.[145,226,1664,1686]" captionTargetBox="[212,1358,198,1631]" captionTargetId="figure-24@17.[209,1361,195,1634]" captionTargetPageId="17" captionText="Figure 10. Velutinidae radulae. A, Marseniopsis mollis, CE 6137. B, Marsenina rhombica, BAU 4229. C, Velutina cf. coriacea, LACM 182870. D, Calyptoconcha pellucida, BAU 3751. E, Variolipallium regium sp. nov., MNHN-IM-2013-69285, holotype. F, Variolipallium sp. (L10), MNHN-IM-2013-69280. G, Pacifica lentiginosa sp. nov., MNHN-IM-2009-16140, holotype. H, Pacifica sp. (L15), MNHN-IM-2013-59969. I, Djiboutia sp. (L22), MNHN-IM-2019-9053. J, Coriocella sp. (L27), MNHN-IM-2013-84230. K, Lamellaria mopsicolor, MNHN-IM-2013-73086. L, Lamellaria latens complex (L41), MNHN-IM-2019-14113. M, Marsenia herberti, MNHN-IM-2019-8503. N, Marsenia sp. (L45), MNHN-IM-2013-59960. O, Marsenia perspicua complex (L55), MNHN-IM-2019-5980. Scale bars: AO, 100 µm. For the sequenced specimens, the corresponding species code (see Fig. 3) is reported in parentheses." figureDoi="http://doi.org/10.5281/zenodo.7814338" httpUri="https://zenodo.org/record/7814338/files/figure.png" pageId="31" pageNumber="955">10J</figureCitation>
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<figureCitation id="9CF72A44FFABFFEC14137288FD3F5DDE" box="[601,643,1417,1442]" captionStart="Figure 11" captionStartId="18.[163,243,1664,1686]" captionTargetBox="[230,1376,198,1631]" captionTargetId="figure-26@18.[227,1379,195,1634]" captionTargetPageId="18" captionText="Figure 11. Velutinidae jaws. A, M, Marseniopsis mollis, CE 6152. B, N, O, Marsenina rhombica, BAU 2339. C, Limneria prolongata, BAU 4228. D, Calyptoconcha pellucida, BAU 3758. E, Variolipallium regium sp. nov., MNHN-IM-2013-69285, holotype. F, Variolipallium sp. (L10), MNHN-IM-2013-69280. G, Pacifica lentiginosa sp. nov., MNHN-IM-2009-16140, holotype.H, Pacifica sp. (L19), MNHN-IM-2009-16136.I,Djiboutia sp.(L22), MNHN-IM-2019-9053.J,Coriocella cf.jayi (L28), MNHN-IM-2009-16119. K, Lamellaria latens complex (L40), BAU 2536.2. L, Marsenia sp. (L45), MNHN-IM-2013-59960. Scale bars: H, MO, 100 µm; AG, IL, 500 µm. For the sequenced specimens, the corresponding species code (see Fig. 3) is reported in parentheses." figureDoi="http://doi.org/10.5281/zenodo.7814342" httpUri="https://zenodo.org/record/7814342/files/figure.png" pageId="31" pageNumber="955">11J</figureCitation>
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<typeStatus id="DB778863FFABFFEC17B072B3FD975DBB" box="[506,555,1458,1479]" pageId="31" pageNumber="955">type</typeStatus>
species
<taxonomicName id="C3CC4D42FFABFFEC14CD72B0FE345D99" ID-CoL="6B4TJ" authority="Blainville, 1824" authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="nigra">
<emphasis id="36B8EAD3FFABFFEC14CD72B0FF735D99" italics="true" pageId="31" pageNumber="955">Coriocella nigra</emphasis>
<bibRefCitation id="605D4B30FFABFFEC169F72D1FE345D99" author="de Blainville HMD" box="[213,392,1488,1510]" pageId="31" pageNumber="955" pagination="1 - 392" refId="ref25490" refString="de Blainville HMD. 1824. Mollusques, Mollusca (Malacozoaires) In: Cuvier F, ed. Dictionnaire des Sciences Naturelles, Vol. 32. Strasbourg &amp; Paris: Levrault &amp; Le Normant, 1 - 392." type="book chapter" year="1824">Blainville, 1824</bibRefCitation>
</taxonomicName>
by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="4CD6654AFFABFFEC16DB72EFFE4B5E78" box="[145,503,1518,1540]" pageId="31" pageNumber="955" type="reference_group">
<paragraph id="047336C1FFABFFEC16DB72EFFE4B5E78" blockId="31.[145,761,1457,1632]" box="[145,503,1518,1540]" pageId="31" pageNumber="955">
