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<mods:title id="556CCA670FBD59517A10EAE71E4565C8">A revision of the Chinese Trigonalyidae (Hymenoptera, Trigonalyoidea)</mods:title>
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<mods:namePart id="3FF3A83F346C056BF992464E7D573D59">Chen, Hua-yan</mods:namePart>
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<mods:namePart id="9B3B9DEFBDEF97CA2A0A636A3F523F12">He, Jun-hua</mods:namePart>
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<subSubSection id="3EAA797E1692F146477BA740C8682997" pageId="109" pageNumber="110" type="nomenclature">
<paragraph id="ABFB386427319B1C4C49EF9F25B0BD29" pageId="109" pageNumber="110">
<taxonomicName id="5FAD6F240C568859AB2C37D01776AA7E" genus="Taeniogonalos" lsidName="Taeniogonalos subtruncata" pageId="109" pageNumber="110" rank="species" species="subtruncata">Taeniogonalos subtruncata</taxonomicName>
<taxonomicNameLabel id="65FA13FC876E74970601A54CC2235F9D" pageId="109" pageNumber="110">nom. n.</taxonomicNameLabel>
Figs 496-517
</paragraph>
</subSubSection>
<subSubSection id="4334C401501D61B9B7F95A81B45A9DC8" pageId="109" pageNumber="110" type="reference_group">
<paragraph id="454196EE6050ADAABFE6886BB7705BA5" pageId="109" pageNumber="110">
<taxonomicName id="4F0787885D36FD2AA93E159ECE1BBE6D" genus="Nanogonalos" lsidName="Nanogonalos flavocincta" pageId="109" pageNumber="110" rank="species" species="flavocincta">Nanogonalos flavocincta</taxonomicName>
Teranishi, 1929: 140;
<bibRefCitation id="8DA1C0332724E391ACEF5ECE0AEB2163" author="Weinstein, P" journalOrPublisher="Journal of Natural History" pageId="141" pageNumber="142" pagination="399 - 433" title="The host relationships of trigonalyid wasps (Hymenoptera: Trigonalyidae), with a review of their biology and catalogue to world species." url="10.1080/00222939100770281" volume="25" year="1991">Weinstein and Austin 1991</bibRefCitation>
: 421.
</paragraph>
<paragraph id="B66CB2C2C659B4D8632609E33269CDCD" pageId="109" pageNumber="110">
<taxonomicName id="29CF061F53B736BFBC4D157163A8E07D" genus="Poecilogonalos" lsidName="Poecilogonalos flavocincta" pageId="109" pageNumber="110" rank="species" species="flavocincta">Poecilogonalos flavocincta</taxonomicName>
;
<bibRefCitation id="8FB276BFC2721E017640E1724C51E9F6" author="Marshakov, VG" editor="Korotyaev, BA" journalOrPublisher="Nauka, Leningrad" pageId="139" pageNumber="140" pagination="100 - 107" title="Morfologia i sistematica nasekomykh Dalnego Vostoka" year="1981">Marshakov 1981</bibRefCitation>
: 107;
<bibRefCitation id="C2AA06F4AFB1849E82B22E61AD1A1618" pageId="109" pageNumber="110">Lelej 1995</bibRefCitation>
: 14. Synonymized with
<taxonomicName id="1EDA1C0B9D21C63813C74FA733B6E870" genus="Nanogonalos" lsidName="Nanogonalos mongolicus" pageId="109" pageNumber="110" rank="species" species="mongolicus">Nanogonalos mongolicus</taxonomicName>
by
<bibRefCitation id="898765433B826F8743E20E56EF3CDA0F" author="Marshakov, VG" editor="Korotyaev, BA" journalOrPublisher="Nauka, Leningrad" pageId="139" pageNumber="140" pagination="100 - 107" title="Morfologia i sistematica nasekomykh Dalnego Vostoka" year="1981">Marshakov 1981</bibRefCitation>
.
