324 lines
47 KiB
XML
324 lines
47 KiB
XML
<document id="CDF2C6C0832DDF903B9B23135DFE11F8" ID-DOI="10.5281/zenodo.184685" ID-GBIF-Dataset="3860ca0e-b126-45dd-819b-ea8bf10450cf" ID-ISSN="1175-5326" ID-Zenodo-Dep="184685" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459998832640" checkinUser="plazi" docAuthor="Zettel, Herbert, Lane, David J. W. & Moore, Stephen" docDate="2008" docId="8C5C87B2FFFE6B36FF13FD68B006FC61" docLanguage="en" docName="zt01920p068.pdf" docOrigin="Zootaxa 1920" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Aphelocheirus Zettel, Lane & Moore, 2008, sp.n." docType="treatment" docVersion="8" lastPageNumber="66" masterDocId="7065FFCAFFFF6B33FF84FFAFB571FFCB" masterDocTitle="Aphelocheirus (s. str.) bruneiensis sp. n., a new benthic water bug (Hemiptera: Heteroptera: Aphelocheiridae) from Brunei, and a key to Bornean Aphelocheirus" masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1698233038423" updateUser="plazi">
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<mods:mods id="E32FF338DCDFD100E80118023326BC52" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4E9DA9BC35443E89825E39AA12EF1F62">
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<mods:title id="7A74B18F53763FEE2C5C493EE614E840">Aphelocheirus (s. str.) bruneiensis sp. n., a new benthic water bug (Hemiptera: Heteroptera: Aphelocheiridae) from Brunei, and a key to Bornean Aphelocheirus</mods:title>
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</mods:titleInfo>
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<mods:name id="DA5F5ACD336FAF71959792D3CB2E3B84" type="personal">
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<mods:role id="AD930A4FBC2E312D48DE80E0063EE1B1">
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<mods:roleTerm id="9B46EF273D06F1880D50874C33CD1332">Author</mods:roleTerm>
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<mods:namePart id="ADB87CB3E4BC692FC6FA604732E07654">Zettel, Herbert</mods:namePart>
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</mods:name>
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<mods:name id="D90BD4B050571F2F4003E9559851F01A" type="personal">
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<mods:roleTerm id="46E01BA1D2413B3368D3FBF06A6B73CF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="841980F5397A297F1639553096420794">Lane, David J. W.</mods:namePart>
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</mods:name>
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<mods:name id="AB9DADEC0A3DEB80F947F01C716B727C" type="personal">
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<mods:role id="4DED195F6CEAC66497CE30A0EC576998">
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<mods:roleTerm id="21366D67F7E0823623A7711FF6631E3A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2A2DA3172AEDBDAA83D28A79CB14A48B">Moore, Stephen</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="688C71BAC6CAB620EAE508C781FDF40B">text</mods:typeOfResource>
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<mods:title id="C5D377233141C0D5C54CEC2A384A9151">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5FB50E9C081B2EAE1DB6C5C064CF29D0">
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<mods:date id="5FE5B7B6C8A0EA9F03F23B0B572151FB">2008</mods:date>
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<mods:detail id="66C501B48FEF400043EA222113D57A13" type="volume">
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<mods:number id="DE8F8AC5B488DC1C2FF74D492682154C">1920</mods:number>
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<mods:end id="E96447D9C62DEA52E1B76C737B76109B">68</mods:end>
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<mods:classification id="FADF40961A9AEF3391E8664382894BF1">journal article</mods:classification>
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<mods:identifier id="49CF3E7AA5E151E2835845751274B556" type="DOI">10.5281/zenodo.184685</mods:identifier>
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<mods:identifier id="AF9F3C99B188789FFABC0B90CB1EE04C" type="GBIF-Dataset">3860ca0e-b126-45dd-819b-ea8bf10450cf</mods:identifier>
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<mods:identifier id="C6C041A7F6BECED46C0329B41850A92D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F29EBC721204320B0FEDF466ACE67E88" type="Zenodo-Dep">184685</mods:identifier>
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<treatment id="8C5C87B2FFFE6B36FF13FD68B006FC61" ID-DOI="http://doi.org/10.5281/zenodo.6227569" ID-GBIF-Taxon="119380595" ID-Zenodo-Dep="6227569" LSID="urn:lsid:plazi:treatment:8C5C87B2FFFE6B36FF13FD68B006FC61" httpUri="http://treatment.plazi.org/id/8C5C87B2FFFE6B36FF13FD68B006FC61" lastPageId="5" lastPageNumber="66" pageId="1" pageNumber="62">
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<subSubSection id="4CEF652FFFFE6B32FF13FD68B457FCC1" pageId="1" pageNumber="62" type="nomenclature">
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<paragraph id="044A36A4FFFE6B32FF13FD68B701FD2A" blockId="1.[151,624,711,778]" box="[151,624,711,737]" pageId="1" pageNumber="62">
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<heading id="5F0281C8FFFE6B32FF13FD68B701FD2A" bold="true" box="[151,624,711,737]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
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<emphasis id="3681EAB6FFFE6B32FF13FD68B701FD2A" bold="true" box="[151,624,711,737]" pageId="1" pageNumber="62">
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<taxonomicName id="C3F54D27FFFE6B32FF13FD68B436FD2A" ID-CoL="62CXH" box="[151,327,711,737]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
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<emphasis id="3681EAB6FFFE6B32FF13FD68B436FD2A" bold="true" box="[151,327,711,737]" italics="true" pageId="1" pageNumber="62">Aphelocheirus</emphasis>
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</taxonomicName>
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(s.str.)