<treatmentCitationGroup id="24DC11EFFFABFFEC16DB72EFFE4B5E78" box="[145,503,1518,1540]" pageId="31" pageNumber="955">
<treatmentCitation id="856D10D0FFABFFEC16DB72EFFE4F5E78" author="Swainson W" box="[145,499,1518,1540]" page="355" pageId="31" pageNumber="955" year="1840">
<taxonomicName id="C3CC4D42FFABFFEC16DB72EFFE4F5E78" ID-CoL="d2fe1387-d0df-477f-88d8-d498ef54d831" authority="Swainson, 1840: 355" authorityName="Swainson" authorityPageNumber="355" authorityYear="1840" box="[145,499,1518,1540]" class="Gastropoda" family="Velutinidae" genus="Chelinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="genus">
<emphasis id="36B8EAD3FFABFFEC16DB72EFFEB75E7F" box="[145,267,1518,1539]" italics="true" pageId="31" pageNumber="955">Chelinotus</emphasis>
<bibRefCitation id="605D4B30FFABFFEC174572EFFE4F5E78" author="Swainson W" box="[271,499,1518,1540]" pageId="31" pageNumber="955" refId="ref28237" refString="Swainson W. 1840. A treatise on malacology or shells and shell-fish. London: Longman." type="book" year="1840">Swainson, 1840: 355</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4CD6654AFFABFFEC17B672EEFD185E5F" pageId="31" pageNumber="955" type="multiple">
<paragraph id="047336C1FFABFFEC17B672EEFD185E5F" blockId="31.[145,761,1457,1632]" pageId="31" pageNumber="955">
<typeStatus id="DB778863FFABFFEC17B672EEFD915E78" box="[508,557,1519,1540]" pageId="31" pageNumber="955">type</typeStatus>
species:
<taxonomicName id="C3CC4D42FFABFFEC14C672EEFDBB5E5E" authority="Quoy &amp; Gaimard, 1832" authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Velutinidae" genus="Sigaretus" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="tonganus">
<emphasis id="36B8EAD3FFABFFEC14C672EEFF465E5F" italics="true" pageId="31" pageNumber="955">Sigaretus tonganus</emphasis>
Quoy &amp; Gaimard, 1832
</taxonomicName>
by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="4CD6654AFFABFFEC16DB712DFD3B5E1C" pageId="31" pageNumber="955" type="reference_group">
<paragraph id="047336C1FFABFFEC16DB712DFD3B5E1C" blockId="31.[145,761,1457,1632]" pageId="31" pageNumber="955">
<treatmentCitationGroup id="24DC11EFFFABFFEC16DB712DFD3B5E1C" pageId="31" pageNumber="955">
<treatmentCitation id="856D10D0FFABFFEC16DB712DFDEB5E3D" author="Herrmannsen AN" box="[145,599,1580,1602]" page="221" pageId="31" pageNumber="955" year="1846">
<taxonomicName id="C3CC4D42FFABFFEC16DB712DFDEB5E3D" ID-CoL="0bc0ce14-c7a4-421e-be93-fa0918080d21" authority="Herrmannsen, 1846: 221" authorityName="Herrmannsen" authorityPageNumber="221" authorityYear="1846" box="[145,599,1580,1602]" class="Gastropoda" family="Velutinidae" genus="Chelyonotus" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="genus">
<emphasis id="36B8EAD3FFABFFEC16DB712DFE9E5E3D" box="[145,290,1580,1601]" italics="true" pageId="31" pageNumber="955">Chelyonotus</emphasis>
<bibRefCitation id="605D4B30FFABFFEC1761712DFDEB5E3D" author="Herrmannsen AN" box="[299,599,1580,1602]" pageId="31" pageNumber="955" refId="ref26614" refString="Herrmannsen AN. 1846. Indicis Generum Malacozoorum primordia, Vol. 1 [i - xxviii + 1 - 232 (1846); 233 - 637 (1847)]. Kassel: Fischer.." type="book" year="1846">Herrmannsen, 1846: 221</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; unnecessary replacement name for
<taxonomicName id="C3CC4D42FFABFFEC17DB714BFD3B5E1C" ID-CoL="d2fe1387-d0df-477f-88d8-d498ef54d831" authority="Swainson." box="[401,647,1610,1632]" class="Gastropoda" family="Velutinidae" genus="Chelinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="genus">
<emphasis id="36B8EAD3FFABFFEC17DB714BFDB75E23" box="[401,523,1610,1631]" italics="true" pageId="31" pageNumber="955">Chelinotus</emphasis>
Swainson.