</paragraph>
<paragraph id="C65FA990AA212DDD8DA8F93AE3F965FA" pageId="109" pageNumber="110">
<taxonomicName id="EF77D940D04DEBC089381F4883C1512F" genus="Taeniogonalos" lsidName="Taeniogonalos flavocincta" pageId="109" pageNumber="110" rank="species" species="flavocincta">Taeniogonalos flavocincta</taxonomicName>
;
<bibRefCitation id="24FD006735B78799A39071510FA73B3D" author="Carmean, D" journalOrPublisher="Systematic Entomology" pageId="138" pageNumber="139" pagination="35 - 76" title="Phylogenetic revision of the parasitoid wasp family Trigonalidae (Hymenoptera)." url="10.1046/j.1365-3113.1998.00042.x" volume="23" year="1998">Carmean and Kimsey 1998</bibRefCitation>
: 67 [not
<taxonomicName id="C997F722FED0D173C53248122837C46F" genus="Taeniogonalos" lsidName="Taeniogonalos flavicincta" pageId="109" pageNumber="110" rank="species" species="flavicincta">Taeniogonalos flavicincta</taxonomicName>
(Bischoff, 1913)].
</paragraph>
</subSubSection>
<subSubSection id="E080ECB5F214BCD51CF1CB9C111CD68D" pageId="109" pageNumber="110" type="type material">
<paragraph id="62532A75AB4EF3A883FC39046D37DD2E" pageId="109" pageNumber="110">Type material.</paragraph>
<paragraph id="9D8BAB8861C83737301343737834FAA1" pageId="109" pageNumber="110">
Holotype, ♂ (OMNH), &quot;Corea, Suigen, V.1928, C.P. Clausen&quot;, &quot;Holotype
<taxonomicName id="F7B6667DE67B3C40A8F90D81028B8668" genus="Nanogonalos" lsidName="Nanogonalos flavicincta" pageId="109" pageNumber="110" rank="species" species="flavicincta">Nanogonalos flavicincta</taxonomicName>
[sic!]&quot;.
</paragraph>
</subSubSection>
<subSubSection id="A46065AEBF26EF1B361677EBB4F3624A" pageId="109" pageNumber="110" type="additional material">
<paragraph id="26AE0B8883644CF1394CC2986057F2F4" pageId="109" pageNumber="110">Additional material.</paragraph>
<paragraph id="B2AF1F8FE0AE3F0C2C02BCF35F2CC931" pageId="109" pageNumber="110">1 ♀ (IZCAS) &quot;[China:] Shaanxi, Zhenping, 21.VII.1983, IOZ(E)1495446&quot;.</paragraph>
</subSubSection>
<subSubSection id="C35E814DEA59253A3363029E4D4948BA" lastPageId="110" lastPageNumber="111" pageId="109" pageNumber="110" type="diagnosis">
<paragraph id="7A5D9E069A7CFC4080CF016E9B2ED98F" pageId="109" pageNumber="110">Diagnosis.</paragraph>
<paragraph id="9E610646A8D94D65F382C5C8C1BD70F1" lastPageId="110" lastPageNumber="111" pageId="109" pageNumber="110">
Supra-antennal elevations 0.4-0.5 times as long as scapus and outer side of elevations oblique, largely smooth (except for sparse punctures) and apically brown (Figs 498, 509); tyloids of male antenna linear; occipital carina distinctly lamelliform, moderately widened (about 0.5 times as wide as diameter of ocellus)
<pageBreakToken id="4D4EC0401CAC5410B06FCA491B6D7EE6" pageId="110" pageNumber="111" start="start">and</pageBreakToken>
smooth medio-dorsally (Fig. 498); head posteriorly with two small ivory spots (Fig. 509; female) or entirely black (male); head dorsally coarsely reticulate-punctate and largely without distinct smooth interspaces (Figs 498, 509); mesosoma black dorsally except for ivory patch(es) on metanotum medially (Fig. 501); mesoscutum and scutellum coarsely punctate-reticulate (Fig. 501); scutellum entirely black (Fig. 501); anterior half of fore wing dark brown (Fig. 506); third submarginal cell of fore wing nearly as long as second submarginal cell (Fig. 506); first discal cell of fore wing comparatively elongate (Fig. 506); metasoma coarsely punctate (Fig. 503), with wide band at apex of second tergite and sternite, and fourth-sixth tergites largely orange or yellowish and remainder of metasoma black (Fig. 503); first tergite about half as long as its apical width; protuberance of second sternite of female subtruncate medio-apically and in lateral view curved and rounded apically (Figs 516, 517); second sternite of male distinctly shallowly depressed medio-posteriorly (Fig. 505); third sternite of male flat medially; third sternite 0.3-0.4 times as long as second sternite.