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<taxonomicName id="C3F54D27FFFE6B32FE26FD68B75FFD2A" box="[418,558,711,737]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="bruneiensis" status="sp. nov.">
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||
<emphasis id="3681EAB6FFFE6B32FE26FD68B75FFD2A" bold="true" box="[418,558,711,737]" italics="true" pageId="1" pageNumber="62">bruneiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2DB257CDFFFE6B32FDB1FD68B701FD2A" box="[565,624,711,737]" pageId="1" pageNumber="62" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="044A36A4FFFE6B32FF13FD5FB457FCC1" blockId="1.[151,624,711,778]" box="[151,294,752,778]" pageId="1" pageNumber="62">
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(
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<figureCitation id="9CCE2A21FFFE6B32FF1BFD5FB46CFCC1" box="[159,285,752,778]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 10" captionStartId-0="2.[151,269,1551,1575]" captionStartId-1="3.[151,269,1561,1585]" captionTargetBox-0="[276,1310,197,1524]" captionTargetBox-1="[278,1308,760,1532]" captionTargetId-0="figure@2.[276,1312,197,1525]" captionTargetId-1="figure@3.[276,1312,757,1534]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." captionText-1="FIGURES 5 – 10. Aphelocheirus bruneiensis sp. n., holotype, male. (5) Mesosternal carina (mstc) and propleuron (ppl), ventrolateral (slightly caudal) aspect. (6) Genitalia, dorsal aspect, after removing the parameres: aed - mainpiece of aedeagus with bulbous outgrowths on both sides; lpa – left parandrium; pyg – pygophore, rpa – right parandrium. (7, 8) Left paramere, approximately dorsal and ventral aspects. (9, 10) Right paramere, approximately ventral and dorsal aspects." httpUri-0="https://zenodo.org/record/184686/files/figure.png" httpUri-1="https://zenodo.org/record/184687/files/figure.png" pageId="1" pageNumber="62">Figs. 1–10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="4CEF652FFFFE6B32FF13FC90B1E4FC91" box="[151,1173,831,858]" pageId="1" pageNumber="62" type="etymology">
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<paragraph id="044A36A4FFFE6B32FF13FC90B1E4FC91" blockId="1.[151,1436,831,1980]" box="[151,1173,831,858]" pageId="1" pageNumber="62">
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<emphasis id="3681EAB6FFFE6B32FF13FC90B452FC92" bold="true" box="[151,291,831,857]" pageId="1" pageNumber="62">Etymology.</emphasis>
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The Latinized adjective
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<taxonomicName id="C3F54D27FFFE6B32FDC3FCEFB7BFFC92" box="[583,718,832,857]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="bruneiensis">
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<emphasis id="3681EAB6FFFE6B32FDC3FCEFB7BFFC92" box="[583,718,832,857]" italics="true" pageId="1" pageNumber="62">bruneiensis</emphasis>
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</taxonomicName>
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refers to the country of origin,
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<collectingCountry id="7CE27634FFFE6B32FBB9FCEFB1E0FC91" box="[1085,1169,832,858]" name="Brunei" pageId="1" pageNumber="62">Brunei</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="4CEF652FFFFE6B32FF41FCC8B49FFB87" pageId="1" pageNumber="62" type="materials_examined">
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<paragraph id="044A36A4FFFE6B32FF41FCC8B6B3FC37" blockId="1.[151,1436,831,1980]" pageId="1" pageNumber="62">
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<typeStatus id="DB4E8806FFFE6B32FF41FCC8B446FC4A" box="[197,311,871,897]" pageId="1" pageNumber="62" type="holotype">
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<emphasis id="3681EAB6FFFE6B32FF41FCC8B446FC4A" bold="true" box="[197,311,871,897]" pageId="1" pageNumber="62">Holotype</emphasis>
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</typeStatus>
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(brachypterous male;
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<collectingCountry id="7CE27634FFFE6B32FDD2FCC7B7D7FC49" box="[598,678,872,898]" name="Brunei" pageId="1" pageNumber="62">Brunei</collectingCountry>
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Museum, Collection No. NHS/BM.INS.
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<date id="704B1064FFFE6B32FB26FCC7B07EFC49" box="[1186,1295,872,898]" pageId="1" pageNumber="62" value="08-01-20">01.20.08</date>
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),
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<typeStatus id="DB4E8806FFFE6B32FAA8FCC8B0EDFC4A" box="[1324,1436,871,897]" pageId="1" pageNumber="62" type="paratype">
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<emphasis id="3681EAB6FFFE6B32FAA8FCC8B0EDFC4A" bold="true" box="[1324,1436,871,897]" pageId="1" pageNumber="62">paratype</emphasis>
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</typeStatus>
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(macropterous male; Natural History Museum Vienna, Collection No. NAUC-187/1), and
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<typeStatus id="DB4E8806FFFE6B32FB45FC20B040FC62" box="[1217,1329,911,937]" pageId="1" pageNumber="62" type="paratype">
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<emphasis id="3681EAB6FFFE6B32FB45FC20B040FC62" bold="true" box="[1217,1329,911,937]" pageId="1" pageNumber="62">paratype</emphasis>
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</typeStatus>
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(nymph, 4th or 5th instar; Universiti
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<collectingCountry id="7CE27634FFFE6B32FE4DFC15B7DFFC1F" box="[457,686,954,980]" name="Brunei" pageId="1" pageNumber="62">Brunei Darussalam</collectingCountry>
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, Collection No.