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4CD6654AFFABFFEC16DB7188FE135F65" pageId="31" pageNumber="955" type="multiple">
<paragraph id="047336C1FFABFFEC16DB7188FE135F65" blockId="31.[145,762,1673,1818]" pageId="31" pageNumber="955">
<emphasis id="36B8EAD3FFABFFEC16DB7188FEE15EE3" box="[145,349,1673,1695]" italics="true" pageId="31" pageNumber="955">Included species:</emphasis>
<taxonomicName id="C3CC4D42FFABFFEC17397188FD4E5EE2" authority="Wellens, 1991" authorityName="Wellens" authorityYear="1991" box="[371,754,1673,1695]" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="hibyae">
<emphasis id="36B8EAD3FFABFFEC17397188FDFF5EE2" box="[371,579,1673,1694]" italics="true" pageId="31" pageNumber="955">Coriocella hibyae</emphasis>
<bibRefCitation id="605D4B30FFABFFEC14067188FD4E5EE2" author="Wellens W" box="[588,754,1673,1695]" pageId="31" pageNumber="955" pagination="73 - 80" refId="ref28653" refString="Wellens W. 1991. Coriocella hibyae n. sp. a new Lamellaria species (Gastropoda: Prosobranchia) from the Republic of the Maldives. Journal of Conchology 34: 73 - 80." type="journal article" year="1991">Wellens, 1991</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C3CC4D42FFABFFEC16DB71A9FDAF5EC2" authority="Wellens, 1996" authorityName="Wellens" authorityYear="1996" box="[145,531,1704,1726]" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="jayi">
<emphasis id="36B8EAD3FFABFFEC16DB71A9FEEC5EC1" box="[145,336,1704,1725]" italics="true" pageId="31" pageNumber="955">Coriocella jayi</emphasis>
Wellens, 1996
</taxonomicName>
,
<taxonomicName id="C3CC4D42FFABFFEC146F71A9FEED5EA0" authority="Blainville, 1824" authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="nigra">
<emphasis id="36B8EAD3FFABFFEC146F71A9FD455EC1" box="[549,761,1704,1725]" italics="true" pageId="31" pageNumber="955">Coriocella nigra</emphasis>
<bibRefCitation id="605D4B30FFABFFEC16DB71C7FEED5EA0" author="de Blainville HMD" box="[145,337,1734,1756]" pageId="31" pageNumber="955" pagination="1 - 392" refId="ref25490" refString="de Blainville HMD. 1824. Mollusques, Mollusca (Malacozoaires) In: Cuvier F, ed. Dictionnaire des Sciences Naturelles, Vol. 32. Strasbourg &amp; Paris: Levrault &amp; Le Normant, 1 - 392." type="book chapter" year="1824">Blainville, 1824</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C3CC4D42FFABFFEC172871C7FD4E5EA0" authority="Wellens, 1999" authorityName="Wellens" authorityYear="1999" box="[354,754,1734,1756]" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="safagae">
<emphasis id="36B8EAD3FFABFFEC172871C7FDFD5EA0" box="[354,577,1734,1756]" italics="true" pageId="31" pageNumber="955">Coriocella safagae</emphasis>
<bibRefCitation id="605D4B30FFABFFEC140071C7FD4E5EA0" author="Wellens W" box="[586,754,1734,1756]" pageId="31" pageNumber="955" pagination="17 - 24" refId="ref28759" refString="Wellens W. 1999. Coriocella safagae n. sp. a new lamellariid (Gastropoda: Prosobranchia) from Safaga, Red Sea, Egypt. Journal of Conchology 36: 17 - 24." type="journal article" year="1999">Wellens, 1999</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C3CC4D42FFABFFEC16DB71E4FDAC5E87" authority="(Bergh, 1875)" baseAuthorityName="Bergh" baseAuthorityYear="1875" box="[145,528,1765,1787]" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="semperi">
<emphasis id="36B8EAD3FFABFFEC16DB71E4FED45E87" box="[145,360,1765,1787]" italics="true" pageId="31" pageNumber="955">Coriocella semperi</emphasis>
(Bergh, 1875)
</taxonomicName>
,