</paragraph>
<caption id="B71534073E7A067AA54C47C6A7EA8D57" pageId="110" pageNumber="111">
<paragraph id="7630CE91CEC8EE722BD6C29C3F2D1DBF" pageId="110" pageNumber="111">
Figures 496-499.
<taxonomicName id="5AE9B899A6D3FF3C424AD25AF90A926D" genus="Taeniogonalos" lsidName="Taeniogonalos flavocincta" pageId="110" pageNumber="111" rank="species" species="flavocincta">Taeniogonalos flavocincta</taxonomicName>
(Teranishi, 1929), holotype, male. 496 Habitus lateral 497 head anterior 498 head dorsal 499 antenna.
</paragraph>
</caption>
<caption id="BE433E4E2831D5770E0A60BCCE2B305F" pageId="110" pageNumber="111">
<paragraph id="B1A4C4D6206A0E249D7DDCF7B78686AC" pageId="110" pageNumber="111">
Figures 500-506.
<taxonomicName id="A0276AAA0587A1BD1E24205DD146C884" genus="Taeniogonalos" lsidName="Taeniogonalos flavocincta" pageId="110" pageNumber="111" rank="species" species="flavocincta">Taeniogonalos flavocincta</taxonomicName>
(Teranishi, 1929), holotype, male. 500 Head lateral 501 mesosoma dorsal 502 mesosoma lateral 503 metasoma dorsal 504 metasoma lateral 505 metasoma ventral 506 fore wing.
</paragraph>
</caption>
<caption id="36AEAFC1B0967059FBDE01554621E6C4" pageId="110" pageNumber="111">
<paragraph id="A7A1C290FB186BF29E62D1A5082B761F" pageId="110" pageNumber="111">
Figures 507-509.
<taxonomicName id="9F5E4EE1E4700DACB9F045109C1CAED6" genus="Taeniogonalos" lsidName="Taeniogonalos subtruncata" pageId="110" pageNumber="111" rank="species" species="subtruncata">Taeniogonalos subtruncata</taxonomicName>
nom. n., female from Shaanxi. 507 Habitus lateral 508 head anterior 509 head dorsal.
</paragraph>
</caption>
<caption id="7E68EB7924420347682E7D4B7C00130C" pageId="110" pageNumber="111">
<paragraph id="AE3E83994EBC41907E463840D66888A9" pageId="110" pageNumber="111">
Figures 510-517.
<taxonomicName id="25EF594AEFE6E8F0D5275DAEAF08F3CF" genus="Taeniogonalos" lsidName="Taeniogonalos subtruncata" pageId="110" pageNumber="111" rank="species" species="subtruncata">Taeniogonalos subtruncata</taxonomicName>
nom. n., female from Shaanxi. 510 Antenna 511 fore and hind wings 512 head lateral 513 mesosoma dorsal 514 mesosoma lateral 515 metasoma dorsal 516 metasoma lateral 517 metasoma ventral.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="735C7D8642D0A913790E6913A5CB9EEA" lastPageId="112" lastPageNumber="113" pageId="110" pageNumber="111" type="description">
<paragraph id="DE30DB2D69CB9098F52D6CF1E956FD4C" pageId="110" pageNumber="111">Description.</paragraph>
<paragraph id="798E31D44C8075DD309AAC661A0A9562" pageId="110" pageNumber="111">Female from Shaanxi, length of body 12.4 mm (of fore wing 9.8 mm).</paragraph>
<paragraph id="39719D68810C65DE91680813226F1F7F" pageId="110" pageNumber="111">Head. Antenna with 25 segments; frons, vertex and temple coarsely reticulate-punctate (Figs 508, 509, 512); head gradually narrowed behind eyes, eye in dorsal view 0.7 times as long as temple (Fig. 509); occipital carina narrowly lamelliform and smooth medio-dorsally; supra-antennal elevations medium-sized (about 0.4 times as long as scapus), outer side oblique and largely smooth except for sparse punctures; clypeus slightly concave and thick medio-ventrally.</paragraph>