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<collectionCode id="62E4AE61FFFE6B32FCF4FC15B6BAFC1F" box="[880,971,954,980]" pageId="1" pageNumber="62">UBDM</collectionCode>
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.3.00005) labelled "
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<collectingCountry id="7CE27634FFFE6B32FB2AFC15B0E5FC1F" box="[1198,1428,954,980]" name="Brunei" pageId="1" pageNumber="62">Brunei Darussalam</collectingCountry>
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\ Sungai Tutong Survey\ site 17B,
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<date id="704B1064FFFE6B32FD98FC4DB7C9FC37" box="[540,696,994,1020]" pageId="1" pageNumber="62" value="2004-06-15">15 June 2004</date>
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\ leg. Stephen Moore".
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</paragraph>
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<paragraph id="044A36A4FFFE6B32FF41FBA5B49FFB87" blockId="1.[151,1436,831,1980]" pageId="1" pageNumber="62">
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<emphasis id="3681EAB6FFFE6B32FF41FBA5B41BFBEF" bold="true" box="[197,362,1034,1060]" pageId="1" pageNumber="62">
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<typeStatus id="DB4E8806FFFE6B32FF41FBA5B473FBEF" box="[197,258,1034,1060]" pageId="1" pageNumber="62">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="7CE27634FFFE6B32FEF6FBA5B727FBEF" box="[370,598,1034,1060]" name="Brunei" pageId="1" pageNumber="62">Brunei Darussalam</collectingCountry>
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, Sungai Tutong at Rambai road bridge (
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<geoCoordinate id="61C15063FFFE6B32FBAEFBA5B1E9FBEF" box="[1066,1176,1034,1060]" direction="north" orientation="latitude" pageId="1" pageNumber="62" precision="55" value="4.629">4.629° N</geoCoordinate>
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,
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<geoCoordinate id="61C15063FFFE6B32FB20FBA5B05BFBEF" box="[1188,1322,1034,1060]" direction="east" orientation="longitude" pageId="1" pageNumber="62" precision="55" value="114.651">114.651° E</geoCoordinate>
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); see also ecological description below.
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</paragraph>
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</subSubSection>
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<subSubSection id="4CEF652FFFFE6B36FF41FBF5B006FC61" lastPageId="5" lastPageNumber="66" pageId="1" pageNumber="62" type="description">
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<paragraph id="044A36A4FFFE6B32FF41FBF5B406FB57" blockId="1.[151,1436,831,1980]" pageId="1" pageNumber="62">
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<emphasis id="3681EAB6FFFE6B32FF41FBF5B708FBBF" bold="true" box="[197,633,1114,1140]" pageId="1" pageNumber="62">Description of brachypterous male</emphasis>
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(
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<typeStatus id="DB4E8806FFFE6B32FD0EFBF5B786FBBF" box="[650,759,1114,1140]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
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). Size. Body length
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<quantity id="C30D9B41FFFE6B32FC62FBF5B135FBBF" box="[998,1092,1114,1140]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="1" pageNumber="62" unit="mm" value="6.1">6.1 mm</quantity>
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. Body width 4.0 mm. Pronotum width
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<quantity id="C30D9B41FFFE6B32FE92FB2DB405FB57" box="[278,372,1154,1180]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="62" unit="mm" value="3.3">3.3 mm</quantity>
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.
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</paragraph>
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<paragraph id="044A36A4FFFE6B32FF41FB05B466FB27" blockId="1.[151,1436,831,1980]" pageId="1" pageNumber="62">
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Colour. Dorsal colour pattern see
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<figureCitation id="9CCE2A21FFFE6B32FDCAFB05B7C3FB0F" box="[590,690,1194,1220]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1551,1575]" captionTargetBox="[276,1310,197,1524]" captionTargetId="figure@2.[276,1312,197,1525]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="1" pageNumber="62">Figure 1</figureCitation>
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. Ventral side, including appendages, yellowish, partly brownish infuscated.
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</paragraph>
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<paragraph id="044A36A4FFFE6B32FF41FB55B585FA7F" blockId="1.[151,1436,831,1980]" pageId="1" pageNumber="62">
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Head (
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<figureCitation id="9CCE2A21FFFE6B32FE9DFB55B412FADF" box="[281,355,1274,1300]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1551,1575]" captionTargetBox="[276,1310,197,1524]" captionTargetId="figure@2.[276,1312,197,1525]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
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) moderately long, length 1.05 times head width; finely, uniformly punctured, distance between punctures mostly (except anteriorly) smaller than their diameter, medioposteriorly punctures confluent. Anterior section of head (anterior of anterior eye margin) short, 0.5 times eye length (eye length measured parallel with median line of head). Rostrum 1.7 times as long as profemur, tip reaching distal end of mesotrochanter.