<taxonomicName id="C3CC4D42FFABFFEC145771E4FE175F65" authority="(Quoy &amp; Gaimard, 1832)" baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1832" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="tongana">
<emphasis id="36B8EAD3FFABFFEC145771E4FD455E87" box="[541,761,1765,1787]" italics="true" pageId="31" pageNumber="955">Coriocella tongana</emphasis>
(Quoy &amp; Gaimard, 1832)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="4CD6654AFFABFFEC16DB7042FC4B5BA6" pageId="31" pageNumber="955" type="description">
<paragraph id="047336C1FFABFFEC16DB7042FC43594B" blockId="31.[145,760,1858,1911]" lastBlockId="31.[809,1425,197,986]" pageId="31" pageNumber="955">
<emphasis id="36B8EAD3FFABFFEC16DB7042FE975F24" box="[145,299,1859,1880]" italics="true" pageId="31" pageNumber="955">Description:</emphasis>
Body of small to medium size for the subfamily,
<quantity id="C3349B24FFABFFEC17017060FE695F0B" box="[331,469,1889,1911]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="8.5" metricValueMin="1.5" pageId="31" pageNumber="955" unit="cm" value="5.0" valueMax="8.5" valueMin="1.5">1.58.5 cm</quantity>
. Shell thin, strongly to moderately calcified; ear shaped, low spire, with expanded aperture; smooth or weakly sculptured by axial growth lines; completely enclosed by the mantle. Periostracum thin.
</paragraph>
<paragraph id="047336C1FFABFFEC150B7641FBB459AC" blockId="31.[809,1425,197,986]" pageId="31" pageNumber="955">Protoconch of 1.01.3 whorls; protoconch I 0.461.10 whorls, nucleus diameter 122150 μm, smooth, with subsutural axial folds; protoconch II with or without axial growth lines; protoconchteleoconch boundary not always distinct.</paragraph>
<paragraph id="047336C1FFABFFEC150B76D8FB1C5AF4" blockId="31.[809,1425,197,986]" pageId="31" pageNumber="955">Mantle dome shaped, outline rounded; thick, with two to six warts on the dorsum; with anterior siphon folds; smooth, wrinkled, often velvet-like in texture; colour highly variable, beige, yellow, red, violet, blue, dark green, brown, black, often patterned with irregular colour patches or spots.</paragraph>
<paragraph id="047336C1FFABFFEC150B7590FA845A98" blockId="31.[809,1425,197,986]" pageId="31" pageNumber="955">Penis to the right of the right cephalic tentacle; with tip of the seminal duct protruded from the penis tip. Vas deferens without a free loop in haemocoel.</paragraph>
<paragraph id="047336C1FFABFFEC150B75ECFC6C5BC7" blockId="31.[809,1425,197,986]" pageId="31" pageNumber="955">Radula reduced taenioglossate, with formula 0:1:1:1:0; rachidian tooth base bifurcated; rachidian cusp with several external small denticles; lateral teeth elongated, with one external triangular pointed and bold cusp with small denticles on the external side, plus one truncated projection with small denticles on the distal side.</paragraph>
<paragraph id="047336C1FFABFFEC150B74C5FC4B5BA6" blockId="31.[809,1425,197,986]" box="[833,1015,964,986]" pageId="31" pageNumber="955">Jaws elongated.</paragraph>
</subSubSection>
<subSubSection id="4CD6654AFFABFFEC15637303FBA65C08" pageId="31" pageNumber="955" type="distribution">
<paragraph id="047336C1FFABFFEC15637303FBA65C08" blockId="31.[809,1425,1026,1140]" pageId="31" pageNumber="955">
<emphasis id="36B8EAD3FFABFFEC15637303FC6E5C6B" box="[809,978,1026,1047]" italics="true" pageId="31" pageNumber="955">Distribution:</emphasis>
Indo-West Pacific (
<collectingCountry id="7CDB7651FFABFFEC12A77303FA365C64" box="[1261,1418,1026,1048]" name="Madagascar" pageId="31" pageNumber="955">Madagascar</collectingCountry>
,
