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<document id="FB848F7EBA51C4C0B6A028CE75DF614F" ID-CLB-Dataset="37304" ID-DOI="10.5281/zenodo.247616" ID-GBIF-Dataset="fcca9f27-99bf-4ec7-9992-386e79a37cd7" ID-ISSN="0013-8835" ID-Zenodo-Dep="247616" IM.bibliography_approvedBy="admin" IM.materialsCitations_approvedBy="admin" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="admin" checkinTime="1484588307211" checkinUser="plazi" docAuthor="Charles Huber &amp; Hannes Baur" docDate="2016" docId="8C13571B821C5D76FF16FA8A772FAE28" docLanguage="en" docName="entomologischeBlätterColeoptera.112.1.203.pdf" docOrigin="Entomologische Blätter und Coleoptera 112 (1)" docStyle="DocumentStyle{}" docTitle="Nebria (Patrobonebria) desgodinsi Oberthur, G. Ledoux 1883" docType="treatment" docVersion="10" lastPageNumber="207" masterDocId="702A2F63821E5D72FFF0FFEE741DA81D" masterDocTitle="Nebria (Patrobonebria) incognita n. sp. and Nebria (P.) hiekeiana n. sp., two new species from the Western Himalaya, with remarks on Nebria (P.) desgodinsi (Coleoptera, Carabidae, Nebriinae)" masterLastPageNumber="214" masterPageNumber="203" pageNumber="205" updateTime="1693850000913" updateUser="ExternalLinkService">
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<mods:title id="AFC3A07F01998FC2CF0E4F5BCD442BDD">Nebria (Patrobonebria) incognita n. sp. and Nebria (P.) hiekeiana n. sp., two new species from the Western Himalaya, with remarks on Nebria (P.) desgodinsi (Coleoptera, Carabidae, Nebriinae)</mods:title>
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<mods:namePart id="BF6E5B0B1B7634D88B205A29E77FEA35">Charles Huber</mods:namePart>
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<mods:namePart id="57BD909A13D5936AE6F942BBE113D526">Hannes Baur</mods:namePart>
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<treatment id="8C13571B821C5D76FF16FA8A772FAE28" ID-DOI="http://doi.org/10.5281/zenodo.6074276" ID-GBIF-Taxon="127601525" ID-Zenodo-Dep="6074276" LSID="urn:lsid:plazi:treatment:8C13571B821C5D76FF16FA8A772FAE28" httpUri="http://treatment.plazi.org/id/8C13571B821C5D76FF16FA8A772FAE28" lastPageId="4" lastPageNumber="207" pageId="2" pageNumber="205">
<paragraph id="0405E60D821C5D70FF16FA8A7539ADBD" blockId="2.[230,833,1380,1440]" pageId="2" pageNumber="205">
<subSubSection id="4CA0B586821C5D70FF16FA8A775CAD9C" box="[230,833,1380,1409]" pageId="2" pageNumber="205" type="nomenclature">
<taxonomicName id="C3BA9D8E821C5D70FF16FA8A775CAD9C" ID-CoL="8MMGC" ID-ENA="2865156" authority="OBERTHUR, 1883" authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[230,833,1380,1409]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="205" phylum="Arthropoda" rank="species" species="desgodinsi" subGenus="Patrobonebria">
<emphasis id="36CE3A1F821C5D70FF16FA8A775CAD9C" bold="true" box="[230,833,1380,1409]" pageId="2" pageNumber="205">
<emphasis id="36CE3A1F821C5D70FF16FA8A7667AD9C" bold="true" box="[230,634,1380,1409]" italics="true" pageId="2" pageNumber="205">Nebria (Patrobonebria) desgodinsi</emphasis>
<bibRefCitation id="602B9BFC821C5D70FD70FA8A76E6AD62" author="OBERTHUR, R." box="[640,763,1380,1409]" firstAuthor="OBERTHUR" journalOrPublisher="Coleopterorum novitates" pageId="2" pageNumber="205" pagination="47 - 49" part="1" refId="ref8817" refString="OBERTHUR, R. (1883): Trois Nebria nouvelles. - Coleopterorum novitates 1: 47 - 49." title="Trois Nebria nouvelles" type="journal article" year="1883">OBERTHÜR</bibRefCitation>
, 1883
</emphasis>
</taxonomicName>
</subSubSection>
<subSubSection id="4CA0B586821C5D70FF16FA667539ADBD" box="[230,292,1416,1440]" pageId="2" pageNumber="205" type="description">
<figureCitation id="9C81FA88821C5D70FF16FA667539ADBD" box="[230,292,1416,1440]" captionStart="Fig. 2" captionStartId="3.[230,265,1247,1269]" captionTargetBox="[230,831,288,1203]" captionTargetId="figure@3.[230,831,283,1203]" captionText="Fig. 2. Habitus of the male type specimen of Nebria (Patrobonebria) desgodinsi OBERTHÜR, 1883 (MNHP). Photograph: C. GERMANN." httpUri="https://zenodo.org/record/247618/files/figure.png" pageId="2" pageNumber="205">Fig. 2</figureCitation>
</subSubSection>
</paragraph>
<subSubSection id="4CA0B586821C5D70FF16FA297759AE7D" pageId="2" pageNumber="205" type="materials_examined">
<paragraph id="0405E60D821C5D70FF16FA297759AE7D" blockId="2.[230,837,1479,1632]" pageId="2" pageNumber="205">
<emphasis id="36CE3A1F821C5D70FF16FA2975DBADC2" bold="true" box="[230,454,1479,1503]" pageId="2" pageNumber="205">Material examined.</emphasis>
Material mislabelled as type material: 1 Male, labelled: Type [red label] // British Bootang, Maria Basti, L. Durel [printed] // Museum Paris, Ex. Coll. M. Maindron, Coll. G. Baubault 1930 [printed] //