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</paragraph>
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<paragraph id="044A36A4FFFE6B32FF41FA6DB067F907" blockId="1.[151,1436,831,1980]" pageId="1" pageNumber="62">
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Thorax: Pronotum (
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<figureCitation id="9CCE2A21FFFE6B32FE31FA6DB48DFA17" box="[437,508,1474,1500]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1551,1575]" captionTargetBox="[276,1310,197,1524]" captionTargetId="figure@2.[276,1312,197,1525]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
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) only slightly convex, medially finely coriaceous, laterally finely granulate. Median length of pronotum 0.6 times median head length. Maximum width at posterior corners, 2.75 times its median length. Lateral margins of pronotum strongly convex and convergent anteriorly; width at anterior corners 0.43 times maximum width. Propleuron (
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<figureCitation id="9CCE2A21FFFE6B32FD31F995B78AF99F" box="[693,763,1594,1620]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,269,1561,1585]" captionTargetBox="[278,1308,760,1532]" captionTargetId="figure@3.[276,1312,757,1534]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Aphelocheirus bruneiensis sp. n., holotype, male. (5) Mesosternal carina (mstc) and propleuron (ppl), ventrolateral (slightly caudal) aspect. (6) Genitalia, dorsal aspect, after removing the parameres: aed - mainpiece of aedeagus with bulbous outgrowths on both sides; lpa – left parandrium; pyg – pygophore, rpa – right parandrium. (7, 8) Left paramere, approximately dorsal and ventral aspects. (9, 10) Right paramere, approximately ventral and dorsal aspects." httpUri="https://zenodo.org/record/184687/files/figure.png" pageId="1" pageNumber="62">Fig. 5</figureCitation>
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) medially with very stout process, demarcated by almost rectangular incision. Mesosternum with extremely high median carina, ending abruptly anteriorly (
|
||
<figureCitation id="9CCE2A21FFFE6B32FA9FF9CDB010F9B7" box="[1307,1377,1634,1660]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,269,1561,1585]" captionTargetBox="[278,1308,760,1532]" captionTargetId="figure@3.[276,1312,757,1534]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Aphelocheirus bruneiensis sp. n., holotype, male. (5) Mesosternal carina (mstc) and propleuron (ppl), ventrolateral (slightly caudal) aspect. (6) Genitalia, dorsal aspect, after removing the parameres: aed - mainpiece of aedeagus with bulbous outgrowths on both sides; lpa – left parandrium; pyg – pygophore, rpa – right parandrium. (7, 8) Left paramere, approximately dorsal and ventral aspects. (9, 10) Right paramere, approximately ventral and dorsal aspects." httpUri="https://zenodo.org/record/184687/files/figure.png" pageId="1" pageNumber="62">Fig. 5</figureCitation>
|
||
) and bearing some ill-defined wrinkles; in lateral view anterior corner almost rectangular (slightly acute). Mesoscutellum finely granulate; anterior width 2.5 times median length, its centre only slightly convex.
|
||
</paragraph>
|
||
<paragraph id="044A36A4FFFE6B32FF41F975B434F8A7" blockId="1.[151,1436,831,1980]" pageId="1" pageNumber="62">
|
||
Hemielytra (
|
||
<figureCitation id="9CCE2A21FFFE6B32FEDAF975B4D7F93F" box="[350,422,1754,1780]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1551,1575]" captionTargetBox="[276,1310,197,1524]" captionTargetId="figure@2.[276,1312,197,1525]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
|
||
) delicately granulate, small, posteriorly not reaching posterior margin of tergite 2, medially with gap measuring 0.28 times hemielytron width. Each hemielytron slightly wider than long (ca. 1.1 times), laterally with prominent triangular embolium process (process width ca. 0.2 times total hemielytron width).
|
||
</paragraph>
|
||
<paragraph id="044A36A4FFFE6B32FF41F8D5B638F877" blockId="1.[151,1436,831,1980]" pageId="1" pageNumber="62">Legs relatively short and stout, femora laterally not much exceeding margin of body. Profemur longer than both meso- and metafemur; 2.8 times as long as wide.</paragraph>
|
||
<caption id="508A662CFFFD6B31FF13F9A0B7E4F944" httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="2" pageNumber="63" targetBox="[276,1310,197,1524]" targetPageId="2">
|
||
<paragraph id="044A36A4FFFD6B31FF13F9A0B7E4F944" blockId="2.[151,1436,1551,1679]" pageId="2" pageNumber="63">
|
||
<emphasis id="3681EAB6FFFD6B31FF13F9A0B435F9EC" bold="true" box="[151,324,1551,1575]" pageId="2" pageNumber="63">FIGURES 1–4.</emphasis>
|
||
<taxonomicName id="C3F54D27FFFD6B31FEC9F9BFB71AF9EC" box="[333,619,1552,1575]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="bruneiensis" status="sp. nov.">
|
||
<emphasis id="3681EAB6FFFD6B31FEC9F9BFB71AF9EC" box="[333,619,1552,1575]" italics="true" pageId="2" pageNumber="63">Aphelocheirus bruneiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2DB257CDFFFD6B31FDF0F9A0B7C0F9EC" box="[628,689,1551,1575]" pageId="2" pageNumber="63" rank="species">
|
||
<emphasis id="3681EAB6FFFD6B31FDF0F9A0B7C0F9EC" bold="true" box="[628,689,1551,1575]" pageId="2" pageNumber="63">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1–3: © NHMW
|
||
<taxonomicName id="C3F54D27FFFD6B31FC77F9FAB1DEF9A7" authority="Image" authorityName="Image" box="[1011,1199,1621,1644]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="order">Hemiptera Image</taxonomicName>
|
||
Collection, published with permission. 4: photo by © Stephen Moore.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="044A36A4FFFD6B31FF41F912B02EF934" blockId="2.[151,1436,1725,2031]" pageId="2" pageNumber="63">
|
||
Abdomen (
|
||
<figureCitation id="9CCE2A21FFFD6B31FECEF912B4FEF91C" box="[330,399,1725,1751]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1551,1575]" captionTargetBox="[276,1310,197,1524]" captionTargetId="figure@2.[276,1312,197,1525]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="2" pageNumber="63">Fig. 1</figureCitation>
|
||
). Tergites 2–7 with acute posterior corners, but corners of tergite 2 very small. Sternites only slightly asymmetrical, without peg-like setae. Sternites 4–5 with very distinct, sharp median carina.