<collectingCountry id="7CDB7651FFABFFEC15637320FC315C4B" box="[809,909,1057,1079]" name="Réunion" pageId="31" pageNumber="955">Reunion</collectingCountry>
,
<collectingCountry id="7CDB7651FFABFFEC15D37320FBAC5C4B" box="[921,1040,1057,1079]" name="Mauritius" pageId="31" pageNumber="955">Mauritius</collectingCountry>
, Red Sea,
<collectingCountry id="7CDB7651FFABFFEC12C67320FB445C4B" box="[1164,1272,1057,1079]" name="Maldives" pageId="31" pageNumber="955">Maldives</collectingCountry>
,
<collectingCountry id="7CDB7651FFABFFEC134F7320FA305C4B" box="[1285,1420,1057,1079]" name="Philippines" pageId="31" pageNumber="955">Philippines</collectingCountry>
, northern
<collectingCountry id="7CDB7651FFABFFEC15DE7341FBBE5C2A" box="[916,1026,1088,1110]" name="Australia" pageId="31" pageNumber="955">Australia</collectingCountry>
,
<collectingCountry id="7CDB7651FFABFFEC12477341FB005C2A" box="[1037,1212,1088,1110]" name="New Caledonia" pageId="31" pageNumber="955">New Caledonia</collectingCountry>
,
<collectingCountry id="7CDB7651FFABFFEC128C7341FA315C2A" box="[1222,1421,1088,1110]" name="French Polynesia" pageId="31" pageNumber="955">French Polynesia</collectingCountry>
,
<collectingCountry id="7CDB7651FFABFFEC1563735FFCCD5C0F" box="[809,881,1118,1139]" name="Tonga" pageId="31" pageNumber="955">Tonga</collectingCountry>
);
<quantity id="C3349B24FFABFFEC15C9735FFC6B5C08" box="[899,983,1118,1140]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="18.0" metricValueMin="0.0" pageId="31" pageNumber="955" unit="m" value="9.0" valueMax="18.0" valueMin="0.0">018 m</quantity>
deep.
</paragraph>
</subSubSection>
<subSubSection id="4CD6654AFFABFFEC1563739CFC4D5EC0" pageId="31" pageNumber="955" type="discussion">
<paragraph id="047336C1FFABFFEC1563739CFC4D5EC0" blockId="31.[809,1425,1181,1724]" pageId="31" pageNumber="955">
<emphasis id="36B8EAD3FFABFFEC1563739CFB905CCE" box="[809,1068,1181,1202]" italics="true" pageId="31" pageNumber="955">
Remarks:
<taxonomicName id="C3CC4D42FFABFFEC15F9739CFB905CCE" authorityName="Blainville" authorityYear="1824" box="[947,1068,1181,1202]" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="genus">Coriocella</taxonomicName>
</emphasis>
is probably the lamellarine genus that can be recognized best from the external morphology only, thanks to the presence of a variable, species-specific number of warts, well evident on the dorsum of living animals (e.g. three for
<taxonomicName id="C3CC4D42FFABFFEC15D17237FBE65D30" authorityName="Blainville" authorityYear="1824" box="[923,1114,1334,1356]" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="nigra">
<emphasis id="36B8EAD3FFABFFEC15D17237FBE65D30" box="[923,1114,1334,1356]" italics="true" pageId="31" pageNumber="955">Coriocella nigra</emphasis>
</taxonomicName>
; five for
<taxonomicName id="C3CC4D42FFABFFEC12837237FC705D16" authority="Wellens, 1999" authorityName="Wellens" authorityYear="1999" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="safage">
<emphasis id="36B8EAD3FFABFFEC12837237FA2D5D37" box="[1225,1425,1334,1355]" italics="true" pageId="31" pageNumber="955">Coriocella safage</emphasis>
<bibRefCitation id="605D4B30FFABFFEC15637254FC705D16" author="Wellens W" box="[809,972,1365,1387]" pageId="31" pageNumber="955" pagination="17 - 24" refId="ref28759" refString="Wellens W. 1999. Coriocella safagae n. sp. a new lamellariid (Gastropoda: Prosobranchia) from Safaga, Red Sea, Egypt. Journal of Conchology 36: 17 - 24." type="journal article" year="1999">Wellens, 1999</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C3CC4D42FFABFFEC15967254FB145D16" authorityName="Wellens" authorityYear="1991" box="[988,1192,1365,1386]" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="hibyae">