<paragraph id="60BF879EE506C6255AEEBB614A172632" pageId="110" pageNumber="111">Mesosoma. Length of mesosoma 1.5 times its height (Fig. 514); mesopleuron coarsely reticulate-punctate anteriorly, transversely rugulose posteriorly; transverse mesopleural groove narrow, deep and moderately crenulate; notauli narrow, deep and finely crenulate; mesoscutum coarsely reticulate-punctate (Fig. 513); scutellar sulcus complete, medium-sized and crenulate medially and wider laterally; scutellum coarsely reticulate-punctate, convex medially and anteriorly slightly above level of mesoscutum; metanotum medially slightly convex, not protruding and rugose (Fig. 513); propodeum coarsely rugose (Fig. 513); posterior propodeal carina thick lamelliform and strongly arched, foramen medially 0.4 times higher than wide basally.</paragraph>
<paragraph id="DD14203DCB8016D17C55270C07F7B53B" pageId="110" pageNumber="111">Wings. Fore wing: length of vein 1-M 1.8 times as long as vein 1-SR (Fig. 511).</paragraph>
<paragraph id="6539B940E4F281AE4444A6105935E40C" pageId="110" pageNumber="111">
Metasoma. First tergite 0.6 times as long as apically wide, largely smooth and with shallow elliptical depression medially (Fig. 515);
<normalizedToken id="9300D1267704923A09A5C0F66045965C" originalValue="secondsixth">second-sixth</normalizedToken>
tergites densely and coarsely punctate; sternites densely and coarsely punctate; second sternite distinctly convex in lateral view, its medio-apical protuberance medium-sized and blunt medio-apically (Fig. 517); third sternite about 0.4 times as long as second sternite, with two close large triangular protuberances medially (Fig. 517); hypopygium triangular in ventral view (Fig. 517).
</paragraph>
<paragraph id="6F5DA3BF4626A365AED0A9E2A15F48E0" pageId="111" pageNumber="112">
<pageBreakToken id="53EF6AB9330D3BBE42999AFD5926A16C" pageId="111" pageNumber="112" start="start">Colour</pageBreakToken>
. Black; palpi dark brown; mandible black with teeth dark brown; antenna dark brown; frons with small yellowish brown spots along inter orbita, remainder of head black except for posterior vertex with small yellowish brown spots and narrow yellowish brown stripes along outer orbita; apex of supra-antennal elevations yellowish brown; tegulae dark brown; mesosoma laterally black except for antero-dorsal yellowish brown patch of pronotal side; metanotum with one medium-sized medial yellowish brown patch; hind trochanter, trochantellus and base of hind femur ivory, remainder of legs dark brown to black with tarsi paler; anterior half of fore wing dark brown, posterior half subhyaline; metasoma with moderate wide orange-brown apical band at apex of second tergite and sternite, third tergite with narrow orange-brown band at posterior margin, fourth to sixth tergites with large orange-brown patches posteriorly.
</paragraph>
<paragraph id="323F88AD928CB1F2B21C3AC810853E18" pageId="112" pageNumber="113">
<pageBreakToken id="6BC4F50230449A1BB63983CB8957FCD7" pageId="112" pageNumber="113" start="start">Male</pageBreakToken>
. Holotype male is very similar to the redescribed female except shape of second sternite. It has length of body 10.7 mm (of fore wing 8.1 mm); brown antenna with 23 segments with longitudinal tyloids on 10th-16th segments; outer orbita largely and pronotum dorso-posterioly ivory, head posteriorly and third tergite apically black; second sternite of male distinctly shallowly depressed medio-posteriorly; third sternite of male flat medially;
</paragraph>
</subSubSection>
<subSubSection id="CB23264D313CA8ACD6EE4537EF6BE08E" pageId="112" pageNumber="113" type="biology">
<paragraph id="ABDB66646096C1051745F6CC7304920E" pageId="112" pageNumber="113">Biology.</paragraph>
<paragraph id="B7B4954739240C1AB9778205CAD557A7" pageId="112" pageNumber="113">Unknown. Collected in May and July.</paragraph>
</subSubSection>
<subSubSection id="0EDAB7DAD0FD442C9040FDA36267504C" pageId="112" pageNumber="113" type="distribution">
<paragraph id="EFA9EF7B0C01A296926D988339FBFF44" pageId="112" pageNumber="113">Distribution.</paragraph>
<paragraph id="3339B762BA541CAFA59730D4BCDE757F" pageId="112" pageNumber="113">China (Shaanxi), Korea.</paragraph>
</subSubSection>
<subSubSection id="338E84DF87C8DCC2D5185493132EA571" pageId="112" pageNumber="113" type="etymology">
<paragraph id="F08B2760E14424146A2EA866B419AE85" pageId="112" pageNumber="113">Etymology.</paragraph>
<paragraph id="888E8558351203059A59F1C63D872D27" pageId="112" pageNumber="113">
Renamed after the blunt (subtruncate) protuberance of the second sternite of the female: from
<normalizedToken id="14051870740FE700C935E7BF05424EAC" originalValue="“sub”">&quot;sub&quot;</normalizedToken>
(Latin for &quot;under, less&quot;) and
<normalizedToken id="41FDA9766DA4B19441547E0E758FEE6F" originalValue="“truncus”">&quot;truncus&quot;</normalizedToken>
(Latin for &quot;cut off&quot;).
</paragraph>
</subSubSection>
<subSubSection id="59C792F1332F9440C0E5417BD643DC60" pageId="112" pageNumber="113" type="notes">
<paragraph id="613E9F93F284A80358B3FB739F8D1603" pageId="112" pageNumber="113">Notes.</paragraph>
<paragraph id="AC92574720180C742B28B8E6FB29FE74" pageId="112" pageNumber="113">
The replacement name of
<taxonomicName id="F36614E09D39E017695B41D2AADE68FF" genus="Taeniogonalos" lsidName="Taeniogonalos subtruncata" pageId="112" pageNumber="113" rank="species" species="subtruncata">Taeniogonalos subtruncata</taxonomicName>
is necessary after transferring flavicincta Bischoff, 1913 from the genus
<taxonomicName id="AD8A6206318E9B47A4731AC90E68C1C2" genus="Lycogonalos" lsidName="Lycogonalos" pageId="112" pageNumber="113" rank="genus">Lycogonalos</taxonomicName>
Bischoff, 1913 and flavocincta Teranishi, 1929 from the genus
<taxonomicName id="3ACDC20B362EBDD0DA3A641704DF8A30" genus="Nanogonalos" lsidName="Nanogonalos" pageId="112" pageNumber="113" rank="genus">Nanogonalos</taxonomicName>
Schulz, 1906 to the genus
<taxonomicName id="F18F24995E964D9C0AE2073136943520" genus="Taeniogonalos" lsidName="Taeniogonalos" pageId="112" pageNumber="113" rank="genus">Taeniogonalos</taxonomicName>
Schulz, 1906 (
<bibRefCitation id="28FF9D3604A993680CFE6F8ACC5F6F7A" author="Carmean, D" journalOrPublisher="Systematic Entomology" pageId="138" pageNumber="139" pagination="35 - 76" title="Phylogenetic revision of the parasitoid wasp family Trigonalidae (Hymenoptera)." url="10.1046/j.1365-3113.1998.00042.x" volume="23" year="1998">Carmean and Kimsey 1998</bibRefCitation>
). Both became secondary homonyms in spite that the names differ in one letter (articles 57.3.1, 58.12 of
<bibRefCitation id="940E3C919C93C2A094B54BA9D9642271" author="International Code Commission on Zoological Nomenclature," journalOrPublisher="The International Turst for Zoological Nomenclature, London" pageId="138" pageNumber="139" title="International Code of Zoological Nomenclature" url="http://iczn.org/iczn/index.jsp" year="1999">ICZN 1999</bibRefCitation>
;
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).
</paragraph>
</subSubSection>
</treatment>
</document>