|
||
</paragraph>
|
||
<paragraph id="044A36A4FFFD6B30FF41F8A2B764FF79" blockId="2.[151,1436,1725,2031]" lastBlockId="3.[151,1436,152,698]" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
|
||
Genitalia: Pygophore (
|
||
<figureCitation id="9CCE2A21FFFD6B31FE4BF8A2B765F8EC" box="[463,532,1805,1831]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,269,1561,1585]" captionTargetBox="[278,1308,760,1532]" captionTargetId="figure@3.[276,1312,757,1534]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Aphelocheirus bruneiensis sp. n., holotype, male. (5) Mesosternal carina (mstc) and propleuron (ppl), ventrolateral (slightly caudal) aspect. (6) Genitalia, dorsal aspect, after removing the parameres: aed - mainpiece of aedeagus with bulbous outgrowths on both sides; lpa – left parandrium; pyg – pygophore, rpa – right parandrium. (7, 8) Left paramere, approximately dorsal and ventral aspects. (9, 10) Right paramere, approximately ventral and dorsal aspects." httpUri="https://zenodo.org/record/184687/files/figure.png" pageId="2" pageNumber="63">Fig. 6</figureCitation>
|
||
) apically with short, roundish tubercle; left parandrium distinctly longer than right one, its apex hook-shaped and slender; right parandrium wide, its apex obliquely truncate. Both parameres stout and short, approximately
|
||
<quantity id="C30D9B41FFFD6B31FD02F8F2B791F8BC" box="[646,736,1885,1911]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="63" unit="mm" value="0.4">0.4 mm</quantity>
|
||
in length, scaly microstructures in distal part very delicate, distinct only under very high magnification. Left paramere (
|
||
<figureCitation id="9CCE2A21FFFD6B31FCF4F82AB694F854" box="[880,997,1925,1951]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,269,1561,1585]" captionTargetBox="[278,1308,760,1532]" captionTargetId="figure@3.[276,1312,757,1534]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Aphelocheirus bruneiensis sp. n., holotype, male. (5) Mesosternal carina (mstc) and propleuron (ppl), ventrolateral (slightly caudal) aspect. (6) Genitalia, dorsal aspect, after removing the parameres: aed - mainpiece of aedeagus with bulbous outgrowths on both sides; lpa – left parandrium; pyg – pygophore, rpa – right parandrium. (7, 8) Left paramere, approximately dorsal and ventral aspects. (9, 10) Right paramere, approximately ventral and dorsal aspects." httpUri="https://zenodo.org/record/184687/files/figure.png" pageId="2" pageNumber="63">Figs. 7, 8</figureCitation>
|
||
) broad, leaf-shaped, apically almost truncate, proximally with very prominent process, ventrally with three long setae, otherwise with short and inconspicuous pilosity. Right paramere (
|
||
<figureCitation id="9CCE2A21FFFD6B31FDF0F87AB785F824" box="[628,756,2005,2031]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,269,1561,1585]" captionTargetBox="[278,1308,760,1532]" captionTargetId="figure@3.[276,1312,757,1534]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Aphelocheirus bruneiensis sp. n., holotype, male. (5) Mesosternal carina (mstc) and propleuron (ppl), ventrolateral (slightly caudal) aspect. (6) Genitalia, dorsal aspect, after removing the parameres: aed - mainpiece of aedeagus with bulbous outgrowths on both sides; lpa – left parandrium; pyg – pygophore, rpa – right parandrium. (7, 8) Left paramere, approximately dorsal and ventral aspects. (9, 10) Right paramere, approximately ventral and dorsal aspects." httpUri="https://zenodo.org/record/184687/files/figure.png" pageId="2" pageNumber="63">Figs. 9, 10</figureCitation>
|
||
) broad sickle-shaped, medially with several longer setae, laterally with very short pilosity.
|
||
</paragraph>
|
||
<paragraph id="044A36A4FFFC6B30FF41FF10B406FEC9" blockId="3.[151,1436,152,698]" pageId="3" pageNumber="64">
|
||
<emphasis id="3681EAB6FFFC6B30FF41FF10B703FF12" bold="true" box="[197,626,191,217]" pageId="3" pageNumber="64">Description of macropterous male</emphasis>
|
||
(
|
||
<typeStatus id="DB4E8806FFFC6B30FD00FF6FB781FF11" box="[644,752,192,218]" pageId="3" pageNumber="64" type="paratype">paratype</typeStatus>
|
||
). Size. Body length
|
||
<quantity id="C30D9B41FFFC6B30FC65FF6FB131FF11" box="[993,1088,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="3" pageNumber="64" unit="mm" value="6.1">6.1 mm</quantity>
|
||
. Body width
|
||
<quantity id="C30D9B41FFFC6B30FB65FF6FB04EFF11" box="[1249,1343,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="64" unit="mm" value="3.9">3.9 mm</quantity>
|
||
. Pronotum width
|
||
<quantity id="C30D9B41FFFC6B30FE92FF47B405FEC9" box="[278,372,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="64" unit="mm" value="3.2">3.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="044A36A4FFFC6B30FF41FEBFB4E4FE69" blockId="3.[151,1436,152,698]" pageId="3" pageNumber="64">
|
||
<collectingCountry id="7CE27634FFFC6B30FF41FEBFB594FEE1" box="[197,229,272,298]" name="American Samoa" pageId="3" pageNumber="64">As</collectingCountry>
|
||
usual in
|
||
<taxonomicName id="C3F54D27FFFC6B30FEDDFEBFB773FEE2" box="[345,514,272,297]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3681EAB6FFFC6B30FEDDFEBFB773FEE2" box="[345,514,272,297]" italics="true" pageId="3" pageNumber="64">Aphelocheirus</emphasis>
|
||
</taxonomicName>
|
||
, macropterous specimens are slightly darker than brachypterous individuals, with slightly coarser sculpturing of head, pronotum, and mesoscutellum, slightly larger eyes, a differently shaped pronotum, a strongly enlarged mesoscutellum, and forewings reaching approximately the tip of the abdomen (see
|
||
<figureCitation id="9CCE2A21FFFC6B30FEC4FE27B4F6FE69" box="[320,391,392,418]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1551,1575]" captionTargetBox="[276,1310,197,1524]" captionTargetId="figure@2.[276,1312,197,1525]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="3" pageNumber="64">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="044A36A4FFFC6B30FF41FE1FB026FD71" blockId="3.[151,1436,152,698]" pageId="3" pageNumber="64">