<emphasis id="36B8EAD3FFABFFEC15967254FB145D16" box="[988,1192,1365,1386]" italics="true" pageId="31" pageNumber="955">Coriocella hibyae</emphasis>
</taxonomicName>
,
<taxonomicName id="C3CC4D42FFABFFEC12F27254FA2D5D17" baseAuthorityName="Bergh" baseAuthorityYear="1875" box="[1208,1425,1365,1387]" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="semperi">
<emphasis id="36B8EAD3FFABFFEC12F27254FA2D5D17" box="[1208,1425,1365,1387]" italics="true" pageId="31" pageNumber="955">Coriocella semperi</emphasis>
</taxonomicName>
and
<taxonomicName id="C3CC4D42FFABFFEC15287275FA375DF5" authority="(Quoy &amp; Gaimard, 1832)" baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1832" box="[866,1419,1396,1418]" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="tongana">
<emphasis id="36B8EAD3FFABFFEC15287275FBF55DF5" box="[866,1097,1396,1417]" italics="true" pageId="31" pageNumber="955">Coriocella tongana</emphasis>
(Quoy &amp; Gaimard, 1832)
</taxonomicName>
; and six for
<taxonomicName id="C3CC4D42FFABFFEC15F57293FBCD5DD4" authorityName="Wellens" authorityYear="1996" box="[959,1137,1426,1448]" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="species" species="jayi">
<emphasis id="36B8EAD3FFABFFEC15F57293FBCD5DD4" box="[959,1137,1426,1448]" italics="true" pageId="31" pageNumber="955">Coriocella jayi</emphasis>
</taxonomicName>
), and the typical light brown or dark velvet-like coloration. However, the warts can become barely visible and the colour often vanishes once the specimen is preserved in alcohol. The shell shape is hardly distinguishable from the other low-spired lamellariine genera (
<taxonomicName id="C3CC4D42FFABFFEC1578714BFC075E23" authorityName="Montagu" authorityYear="1816" box="[818,955,1610,1631]" class="Gastropoda" family="Velutinidae" genus="Lamellaria" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="genus">
<emphasis id="36B8EAD3FFABFFEC1578714BFC075E23" box="[818,955,1610,1631]" italics="true" pageId="31" pageNumber="955">Lamellaria</emphasis>
</taxonomicName>
,
<taxonomicName id="C3CC4D42FFABFFEC159B714AFBF85E1C" authorityName="Oken" authorityYear="1823" box="[977,1092,1611,1632]" class="Gastropoda" family="Velutinidae" genus="Marsenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="genus">
<emphasis id="36B8EAD3FFABFFEC159B714AFBF85E1C" box="[977,1092,1611,1632]" italics="true" pageId="31" pageNumber="955">Marsenia</emphasis>
</taxonomicName>
and
<taxonomicName id="C3CC4D42FFABFFEC12C4714BFABE5E23" authorityName="Vayssiere" authorityYear="1912" box="[1166,1282,1610,1631]" class="Gastropoda" family="Velutinidae" genus="Djiboutia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="genus">
<emphasis id="36B8EAD3FFABFFEC12C4714BFABE5E23" box="[1166,1282,1610,1631]" italics="true" pageId="31" pageNumber="955">Djiboutia</emphasis>
</taxonomicName>
). However, most adult specimens of
<taxonomicName id="C3CC4D42FFABFFEC12157168FB6A5E02" authorityName="Blainville" authorityYear="1824" box="[1119,1238,1641,1662]" class="Gastropoda" family="Velutinidae" genus="Coriocella" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="955" phylum="Mollusca" rank="genus">
<emphasis id="36B8EAD3FFABFFEC12157168FB6A5E02" box="[1119,1238,1641,1662]" italics="true" pageId="31" pageNumber="955">Coriocella</emphasis>
</taxonomicName>
show a higher level of shell calcification compared with the rest of the subfamily.
</paragraph>
</subSubSection>
</treatment>
</document>