<taxonomicName id="C3BA9D8E821C5D70FD0BF9C675DFAE7D" authority="Desgodinsi Oberthur" authorityName="desgodinsi OBERTHUR" authorityYear="1883" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="36CE3A1F821C5D70FD0BF9C67540AE7D" italics="true" pageId="2" pageNumber="205">Nebria Desgodinsi</emphasis>
Oberthur
</taxonomicName>
type [handwritten] (MNHN);
<figureCitation id="9C81FA88821C5D70FCF1F9A6775DAE7D" box="[769,832,1608,1632]" captionStart="Fig. 3" captionStartId="3.[867,902,1247,1269]" captionTargetBox="[933,1336,283,1205]" captionTargetId="figure@3.[933,1336,283,1205]" captionText="Fig. 3. Original labels of the (putative) male type specimen of Nebria (Patrobonebria) desgodinsi OBERTHÜR, 1883 (MNHP). Photograph: C. GERMANN." httpUri="https://zenodo.org/record/247619/files/figure.png" pageId="2" pageNumber="205">Fig. 3</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4CA0B586821C5D71FC78FC6977D4AFB8" lastPageId="3" lastPageNumber="206" pageId="2" pageNumber="205" type="discussion">
<paragraph id="0405E60D821C5D70FC78FC69709DABC2" blockId="2.[867,1470,903,1663]" pageId="2" pageNumber="205">
<emphasis id="36CE3A1F821C5D70FC78FC6977EAAB82" bold="true" box="[904,1015,903,927]" pageId="2" pageNumber="205">Remarks.</emphasis>
After
<bibRefCitation id="602B9BFC821C5D70FBC9FC667122AB82" author="LEDOUX, G. &amp; ROUX, P." box="[1081,1343,903,927]" firstAuthor="LEDOUX" journalOrPublisher="Nippononebria et Oreonebria du MNHN a Paris. - Revue francaise d'Entomologie (N. S.)" pageId="2" pageNumber="205" pagination="29 - 48" part="14" refId="ref8581" refString="LEDOUX, G. &amp; ROUX, P. (1992 b): Les types de Nebria, Nippononebria et Oreonebria du MNHN a Paris. - Revue francaise d'Entomologie (N. S.) 14: 29 - 48." title="Les types de Nebria" type="journal article" year="1992" yearSuffix="b">LEDOUX &amp; ROUX (1992b)</bibRefCitation>
the type series of
<taxonomicName id="C3BA9D8E821C5D70FC59FC467024ABA2" authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[937,1081,935,959]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="205" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821C5D70FC59FC467024ABA2" box="[937,1081,935,959]" italics="true" pageId="2" pageNumber="205">N. desgodinsi</emphasis>
</taxonomicName>
of the OBERTHÜR collection (MNHN) consists of two specimens.
</paragraph>
<paragraph id="0405E60D821C5D70FC78FC0977F6ADA2" blockId="2.[867,1470,903,1663]" pageId="2" pageNumber="205">
<bibRefCitation id="602B9BFC821C5D70FC78FC09705DABE2" author="OBERTHUR, R." box="[904,1088,999,1023]" firstAuthor="OBERTHUR" journalOrPublisher="Coleopterorum novitates" pageId="2" pageNumber="205" pagination="47 - 49" part="1" refId="ref8817" refString="OBERTHUR, R. (1883): Trois Nebria nouvelles. - Coleopterorum novitates 1: 47 - 49." title="Trois Nebria nouvelles" type="journal article" year="1883">OBERTHÜR (1883)</bibRefCitation>
dedicated the species to the French missionary in Tibet AUGUSTE DESGODINS (1826-1914), who was known to collect butterFies for OBERTHÜR. Describing the species, OBERTHÜR mentioned as type locality the surroundings of Darjeeling, India (“les environs même de Darjeeling”); DESGODINS had lived in Darjeeling. But OBERTHÜR noted neither a locality named Maria Basti nor a collector named DUREL nor a calendar date of collecting.
<bibRefCitation id="602B9BFC821C5D70FC2AFB09708AACE2" author="ANDREWES, H. E." bookContentInfo="431 pp." box="[986,1175,1255,1279]" firstAuthor="ANDREWES" journalOrPublisher="London" pageId="2" pageNumber="205" refId="ref8137" refString="ANDREWES, H. E. (1929): The fauna of British India, including Ceylon and Burma. Coleoptera, Carabidae, Vol. 1 - Carabinae. - Taylor and Francis, London, 431 pp., 10 pl. + 1 map." title="The fauna of British India, including Ceylon and Burma. Coleoptera, Carabidae, Vol. 1 - Carabinae. - Taylor and Francis" type="book" year="1929">ANDREWES (1929)</bibRefCitation>
additionally indicated Maria Basti, the locality for the type specimen mentioned above. Maria Basti is a Christian church near Pedong and Kalimpong, 40 km east of the city of Darjeeling. The Maria Basti parish was established in 1891 (
<bibRefCitation id="602B9BFC821C5D70FAB8FA867781AD82" accessDate="invalid" author="RAI GOMES, F." firstAuthor="RAI GOMES" journalOrPublisher="The Herald" pageId="2" pageNumber="205" pagination="67" part="11" publicationUrl="http://calcuttaherald.wordpress.com/2012/07/11/67/" refId="ref8838" refString="RAI GOMES, F. (2012): 125 Years of Maria Busty Church. - The Herald, July 11, 2012 (http: // calcuttaherald. wordpress. com / 2012 / 07 / 11 / 67 /)" title="125 Years of Maria Busty Church" type="url" year="2012">RAI GOMES 2012</bibRefCitation>
), several years after OBERTHÜR described the species in 1883.
</paragraph>
<paragraph id="0405E60D821C5D70FC78FA2670D4AE62" blockId="2.[867,1470,903,1663]" pageId="2" pageNumber="205">
L. DUREL seems to have been an active collector in Maria Basti between 1898 (
<bibRefCitation id="602B9BFC821C5D70FB52FA09715EADE2" author="GRESSITT, J. L." box="[1186,1347,1511,1535]" firstAuthor="GRESSITT" journalOrPublisher="Proceedings of the Hawaiian Entomological Society" pageId="2" pageNumber="205" pagination="317 - 326" part="15" refId="ref8304" refString="GRESSITT, J. L. (1954): The Longicorn Genus Thranius (Coleoptera: Cerambycidae) - Proceedings of the Hawaiian Entomological Society 15: 317 - 326." title="The Longicorn Genus Thranius (Coleoptera: Cerambycidae)" type="journal article" year="1954">GRESSITT 1954</bibRefCitation>
,
<bibRefCitation id="602B9BFC821C5D70FAA4FA0677BBAE02" author="LIN, M. &amp; MONTREUIL, O. &amp; TAVAKILIAN, G. &amp; YANG X." etAl="et al." firstAuthor="LIN" journalOrPublisher="Zootaxa" pageId="2" pageNumber="205" pagination="1 - 22" part="2137" refId="ref8647" refString="LIN, M., MONTREUIL, O., TAVAKILIAN, G. &amp; YANG X. (2009 a): Reinstatement of the genus Heteroglenea GAHAN, with four new combinations, four new synonyms and three new species (Coleoptera: Cerambycidae: Lamiinae: Saperdini). - Zootaxa, 2137: 1 - 22." title="Reinstatement of the genus Heteroglenea GAHAN, with four new combinations, four new synonyms and three new species (Coleoptera: Cerambycidae: Lamiinae: Saperdini)" type="journal article" year="2009" yearSuffix="a">LIN et al. 2009a</bibRefCitation>
) and 1914 (
<bibRefCitation id="602B9BFC821C5D70FBC2F9E670F6AE02" author="LIN, M. &amp; MONTREUIL, O. &amp; TAVAKILIAN, G. &amp; YANG X." box="[1074,1259,1543,1567]" etAl="et al." firstAuthor="LIN" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="2" pageNumber="205" pagination="157 - 176" part="45" refId="ref8708" refString="LIN, M., MONTREUIL, O., TAVAKILIAN, G. &amp; YANG X. (2009 b): A study on the indiana &amp; galathea species-group of the genus Glenea, with descriptions of four new species (Coleoptera: Cerambycidae: Lamiinae: Saperdini). - Annales de la Societe Entomologique de France (N. S.) 45 (2): 157 - 176." title="A study on the indiana &amp; galathea species-group of the genus Glenea, with descriptions of four new species (Coleoptera: Cerambycidae: Lamiinae: Saperdini)" type="journal article" year="2009" yearSuffix="b">LIN et al. 2009b</bibRefCitation>
); an identically labelled
<taxonomicName id="C3BA9D8E821C5D70FC59F9C97006AE22" authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[937,1051,1575,1599]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="205" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821C5D70FC59F9C97006AE22" box="[937,1051,1575,1599]" italics="true" pageId="2" pageNumber="205">desgodinsi</emphasis>
</taxonomicName>
specimen at the BMNH (“British Bootang, Maria Basti, L. Durel”) is annotated by OBERTHÜR himself as “compared with type”.
</paragraph>
<caption id="50C5B685821C5D70FF16F95F71AAAEFF" pageId="2" pageNumber="205">
<paragraph id="0405E60D821C5D70FF16F95F71AAAEFF" pageId="2" pageNumber="205">
<emphasis id="36CE3A1F821C5D70FF16F95F7536AEDB" bold="true" box="[230,299,1713,1734]" pageId="2" pageNumber="205">Tab. 2.</emphasis>
First and second-best ratios found by the LDA ratio extractor for separating various groupings of
<taxonomicName id="C3BA9D8E821C5D70FB23F95F71ABAEDB" authority="(Patrobonebria)" authorityName="Patrobonebria" box="[1235,1462,1713,1734]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="36CE3A1F821C5D70FB23F95F7108AEDB" box="[1235,1301,1713,1734]" italics="true" pageId="2" pageNumber="205">Nebria</emphasis>
(
<emphasis id="36CE3A1F821C5D70FAD3F95F71B3AEDB" box="[1315,1454,1713,1734]" italics="true" pageId="2" pageNumber="205">Patrobonebria</emphasis>
)
</taxonomicName>
species. Ratios marked with * have very little or no overlap and were thus eligible for use in the identiFcation key and the diagnoses.
</paragraph>
</caption>
<caption id="50C5B685821D5D71FF16FB31757FAD37" httpUri="https://zenodo.org/record/247618/files/figure.png" pageId="3" pageNumber="206" targetBox="[230,831,288,1203]" targetPageId="3">
<paragraph id="0405E60D821D5D71FF16FB31757FAD37" pageId="3" pageNumber="206">
<emphasis id="36CE3A1F821D5D71FF16FB317535ACE9" bold="true" box="[230,296,1247,1269]" pageId="3" pageNumber="206">Fig. 2.</emphasis>
Habitus of the male type specimen of
<taxonomicName id="C3BA9D8E821D5D71FD5EFB3175A8AD12" authorityName="Oberthur, G. Ledoux" authorityYear="1883" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi" subGenus="Patrobonebria">
<emphasis id="36CE3A1F821D5D71FD5EFB3176ECACE9" box="[686,753,1247,1268]" italics="true" pageId="3" pageNumber="206">Nebria</emphasis>
(
<emphasis id="36CE3A1F821D5D71FCF1FB0E7520AD12" italics="true" pageId="3" pageNumber="206">Patrobonebria</emphasis>
)
<emphasis id="36CE3A1F821D5D71FEBEFB1475A8AD12" box="[334,437,1274,1295]" italics="true" pageId="3" pageNumber="206">desgodinsi</emphasis>
</taxonomicName>
<bibRefCitation id="602B9BFC821D5D71FE4EFB147641AD12" author="OBERTHUR, R." box="[446,604,1274,1295]" firstAuthor="OBERTHUR" journalOrPublisher="Coleopterorum novitates" pageId="3" pageNumber="206" pagination="47 - 49" part="1" refId="ref8817" refString="OBERTHUR, R. (1883): Trois Nebria nouvelles. - Coleopterorum novitates 1: 47 - 49." title="Trois Nebria nouvelles" type="journal article" year="1883">OBERTHÜR, 1883</bibRefCitation>
(MNHP). Photograph: C. GERMANN.
</paragraph>
</caption>
<caption id="50C5B685821D5D71FC93FB31707DAD37" httpUri="https://zenodo.org/record/247619/files/figure.png" pageId="3" pageNumber="206" targetBox="[933,1336,283,1205]" targetPageId="3">
<paragraph id="0405E60D821D5D71FC93FB31707DAD37" pageId="3" pageNumber="206">
<emphasis id="36CE3A1F821D5D71FC93FB3177BEACE9" bold="true" box="[867,931,1247,1269]" pageId="3" pageNumber="206">Fig. 3.</emphasis>
Original labels of the (putative) male type specimen of
<taxonomicName id="C3BA9D8E821D5D71FC93FB1470A5AD12" authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[867,1208,1274,1295]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi" subGenus="Patrobonebria">
<emphasis id="36CE3A1F821D5D71FC93FB1477BBAD12" box="[867,934,1274,1295]" italics="true" pageId="3" pageNumber="206">Nebria</emphasis>
(
<emphasis id="36CE3A1F821D5D71FC45FB14705CAD12" box="[949,1089,1274,1295]" italics="true" pageId="3" pageNumber="206">Patrobonebria</emphasis>
)
<emphasis id="36CE3A1F821D5D71FBA1FB1470A5AD12" box="[1105,1208,1274,1295]" italics="true" pageId="3" pageNumber="206">desgodinsi</emphasis>
</taxonomicName>
<bibRefCitation id="602B9BFC821D5D71FB4FFB147146AD12" author="OBERTHUR, R." box="[1215,1371,1274,1295]" firstAuthor="OBERTHUR" journalOrPublisher="Coleopterorum novitates" pageId="3" pageNumber="206" pagination="47 - 49" part="1" refId="ref8817" refString="OBERTHUR, R. (1883): Trois Nebria nouvelles. - Coleopterorum novitates 1: 47 - 49." title="Trois Nebria nouvelles" type="journal article" year="1883">OBERTHÜR, 1883</bibRefCitation>
(MNHP). Photograph: C. GERMANN.
</paragraph>
</caption>
<paragraph id="0405E60D821D5D71FEFBFAA375DDAE98" blockId="3.[230,831,1357,2117]" pageId="3" pageNumber="206">
LÉON MARTIN DUREL (1869-1940), the collector of the specimen in question, was born in Bernac (Tarn
<emphasis id="36CE3A1F821D5D71FD2BFA8276F9AD99" bold="true" box="[731,740,1388,1412]" pageId="3" pageNumber="206">)</emphasis>
, France, was a priest of the Missions Étrangères de Paris (see
<bibRefCitation id="602B9BFC821D5D71FF16FA437540ADD8" author="MEP" box="[230,349,1453,1477]" firstAuthor="MEP" journalOrPublisher="Missions Etrangeres de Paris" pageId="3" pageNumber="206" pagination="2015" part="10" publicationUrl="http://archives.mepasie.org/notices/notices-biographiques" refId="ref8783" refString="MEP (2015): Missions Etrangeres de Paris. http: // archives. mepasie. org / notices / notices-biographiques / durel (visited: 8. 10. 2015)." type="journal article" year="2015">MEP 2015</bibRefCitation>
). DUREL was ordained in February 1894, and he left Paris immediately in March 1894 for the “Mission du Tibet” with his destination being Maria Basti in Sikkim. Hence, DUREL could not have been the collector of OBERTHÜR s type material in (or before) 1883. However, DUREL probably met DESGODINS in Pedong, where DESGODINS died in 1913.
</paragraph>
<paragraph id="0405E60D821D5D71FEFBF963769EAF38" blockId="3.[230,831,1357,2117]" pageId="3" pageNumber="206">
For these reasons, the
<taxonomicName id="C3BA9D8E821D5D71FDF0F963766FAEB8" authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[512,626,1677,1701]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821D5D71FDF0F963766FAEB8" box="[512,626,1677,1701]" italics="true" pageId="3" pageNumber="206">desgodinsi</emphasis>
</taxonomicName>
specimen on hand from the OBERTHÜR collection of the MNHN from Maria Basti and collected by DUREL does not belong to the type series. This specimen was erroneously listed as a type specimen by
<bibRefCitation id="602B9BFC821D5D71FE83F8E37662AF38" author="LEDOUX, G. &amp; ROUX, P." box="[371,639,1805,1829]" firstAuthor="LEDOUX" journalOrPublisher="Nippononebria et Oreonebria du MNHN a Paris. - Revue francaise d'Entomologie (N. S.)" pageId="3" pageNumber="206" pagination="29 - 48" part="14" refId="ref8581" refString="LEDOUX, G. &amp; ROUX, P. (1992 b): Les types de Nebria, Nippononebria et Oreonebria du MNHN a Paris. - Revue francaise d'Entomologie (N. S.) 14: 29 - 48." title="Les types de Nebria" type="journal article" year="1992" yearSuffix="b">LEDOUX &amp; ROUX (1992b)</bibRefCitation>
.
</paragraph>
<paragraph id="0405E60D821D5D71FEFBF8C377D4AFB8" blockId="3.[230,831,1357,2117]" lastBlockId="3.[867,1470,1357,2117]" pageId="3" pageNumber="206">
<emphasis id="36CE3A1F821D5D71FEFBF8C375ECAF58" bold="true" box="[267,497,1837,1861]" pageId="3" pageNumber="206">Additional material.</emphasis>
1 female, British Bootang, Maria Basti, L. Durel [printed], 1901-193 [on the underside of the label] //
<taxonomicName id="C3BA9D8E821D5D71FE62F88376ABAF98" authority="R. Obt." authorityName="R. Obt." box="[402,694,1901,1925]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821D5D71FE62F8837644AF98" box="[402,601,1901,1925]" italics="true" pageId="3" pageNumber="206">Nebria desgodinsi</emphasis>
R. Obt.
</taxonomicName>
comparé au TYPE par R. Oberthür [printed] //
<taxonomicName id="C3BA9D8E821D5D71FD87F86375A4AFD8" authority="Oberth. G. Ledoux" authorityName="Oberth. G. Ledoux" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821D5D71FD87F8637722AFB8" box="[631,831,1933,1957]" italics="true" pageId="3" pageNumber="206">Nebria desgodinsi</emphasis>
Oberth. G. Ledoux
</taxonomicName>
det. 82 [handwritten] (BMNH); 1 male, 2 females, Gopaldhara, Rungbong Valley, Darjiling, 19-xi-1919, 3500 ft., H. Stevens //
<taxonomicName id="C3BA9D8E821D5D71FD54F80375E5A038" authority="Oberth. G. Ledoux" authorityName="Oberth. G. Ledoux" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821D5D71FD54F8037507A038" italics="true" pageId="3" pageNumber="206">Nebria desgodinsi</emphasis>
Oberth. G. Ledoux
</taxonomicName>
det. 1982 (BMNH); 1 male, Gopaldhara, Rungbong Valley, Darjiling, 23-25-xi-1919, 3500 ft., H. Stevens //
<taxonomicName id="C3BA9D8E821D5D71FBADFAA377D2AD98" authority="Oberth. H. E. Andrewes" authorityName="Oberth. H. E. Andrewes" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821D5D71FBADFAA3713FAD78" box="[1117,1314,1357,1381]" italics="true" pageId="3" pageNumber="206">Nebria desgodinsi</emphasis>
Oberth. H. E. Andrewes
</taxonomicName>
det. (BMNH); 3 males, 1 female, Gopaldhara, Rungbong Valley, Darjiling, Dec-1920, 3500 ft., H. Stevens //
<taxonomicName id="C3BA9D8E821D5D71FC44FA437152ADD8" authority="Oberth. G. Ledoux" authorityName="Oberth. G. Ledoux" box="[948,1359,1453,1477]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821D5D71FC44FA437064ADD8" box="[948,1145,1453,1477]" italics="true" pageId="3" pageNumber="206">Nebria desgodinsi</emphasis>
Oberth. G. Ledoux
</taxonomicName>
det. 1982 (BMNH); 1 female, Soondrijal, Nepal [printed] // Indian Mus. Calcutta [printed] //
<taxonomicName id="C3BA9D8E821D5D71FB50FA0371A3AE18" authority="Oberth." authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[1184,1470,1517,1541]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821D5D71FB50FA037179AE18" box="[1184,1380,1517,1541]" italics="true" pageId="3" pageNumber="206">Nebria desgodinsi</emphasis>
Oberth.
</taxonomicName>
[handwritten] // H.E. Andrewes coll. (BMNH); 1 male, Nepal, Himalaya, S Khumbu-Himal, Junbesi Umgeb., Junbesi Khola, 2600 m, 24. 4. 1993, leg. Kleeberg,
<taxonomicName id="C3BA9D8E821D5D71FA67F9A07021AE98" authority="Ob." authorityName="Oberthur, G. Ledoux" authorityYear="1883" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821D5D71FA67F9A07017AE98" italics="true" pageId="3" pageNumber="206">Nebria desgodinsi</emphasis>
Ob.
</taxonomicName>
det. J. Schmidt 2004 (CSCHM); 1 male Nepal, Himalaya, S Khumbu-Himal, Umgeb. Jiri, Shivalaya, 1707 m, 1. 5. 1993, leg. Kleeberg //
<taxonomicName id="C3BA9D8E821D5D71FA83F943703DAEF8" authority="Obth." authorityName="Oberthur, G. Ledoux" authorityYear="1883" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821D5D71FA83F94377C8AEF8" italics="true" pageId="3" pageNumber="206">Nebria desgodinsi</emphasis>
Obth.
</taxonomicName>
det. J. Schmidt (CSCHM); 1 male Ost-Nepal, Umgebung Shivalaya, Ufer Kimti Khola, 2. 5. 1993, leg. Kleeberg //
<taxonomicName id="C3BA9D8E821D5D71FB74F8E371A7AF38" authority="Oberthur" authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[1156,1466,1805,1829]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821D5D71FB74F8E37151AF38" box="[1156,1356,1805,1829]" italics="true" pageId="3" pageNumber="206">Nebria desgodinsi</emphasis>
Oberthür
</taxonomicName>
, det. C. Huber (NMBE); 1 male Nepal, Ramechap distr., Khimti Khola near Shivalaya, 1800 m, 5. 5. 1997, leg. M. Hauser //
<taxonomicName id="C3BA9D8E821D5D71FC05F883713FAF98" authority="Oberthur" authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[1013,1314,1901,1925]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821D5D71FC05F88370A4AF98" box="[1013,1209,1901,1925]" italics="true" pageId="3" pageNumber="206">Nebria desgodinsi</emphasis>
Oberthür
</taxonomicName>
det. C. Huber (NMBE).
</paragraph>
</subSubSection>
<subSubSection id="4CA0B586821D5D71FC78F843710EA038" pageId="3" pageNumber="206" type="materials_examined">
<paragraph id="0405E60D821D5D71FC78F843710EA038" blockId="3.[867,1470,1357,2117]" pageId="3" pageNumber="206">
<emphasis id="36CE3A1F821D5D71FC78F84370FEAFD8" bold="true" box="[904,1251,1965,1989]" pageId="3" pageNumber="206">Unexamined type material.</emphasis>
Lectotype: Male (MNHN), labelled with “Dargeeling // A. Desgodins” and “
<taxonomicName id="C3BA9D8E821D5D71FC6BF803707FA018" authority="Desgodinsi" authorityName="desgodinsi OBERTHUR" authorityYear="1883" box="[923,1122,2029,2053]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="genus">
<emphasis id="36CE3A1F821D5D71FC6BF803707FA018" box="[923,1122,2029,2053]" italics="true" pageId="3" pageNumber="206">Nebria Desgodinsi</emphasis>
</taxonomicName>
// R. Oberthür, type // Col. novit. I. p. 47” (
<bibRefCitation id="602B9BFC821D5D71FC3AF7E370D9A038" author="LEDOUX, G. &amp; ROUX, P." box="[970,1220,2061,2085]" firstAuthor="LEDOUX" journalOrPublisher="Nippononebria et Oreonebria du MNHN a Paris. - Revue francaise d'Entomologie (N. S.)" pageId="3" pageNumber="206" pagination="29 - 48" part="14" refId="ref8581" refString="LEDOUX, G. &amp; ROUX, P. (1992 b): Les types de Nebria, Nippononebria et Oreonebria du MNHN a Paris. - Revue francaise d'Entomologie (N. S.) 14: 29 - 48." title="Les types de Nebria" type="journal article" year="1992" yearSuffix="b">LEDOUX &amp; ROUX 1992b</bibRefCitation>
, p. 34).
</paragraph>
</subSubSection>
<subSubSection id="4CA0B586821D5D76FC78F7C3753EAC88" lastPageId="4" lastPageNumber="207" pageId="3" pageNumber="206" type="discussion">
<paragraph id="0405E60D821D5D76FC78F7C3769DA968" blockId="3.[867,1470,1357,2117]" lastBlockId="4.[230,837,285,1589]" lastPageId="4" lastPageNumber="207" pageId="3" pageNumber="206">
Even though
<bibRefCitation id="602B9BFC821D5D71FBE6F7C370D0A058" author="OBERTHUR, R." box="[1046,1229,2093,2117]" firstAuthor="OBERTHUR" journalOrPublisher="Coleopterorum novitates" pageId="3" pageNumber="206" pagination="47 - 49" part="1" refId="ref8817" refString="OBERTHUR, R. (1883): Trois Nebria nouvelles. - Coleopterorum novitates 1: 47 - 49." title="Trois Nebria nouvelles" type="journal article" year="1883">OBERTHÜR (1883)</bibRefCitation>
gave no exact number of type specimens, this specimen originating from Darjeeling as noted by OBERTHÜR was correctly designated as lectotype by
<bibRefCitation id="602B9BFC821A5D76FE80FEB07661A968" author="LEDOUX, G. &amp; ROUX, P." box="[368,636,349,373]" firstAuthor="LEDOUX" journalOrPublisher="Nippononebria et Oreonebria du MNHN a Paris. - Revue francaise d'Entomologie (N. S.)" pageId="4" pageNumber="207" pagination="29 - 48" part="14" refId="ref8581" refString="LEDOUX, G. &amp; ROUX, P. (1992 b): Les types de Nebria, Nippononebria et Oreonebria du MNHN a Paris. - Revue francaise d'Entomologie (N. S.) 14: 29 - 48." title="Les types de Nebria" type="journal article" year="1992" yearSuffix="b">LEDOUX &amp; ROUX (1992b)</bibRefCitation>
.
</paragraph>
<paragraph id="0405E60D821A5D76FEFBFE9375F7A9E8" blockId="4.[230,837,285,1589]" pageId="4" pageNumber="207">
<emphasis id="36CE3A1F821A5D76FEFBFE937626A988" bold="true" box="[267,571,381,405]" pageId="4" pageNumber="207">Remarks on morphology.</emphasis>
Morphological characters of
<taxonomicName id="C3BA9D8E821A5D76FEC4FE7075D5A9A8" authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[308,456,413,437]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821A5D76FEC4FE7075D5A9A8" box="[308,456,413,437]" italics="true" pageId="4" pageNumber="207">N. desgodinsi</emphasis>
</taxonomicName>
additional to or differing from the descriptions of
<bibRefCitation id="602B9BFC821A5D76FE7FFE537656A9C8" author="OBERTHUR, R." box="[399,587,445,469]" firstAuthor="OBERTHUR" journalOrPublisher="Coleopterorum novitates" pageId="4" pageNumber="207" pagination="47 - 49" part="1" refId="ref8817" refString="OBERTHUR, R. (1883): Trois Nebria nouvelles. - Coleopterorum novitates 1: 47 - 49." title="Trois Nebria nouvelles" type="journal article" year="1883">OBERTHÜR (1883)</bibRefCitation>
,
<bibRefCitation id="602B9BFC821A5D76FDA7FE537708A9C8" author="ANDREWES, H. E." bookContentInfo="431 pp." box="[599,789,445,469]" firstAuthor="ANDREWES" journalOrPublisher="London" pageId="4" pageNumber="207" refId="ref8137" refString="ANDREWES, H. E. (1929): The fauna of British India, including Ceylon and Burma. Coleoptera, Carabidae, Vol. 1 - Carabinae. - Taylor and Francis, London, 431 pp., 10 pl. + 1 map." title="The fauna of British India, including Ceylon and Burma. Coleoptera, Carabidae, Vol. 1 - Carabinae. - Taylor and Francis" type="book" year="1929">ANDREWES (1929)</bibRefCitation>
and
<bibRefCitation id="602B9BFC821A5D76FF16FE3075FBA9E8" author="LEDOUX, G. &amp; ROUX, P." bookContentInfo="976 pp." box="[230,486,477,501]" firstAuthor="LEDOUX" journalOrPublisher="Saint- Just-la-Pendue" pageId="4" pageNumber="207" refId="ref8624" refString="LEDOUX, G. &amp; ROUX, P. (2005): Nebria. - Chirat, Saint-Just-la-Pendue, 976 pp." title="Nebria. - Chirat" type="book" year="2005">LEDOUX &amp; ROUX (2005)</bibRefCitation>
:
</paragraph>
<paragraph id="0405E60D821A5D76FEFBFE137702AAC8" blockId="4.[230,837,285,1589]" pageId="4" pageNumber="207">
Lobus lateralis lobe of the mentum apically deeply incised, incisio apicalis conspicuous (
<figureCitation id="9C81FA88821A5D76FDA8FDF3768AAA28" box="[600,663,541,565]" captionStart="Fig. 4" captionStartId="4.[232,267,2080,2102]" captionTargetBox="[233,835,1653,2018]" captionTargetId="figure@4.[228,835,1651,2023]" captionText="Fig. 4. Anatomy (schematic) of the mentum in Patrobonebria." httpUri="https://zenodo.org/record/247620/files/figure.png" pageId="4" pageNumber="207">Fig. 4</figureCitation>
,
<figureCitation id="9C81FA88821A5D76FD51FDF376ADAA28" box="[673,688,541,565]" captionStart="Fig. 5" captionStartId="5.[230,265,1689,1711]" captionTargetBox="[230,1470,283,1650]" captionTargetId="figure@5.[230,1470,283,1650]" captionText="Fig. 5. Menta of Nebria (Patrobonebria) species. A N. desgodinsi OBERTHÜR, 1883, (putative) male type specimen (MHNP), the right part of the mentum is white-bordered; B N. himalayica BATES, 1889, type specimen (MNHN); C N. capillosa LEDOUX &amp; ROUX, 1992; D N. incognita n. sp.; E N. hiekeiana n. sp. All photographs C. GERMANN." httpUri="https://zenodo.org/record/247621/files/figure.png" pageId="4" pageNumber="207">5</figureCitation>
). Spina apicalis short or stumpy or even absent; if present, the tip of the apex does not overtop the lobus lateralis anteriorly. Submentum with a row of 8-10 setae. Eyes very prominent (
<tableCitation id="4938D3B6821A5D76FF1DFD737528AAA8" box="[237,309,669,693]" captionStart="Tab. 3" captionStartId="9.[230,271,1491,1512]" captionText="Tab. 3. Ratio and range of maximal head width / frons width in species of the subgenus Patrobonebria. N = number of examined specimens." pageId="4" pageNumber="207">Tab. 3</tableCitation>
): Ratio maximal head width/frons width = 1.52 (1.48-1.55). Supraantennal edge narrowly rebordered.
</paragraph>
<paragraph id="0405E60D821A5D76FEFBFD337509ABA8" blockId="4.[230,837,285,1589]" pageId="4" pageNumber="207">
Lateral groove of the pronotum large, punctate. Lateral margin regularly rounded, with an inconspicuous outline break at the insertion site of the lateral seta (the distinct outline break mentioned by
<bibRefCitation id="602B9BFC821A5D76FD94FCD37702AB48" author="ANDREWES, H. E." bookContentInfo="431 pp." box="[612,799,829,853]" firstAuthor="ANDREWES" journalOrPublisher="London" pageId="4" pageNumber="207" refId="ref8137" refString="ANDREWES, H. E. (1929): The fauna of British India, including Ceylon and Burma. Coleoptera, Carabidae, Vol. 1 - Carabinae. - Taylor and Francis, London, 431 pp., 10 pl. + 1 map." title="The fauna of British India, including Ceylon and Burma. Coleoptera, Carabidae, Vol. 1 - Carabinae. - Taylor and Francis" type="book" year="1929">ANDREWES (1929)</bibRefCitation>
for
<taxonomicName id="C3BA9D8E821A5D76FF16FCB0756AAB68" authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[230,375,861,885]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821A5D76FF16FCB0756AAB68" box="[230,375,861,885]" italics="true" pageId="4" pageNumber="207">N. desgodinsi</emphasis>
</taxonomicName>
and by
<bibRefCitation id="602B9BFC821A5D76FE3DFCB076D7AB68" author="LEDOUX, G. &amp; ROUX, P." bookContentInfo="976 pp." box="[461,714,861,885]" firstAuthor="LEDOUX" journalOrPublisher="Saint- Just-la-Pendue" pageId="4" pageNumber="207" refId="ref8624" refString="LEDOUX, G. &amp; ROUX, P. (2005): Nebria. - Chirat, Saint-Just-la-Pendue, 976 pp." title="Nebria. - Chirat" type="book" year="2005">LEDOUX &amp; ROUX (2005)</bibRefCitation>
for
<taxonomicName id="C3BA9D8E821A5D76FD06FCB0753CAB88" authorityName="Ledoux &amp; Roux" authorityYear="1992" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="capillosa">
<emphasis id="36CE3A1F821A5D76FD06FCB0753CAB88" italics="true" pageId="4" pageNumber="207">N. capillosa</emphasis>
</taxonomicName>
respectively refers to specimens of
<taxonomicName id="C3BA9D8E821A5D76FD55FC907734AB88" authority="Charles Huber &amp; Hannes Baur, 2016" authorityName="Charles Huber &amp; Hannes Baur" authorityYear="2016" box="[677,809,894,917]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="incognita" status="n. sp.">
<emphasis id="36CE3A1F821A5D76FD55FC907734AB88" box="[677,809,894,917]" italics="true" pageId="4" pageNumber="207">N. incognita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DFD8764821A5D76FCC0FC90751AABA8" pageId="4" pageNumber="207" rank="species">n. sp.</taxonomicNameLabel>
).
</paragraph>
<paragraph id="0405E60D821A5D76FEFBFC5375BDAC48" blockId="4.[230,837,285,1589]" pageId="4" pageNumber="207">Prosternal process slender, triangular, longitudinally bulged, coarsely punctate, unmargined. 3rd interval of the elytra with 7-12 small setae. Metacoxa with 4-6 basal and 1(-2) apical setae. Sterna 4-6 each with 2-3 posterior paramedial setae.</paragraph>
<paragraph id="0405E60D821A5D76FEFBFBB3753EAC88" blockId="4.[230,837,285,1589]" pageId="4" pageNumber="207">Aedeagus large, midshaft to apex straight; apical tip short.</paragraph>
</subSubSection>
<subSubSection id="4CA0B586821A5D76FEFBFB7376EAAD88" pageId="4" pageNumber="207" type="diagnosis">
<paragraph id="0405E60D821A5D76FEFBFB7376EAAD88" blockId="4.[230,837,285,1589]" pageId="4" pageNumber="207">
<emphasis id="36CE3A1F821A5D76FEFBFB73759DACA8" bold="true" box="[267,384,1181,1205]" pageId="4" pageNumber="207">Diagnosis.</emphasis>
<taxonomicName id="C3BA9D8E821A5D76FE75FB707608ACA8" authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[389,533,1181,1205]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821A5D76FE75FB707608ACA8" box="[389,533,1181,1205]" italics="true" pageId="4" pageNumber="207">N. desgodinsi</emphasis>
</taxonomicName>
belongs to the species group with strongly protruding eyes (
<tableCitation id="4938D3B6821A5D76FDDEFB53766EACC8" box="[558,627,1213,1237]" captionStart="Tab. 3" captionStartId="9.[230,271,1491,1512]" captionText="Tab. 3. Ratio and range of maximal head width / frons width in species of the subgenus Patrobonebria. N = number of examined specimens." pageId="4" pageNumber="207">Tab. 3</tableCitation>
).
<taxonomicName id="C3BA9D8E821A5D76FD77FB507705ACC8" authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[647,792,1213,1237]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821A5D76FD77FB507705ACC8" box="[647,792,1213,1237]" italics="true" pageId="4" pageNumber="207">N. desgodinsi</emphasis>
</taxonomicName>
differs from all species of the subgenus by the apically deeply incised lateral lobe of the mentum (
<figureCitation id="9C81FA88821A5D76FD66FB1376C7AD08" box="[662,730,1277,1301]" captionStart="Fig. 5" captionStartId="5.[230,265,1689,1711]" captionTargetBox="[230,1470,283,1650]" captionTargetId="figure@5.[230,1470,283,1650]" captionText="Fig. 5. Menta of Nebria (Patrobonebria) species. A N. desgodinsi OBERTHÜR, 1883, (putative) male type specimen (MHNP), the right part of the mentum is white-bordered; B N. himalayica BATES, 1889, type specimen (MNHN); C N. capillosa LEDOUX &amp; ROUX, 1992; D N. incognita n. sp.; E N. hiekeiana n. sp. All photographs C. GERMANN." httpUri="https://zenodo.org/record/247621/files/figure.png" pageId="4" pageNumber="207">Fig. 5</figureCitation>
a), by the wide lateral groove of the pronotum, by the densely and coarsely punctate neck, and by the apex of the 4th antennomere without additional short setae. The ratio ant.l/prm.l (see
<tableCitation id="4938D3B6821A5D76FEE6FA937544AD88" box="[278,345,1405,1429]" captionStart="Tab. 2" captionStartId="2.[230,271,1713,1734]" captionText="Tab. 2. First and second-best ratios found by the LDA ratio extractor for separating various groupings of Nebria (Patrobonebria) species. Ratios marked with * have very little or no overlap and were thus eligible for use in the identiFcation key and the diagnoses." pageId="4" pageNumber="207">Tab. 2</tableCitation>
) can be used to separate
<taxonomicName id="C3BA9D8E821A5D76FD90FA9076EDAD88" authorityName="Oberthur, G. Ledoux" authorityYear="1883" box="[608,752,1405,1429]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="desgodinsi">
<emphasis id="36CE3A1F821A5D76FD90FA9076EDAD88" box="[608,752,1405,1429]" italics="true" pageId="4" pageNumber="207">N. desgodinsi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="4CA0B586821A5D76FEFBFA73772FAE28" pageId="4" pageNumber="207" type="distribution">
<paragraph id="0405E60D821A5D76FEFBFA73772FAE28" blockId="4.[230,837,285,1589]" pageId="4" pageNumber="207">
<emphasis id="36CE3A1F821A5D76FEFBFA737586ADA8" bold="true" box="[267,411,1437,1461]" pageId="4" pageNumber="207">Distribution.</emphasis>
Central Nepal (Sundarijal north of Kathmandu Valley), Eastern Nepal, Darjeeling District, West Bengal, India. Specimens reported from the more western portions of the Himalaya are
<taxonomicName id="C3BA9D8E821A5D76FD9FFA107563AE28" authority="Ledoux &amp; Roux, 1992" authorityName="Ledoux &amp; Roux" authorityYear="1992" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="capillosa">
<emphasis id="36CE3A1F821A5D76FD9FFA1076F3AE08" box="[623,750,1533,1557]" italics="true" pageId="4" pageNumber="207">N. capillosa</emphasis>
<bibRefCitation id="602B9BFC821A5D76FD04FA137563AE28" author="LEDOUX, G. &amp; ROUX, P." firstAuthor="Ledoux" journalOrPublisher="Nouvelle Revue d'Entomologie (N. S.)" pageId="4" pageNumber="207" pagination="142" part="9" refId="ref8540" refString="LEDOUX, G. &amp; ROUX, P. (1992 a): Nebria (Patrobonebria) capillosa nouvelle espece himalayenne (Col. Nebriidae) - Nouvelle Revue d'Entomologie (N. S.) 9: 142." title="Nebria (Patrobonebria) capillosa nouvelle espece himalayenne (Col. Nebriidae)" type="journal article" year="1992">Ledoux &amp; Roux, 1992</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C3BA9D8E821A5D76FE79F9F07611AE28" authority="Charles Huber &amp; Hannes Baur, 2016" authorityName="Charles Huber &amp; Hannes Baur" authorityYear="2016" box="[393,524,1566,1589]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="incognita" status="n. sp.">
<emphasis id="36CE3A1F821A5D76FE79F9F07611AE28" box="[393,524,1566,1589]" italics="true" pageId="4" pageNumber="207">N. incognita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DFD8764821A5D76FDE2F9F07656AE28" box="[530,587,1566,1589]" pageId="4" pageNumber="207" rank="species">n. sp.</taxonomicNameLabel>
or
<taxonomicName id="C3BA9D8E821A5D76FD9DF9F076EFAE28" authority="Charles Huber &amp; Hannes Baur, 2016" authorityName="Charles Huber &amp; Hannes Baur" authorityYear="2016" box="[621,754,1565,1589]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="hiekeiana" status="n. sp.">
<emphasis id="36CE3A1F821A5D76FD9DF9F076EFAE28" box="[621,754,1565,1589]" italics="true" pageId="4" pageNumber="207">N. hiekeiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DFD8764821A5D76FD09F9F0772FAE28" box="[761,818,1566,1589]" pageId="4" pageNumber="207" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>