|
||
Head (
|
||
<figureCitation id="9CCE2A21FFFC6B30FE92FE1FB42EFE01" box="[278,351,432,458]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1551,1575]" captionTargetBox="[276,1310,197,1524]" captionTargetId="figure@2.[276,1312,197,1525]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="3" pageNumber="64">Fig. 2</figureCitation>
|
||
) moderately long, length 1.25 times head width; uniformly punctured; anterior section of head short, 0.5 times eye length. Pronotum (
|
||
<figureCitation id="9CCE2A21FFFC6B30FD26FE77B79BFE39" box="[674,746,472,498]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1551,1575]" captionTargetBox="[276,1310,197,1524]" captionTargetId="figure@2.[276,1312,197,1525]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="3" pageNumber="64">Fig. 2</figureCitation>
|
||
) slightly convex, with sinuate hind margin; median length of pronotum 0.75 times median head length; maximum width near posterior corners, 3.3 times its median length; width at anterior corners 0.44 times maximum width. Propleuron with medial process as in brachypterous morph. Mesosternum with similar median carina as in brachypterous morph, but posteriorly less highly raised above the more swollen mesosoma. Embolar margin of hemielytron strongly expanded laterally, forming a sharp wing-shaped process (
|
||
<figureCitation id="9CCE2A21FFFC6B30FDD9FD0FB7D4FD71" box="[605,677,672,698]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1551,1575]" captionTargetBox="[276,1310,197,1524]" captionTargetId="figure@2.[276,1312,197,1525]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="3" pageNumber="64">Fig. 2</figureCitation>
|
||
). Abdomen including genitalia as in brachypterous morph.
|
||
</paragraph>
|
||
<caption id="508A662CFFFC6B30FF13F9B6B0F2F953" httpUri="https://zenodo.org/record/184687/files/figure.png" pageId="3" pageNumber="64" targetBox="[278,1308,760,1532]" targetPageId="3">
|
||
<paragraph id="044A36A4FFFC6B30FF13F9B6B0F2F953" blockId="3.[151,1436,1561,1688]" pageId="3" pageNumber="64">
|
||
<emphasis id="3681EAB6FFFC6B30FF13F9B6B421F9FA" bold="true" box="[151,336,1561,1585]" pageId="3" pageNumber="64">FIGURES 5–10.</emphasis>
|
||
<taxonomicName id="C3F54D27FFFC6B30FED3F9B6B705F9FB" box="[343,628,1561,1584]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="bruneiensis" status="sp. nov.">
|
||
<emphasis id="3681EAB6FFFC6B30FED3F9B6B705F9FB" box="[343,628,1561,1584]" italics="true" pageId="3" pageNumber="64">Aphelocheirus bruneiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2DB257CDFFFC6B30FDFFF9B6B7C6F9FB" box="[635,695,1561,1585]" pageId="3" pageNumber="64" rank="species">
|
||
<emphasis id="3681EAB6FFFC6B30FDFFF9B6B7C0F9FA" bold="true" box="[635,689,1561,1585]" pageId="3" pageNumber="64">sp. n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype, male. (5) Mesosternal carina (mstc) and propleuron (ppl), ventrolateral (slightly caudal) aspect. (6) Genitalia, dorsal aspect, after removing the parameres: aed - mainpiece of aedeagus with bulbous outgrowths on both sides; lpa – left parandrium; pyg – pygophore, rpa – right parandrium. (7, 8) Left paramere, approximately dorsal and ventral aspects. (9, 10) Right paramere, approximately ventral and dorsal aspects.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="044A36A4FFFC6B30FF41F964B5BDF8C5" blockId="3.[151,1436,1739,2006]" pageId="3" pageNumber="64">
|
||
<emphasis id="3681EAB6FFFC6B30FF41F964B4A9F92E" bold="true" box="[197,472,1739,1765]" pageId="3" pageNumber="64">Description of nymph</emphasis>
|
||
(
|
||
<typeStatus id="DB4E8806FFFC6B30FE6DF963B724F92D" box="[489,597,1740,1766]" pageId="3" pageNumber="64" type="paratype">paratype</typeStatus>
|
||
). Size. Body length
|
||
<quantity id="C30D9B41FFFC6B30FCC2F963B6D4F92D" box="[838,933,1740,1766]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="64" unit="mm" value="5.1">5.1 mm</quantity>
|
||
. Body width
|
||
<quantity id="C30D9B41FFFC6B30FBC0F963B1D2F92D" box="[1092,1187,1740,1766]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="64" unit="mm" value="3.6">3.6 mm</quantity>
|
||
. Pronotum width
|
||
<quantity id="C30D9B41FFFC6B30FAF3F963B5B8F8C5" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="64" unit="mm" value="2.8">2.8 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="044A36A4FFFC6B37FF41F8B3B196FEC9" blockId="3.[151,1436,1739,2006]" lastBlockId="4.[151,1436,152,258]" lastPageId="4" lastPageNumber="65" pageId="3" pageNumber="64">
|
||
Colour. Dorsal colour pattern see
|
||
<figureCitation id="9CCE2A21FFFC6B30FDD5F8B3B7C4F8FD" box="[593,693,1820,1846]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1551,1575]" captionTargetBox="[276,1310,197,1524]" captionTargetId="figure@2.[276,1312,197,1525]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="3" pageNumber="64">Figure 3</figureCitation>
|
||
. Ventral side, including all appendages, pale yellowish. Dorsal surface partly smooth and shining but with delicate sculpturing; with distinct, short pilosity; median line of thorax and abdomen with more or less distinct carina. Head short, length 0.9 times head width; anterior section of head 0.5 times eye length. Pronotum (
|
||
<figureCitation id="9CCE2A21FFFC6B30FCB0F83BB60BF865" box="[820,890,1940,1966]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1551,1575]" captionTargetBox="[276,1310,197,1524]" captionTargetId="figure@2.[276,1312,197,1525]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="3" pageNumber="64">Fig. 3</figureCitation>
|
||
) almost flat, maximum width at posterior corners 5.0 times its median length. Sides of pronotum strongly convex and convergent anteriorly; width at anterior corners 0.40 times maximum width. Propleuron medially simply rounded. Mesosternum with distinct median carina, ending abruptly anteriorly, although less high than in adults. Sternites 3–6 with very distinct, sharp median carina. Sternite 3 with two subapical setae (homologous with peg-like bristles?).
|
||
</paragraph>
|
||
<caption id="508A662CFFFB6B37FF13F8A2B745F819" httpUri="https://zenodo.org/record/184688/files/figure.png" pageId="4" pageNumber="65" targetBox="[276,1309,314,1781]" targetPageId="4">
|
||
<paragraph id="044A36A4FFFB6B37FF13F8A2B745F819" blockId="4.[151,1436,1805,2002]" pageId="4" pageNumber="65">
|
||
<emphasis id="3681EAB6FFFB6B37FF13F8A2B42CF8EE" bold="true" box="[151,349,1805,1829]" pageId="4" pageNumber="65">FIGURES 11–14.</emphasis>
|
||
Dorsal aspects of some Bornean species of
|
||
<taxonomicName id="C3F54D27FFFB6B37FCC4F8A1B6ABF8EE" box="[832,986,1806,1829]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3681EAB6FFFB6B37FCC4F8A1B6ABF8EE" box="[832,986,1806,1829]" italics="true" pageId="4" pageNumber="65">Aphelocheirus</emphasis>
|
||
</taxonomicName>
|
||
(all specimens from the collection of the Natural History Museum Vienna). (11)
|
||
<taxonomicName id="C3F54D27FFFB6B37FDCCF89EB797F883" box="[584,742,1841,1864]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="kelabitensis">
|
||
<emphasis id="3681EAB6FFFB6B37FDCCF89EB797F883" box="[584,742,1841,1864]" italics="true" pageId="4" pageNumber="65">A. kelabitensis</emphasis>
|
||
</taxonomicName>
|
||
, brachypterous male paratype from Kelabit Highland, Sarawak (body length 7.2 mm). (12)
|
||
<taxonomicName id="C3F54D27FFFB6B37FE46F8FCB74CF8A1" box="[450,573,1875,1898]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="kinabalu">
|
||
<emphasis id="3681EAB6FFFB6B37FE46F8FCB74CF8A1" box="[450,573,1875,1898]" italics="true" pageId="4" pageNumber="65">A. kinabalu</emphasis>
|
||
</taxonomicName>
|
||
, brachypterous female from Mt.
|
||
<taxonomicName id="C3F54D27FFFB6B37FC26F8FCB177F8A1" box="[930,1030,1875,1898]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="kinabalu">Kinabalu</taxonomicName>
|
||
, Sabah (body length 8.9 mm). (13)
|
||
<taxonomicName id="C3F54D27FFFB6B37FA01F8FCB585F846" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="kinabalu">
|
||
<emphasis id="3681EAB6FFFB6B37FA01F8FCB585F846" italics="true" pageId="4" pageNumber="65">A. kinabalu</emphasis>
|
||
</taxonomicName>
|
||
, macropterous male from Kelabit Highland, Sarawak (body length 8.2 mm; genital capsule dissected). (14)
|
||
<taxonomicName id="C3F54D27FFFB6B37FA01F8D9B59CF87B" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="kodadai">
|
||
<emphasis id="3681EAB6FFFB6B37FA01F8D9B59CF87B" italics="true" pageId="4" pageNumber="65">A. kodadai</emphasis>
|
||
</taxonomicName>
|
||
, brachypterous female paratype from Crocker Range, Sabah (body length 7.6 mm). © NHMW
|
||
<taxonomicName id="C3F54D27FFFB6B37FB66F837B0EDF864" authority="Image" authorityName="Image" box="[1250,1436,1944,1967]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="order">Hemiptera Image</taxonomicName>
|
||
Collection, published with permission.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="044A36A4FFFA6B36FF41FF38B4DAFF79" blockId="5.[151,1437,151,938]" box="[197,427,151,178]" pageId="5" pageNumber="66">
|
||
<emphasis id="3681EAB6FFFA6B36FF41FF38B459FF7A" bold="true" box="[197,296,151,177]" pageId="5" pageNumber="66">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="044A36A4FFFA6B36FF41FF10B02BFE69" blockId="5.[151,1437,151,938]" pageId="5" pageNumber="66">
|
||
<emphasis id="3681EAB6FFFA6B36FF41FF10B7D4FF12" bold="true" box="[197,677,191,217]" pageId="5" pageNumber="66">
|
||
Ecological description of
|
||
<typeStatus id="DB4E8806FFFA6B36FD81FF10B74AFF12" box="[517,571,191,217]" pageId="5" pageNumber="66">type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Site 17 of the Sungai Tutong Survey 2004 (
|
||
<bibRefCitation id="60644B55FFFA6B36FB3CFF6FB0FEFF11" author="Evans" box="[1208,1423,192,218]" pageId="5" pageNumber="68" refString="Evans, C., Moore, S. & Trangmar, T. (2005) Sungai Tutong Catchment Environmental Study. Prepared for the Department of Environment, Parks & Recreation Brunei Darussalam." type="book" year="2005">
|
||
Evans
|
||
<emphasis id="3681EAB6FFFA6B36FA8FFF6FB036FF12" box="[1291,1351,192,217]" italics="true" pageId="5" pageNumber="66">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
), i.e., the
|
||
<typeStatus id="DB4E8806FFFA6B36FF71FF47B459FEC9" box="[245,296,232,258]" pageId="5" pageNumber="66">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="C3F54D27FFFA6B36FE2AFF47B727FECA" box="[430,598,232,257]" class="Insecta" family="Aphelocheiridae" genus="Aphelocheirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="bruneiensis" status="sp. nov.">
|
||
<emphasis id="3681EAB6FFFA6B36FE2AFF47B727FECA" box="[430,598,232,257]" italics="true" pageId="5" pageNumber="66">A. bruneiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2DB257CDFFFA6B36FDDAFF47B7E6FEC9" box="[606,663,232,258]" pageId="5" pageNumber="66" rank="species">sp.n.</taxonomicNameLabel>
|
||
, is in the mid reaches of the river near Kampong Rambai, located
|
||
<quantity id="C30D9B41FFFA6B36FF13FEBFB594FEE1" box="[151,229,272,298]" metricMagnitude="4" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="66" unit="km" value="46.0">46 km</quantity>
|
||
upstream of the river mouth, below the Rambai road bridge (
|
||
<figureCitation id="9CCE2A21FFFA6B36FC5AFEBFB15BFEE1" box="[990,1066,272,298]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,269,1551,1575]" captionTargetBox="[276,1310,197,1524]" captionTargetId="figure@2.[276,1312,197,1525]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Aphelocheirus bruneiensis sp. n. (1) Habitus of holotype, brachypterous male (body length 6.1 mm), dorsal aspect. (2) Habitus of paratype, macropterous male (body length 6.1 mm), dorsal aspect. (3) Habitus of paratype nymph (body length 5.1 mm), dorsal aspect. (4) Type locality. 1 – 3: © NHMW Hemiptera Image Collection, published with permission. 4: photo by © Stephen Moore." httpUri="https://zenodo.org/record/184686/files/figure.png" pageId="5" pageNumber="66">Fig. 4</figureCitation>
|
||
). The site was unusual in that gabion baskets (used to stabilise the riverbed) created an artificially narrow, hard-bedded channel with moderate to fast flow. These habitat conditions differed from those of most of the length of the Sungai Tutong where slow-flowing, sandy-bedded, and grassy-margin habitats were predominant (in 23 of 25 sampling sites).
|
||
</paragraph>
|
||
<paragraph id="044A36A4FFFA6B36FF41FE1FB65BFD59" blockId="5.[151,1437,151,938]" pageId="5" pageNumber="66">
|
||
The Rambai bridge site supported aquatic invertebrates commonly associated with hard beds and moderate to fast currents, including hydropsychid caddisflies (37 % of the individuals found at this site), heptageniid mayflies (23 %), simuliid black flies (2 %), perlid stoneflies (2 %), and psephenid beetles (1 %). These groups were less commonly recorded in the sandy/grassy habitats found along most of the Sungai Tutong where the most common taxa included chironomid midges (subfamilies
|
||
<taxonomicName id="C3F54D27FFFA6B36FCFDFDFFB150FDA1" box="[889,1057,592,618]" class="Insecta" family="Chironomidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Chironominae">Chironominae</taxonomicName>
|
||
and
|
||
<taxonomicName id="C3F54D27FFFA6B36FBDAFDFFB18BFDA1" box="[1118,1274,592,618]" class="Insecta" family="Chironomidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Tanypodinae">Tanypodinae</taxonomicName>
|
||
), baetid mayflies, libellulid dragonflies, and protoneurid damselflies.
|
||
</paragraph>
|
||
<paragraph id="044A36A4FFFA6B36FF41FD0FB006FC61" blockId="5.[151,1437,151,938]" pageId="5" pageNumber="66">
|
||
The general water quality of the Sungai Tutong is typical of many tropical Asian slow-flowing rivers. The river had relatively warm temperatures, with a median from all 25 sites (sampling dates between
|
||
<date id="704B1064FFFA6B36FAB6FD67B425FCC1" pageId="5" pageNumber="66" value="2004-06" valueMax="2004-12-31" valueMin="2004-06-00">June and December 2004</date>
|
||
) of 28.5 °C. The river was always visibly turbid, due primarily to high suspended sediment levels (median from all sites of 18 ppm). The river water was always acidic (median
|
||
<collectingCountry id="7CE27634FFFA6B36FBF4FCB7B1E5FCF9" box="[1136,1172,792,818]" name="Philippines" pageId="5" pageNumber="66">pH</collectingCountry>
|
||
from all sites was 5.7) due to the acidic breakdown products of forest and wetland vegetation. The slow flow at most Sungai Tutong sites contributed to low oxygen saturation levels, with a median from all 25 sampling sites of 60 %; however, the faster flow of the Rambai bridge site produced a higher median dissolved oxygen saturation of 74.5 %.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |