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<document id="C770FBE7F8447ED99C1974C7305F0BFB" ID-DOI="10.3897/zookeys.1213.133171" ID-ZooBank="CCDC5C64-CBF4-4F02-AAD9-979642899A68" ID-publisher-id="133171" URI-arpha="A14DBDE2-4BCF-59C6-96D8-93BB78371946" URI-zoobank="http://zoobank.org/CCDC5C64-CBF4-4F02-AAD9-979642899A68" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1727362524340" checkinUser="pensoft" docAuthor="Iorio, Étienne &amp; Bonato, Lucio" docDate="2024" docId="8BE8362270C55EB4B876324E5316EA3F" docLanguage="en" docName="ZooKeys 1213: 199-224" docOrigin="ZooKeys 1213" docSource="https://zookeys.pensoft.net/article/133171/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Endogeophilus alberti Iorio &amp; Bonato, 2024, sp. nov." docType="treatment" docUuid="D21886AF-5DFE-4248-8532-8D84E673C6DA" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="224" masterDocId="A14DBDE24BCF59C696D893BB78371946" masterDocTitle="A new species of the rare genus Endogeophilus from southern France, with a key to the European genera of Geophilidae s. l. (Chilopoda)" masterLastPageNumber="224" masterPageNumber="199" pageNumber="199" updateTime="1727376973158" updateUser="admin">
<mods:mods id="3B236BBED1FD3FC8D530F156FF7650DA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BCAC110973DE6F9AC1D2DC9D16B037B8">
<mods:title id="0DB42C1CA79AB62A27BC71E5791EE96F">A new species of the rare genus Endogeophilus from southern France, with a key to the European genera of Geophilidae s. l. (Chilopoda)</mods:title>
</mods:titleInfo>
<mods:name id="A8C585740C22076BF902037E0CDAED39" type="personal">
<mods:role id="E99A67C766A648AD033C04FC82E00F14">
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</mods:role>
<mods:namePart id="1770567D89FAE942D4E0F3B6D15D6133">Iorio, Étienne</mods:namePart>
<mods:nameIdentifier id="42B814CEA4DB699F12684260BB15924D" type="ORCID">https://orcid.org/0009-0003-2550-3480</mods:nameIdentifier>
<mods:affiliation id="391CD7B6EF345DD279E8FC34B7EDDE95">EI Entomologie &amp; Myriapodologie, 36 impasse des Acacias, F- 84260 Sarrians, France</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="9AAD5226D3DB49A9F9A102290BD735F0">Bonato, Lucio</mods:namePart>
<mods:nameIdentifier id="884B0AB48366D348F765A90EA5D70EF3" type="ORCID">0000-0002-8312-7570</mods:nameIdentifier>
<mods:affiliation id="5C432281FA30D0F2551D66396E477A2B">Dipartimento di Biologia, Università di Padova, via U. Bassi 58 b, I- 35131 Padova, Italy &amp; National Biodiversity Future Centre, Palermo, Italy</mods:affiliation>
</mods:name>
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<mods:date id="118A49797B954C5399C636C0EEAB21DA">2024</mods:date>
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<mods:number id="E2AD4D7B73712799C891BE6F9F40DA4F">2024-09-26</mods:number>
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<mods:identifier id="8207A628AB1EA7269D8FBC34A21D81EE" type="DOI">10.3897/zookeys.1213.133171</mods:identifier>
<mods:identifier id="578564E521F7AE076570EB5C1ECB9BE1" type="ZooBank">CCDC5C64-CBF4-4F02-AAD9-979642899A68</mods:identifier>
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<treatment id="8BE8362270C55EB4B876324E5316EA3F" ID-arpha="8BE83622-70C5-5EB4-B876-324E5316EA3F" ID-zoobank="https://zoobank.org/D21886AF-5DFE-4248-8532-8D84E673C6DA" LSID="urn:lsid:zoobank.org:act:D21886AF-5DFE-4248-8532-8D84E673C6DA" LSID-ZBK="urn:lsid:zoobank.org:act:D21886AF-5DFE-4248-8532-8D84E673C6DA" httpUri="http://treatment.plazi.org/id/8BE8362270C55EB4B876324E5316EA3F">
<subSubSection id="1DE80F67AAC9BDF404A21DCD8CFAA36F" type="nomenclature">
<paragraph id="47DDC26DE055DB70FF4923136339FB57">
<taxonomicName id="2E6BDD751A902E9710F85B73ACC7DE5B" authorityName="Iorio &amp; Bonato" authorityYear="2024" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
<emphasis id="1CE5752A68B56771F0E046C5726553B9" italics="true">Endogeophilus alberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6811C66951C72983E26B627A551A807" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWKAE" type="Type specimen">
<paragraph id="72317015A81C42A55C74F2CFE32E6D15">
<heading id="C13B403BB7EB14A52195DA60A6E27335" reason="title">Type specimen.</heading>
</paragraph>
<paragraph id="158B0D8F534605B1E937F98CC33ABD30">
<materialsCitation id="C0EFD8868D790BEA595BD330D03CB21B" collectingDate="2023-03-31" collectionCode="MALAT" collectorName="E. Iorio" latitude="43.1795" location="Cavalaire-sur-Mer" longLatPrecision="6" longitude="6.5068" municipality="Malatra" specimenCode="MALAT 3, WGS 84, M 370" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="4308F095E7B5B53068AF6E74338EA258" bold="true" italics="true">
<typeStatus id="DEFDCB9FD90913C1881C581754A05C8A" type="holotype">Holotype</typeStatus>
</emphasis>
(Figs
<figureCitation id="655D09591F9AA65A7A830A3519E3F651" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2</figureCitation>
,
<figureCitation id="172EF039C41AD5B24CA160C589D31E61" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3</figureCitation>
): •
<specimenCount id="D4C2A4725B043CCEEAC25D6BCDE28024" count="1" type="female"></specimenCount>
,
<date id="7D3E202A0A0B4E7C4DC8E5BA8E9BCD1C" value="2023-03-31">
<collectingDate id="E86027BC66AAE61094E7F394BDF8996C" value="2023-03-31">31. III. 2023</collectingDate>
</date>
,
<collectorName id="2482161B7B2D8116887540CE1997C26F">É. Iorio</collectorName>
leg. In ethanol. Body in four pieces: head, maxillae, mandibles, and trunk. Original label:
<location id="69DC75FCB3B62D720B82DB105A8DF746" LSID="urn:lsid:plazi:treatment:8BE8362270C55EB4B876324E5316EA3F:69DC75FCB3B62D720B82DB105A8DF746" latitude="43.1795" longLatPrecision="6" longitude="6.5068" municipality="Malatra" name="Cavalaire-sur-Mer">Cavalaire-sur-Mer</location>
(Var),
<collectingMunicipality id="CB59C38BA87FD511210B17F23564D207">Malatra</collectingMunicipality>
, “
<specimenCode id="E97CA9EE8BDFC0A1070DA7794C7F9628" collectionCode="MALAT">MALAT 3</specimenCode>
”,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMLAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[6.506800,43.179500]}">
<geoCoordinate id="F002D903E7462C2DF44BD660761FC006" degrees="43.1795" direction="north" orientation="latitude" precision="5" value="43.1795">43.1795 ° N</geoCoordinate>
,
<geoCoordinate id="D0A1F2B43A8BC4470CC367B946B65EFA" degrees="6.5068" direction="east" orientation="longitude" precision="5" value="6.5068">6.5068 ° E</geoCoordinate>
</named-content>
(
<specimenCode id="4F88793EEA5C31980C0A560B8AF1E150">WGS 84</specimenCode>
),
<date id="1E9E675FC48E0BAD3B080DE3E2C4431B" value="2023-03-31">31. III. 2023</date>
, É. Iorio leg. Deposited in the Muséum national dHistoire naturelle, Paris (
<taxonomicName id="26164D916D76AA3722EB2BB53FB5550B" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" phylum="Arthropoda" rank="class">Chilopoda</taxonomicName>
collection, number
<specimenCode id="A6951B12B60184EFD0655BCB2225ECC2">M 370</specimenCode>
).
</materialsCitation>
</paragraph>
<caption id="40A0D20AD7A932E771C9B932B5B18E18" ID-DOI="10.3897/zookeys.1213.133171.figure2" ID-arpha="9C813A5D-BF44-5A7B-8134-F5BF939D52B8" httpUri="https://binary.pensoft.net/fig/1142653" startId="F2">
<paragraph id="D9078431AD0B0AC9B7EB6642C9FD299B">
<label id="89968F3DB917252FF540D853E16D7EBF">Figure 2.</label>
</paragraph>
<paragraph id="7C31FB6C9ECBF5623143E1ADD63D9BE2">
<taxonomicName id="8418560D1D8F5F5C436DC87D470E3EF0" authorityName="Iorio &amp; Bonato" authorityYear="2024" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
<emphasis id="BF41FE8CAA4AAE4A1B7412600DAD14A5" italics="true">Endogeophilus alberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2CF1B8C0D35EE9A2C04B4B08E031F483" rank="species">sp. nov.</taxonomicNameLabel>
, holotype
<emphasis id="7B6A21A448A88A83ED6910F276191902" bold="true">A</emphasis>
labrum
<emphasis id="DF51BCA51B380DEF8F875832B9DE4222" bold="true">B</emphasis>
first maxillae
<emphasis id="28ACACCA02A2D6B143F736FD8BF5A078" bold="true">C</emphasis>
right part of the forcipular segment
<emphasis id="F2CED54DCDD54B3AD7956122498C25A6" bold="true">D</emphasis>
<abbrev id="ABBRID0ERMAE" xlink_title="leg-bearing segment(s)">LBS</abbrev>
18
<emphasis id="1D87378A8B7DB5AA797D78F59B41F026" bold="true">E</emphasis>
ultimate
<abbrev id="ABBRID0EXMAE" xlink_title="leg-bearing segment(s)">LBS</abbrev>
without distal part of legs
<emphasis id="8B420B1429872BE86F2D0BC673E865FB" bold="true">F</emphasis>
left part of the ultimate
<abbrev id="ABBRID0E4MAE" xlink_title="leg-bearing segment(s)">LBS</abbrev>
without distal part of leg. Views: ventral (
<emphasis id="D21764AE3EAAF08DC32E837CE61563B8" bold="true">A E</emphasis>
), dorsal (
<emphasis id="4AF910ACD16FD6BDA1934C9BBA0E7160" bold="true">F</emphasis>
). Scale bars: 20 µm (
<emphasis id="F29C1526A66B85272FA3260EA991A6EC" bold="true">A</emphasis>
); 40 µm (
<emphasis id="32F86BF8E731BCB81E94FDCBCC376DCD" bold="true">B</emphasis>
); 100 µm (
<emphasis id="F196A07A87ACF3FE3ACF6DA14ECEC179" bold="true">C F</emphasis>
).
</paragraph>
</caption>
<caption id="7C25C4AB480AC6479B315C092BD7D5CA" ID-DOI="10.3897/zookeys.1213.133171.figure3" ID-arpha="5FDCB4FD-A474-5B9B-A5BB-975EEAFF97BD" httpUri="https://binary.pensoft.net/fig/1142654" startId="F3">
<paragraph id="F8180EE302C9805DC806056427A55AF2">
<label id="D5CA7BEBFC9022DDF739860255D15D6E">Figure 3.</label>
</paragraph>
<paragraph id="E0A28A1CF51853CD597C9046D9E9C5DF">
<taxonomicName id="D3B1A3E233A2618733121F3177F5DBA9" authorityName="Iorio &amp; Bonato" authorityYear="2024" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
<emphasis id="C2B9105C6AA0C6852CBD2B182FEDBA75" italics="true">Endogeophilus alberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60D5751BD979AA29922A48DCB00DDFD6" rank="species">sp. nov.</taxonomicNameLabel>
, holotype
<emphasis id="49B05B114A4000636ADE37528564B3C2" bold="true">A</emphasis>
head and forcipular segment
<emphasis id="AD0F67C36B14E9DCC9271A2C4627B7C4" bold="true">B</emphasis>
antennae
<emphasis id="6B972EC6340C7EC9563EB85E48E21C7D" bold="true">C</emphasis>
distal articles of left antenna
<emphasis id="2B84B5016378630CFDDB59EAD064F0BD" bold="true">D</emphasis>
right telopodite of second maxillae
<emphasis id="26F5D2D2F293DF566F380B51C429CC86" bold="true">E</emphasis>
head and forcipular segment
<emphasis id="638E265DAFE52A5A02A40E3D00DAB284" bold="true">F</emphasis>
right forcipule
<emphasis id="2B746DABF32D06AABC9BDB2FA0AD1AB0" bold="true">G</emphasis>
<abbrev id="ABBRID0EROAE" xlink_title="leg-bearing segment(s)">LBS</abbrev>
15 and anterior part of 16
<emphasis id="80FC10D3B2789BBA3A871DDCD18BD8AE" bold="true">H</emphasis>
ultimate
<abbrev id="ABBRID0EXOAE" xlink_title="leg-bearing segment(s)">LBS</abbrev>
without right leg. Views: ventral (
<emphasis id="BFE8A0AF9AFC778EABBA811CB542D1F7" bold="true">D H</emphasis>
), dorsal (
<emphasis id="C0627FC9172F727866F67490523CEE9B" bold="true">A C</emphasis>
). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (
<emphasis id="A608893F88C2196997E07EC8A1D4D2DC" bold="true">A B, E, G H</emphasis>
); 50 µm (
<emphasis id="EFC290FD8449CF3704C3C0642BB2C662" bold="true">C D, F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ERPAE" type="type locality">
<paragraph id="04235FBF6D3F7D37953C28582977FFE7">
<heading id="C51C6BE387AE2A6069F7C473BBC91628" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="850DA879609E5EDD043260C138F0CEF4">
<collectingCountry id="5A87606950904AFCBCABB36D292681EF" name="France">France</collectingCountry>
: Var department: Cavalaire-sur-Mer: near Malatra:
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1PAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[6.506800,43.179500]}">
<geoCoordinate id="6726EEBD392851E20C42CC25212429F2" degrees="43.1795" direction="north" orientation="latitude" precision="5" value="43.1795">43.1795 ° N</geoCoordinate>
,
<geoCoordinate id="B631BEE9158951316A1A0DDFF94CF93A" degrees="6.5068" direction="east" orientation="longitude" precision="5" value="6.5068">6.5068 ° E</geoCoordinate>
</named-content>
(WGS 84),
<quantity id="7A25FA5BA332F8FD2845D2187E0E25C7" metricMagnitude="2" metricUnit="m" metricValue="2.15" unit="m" value="215.0">215 m</quantity>
a. s. l., north-eastern slope (Fig.
<figureCitation id="DD211FE77009969885229902F9DE49B6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of all known records of Endogeophilus." figureDoi="10.3897/zookeys.1213.133171.figure1" httpUri="https://binary.pensoft.net/fig/1142652">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDQAE" type="diagnosis">
<paragraph id="56C5189D75193F7126E485B8EDEBCAFD">
<heading id="EC1C5F884F5AB048DC4CCC47AF2BD5FF" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="FA4CF23E15E9A3A795D3887C5921483C">
An
<taxonomicName id="CE4B7EE949BE5CE1A80188C889D34A08">
<emphasis id="CA003816242801760BABB30E8574BF1D" italics="true">Endogeophilus</emphasis>
species
</taxonomicName>
with claw of the second maxillae slender and hooked at its tip; forcipular trochanteroprefemur ~ 1.1 × as long as wide; forcipular tarsungulum&gt; 2.0 × as long as wide, almost as long as the trochanteroprefemur, distinctly curved, fairly slender, and gradually narrowing. See also Table
<tableCitation id="D9FCB7ACD6BB9BC0BBEA025A020D07BD" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Differences between the single specimen of Endogeophilus alberti sp. nov. and the three specimens of E. ichnusae. For each character, each possible explanation (intraspecific variability, errors in character evaluation or divergence between species) has been evaluated as probable (x), possible (?) or improbable (), based on what is known from other Geophilidae. Data are from Bonato et al. (2016) and this study." httpUri="http://table.plazi.org/id/286526C3D6979A25E47B02119AAC51B5" tableUuid="286526C3D6979A25E47B02119AAC51B5">1</tableCitation>
, Figs
<figureCitation id="987A5D6B3C77AA7D816754D4199A9664" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2</figureCitation>
<figureCitation id="CDF1D28389C3B09DB72ACD04D6C0FD6A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Endogeophilus ichnusae, paratype PD 1373, ventral A head and forcipular segment B right forcipule and apical claw of the second maxillae. Abbreviation: cmx claw of the second maxillae. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1213.133171.figure4" httpUri="https://binary.pensoft.net/fig/1142655">4</figureCitation>
, and Discussion.
</paragraph>
<caption id="D3B826B6822ADCE3DB0A1492203023CD" ID-DOI="10.3897/zookeys.1213.133171.figure4" ID-arpha="986DAFFD-75A0-5434-9DDC-48FC7925F136" httpUri="https://binary.pensoft.net/fig/1142655" startId="F4">
<paragraph id="471631684869FFF88D1FC96083F9563C">
<label id="EECE9D2A65E67E5E0B1720B8D49CDAF3">Figure 4.</label>
</paragraph>
<paragraph id="8E0FCA5879693E0AE71E1ABE0767E627">
<taxonomicName id="459A7A6D1CFF7E050502825A5180996D" authorityName="Bonato, Zapparoli, Drago &amp; Minelli" authorityYear="2016" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="ichnusae">
<emphasis id="9620F53B87B0CAAD05B7069B7E2B473E" italics="true">Endogeophilus ichnusae</emphasis>
</taxonomicName>
, paratype PD 1373, ventral
<emphasis id="10023C0A6412547645D5869D241EA192" bold="true">A</emphasis>
head and forcipular segment
<emphasis id="F5B70569FE3C23124CF24577362709FC" bold="true">B</emphasis>
right forcipule and apical claw of the second maxillae. Abbreviation: cmx claw of the second maxillae. Scale bars: 100 µm.
</paragraph>
</caption>
<caption id="286526C3D6979A25E47B02119AAC51B5" ID-Table-UUID="286526C3D6979A25E47B02119AAC51B5" httpUri="http://table.plazi.org/id/286526C3D6979A25E47B02119AAC51B5" startId="T1" targetIsTable="true">
<paragraph id="877D5D43266DCAB6B75405719332F493">
<label id="6381A91D6114E2ED77A4477511711107">Table 1.</label>
</paragraph>
<paragraph id="9CB8856251689D917AA91DE026D15919">
Differences between the single specimen of
<taxonomicName id="BEF17E3D9CC05659D4CAC2C7D52BB6F7" authorityName="Iorio &amp; Bonato" authorityYear="2024" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
<emphasis id="646936880278A1B8771A050D2510C82E" italics="true">Endogeophilus alberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="14969A6C8EAB012A54B3025DF25DDCE1" rank="species">sp. nov.</taxonomicNameLabel>
and the three specimens of
<taxonomicName id="DC9B09890AE3BFCB3FAFA3C033D68E47" authorityName="Bonato, Zapparoli, Drago &amp; Minelli" authorityYear="2016" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="ichnusae">
<emphasis id="661E78F65D073D5EA293CDAD1C34342E" italics="true">E. ichnusae</emphasis>
</taxonomicName>
. For each character, each possible explanation (intraspecific variability, errors in character evaluation or divergence between species) has been evaluated as probable (x), possible (?) or improbable (), based on what is known from other
<taxonomicName id="0500444A1F37613B680E8ABCB961D096" class="Chilopoda" family="Geophilidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Geophilidae</taxonomicName>
. Data are from
<bibRefCitation id="A972984C9CD757B995473328CD56EE1B" DOI="10.5852/ejt.2016.231" author="Bonato L &amp; Zapparoli M &amp; Drago L &amp; Minelli A" etAl="et al." firstAuthor="Bonato" journalOrPublisher="European Journal of Taxonomy" pagination="1-19" refId="B14" refString="Bonato L, Zapparoli M, Drago L, Minelli A (2016) An unusually elongate endogeic centipede from Sardinia (Chilopoda: Geophilidae). European Journal of Taxonomy 231: 119. https://doi.org/10.5852/ejt.2016.231" title="An unusually elongate endogeic centipede from Sardinia (Chilopoda: Geophilidae)." volume="231" year="2016">Bonato et al. (2016)</bibRefCitation>
and this study.
</paragraph>
</caption>
<paragraph id="9C67D064184116D3C1C6DED4DBAEB3F3">
<table id="TID0E1DAI" rules="all">
<tbody id="90926F087A631C4984F05F9040944A57">
<tr id="6DF00B136B5851C046EA333D46C80C05">
<th id="EC18135F469CAB0BEC6C8BBEE8016706" colspan="1" rowspan="1">Species</th>
<th id="08DAD4125B64843BCE5D9D6C87A32053" colspan="1" rowspan="1">
<taxonomicName id="4C116E6FD244B08426756DB98E7CB563" authorityName="Iorio &amp; Bonato" authorityYear="2024" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
<emphasis id="65EC8FA883305AF2123192B027EA4E3E" italics="true">E. alberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="20020EB86EF5717703B617CB8406D52D" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<taxonomicName id="3AFE0BF437D5BB451607BAC3C53CE34D" authority="Explanations" authorityName="Explanations" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="ichnusae">
<th id="A21C8D3C1B434D66000DD7587D92F196" colspan="1" rowspan="1">
<emphasis id="667C7326551DB1B0E8551716053214A2" italics="true">E. ichnusae</emphasis>
</th>
<th id="9B195AD38F5ECBCB92CB37864F06B2B0" colspan="5" rowspan="1">Explanations</th>
</taxonomicName>
</tr>
<tr id="5CCDAB7D1400AE395A97DDBF18B8646C">
<th id="D3C3F10FBE4B075768399895FAB246F7" colspan="1" rowspan="1">Specimens examined</th>
<th id="FF244EF7DB3670541B69F23BC89B31D1" colspan="1" rowspan="1">1, female, 24.5 mm</th>
<th id="33898A9CB787CF257B8DE8A42DF4DA01" colspan="1" rowspan="1">3, both sexes, 3134 mm</th>
<th id="48EA3106A39D00052660FE7FF9243E8D" colspan="1" rowspan="1">Differences in body size</th>
<th id="704CD728F47B1B397999A5263CE43559" colspan="1" rowspan="1">Differences between sexes</th>
<th id="B913B5BADF86EC78DE1AEEDB0D22FE45" colspan="1" rowspan="1">Variability between individuals</th>
<th id="678DF6C5EACDE93CEFAF1309BBB6AF2E" colspan="1" rowspan="1">Errors in character evaluation</th>
<th id="9F78C9EC1B346E3D4435E0A06A8696B4" colspan="1" rowspan="1">Differences between species</th>
</tr>
<tr id="847724C7C226C17E3F7532A0EF14CC22">
<td id="963F7D51689F4D05F273CFDDD537BA6E" colspan="1" rowspan="1">Head: setae: max length</td>
<td id="4E83799744BCA9E3173CA3AC3AF78A5C" colspan="1" rowspan="1">22 μm</td>
<td id="7136A6519F3E7B5F4EE636B02B67744B" colspan="1" rowspan="1">15 μm</td>
<td id="C6771563A1633995003668AF38B70388" colspan="1" rowspan="1"></td>
<td id="3186BCA6CC73B25CFE6755E4253EE86C" colspan="1" rowspan="1"></td>
<td id="56B94B73D20B85E79D08D460F32111B5" colspan="1" rowspan="1">?</td>
<td id="803C33BBFEEE131272830BAC57F38D34" colspan="1" rowspan="1">x</td>
<td id="AD3AEA7E6147EFB11BF887A06D550280" colspan="1" rowspan="1">?</td>
</tr>
<tr id="78A8C8C9C113D2A7596176D0DED073F1">
<td id="36176284485DCFEFCCE3633B3D998769" colspan="1" rowspan="1">Labrum: distinction between side-pieces and intermediate part</td>
<td id="569278C70E5E2DDECF159FA3B851CA34" colspan="1" rowspan="1">faint</td>
<td id="9503B3507F32E9F57A2A2DC8C2E16203" colspan="1" rowspan="1">distinct</td>
<td id="3E88296F736D03F895B765BA41B45723" colspan="1" rowspan="1">x</td>
<td id="97BF85CAEDCEB95DCF99566041D79474" colspan="1" rowspan="1"></td>
<td id="8B20E1394E28AEE1AC84CBEA5C5D877E" colspan="1" rowspan="1">?</td>
<td id="48536FFF007861409A21F078DEC44091" colspan="1" rowspan="1">?</td>
<td id="149BC9C25F8CB689E5376755ABFDF24F" colspan="1" rowspan="1"></td>
</tr>
<tr id="B907688DCC7559C13F92BFE5F5090E70">
<td id="BEBC92C6AB51436BE0F5AB52FCE62519" colspan="1" rowspan="1">Labrum: number of tubercles</td>
<td id="1331F16A8EBA7C833DF7894162F04100" colspan="1" rowspan="1">1</td>
<td id="35C5BFE798FDC6D8FB2438C650A35424" colspan="1" rowspan="1">2</td>
<td id="657853532D5E7ACDFEDD18BACC9B98FF" colspan="1" rowspan="1">?</td>
<td id="C526BDECBDD87AED2DDE36D82DEE4DBE" colspan="1" rowspan="1"></td>
<td id="4E9DF23C1C7B34E4FB4B4AF119BAE075" colspan="1" rowspan="1">x</td>
<td id="8C7CA761F801AFEB953D309E73DF8FE3" colspan="1" rowspan="1"></td>
<td id="EC4DBE6793DD8409732E0FF88B58FB51" colspan="1" rowspan="1"></td>
</tr>
<tr id="5915A45F245F6BB672925A2D3291555F">
<td id="D67702BE89216EBAE1310D821A97E1DA" colspan="1" rowspan="1">Antenna: length / width</td>
<td id="3CE736AA567179D5C17A4F0E7E8CE243" colspan="1" rowspan="1">3.2</td>
<td id="3DE601D4012511AB01ABA443CA47636C" colspan="1" rowspan="1">3.6</td>
<td id="8B82BA578BF5AE1212B5189803DB5A9D" colspan="1" rowspan="1">?</td>
<td id="74BEC00BF58E72A282756ECB60DA1060" colspan="1" rowspan="1"></td>
<td id="5C24AFD43970913AA44D72CCBF3F4D10" colspan="1" rowspan="1">x</td>
<td id="3B9CAEEB04079B3B1B0C8A2649ED0D34" colspan="1" rowspan="1">?</td>
<td id="18D7C9A87E9C345030BD5DDDDF75235F" colspan="1" rowspan="1"></td>
</tr>
<tr id="67135F4ACEF6F1FA72E120C226F6F2BD">
<td id="45D157579927373372D22C48D2FCF525" colspan="1" rowspan="1">First maxillae: coxosternal lappet</td>
<td id="3B4C80E8D88ECC7A85010DAC5D8D14AC" colspan="1" rowspan="1">absent</td>
<td id="E87870DDA524F4B06DB4A5179BD51FB4" colspan="1" rowspan="1">small</td>
<td id="CE934709FB8E67467A2D5A2C1B2DD4EB" colspan="1" rowspan="1">?</td>
<td id="E68521D3D057AF9078C9C802C56923C8" colspan="1" rowspan="1"></td>
<td id="203F2B04E9373F2ED5C208955945022D" colspan="1" rowspan="1">x</td>
<td id="129DDA8053C91FBBF9ACC5A7ABB8B7AF" colspan="1" rowspan="1"></td>
<td id="E5FCE8CF8B56DB7FEC714B50A2A2E66F" colspan="1" rowspan="1">?</td>
</tr>
<tr id="7B7850FBCD9E939F72FC55A47576B63D">
<td id="1887B13249278C08496D4A98654D12F4" colspan="1" rowspan="1">First maxillae: telopodital lappet</td>
<td id="81D561D77FF92B5B1FD043298D547647" colspan="1" rowspan="1">large</td>
<td id="1641F1EE0B9E536E020D7BCAA4B9D134" colspan="1" rowspan="1">small</td>
<td id="C3C1E3A897FAB452A9461E7837A94308" colspan="1" rowspan="1">?</td>
<td id="9C38EA7AABF3BCE7344B00A164245D78" colspan="1" rowspan="1"></td>
<td id="C2E1E41DB3AFFBFAFC22D208E38B239F" colspan="1" rowspan="1">x</td>
<td id="CA65EE11FF8D23B01CF3D173ED706E95" colspan="1" rowspan="1"></td>
<td id="8A39222775AF6B4247B8D9F35F824680" colspan="1" rowspan="1">?</td>
</tr>
<tr id="FE22F6F0F0B76C37C39EF6692DF922ED">
<td id="B2F1DBC0C671D674DE37783D6D23F31B" colspan="1" rowspan="1">Second maxillae: pretarsus</td>
<td id="582E1D75372F315B1A903261FC4EDC9C" colspan="1" rowspan="1">slender, hooked at the tip</td>
<td id="3595A58591D9AC7DB79922ECBF9A8F7B" colspan="1" rowspan="1">stout, not hooked</td>
<td id="24DD182588B52B3B6A124FA65BBDD73A" colspan="1" rowspan="1"></td>
<td id="E2E973E271711CDC62462F8ACB7D8DF6" colspan="1" rowspan="1"></td>
<td id="BB8C20848CD62B49CE1824DFE0696598" colspan="1" rowspan="1"></td>
<td id="F991B676CA556B7749B1FD335EAF05BA" colspan="1" rowspan="1"></td>
<td id="D170A139CE1EEDC9C3DCA2A20AC613A8" colspan="1" rowspan="1">x</td>
</tr>
<tr id="D46CA1EAC94A391E2521476CF8DD649F">
<td id="ABD18893DFBFADF79AF8E4E564190F27" colspan="1" rowspan="1">Forcipular metatergite: width / length</td>
<td id="D812ED30BF4D9ED3831173E61180C562" colspan="1" rowspan="1">2.3</td>
<td id="4954D6B00EDC5BD8124792B62632E875" colspan="1" rowspan="1">1.8</td>
<td id="335C43D94B2F40C61CF7E147A55A53BC" colspan="1" rowspan="1">?</td>
<td id="4871DEFD42ED50997731C2E07239D83D" colspan="1" rowspan="1"></td>
<td id="79DB7490B66007C4478ACF28CBFAAA9D" colspan="1" rowspan="1">x</td>
<td id="6B53CD01639AE94F4A38A6C75FD53B86" colspan="1" rowspan="1">?</td>
<td id="97281D545340F64BDFFBFF426A507466" colspan="1" rowspan="1">?</td>
</tr>
<tr id="3706FE72BF4CA2ABA21496EB4A04E242">
<td id="CFA2AE2486327621946E84A559DF0189" colspan="1" rowspan="1">Forcipular trochanteroprefemur: length / width</td>
<td id="C74D43FDE73AE60AF547DFF0F0770693" colspan="1" rowspan="1">1.1</td>
<td id="C79C00EDF8E4C3DD513FD176DCC47C02" colspan="1" rowspan="1">1.3</td>
<td id="641C7272F5AC2866E5865D64304D6298" colspan="1" rowspan="1"></td>
<td id="BADB9142172E763C54C7BCE547354D16" colspan="1" rowspan="1"></td>
<td id="CF8D15A67946CA500990A1435974C61B" colspan="1" rowspan="1"></td>
<td id="F5D4FC65D4ECFB7FA044FBF0CBA28FEC" colspan="1" rowspan="1"></td>
<td id="0C8F388517949950ED2FFE2D89A67E66" colspan="1" rowspan="1">x</td>
</tr>
<tr id="E88896D308561DCE96B4CD3CF0ACC90F">
<td id="56D4B054E2F304B899E4F0CD0EDA9FD6" colspan="1" rowspan="1">Forcipular tarsungulum: length / width</td>
<td id="7303534B9FB4BB979DDE0CBCDE233ACB" colspan="1" rowspan="1">2.2</td>
<td id="8D7926F7E8FDCDF1A4511F87F3E1FE39" colspan="1" rowspan="1">1.9</td>
<td id="60FBEEDCEC733D27D8C4C55BC5AAB4F0" colspan="1" rowspan="1"></td>
<td id="65348085B3CFD9C99F03F6C4491B1F0C" colspan="1" rowspan="1"></td>
<td id="7DE80D7B1129E8F9A1F2BBF3854BFD49" colspan="1" rowspan="1"></td>
<td id="E8F948023CD417EE094390D8E3B2DC47" colspan="1" rowspan="1"></td>
<td id="5D8AA64CF7D420C1163B753BB7F38683" colspan="1" rowspan="1">x</td>
</tr>
<tr id="7C41CA3A9CBF8184B96B82722713E69D">
<td id="7C38019D071F563D165FBD264B102D67" colspan="1" rowspan="1">Forcipule / coxosternite length</td>
<td id="A1FB0B74392AABFEBBCBC3364636F4AE" colspan="1" rowspan="1">0.91.0</td>
<td id="3E86086B0C732F402D32036DB7D92D0A" colspan="1" rowspan="1">0.70.8</td>
<td id="3FEEF6D3500722DCE1D01115FE717186" colspan="1" rowspan="1"></td>
<td id="A04CEC85E3D8BB72FC3F834256863C1B" colspan="1" rowspan="1"></td>
<td id="CA11768DA250B37712B20B18CD642EB1" colspan="1" rowspan="1"></td>
<td id="83C8EF2CA2F8F5BE4CA20B218B34BFE6" colspan="1" rowspan="1"></td>
<td id="17720F2C5B2BEA1B51930DDCB24E9E59" colspan="1" rowspan="1">x</td>
</tr>
<tr id="71C7D5CE5F6CB1835E237973FDD89075">
<td id="050D8D80B34690D9608F659A4B6EC1AA" colspan="1" rowspan="1">Forcipular tarsungulum: shape</td>
<td id="AE5385688D1D4D0FC6DCA7C3BEA02A49" colspan="1" rowspan="1">narrowing gradually</td>
<td id="F2C621EC51AD006C4CCAA15564759C20" colspan="1" rowspan="1">narrowing abruptly</td>
<td id="BD66BDEA46B715AF7AC29FB9FC280B79" colspan="1" rowspan="1"></td>
<td id="FB7E750166A39E9D6FA9304281BBADE4" colspan="1" rowspan="1"></td>
<td id="91FE1839F9772D717F56025D44FF1083" colspan="1" rowspan="1"></td>
<td id="C467EEB19E6D44E17EC667AAA68ED371" colspan="1" rowspan="1"></td>
<td id="296C2D8595D485775F2642BC98534DC5" colspan="1" rowspan="1">x</td>
</tr>
<tr id="A32028FEDE24BB2FE3A039A8EC66D211">
<td id="5A86D4DB3CD7506B0B0CE59EE6A55B97" colspan="1" rowspan="1">Anterior part of trunk: metasternite: carpophagus pit</td>
<td id="536E4A3135E110BF99F9FE3254EDFCA5" colspan="1" rowspan="1">distinct, fairly deep</td>
<td id="0C2C44BDE85ACB2E46A69E8BB54EA1E1" colspan="1" rowspan="1">faint, shallow</td>
<td id="4EC24183A66BD51445A8AA4E50F60477" colspan="1" rowspan="1">?</td>
<td id="CAD7EBC1154BBD95DD9BCD00429F1715" colspan="1" rowspan="1"></td>
<td id="C5D39B865FC2DBA852574AEE78F54535" colspan="1" rowspan="1">x</td>
<td id="0C2607588B623F927DCD7ED4CFBBF502" colspan="1" rowspan="1"></td>
<td id="DBFEE31D0201A7F52A1AB79A9087BDD6" colspan="1" rowspan="1">?</td>
</tr>
<tr id="A6C4F0F81DA3924FF2C60DFD68672206">
<td id="5DEC6F94707E452D1883AE9B0F987415" colspan="1" rowspan="1">Legs in female: number</td>
<td id="C5381548AC7D45576DF487BB054B5274" colspan="1" rowspan="1">99</td>
<td id="86673FA1AD6088BA02B18DBC729F09D9" colspan="1" rowspan="1">107</td>
<td id="8F8A42B20C372EC2762E3E5FF76AB984" colspan="1" rowspan="1"></td>
<td id="421C0CF5B241796EE1C969B40689EC4C" colspan="1" rowspan="1"></td>
<td id="4E3A6339109C6CB55E28EC60AD18F51C" colspan="1" rowspan="1">x</td>
<td id="BC2CE7B82B5D71ADC202C9E757B41868" colspan="1" rowspan="1"></td>
<td id="D07B3BFB9EEEE50A84E8D382516455C7" colspan="1" rowspan="1">?</td>
</tr>
<tr id="4F1F5A0DFBB59D6F4A8096A96F9CE449">
<td id="D292558EB95EBE3BC973A3982D534B76" colspan="1" rowspan="1">Coxopleuron: pores: number</td>
<td id="A0330561773EA2D4FA64523D2EF95A9A" colspan="1" rowspan="1">5 or 6</td>
<td id="589E9E237AD84BE8570ED6A34498E9C3" colspan="1" rowspan="1">915</td>
<td id="01C503BB964A47BEDC36CFCDB92B3972" colspan="1" rowspan="1">x</td>
<td id="6B284E916B8C1F78BBB262E5B91DBF97" colspan="1" rowspan="1"></td>
<td id="C470130B70FC15BE2B3FFA9BC169D379" colspan="1" rowspan="1"></td>
<td id="77E52C40CFBF35836E35750476F86DBB" colspan="1" rowspan="1"></td>
<td id="FB8BE00AB07A959DC51B560F87F656D6" colspan="1" rowspan="1">?</td>
</tr>
<tr id="B58582C202AD7553C5E1DE44B18D2C15">
<td id="4828A58812878469530AD4DA1C070E8A" colspan="1" rowspan="1">Ultimate / penultimate telopodite length</td>
<td id="3E1940AD1F753672D65F3C0EE7C64CDE" colspan="1" rowspan="1">1.7</td>
<td id="7432747B713C2F30D2B4DF255CEE256A" colspan="1" rowspan="1">2.0</td>
<td id="F829B1E09A1496BAF48F02606BF3597F" colspan="1" rowspan="1">x</td>
<td id="D1F0A9FDE4C54B573B74EE9A74CB6B91" colspan="1" rowspan="1"></td>
<td id="F243E0A2D687748303724E9465E9DD69" colspan="1" rowspan="1"></td>
<td id="0027F53A692E805C20C70C410D7407ED" colspan="1" rowspan="1">?</td>
<td id="60978C604E385886AD3724C5FE0C7078" colspan="1" rowspan="1">?</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4BAG" type="description">
<paragraph id="1D9D66DA9F95F85BD7E71D8998F2AEE7">
<heading id="70D69B5F3B4AE4EC0708FB475CE95FED" reason="title">
Description of the
<typeStatus id="1839BAC597230C57CFDB29F4293B0636" type="holotype">holotype</typeStatus>
</heading>
</paragraph>
<paragraph id="9B5D9655BDA1D0DCBCA84FAE69E345B5">
(Figs
<figureCitation id="4536298F4DD01F3CB9E0C2E8773842BC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2</figureCitation>
,
<figureCitation id="85AB4372F9B692A97A034F76C8F15DA6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3</figureCitation>
).
<emphasis id="578DC17A6217E185B7ACAA6F355D25E5" bold="true">
<emphasis id="D0882ACF8B92AF3B9228BFBFDBB7E6EE" bold="true" italics="true">General features</emphasis>
.
</emphasis>
Body remarkably slender,
<quantity id="46F2EA31595D81668A3639A6957045C7" metricMagnitude="-2" metricUnit="m" metricValue="2.45" unit="mm" value="24.5">24.5 mm</quantity>
long, uniformly ~
<quantity id="88ECEF214F488D76D7A4DB350A0FD6DE" metricMagnitude="-4" metricUnit="m" metricValue="3.0" unit="mm" value="0.3">0.3 mm</quantity>
wide for most part of the trunk, only very slightly narrowing anteriorly along ~ 20 most anterior leg-bearing segments and backwards along approximately the five most posterior leg-bearing segments. Legs relatively short. Colour almost uniformly pale yellow, only the forcipules and the head slightly darker, both pale orange.
</paragraph>
<paragraph id="1771A2378EEBBA919D6295BA5C4D28AD">
<emphasis id="157A7FF36A53CC0464822363727B6042" bold="true">
<emphasis id="896892B3D44C413CF8AC4E5F16FB9BAD" bold="true" italics="true">Cephalic capsule and antennae</emphasis>
.
</emphasis>
Head
<quantity id="D463D903A034AC111D0BB452B7D8AEF3" metricMagnitude="-4" metricUnit="m" metricValue="4.0" unit="mm" value="0.4">0.4 mm</quantity>
long, sub-rectangular, 1.15 × longer than wide (Fig.
<figureCitation id="A1BB1CC6584285683C2D7FB1746ECA99" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 A</figureCitation>
); the anterior margin slightly angulated with a small medial notch. Transverse suture absent. Approximately 40 setae on the head, the majority being short except those of the lateral margins, longer, reaching up to ~ 22 μm. Clypeus uniformly areolate, without clypeal areas and without plagulae, with six setae, arranged in a longitudinal series of three pairs on the anterior median part of the clypeus. Pleurites uniformly areolate, with some setae. Labrum composed of an intermediate part narrow and almost negligible, having one tubercle (Fig.
<figureCitation id="001FAC0532474EBA78510F0497D990CB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 A</figureCitation>
); side pieces not clearly distinct from the intermediate part of labrum, each with four bristles. Antennae with 14 articles,
<quantity id="40D8CB50980E583D7E3995B6B471CBD7" metricMagnitude="-3" metricUnit="m" metricValue="1.3" unit="mm" value="1.3">1.3 mm</quantity>
long, 3.2 × longer than the head (Fig.
<figureCitation id="0019D1850AA137486665460D2F7A92A5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 B</figureCitation>
). article I ~ 0.7 × longer than wide, articles II VII up to 1.7 × longer than wide, article VIII ~ 1.2 × longer than wide, articles IX XIII approximately as long as wide; setae gradually denser and shorter from basal to distal articles, both ventrally and dorsally. Article XIV 2.0 × longer than wide, with numerous setae and with club-like sensilla grouped on the distal parts of both internal and external sides (Fig.
<figureCitation id="4F07EDEC9F503B1A96E05E5AEC0FE072" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 C</figureCitation>
).
</paragraph>
<paragraph id="A9F4BADAE763EDFA68D12505699FC8C9">
<emphasis id="51FDB0A06D7D58A148C5868600117967" bold="true">
<emphasis id="E073E395E723626346FE6B09D3DD7FD5" bold="true" italics="true">Mandibles and maxillae</emphasis>
.
</emphasis>
A single pectinate lamella on each mandible. Coxosternite of the first maxillae entire, without mid-longitudinal sulcus. Coxal projection sub-triangular, longer than wide, bearing one or two basal setae and some more distal spine-like sensilla. Telopodite comprises two articles, the basal one without setae, the distal one with three setae and five or six spine-like sensilla. No coxosternal lappets; telopodital lappets present and pointed (Fig.
<figureCitation id="F3EF2B07EB5FC37BF76A922B5045B535" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 B</figureCitation>
). Coxosternite of the second maxillae entire, the intermediate part uniformly sclerotised as the remaining parts, its anterior margin concave; no sclerotised ridges. Telopodite of the second maxillae composed of three articles: article 1 with two or three very short external setae, article 2 with one or two very short external setae, article 3 with five or six long distal setae; a simple long apical claw, subconical but hook-shaped at its extremity, with a dorsal bulge at its mid length (Fig.
<figureCitation id="59EB2CA2C5DE20F63B1B22FA320ED1FD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 D</figureCitation>
).
</paragraph>
<paragraph id="65D64DA3901368E3D37C319203FAFFEC">
<emphasis id="2CA7155E5B55155C21A6CAEF0C1A6B21" bold="true">
<emphasis id="D6F3BBB4215DA7F8F151BCEF0161C78F" bold="true" italics="true">Forcipular segment</emphasis>
.
</emphasis>
Tergite trapezoid, the lateral margins distinctly converging anteriorly, ~ 2.3 × wider than long, posteriorly almost as wide as the subsequent metatergite (Fig.
<figureCitation id="43DD6366F7EE29F1581FEACA71E54DF6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 A</figureCitation>
). Exposed part of the coxosternite ~ 1.1 × wider than long; anterior margin with a medial shallow concavity, without denticles. Coxopleural sutures complete, entirely ventral or almost, only very slightly sinuous, and strongly converging posteriorly. Chitin-lines well distinct, reaching the condyles, moderately curved and converging posteriorly (Figs
<figureCitation id="B9E842D6FE4705D1CF27E16BE9847835" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 C</figureCitation>
,
<figureCitation id="872920D5C7B8A10B3E9BD74A79738FEA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 E</figureCitation>
). Trochanteroprefemur ~ 1.1 × as wide as long, the external side ~ 2.2 × longer than the internal side, without denticles and with some setae (Figs
<figureCitation id="A00F7841CB97814406F45195BDB087D1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 C</figureCitation>
,
<figureCitation id="5009CCFCB4ACB2F83BA7F80DEE688E0B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 F</figureCitation>
). Forcipular intermediate articles distinct, without denticles, each with some long setae. Tarsungulum curved, gradually narrowing, ~ 2.2 × as long as wide and with a basal sub-conic tubercle; slightly crenulated in its concavity, with 36 shallow projections, less pronounced on the left than on the right (Figs
<figureCitation id="8408F6C17502D2DAD0B6F40E54E310A0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 C</figureCitation>
,
<figureCitation id="928DF0C4080B7E3A89AB9272C8BB7A14" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 F</figureCitation>
).
</paragraph>
<paragraph id="E47B2537BABFB991C8CF6781F424CC29">
<emphasis id="7DBF88D28F88715CE5488C14AFF42E85" bold="true">
<emphasis id="F794033F7A579AD7A398F3F019EF21AC" bold="true" italics="true">Leg-bearing segments</emphasis>
.
</emphasis>
99
<abbrev id="ABBRID0E6EAG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
. No paratergites. Metatergite 1 wider than the subsequent one, lateral margins converging posteriorly, without pretergite. Metatergites with two paramedian sulci. Metasternites slightly longer than wide (length / width ratio ~ 1.1 at ~ 20 % of the
<abbrev id="ABBRID0EDFAG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
), uniformly areolate; setae very sparse. Metasternites of
<abbrev id="ABBRID0EHFAG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
1020 with a carpophagus pit on the anterior margin, fairly deep on
<abbrev id="ABBRID0EUFAG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
1419, ~ 0.6 × as wide as the margin of metasternite (Figs
<figureCitation id="1558FD445EEE94C91397CFC9FF6333ED" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 D</figureCitation>
,
<figureCitation id="761548C25B40C4512483142587BDA908" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 G</figureCitation>
). Glandular pores on the metasternites from the 2
<superScript id="DEF5F72EDEF0414CFED5D31EC000BC67">nd</superScript>
to the penultimate, up to 3035 pores and loosely arranged in a sub-ovoid / sub-triangular pore-field on the posterior half of each metasternite in the anterior part of the trunk (Fig.
<figureCitation id="0A5D1466086A1E8AB7E44588EFEA0CB8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 D</figureCitation>
); the pore-fields being much less visible in the posterior half of the trunk, loosely arranged in a transverse band of pores in the ~ 510 penultimate
<abbrev id="ABBRID0EGGAG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
. Length / width ratio of leg tarsus ~ 2.3 at ~ 20 % of the
<abbrev id="ABBRID0EKGAG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
. Leg claws simple.
</paragraph>
<paragraph id="E8A41264DA8F7A6E9788801FCFECBC46">
<emphasis id="6C31D4C23181006A2C5607253409F89C" bold="true">
<emphasis id="C249C9139BB30630CC6C7AEA57FD3D6A" bold="true" italics="true">Ultimate leg-bearing segment</emphasis>
.
</emphasis>
Setae uniformly sparse. Pleuropretergite entire, lacking sutures or sulci. Metatergite sub-trapezoid, ~ 1.5 × wider than long, lateral margins very slightly convex and distinctly converging posteriorly, posterior margin convex. Presternite not medially constricted. Metasternite trapezoid, ~ 1.3 × wider than long, anteriorly 1.4 × wider than posteriorly (Fig.
<figureCitation id="74E870F2ACA5E1B78D88E9F1CBBED88E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 E</figureCitation>
); lateral margins slightly convex, converging backwards; posterior margin straight. Coxal organs of each coxopleuron opening through five or six independent pores: two dorsal pores, one covered by the pleuropretergite and one exposed on the dorso-lateral side (Fig.
<figureCitation id="E919E3A4B60F7D763D54EF146AA9BCD1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 F</figureCitation>
); three or four ventral pores including two or three more or less aligned under the edge of the corresponding metasternite and a single pore isolated on the postero-central part of the ventral side (Figs
<figureCitation id="E10C28D09D9F1038DA0BC42BFA93C8FC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 E</figureCitation>
,
<figureCitation id="FC9B5530A951C235F02B6B745130BE0D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 H</figureCitation>
). Telopodite
<quantity id="C1C7ECCB9321F64B9C00457EC31DD91E" metricMagnitude="-4" metricUnit="m" metricValue="4.3" unit="mm" value="0.43">0.43 mm</quantity>
long, 7.8 × as long as wide, ~ 1.7 × as long as the penultimate telopodite; six articles, all covered dorsally with sparse long setae, ventrally with dense shorter setae; a fairly well-developed apical claw (Fig.
<figureCitation id="250443FDB9F1BF90E10818042FA093FB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 H</figureCitation>
).
</paragraph>
<paragraph id="78A1BCE8418F4BBE0DD6DBC4C3ECCAAE">
<emphasis id="EE050F304535EDE13A4F95F3EAE5F34D" bold="true">
<emphasis id="5B0CE72FE4FF74FAFD2A2E505E6AE8E1" bold="true" italics="true">Postpedal segments</emphasis>
.
</emphasis>
Gonopods in the shape of a short, slightly bilobate lamina. A pair of anal pores.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQHAG" type="distribution">
<paragraph id="188AE1A79D7DA8D20C8E48D4F8E71BCF">
<heading id="81700948E0698A6291A561BB83EF75A2" reason="title">Distribution and ecology.</heading>
</paragraph>
<paragraph id="91586E96B262F35F42750743408D167C">
The new species is only known from the type locality (see above; Fig.
<figureCitation id="33CA2B43BA24AEF3E3E2B659A7BB3320" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of all known records of Endogeophilus." figureDoi="10.3897/zookeys.1213.133171.figure1" httpUri="https://binary.pensoft.net/fig/1142652">1</figureCitation>
). The single specimen was collected in a shaded maquis of old
<taxonomicName id="DE6C120A438594D59F5876E62674ED0F" class="Magnoliopsida" family="Ericaceae" genus="Arbutus" kingdom="Plantae" order="Ericales" phylum="Tracheophyta" rank="species" species="unedo">
<emphasis id="5E598C011CFC5890496E999CCA4DDEC6" italics="true">Arbutus unedo</emphasis>
</taxonomicName>
L., 1753 with also some large
<taxonomicName id="70630B9D6F51C3299E6021DEC2D3184F" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="species" species="suber">
<emphasis id="093E20B7AC1662F18A58CD0CA14CAFF9" italics="true">Quercus suber</emphasis>
</taxonomicName>
L., 1753 (Fig.
<figureCitation id="D85EE3609EE27E5563AEA00B57E1FFF5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitat of Endogeophilus alberti sp. nov., near Malatra, Cavalaire-sur-Mer, France (Fig. 1)." figureDoi="10.3897/zookeys.1213.133171.figure5" httpUri="https://binary.pensoft.net/fig/1142656">5</figureCitation>
). It was found after sieving with a Winkler apparatus, at a depth of
<quantity id="B3A552CBF8B0791CBC2550B0591E5F32" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" unit="cm" value="15.0" valueMax="20.0" valueMin="10.0">1020 cm</quantity>
in the soil.
</paragraph>
<caption id="1FDA533489301D1DC109C396485BF44B" ID-DOI="10.3897/zookeys.1213.133171.figure5" ID-arpha="4F206106-0370-5E28-B022-F1A56F60289A" httpUri="https://binary.pensoft.net/fig/1142656" startId="F5">
<paragraph id="288C65604ABB281595A89617EBE2BFD1">
<label id="8A97193596F0736E66E1CCE2F8C1C883">Figure 5.</label>
</paragraph>
<paragraph id="32DCC84B6AC43ADA2CB11907DBBE9362">
Habitat of
<taxonomicName id="DDDFBA62DDB8F9207E4C610CA5D1BD5A" authorityName="Iorio &amp; Bonato" authorityYear="2024" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
<emphasis id="30F097259E0A245ADF21CDF5572B38F1" italics="true">Endogeophilus alberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60D018AAB30975695C84976A98E77137" rank="species">sp. nov.</taxonomicNameLabel>
, near Malatra, Cavalaire-sur-Mer, France (Fig.
<figureCitation id="92C4D120E585F41CF7763D314B86F9BF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of all known records of Endogeophilus." figureDoi="10.3897/zookeys.1213.133171.figure1" httpUri="https://binary.pensoft.net/fig/1142652">1</figureCitation>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EQJAG" type="etymology">
<paragraph id="DA68D4513FE7740285032A98A5ACA6C1">
<heading id="7932838BC6A4B54664DCD1858802D4C6" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="3D05D441D2414994C1BAD5C04E699607">
This species is dedicated to Prince Albert II de
<collectingCountry id="2A0F36093FFDE2436F382CC4DD8FF4C0" name="Monaco">Monaco</collectingCountry>
, because the Foundation Prince Albert II de
<collectingCountry id="317C4F4EF606F556C2A3A5937693104A" name="Monaco">Monaco</collectingCountry>
has supported the field work of this study. The epithet
<taxonomicName id="F8486AD3DDF77C4244C3C4AD2FC638E2" authorityName="Iorio &amp; Bonato" authorityYear="2024" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="alberti">
<emphasis id="3AF1D8DD276F5E401A51158A3324F54E" italics="true">alberti</emphasis>
</taxonomicName>
is intended as a noun in the genitive case.
</paragraph>
<paragraph id="439BB53A96807FC29BF5DD85A2E2EE20">
<title id="CD3B4D288C1ACA5D95874A082FEED921">
Identification key to the European genera of
<taxonomicName id="E46F0AF5D59E7D6C75E84D7C986FCB9D" class="Chilopoda" family="Geophilidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Geophilidae</taxonomicName>
s. l.
</title>
</paragraph>
<paragraph id="BFD85FDAC9B2A2AEA6C8FD23CEFEEDB3">
Within Europe, the
<taxonomicName id="DEF9B346F6DF37B2E5CA88C11BE05E94" class="Chilopoda" family="Geophilidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Geophilidae</taxonomicName>
s. l. can be distinguished from all other
<taxonomicName id="1AA4537C70EE20F47F6AF42D72A564ED" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" phylum="Arthropoda" rank="class">Chilopoda</taxonomicName>
by means of the combination of the following two characters:&gt; 25 pairs of legs and second maxillary pretarsi in shape of either subconical, non-spatulate, pointed claw or a tubercle with only one or two tiny spines. A total of 20 genera of
<taxonomicName id="BD6575717237472E6A828038A9BFE7B9" class="Chilopoda" family="Geophilidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Geophilidae</taxonomicName>
s. l. are recorded in Europe (Table
<tableCitation id="F6691C137242F8C1E886679B406B0A37" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Genera of Geophilidae s. l. and approximate number of species recognised in Europe. Only genera recorded within the geographic boundaries adopted by Fauna Europaea are considered (see Materials and methods). Occurrences are also indicated for the biogeographical subregions of SW Europe and the two countries where Endogeophilus occurs." httpUri="http://table.plazi.org/id/14CD0D2CBE447841518462A781172BB1" tableUuid="14CD0D2CBE447841518462A781172BB1">2</tableCitation>
).
</paragraph>
<caption id="14CD0D2CBE447841518462A781172BB1" ID-Table-UUID="14CD0D2CBE447841518462A781172BB1" httpUri="http://table.plazi.org/id/14CD0D2CBE447841518462A781172BB1" startId="T2" targetIsTable="true">
<paragraph id="2B37B0F1B8780D7FB12C1377369E1245">
<label id="9CF775AF0ACD2833F3CEDD52338E2A9D">Table 2.</label>
</paragraph>
<paragraph id="7E39C29C4EA69B4C311A0979A2DD904D">
Genera of
<taxonomicName id="B531D611AD71340F9C2A0CC6CE3DF7C7" class="Chilopoda" family="Geophilidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Geophilidae</taxonomicName>
s. l. and approximate number of species recognised in Europe. Only genera recorded within the geographic boundaries adopted by Fauna Europaea are considered (see Materials and methods). Occurrences are also indicated for the biogeographical subregions of SW Europe and the two countries where
<taxonomicName id="13178045664745371E5AF4259D28FDAE" authorityName="Bonato, Zapparoli, Drago &amp; Minelli" authorityYear="2016" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="4D78B538B37C5BD3DD9DF89FC1C413FF" italics="true">Endogeophilus</emphasis>
</taxonomicName>
occurs.
</paragraph>
</caption>
<paragraph id="6FC89901B55B69282CE3551FF58EC4C0">
<table id="TID0EWDBI" rules="all">
<tbody id="9C28495AF78303BBED174F85CF2F4034">
<tr id="0078676AD29BB07E466D3FD992EA90FF">
<th id="B8871BBD12E2B21CA28CE80EE93A2ED9" colspan="1" rowspan="2">Genus</th>
<th id="0EB9BF85157F160FE8C3881CB5DC6AB0" colspan="1" rowspan="2">Approximate number of species in Europe</th>
<th id="7D7ED1E6B0818EE2404D4081C482C48E" colspan="3" rowspan="1">Occurrence</th>
</tr>
<tr id="20BF01FA5BDE6E5FA5A3E8EBDEDC194F">
<th id="3626365D52E1A5C91FAE651877687440" colspan="1" rowspan="1">South-western Europe</th>
<th id="9D2E2662393714BD3906609EBAC66719" colspan="1" rowspan="1">France</th>
<th id="F4DFE40FE1326FBDA024AA41569F65B9" colspan="1" rowspan="1">Italy</th>
</tr>
<tr id="D77BF2A54B16A0B2A5572F640AB20546">
<td id="460F66C2710A31DFC7456EF9576E12E7" colspan="1" rowspan="1">
<taxonomicName id="545AE1AB462C25082AE41659AE667108" authority="Minelli, 1982" authorityName="Minelli" authorityYear="1982" class="Chilopoda" family="Geophilidae" genus="Acanthogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="C51F04FDCE865E13E3E65109AF70C021" italics="true">Acanthogeophilus</emphasis>
<bibRefCitation id="A805B21B8C5F00EADDE8F3701F8B0D22" author="Minelli A" journalOrPublisher="Bollettino del Museo Civico di Storia Naturale di Verona" pagination="235-240" refId="B40" refString="Minelli A (1982) Un nuovo chilopodo italiano Acanthogeophilus dentifer, n. g. n. sp. (Geophilomorpha Geophilidae). Bollettino del Museo Civico di Storia Naturale di Verona 8: 235 - 240." title="Un nuovo chilopodo italiano Acanthogeophilus dentifer, n. g. n. sp. (Geophilomorpha Geophilidae)." volume="8" year="1982">Minelli, 1982</bibRefCitation>
</taxonomicName>
</td>
<td id="F35C3A6D6BF412302EBA07733828C389" colspan="1" rowspan="1">1</td>
<td id="4190244203A652B7F47F92F015ACC41C" colspan="1" rowspan="1">x</td>
<td id="8A2E83D1952DDE996FEA17F80F6ED0A1" colspan="1" rowspan="1"></td>
<td id="DABE834C968C77D3765BD1931B7CEF29" colspan="1" rowspan="1">x</td>
</tr>
<tr id="89AA571E3186BBE87C379A8692E0B3A4">
<td id="B15F658CE4DE4EFA60B7DDBB23E3E5FD" colspan="1" rowspan="1">
<taxonomicName id="5A0BED53FFB93EEEA944A45E3AEC405B" authority="Brolemann, 1925" authorityName="Brolemann" authorityYear="1925" class="Chilopoda" family="Geophilidae" genus="Algerophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="CBBB88280BCAD222A4D23582533D7446" italics="true">Algerophilus</emphasis>
Brolemann, 1925
</taxonomicName>
</td>
<td id="62951FFF66ABD5D3602CC3E21CC4CB8D" colspan="1" rowspan="1">1</td>
<td id="4EC861A183DDF25478B1B9462ECA9A7D" colspan="1" rowspan="1">x</td>
<td id="F49DD50184E53B51E9CCE7B3B4CAFFD4" colspan="1" rowspan="1"></td>
<td id="2DF0E2651052357D50137DA3C51717BF" colspan="1" rowspan="1"></td>
</tr>
<tr id="7AED1E87B6BA1B7CAD17E70662BF49BF">
<td id="26C486F529B7A2CA551A9917B21B44DD" colspan="1" rowspan="1">
<taxonomicName id="79ECC710311C78B5FB5BA066DF31CCFF" authority="Attems, 1909" authorityName="Attems" authorityYear="1909" class="Chilopoda" family="Geophilidae" genus="Arctogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="7B002A48D77476C679FBA802398A954F" italics="true">Arctogeophilus</emphasis>
Attems, 1909
</taxonomicName>
</td>
<td id="03FCBBE6E7B7767B8D18D1C9A8C7ED58" colspan="1" rowspan="1">3</td>
<td id="0F43D6F051CD7E53B3BA0BE1D333AD15" colspan="1" rowspan="1"></td>
<td id="988E4408528DFE02E865E4CEAB30F51C" colspan="1" rowspan="1">x</td>
<td id="3FD68C8D8CF89D6180364711D111A10E" colspan="1" rowspan="1"></td>
</tr>
<tr id="89FA89289A5CAD4AF728262C40AB172F">
<td id="781FFA5D4B7BBABDFD7C0D0FC29DC7D5" colspan="1" rowspan="1">
<taxonomicName id="039C65CA76FE943949A4DF57AAB909BA" authority="Chamberlin, 1912" authorityName="Chamberlin" authorityYear="1912" class="Chilopoda" family="Geophilidae" genus="Arenophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="C0621EB0689DFD57E92D7AD052333B04" italics="true">Arenophilus</emphasis>
Chamberlin, 1912
</taxonomicName>
</td>
<td id="38DD64BA6C1B753CCE5C026491829831" colspan="1" rowspan="1">1</td>
<td id="19BE99DA172E311CB106FA9910335B89" colspan="1" rowspan="1">x</td>
<td id="3A094DD692225AE91CBFDB26CDAF9256" colspan="1" rowspan="1">x</td>
<td id="3B1CA27DED914CB0756296CC76053D40" colspan="1" rowspan="1"></td>
</tr>
<tr id="4ED2831DEF4DFFDCA0771D698C7AFCAC">
<td id="6CB60A0DBFF128D215A597D0A245B5F1" colspan="1" rowspan="1">
<taxonomicName id="5FE529622ACAA64B59691CC685EFBB40" authority="Verhoeff, 1941" authorityName="Verhoeff" authorityYear="1941" class="Chilopoda" family="Geophilidae" genus="Bebekium" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="FBE7979CD454D3C8CA702DB7830D5C99" italics="true">Bebekium</emphasis>
<bibRefCitation id="54A75967AC921C26E58AA8E7452A26E8" author="Verhoeff KW" journalOrPublisher="Zoologischer Anzeiger" pagination="35-41" refId="B45" refString="Verhoeff KW (1941) Untersuchungen uber die Fauna des Hologaeum: Chilopoden, Diplopoden und Land-Isopoden. Zoologischer Anzeiger 135: 35 - 41." title="Untersuchungen über die Fauna des Hologaeum: Chilopoden, Diplopoden und Land-Isopoden." volume="135" year="1941">Verhoeff, 1941</bibRefCitation>
</taxonomicName>
</td>
<td id="A0411E4AAB0B4A3DD5A0E75D8CFD2CA0" colspan="1" rowspan="1">1</td>
<td id="78D07790827C12F1A8472191B73F0648" colspan="1" rowspan="1"></td>
<td id="C7596A0516A4A0FF0FD2001DFDC06E57" colspan="1" rowspan="1"></td>
<td id="D3914CFE378F9F59B8B897DAB6CC2000" colspan="1" rowspan="1"></td>
</tr>
<tr id="8E1EEE5CD8AD3885AA4C7153BD3809B2">
<td id="CCE4A950666B4242312610763A3E48B4" colspan="1" rowspan="1">
<taxonomicName id="568194D87B857369CE8A602F7305F7E1" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="576666328111A0721DA2BC7FBD73D347" italics="true">Clinopodes</emphasis>
C. L. Koch, 1847
</taxonomicName>
</td>
<td id="AFF1BC5DAF0B81864453DC1E555536CE" colspan="1" rowspan="1">9</td>
<td id="F45D67AA3C7CE413DE6013B8FBF4519D" colspan="1" rowspan="1">x</td>
<td id="BE7B11709D0CC784A1F3075EEA07A412" colspan="1" rowspan="1">x</td>
<td id="7ED2A50A9CBAF8DF035AE953C0C1E3A4" colspan="1" rowspan="1">x</td>
</tr>
<tr id="06C96C562E51664AFDAD3C5CB373BA6C">
<td id="33F300F6EC3D83502C18A707B3C7FFF3" colspan="1" rowspan="1">
<taxonomicName id="BF5BE48234DE76FA8F2EB88431C98D09" authority="Meinert, 1870" authorityName="Meinert" authorityYear="1870" class="Chilopoda" family="Dignathodontidae" genus="Dignathodon" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="A9D29A48527A0EA0C4324A49F587603A" italics="true">Dignathodon</emphasis>
Meinert, 1870
</taxonomicName>
</td>
<td id="D874ADC0E42D5300B7485589F1F69F26" colspan="1" rowspan="1">2</td>
<td id="4565527246DD449F7A1471005A8473BF" colspan="1" rowspan="1">x</td>
<td id="880E98C605F59740731BCA62605C8F94" colspan="1" rowspan="1">x</td>
<td id="F99373F2E39C9E0AD7A9CA58308D97B0" colspan="1" rowspan="1">x</td>
</tr>
<tr id="51E01F7F7C190AD0A3A71A48F079B0B6">
<td id="71ADEFE7DABED20767649ECCFD50F417" colspan="1" rowspan="1">
<taxonomicName id="CE0644443D3D56E95193760F1C090095" authority="Bonato, Zapparoli &amp; Minelli, 2008" authorityName="Bonato, Zapparoli &amp; Minelli" authorityYear="2008" class="Chilopoda" family="Geophilidae" genus="Diphyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="2A285FAAE77C994776C384282467281E" italics="true">Diphyonyx</emphasis>
Bonato, Zapparoli &amp; Minelli, 2008
</taxonomicName>
</td>
<td id="ED727FB482D77E5296924ED5B79E110E" colspan="1" rowspan="1">2</td>
<td id="2356C4DAC39F0C79A36AC13C1A7FF3FB" colspan="1" rowspan="1"></td>
<td id="1D12E1212568EFE7A26F0298C360F37A" colspan="1" rowspan="1"></td>
<td id="9FD3794855C5E4939D3E9F17C883C2BC" colspan="1" rowspan="1"></td>
</tr>
<tr id="F69795F9C5C4A2ED62208A37D7ED14E0">
<td id="9E00BE88EECB1061EAC2AD1F6F968535" colspan="1" rowspan="1">
<taxonomicName id="CB551C2AA9308FF0E8096F3311E37CA9" authority="Bonato, Zapparoli, Drago &amp; Minelli, 2016" authorityName="Bonato, Zapparoli, Drago &amp; Minelli" authorityYear="2016" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="EF6227394335BBF27D53FE743D845455" italics="true">Endogeophilus</emphasis>
<bibRefCitation id="4B45A11395607BD06369FB7BB99CD22A" DOI="10.5852/ejt.2016.231" author="Bonato L &amp; Zapparoli M &amp; Drago L &amp; Minelli A" journalOrPublisher="European Journal of Taxonomy" pagination="1-19" refId="B14" refString="Bonato L, Zapparoli M, Drago L, Minelli A (2016) An unusually elongate endogeic centipede from Sardinia (Chilopoda: Geophilidae). European Journal of Taxonomy 231: 1 - 19. https: // doi. org / 10.5852 / ejt. 2016.231" title="An unusually elongate endogeic centipede from Sardinia (Chilopoda: Geophilidae)." volume="231" year="2016">Bonato, Zapparoli, Drago &amp; Minelli, 2016</bibRefCitation>
</taxonomicName>
</td>
<td id="D16B12B3543B2DF862F0B9183FB08392" colspan="1" rowspan="1">2</td>
<td id="A7C774912F33A436AECECC242981D96B" colspan="1" rowspan="1">x</td>
<td id="B03963F29A2003683AD0D37404C8A025" colspan="1" rowspan="1">x</td>
<td id="9961BD67EA0E7115FE5C85DA933570D6" colspan="1" rowspan="1">x</td>
</tr>
<tr id="AC6C28A184AA9D626D20C6198B31D25C">
<td id="D214E0915EF4E66F1416196B79FF5611" colspan="1" rowspan="1">
<taxonomicName id="3ECD48F1641E4F827AA0831570A5DAEA" authority="Verhoeff, 1899" authorityName="Verhoeff" authorityYear="1899" class="Chilopoda" family="Geophilidae" genus="Eurygeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="278CD3F65534B7DE97399A40C6F16168" italics="true">Eurygeophilus</emphasis>
Verhoeff, 1899
</taxonomicName>
</td>
<td id="F66C2E1A13EA7202F3162160F8881F7C" colspan="1" rowspan="1">2</td>
<td id="A55E98ACABF1A902F726FDDD1AEEF954" colspan="1" rowspan="1">x</td>
<td id="C85269D83DE695BFA81D98972CB90645" colspan="1" rowspan="1">x</td>
<td id="1DD9A407916ED69DD71DE6EA862CC9A5" colspan="1" rowspan="1">x</td>
</tr>
<tr id="007CF51E21D7AF5AEBC2580D40C41F54">
<td id="513E0582FCDADDDB488F24EF8EA64669" colspan="1" rowspan="1">
<taxonomicName id="A32F9A2BEADE28DE5493295EAB50B720" authority="Ribaut &amp; Brolemann, 1927" authorityName="Ribaut &amp; Brolemann" authorityYear="1927" class="Chilopoda" family="Geophilidae" genus="Galliophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="6615DBC6D886D306C333BF9E19FCC205" italics="true">Galliophilus</emphasis>
Ribaut &amp; Brolemann, 1927
</taxonomicName>
</td>
<td id="78466F49B465A5AAD41263BC55A0958A" colspan="1" rowspan="1">1</td>
<td id="B1D7C3D323B4CACC290D181B37677A49" colspan="1" rowspan="1">x</td>
<td id="71F4F40A4B28413F36DF757584CD4F44" colspan="1" rowspan="1">x</td>
<td id="F63B84D893FEF995EF567FE4D272E4D8" colspan="1" rowspan="1"></td>
</tr>
<tr id="427D36396601547A62F8F56286C0B0F4">
<td id="1EE94A275CB9795AE9884F7731C3A0EC" colspan="1" rowspan="1">
<taxonomicName id="08924604CB2E0BB8092EEEC083D37D5D" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="D69BD4EE57A05E39A557E430B34C229D" italics="true">Geophilus</emphasis>
Leach, 1814
</taxonomicName>
</td>
<td id="3F829C95337E008015BFFD1EC612F21F" colspan="1" rowspan="1">49</td>
<td id="221AB7C73B05A8FB6F534B112D4CD04B" colspan="1" rowspan="1">x</td>
<td id="9C9ABBFC5D8C76EA11728EE8F8834F14" colspan="1" rowspan="1">x</td>
<td id="65D020318316146FD09B8895B8EE5FFA" colspan="1" rowspan="1">x</td>
</tr>
<tr id="D6BD21DAD34268BC612AA665ED33920C">
<td id="7D67FE7F21C8C5992C9553A04DE1E766" colspan="1" rowspan="1">
<taxonomicName id="3F3A121FF148FE17D8A20990207E91BE" authority="Brolemann, 1909" authorityName="Brolemann" authorityYear="1909" class="Chilopoda" family="Geophilidae" genus="Gnathoribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="52403007303271AFA5EDB6429803FF5F" italics="true">Gnathoribautia</emphasis>
Brolemann, 1909
</taxonomicName>
</td>
<td id="95B15D824324B65E9AA521CA19D4611E" colspan="1" rowspan="1">2</td>
<td id="99F4C5F4733EF4A88D95AFB020B5C30E" colspan="1" rowspan="1">x</td>
<td id="1A2AD09269CA65E211B564E008481012" colspan="1" rowspan="1"></td>
<td id="A632D6D8354AF62829B273A9D3EA0195" colspan="1" rowspan="1">x</td>
</tr>
<tr id="43073034F90AB4A9FA7B5941DB59B3E5">
<td id="68FDBEB72ACCB0255786C0A87BB19E5A" colspan="1" rowspan="1">
<taxonomicName id="70FC491A38AEC71B4A6DA0BE0EB5D16D" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" class="Chilopoda" family="Dignathodontidae" genus="Henia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="30C2D468C70B8CCD1BF6F9C1B0895A68" italics="true">Henia</emphasis>
C. L. Koch, 1847
</taxonomicName>
</td>
<td id="32AC8872D4B05A1A1AD27CF0B3D5218F" colspan="1" rowspan="1">17</td>
<td id="55B29377895A8CFA12D22D0D488FE357" colspan="1" rowspan="1">x</td>
<td id="FDF311D9AAF276B14AFC7FA5808A7FF9" colspan="1" rowspan="1">x</td>
<td id="3E71F52449FC3B511239A9889EF09708" colspan="1" rowspan="1">x</td>
</tr>
<tr id="1A482F6CAE888017A203EBA8BE3EC5F9">
<td id="2FF993AEB1146A6D95F50BFC5F9C13EB" colspan="1" rowspan="1">
<taxonomicName id="CAD9A1FBDA9E10454D1840C6E91D5EEB" authority="Lewis, Jones &amp; Keay, 1988" authorityName="Lewis, Jones &amp; Keay" authorityYear="1988" class="Chilopoda" family="Geophilidae" genus="Nothogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="8284CFBA413BCA6E3722D50932A170D0" italics="true">Nothogeophilus</emphasis>
<bibRefCitation id="80277EE70724BFD3ED75171890BB6182" DOI="10.1080/00222938800771031" author="Lewis JGE &amp; Jones RE &amp; Keay AN" journalOrPublisher="Journal of Natural History" pagination="1657-1663" refId="B36" refString="Lewis JGE, Jones RE, Keay AN (1988) On a new genus and species of centipede (Chilopoda Geophilomorpha Chilenophilidae) from the British Isles. Journal of Natural History 22: 1657 - 1663. https: // doi. org / 10.1080 / 00222938800771031" title="On a new genus and species of centipede (Chilopoda Geophilomorpha Chilenophilidae) from the British Isles." volume="22" year="1988">Lewis, Jones &amp; Keay, 1988</bibRefCitation>
</taxonomicName>
</td>
<td id="6790D861F9DF009663C5EF4ECFDDB2E1" colspan="1" rowspan="1">1</td>
<td id="3AFF261F4D93E716AE9358336AE62302" colspan="1" rowspan="1"></td>
<td id="0D63E8FAE7735F5A6FD23A382F027A59" colspan="1" rowspan="1"></td>
<td id="6CED6593089A036929ECCD71C9538B71" colspan="1" rowspan="1"></td>
</tr>
<tr id="3A05487B6586A3CDB283E8B16C45C567">
<td id="82F4226FCE7502FCC5CFF6883EA06B88" colspan="1" rowspan="1">
<taxonomicName id="FF5BDE77C25A2F3DF81F4AB3DA9CACAC" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="82373652C71F84A15C1966B92643D604" italics="true">Pachymerium</emphasis>
C. L. Koch, 1847
</taxonomicName>
</td>
<td id="71F4C8AFB6AA1674E94CFF04416D079F" colspan="1" rowspan="1">6</td>
<td id="D38BD32D1A9EA2F2DD1DC37AAE56E4EC" colspan="1" rowspan="1">x</td>
<td id="F871F2A14DF291DCE67D9DCFC6CCCFC7" colspan="1" rowspan="1">x</td>
<td id="25FE38FCAD33AB37D3C2B1C5A99A22DA" colspan="1" rowspan="1">x</td>
</tr>
<tr id="67062B715BEB9D8694DBA47B4492172E">
<td id="09FF46D2879B801B9AA31911D0C2B982" colspan="1" rowspan="1">
<taxonomicName id="E6615F01BE5294F73DAB4E38EBA74EBF" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" class="Chilopoda" family="Geophilidae" genus="Pleurogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="8A809AE4288BB4B46B5346B6B408E580" italics="true">Pleurogeophilus</emphasis>
Verhoeff, 1901
</taxonomicName>
</td>
<td id="D388503550D58137DE41296005F72F40" colspan="1" rowspan="1">3</td>
<td id="6743CE6478F869C228C3063868EC28AF" colspan="1" rowspan="1">x</td>
<td id="ED8CEC4BD2B44DB1ABACDD7A27ADD449" colspan="1" rowspan="1">x</td>
<td id="553FCA145F2134F5E13B829C71D75CFC" colspan="1" rowspan="1">x</td>
</tr>
<tr id="7BEAB396DD0EC5A9C14CBD52D08AC445">
<td id="953C9CD4FE558703B97222C8A24E76FB" colspan="1" rowspan="1">
<taxonomicName id="A6B998FF1B17C0454522A987098DB152" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" class="Chilopoda" family="Geophilidae" genus="Stenotaenia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="581DC75815A13A203C1B70224A7F23C9" italics="true">Stenotaenia</emphasis>
C. L. Koch, 1847
</taxonomicName>
</td>
<td id="F4E24882C4855AF362516DCAE597D58F" colspan="1" rowspan="1">9</td>
<td id="42CEDFBF8F7E9371AFB2AD66F42C93C2" colspan="1" rowspan="1">x</td>
<td id="6FFF3C455101B149F2ACEA92207C2D8E" colspan="1" rowspan="1">x</td>
<td id="46AFE7A94751E2915B943C718CB7D371" colspan="1" rowspan="1">x</td>
</tr>
<tr id="382D843FCCD4B4A611832E6ED7F5BA67">
<td id="5C2B6E97E3E3B766753388EAC140E446" colspan="1" rowspan="1">
<taxonomicName id="438A4CE2DEB15EE4F3565A56A96C0651" authority="Gray, 1843" authorityName="Gray" authorityYear="1843" class="Chilopoda" family="Linotaeniidae" genus="Strigamia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="0FB44EC8580E14020E4D64BCC9C985D5" italics="true">Strigamia</emphasis>
Gray, 1843
</taxonomicName>
</td>
<td id="5BFFAC360FB86BA51702B2812F25E01F" colspan="1" rowspan="1">7</td>
<td id="3C19E350E4CC522304FE17FD80D63841" colspan="1" rowspan="1">x</td>
<td id="2F494AD5EAC5C8A98E3173371A1349E0" colspan="1" rowspan="1">x</td>
<td id="098468D66343413BB32C65BFB5A23B33" colspan="1" rowspan="1">x</td>
</tr>
<tr id="CA1757479DEE31699EDD30ACB25ECA9B">
<td id="9F77969E1CD758AA8D0562915EA3A637" colspan="1" rowspan="1">
<taxonomicName id="2EA9D1511D26F9BD88F7BEAB36678E98" authority="Chamberlin, 1920" authorityName="Chamberlin" authorityYear="1920" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="58C3F23D7DD31693384F4660E14F65A0" italics="true">Tuoba</emphasis>
Chamberlin, 1920
</taxonomicName>
</td>
<td id="4622EFEAFFCE3C0C492CBC93712F7410" colspan="1" rowspan="1">2</td>
<td id="87CDFDE85553672FB5B86916DBE86DBA" colspan="1" rowspan="1">x</td>
<td id="90BE9F7C23AB04831CD53BDBA323532C" colspan="1" rowspan="1">x</td>
<td id="5A4D5CADD051C991D2F33D6F36158E15" colspan="1" rowspan="1">x</td>
</tr>
<tr id="EC755E2BFAC39AC2A1C3DE84E2FAC31D">
<td id="7D11082254FDF656698DD46CAAD5048C" colspan="1" rowspan="1">Total</td>
<td id="9666D10A80B07C7806386D939F864E25" colspan="1" rowspan="1">121 species</td>
<td id="45706A8BEF614F5F693D31076BD7CABA" colspan="1" rowspan="1">16 genera</td>
<td id="913E4BFE2224798161E3B42226A2DE7E" colspan="1" rowspan="1">14 genera</td>
<td id="166799F954FD1EF83B0FF7C433C3BD0C" colspan="1" rowspan="1">13 genera</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="502D028B73AB93BD103664E23E85C2A1">
The key should be applied by examining any specimen through a microscope. A magnification of 50 × is recommended, even 100 × for examining labrum and pore-fields. For characters defined on the leg-bearing segments, examination between the 5
<superScript id="220DBC24D689F5D4D71453D9EE870D6A">th</superScript>
and the 20
<superScript id="76CE44283007E10909215097D3C180DA">th</superScript>
<abbrev id="ABBRID0EC3AG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
is recommended.
</paragraph>
<paragraph id="D0415854A96B350555F50FEAAC1FC9D7">In addition to the dichotomic characters included in the key, additional information is given for each genus (number of species recorded in Europe, geographical distribution within Europe, and additional morphological characters), between square brackets, to assist in the identification.</paragraph>
<paragraph id="8FE2426D2E40BAF5EA707EC96C2CED2A">
<table id="TID0E5MAC" rules="all">
<tbody id="355AB1DB2FC35A936CE5639411E32EAD">
<table-wrap id="9E2AAD3C1DCED83813B25808C43B6775" content-type="key" orientation="portrait" position="anchor">
<tr id="A9F64F23927258E5A4465B42EAFA50C7">
<td id="B34B0B0F099862071E54C25733A62AF6" colspan="1" rowspan="1">1</td>
<td id="4B302ACC51857344A2A58CE805CD8906" colspan="1" rowspan="1">
Legs of the ultimate pair with an apical tubercle having minute spines (Fig.
<figureCitation id="EAD8283FC2B34932A345DBD60044BA6A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Legs of the ultimate pair A, C right leg B, D distal articles of the right leg E G both legs H left leg. Species A Arctogeophilus inopinatus male B Arenophilus peregrinus female C Eurygeophilus pinguis female D Geophilus fucorum female E Geophilus richardi female F Pachymerium ferrugineum female G Pleurogeophilus mediterraneus male H Pleurogeophilus mediterraneus female I Acanthogeophilus spiniger female. Views: ventral (A-E, I), lateral (F) and dorsal (G-H). Abbreviations: c claw, t tubercle with spines. Scale bars: 300 µm (A, C, F); 50 µm (B); 100 µm (D, E, I); 1 mm (G H)." figureDoi="10.3897/zookeys.1213.133171.figure10" httpUri="https://binary.pensoft.net/fig/1142661">10 B</figureCitation>
: t)
</td>
<td id="D1ACF6F5203DBF05C27938EE3336F6F8" colspan="1" rowspan="1">
<taxonomicName id="E831B874F3A91791B40592514AFD32FB" authority="Chamberlin, 1912" authorityName="Chamberlin" authorityYear="1912" class="Chilopoda" family="Geophilidae" genus="Arenophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="D47F90712C7422ED819153C5C049C4BD" bold="true">
<emphasis id="733ECFB5F9A815CCBBBDD463ACE98289" bold="true" italics="true">Arenophilus</emphasis>
Chamberlin, 1912
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="EC71206A65F1C8F51F0AE5963597CB61">
<td id="46131AB3627B365BD4FBAC619367F5A4" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="87E9C4BDF669A6C188CE9E66E1B9431E" colspan="1" rowspan="1">
[In Europe: a single species,
<taxonomicName id="EF54C52C7E205A27FE2E47EFD59D9FFA" authority="Jones, 1989" authorityName="Jones" authorityYear="1989" class="Chilopoda" family="Geophilidae" genus="Arenophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="753FE3AEC4D0CB5742849BFC4B88A24C" italics="true">A. peregrinus</emphasis>
Jones, 1989
</taxonomicName>
; recorded in very few localities on and near the coasts of south-western England, western France, and western Iberian peninsula. Always with 45
<abbrev id="ABBRID0EZ4AG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
.]
</td>
<td id="57CF1E4F333D0874429C6752C9A11B29" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="12B716E35CAF17E41C0CF51CCD0B7583">
<td id="5DCB51B1615A37B7E84E288E953BEC9D" colspan="1" rowspan="1"></td>
<td id="4ECFA0EC6D850CA9D3F50BC11BFC505C" colspan="1" rowspan="1">
Legs of the ultimate pair with an apical claw or missing the pretarsus at all (Figs
<figureCitation id="FE57533D6AB58B36EAAA35F96364A945" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 H</figureCitation>
,
<figureCitation id="A6C8F07546C76D802D6579149E8AA569" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Legs of the ultimate pair A, C right leg B, D distal articles of the right leg E G both legs H left leg. Species A Arctogeophilus inopinatus male B Arenophilus peregrinus female C Eurygeophilus pinguis female D Geophilus fucorum female E Geophilus richardi female F Pachymerium ferrugineum female G Pleurogeophilus mediterraneus male H Pleurogeophilus mediterraneus female I Acanthogeophilus spiniger female. Views: ventral (A-E, I), lateral (F) and dorsal (G-H). Abbreviations: c claw, t tubercle with spines. Scale bars: 300 µm (A, C, F); 50 µm (B); 100 µm (D, E, I); 1 mm (G H)." figureDoi="10.3897/zookeys.1213.133171.figure10" httpUri="https://binary.pensoft.net/fig/1142661">10 A, C I</figureCitation>
)
</td>
<td id="F170D3DD709E0D77915CB09836C013B3" colspan="1" rowspan="1">
<emphasis id="380E33B162A7C42DD53AD612C5DA69DB" bold="true">2</emphasis>
</td>
</tr>
<tr id="4F7A34A50329D6323CE09BA0C1E55904">
<td id="E72D02943658C7D5317F280225C711A2" colspan="1" rowspan="1">2</td>
<td id="8BA69302F3C66BCBF5BA1785C7FDBDD3" colspan="1" rowspan="1">
Forcipular tergite sub-rectangular, more or less headband-shaped (Fig.
<figureCitation id="3D0D869D60D61CD280E36BC82A3A9236" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 D, E</figureCitation>
: ft)
</td>
<td id="3E7BE7879FCC1CF83DB514577FB991A8" colspan="1" rowspan="1">
<emphasis id="E60E23BD7F2A88565B90639575A96CF6" bold="true">3</emphasis>
</td>
</tr>
<tr id="23D89DCA674191E6BD2AC0AE48EF9759">
<td id="0B7B0C540F01AA38FB7F54293E6ED775" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="9D3D3E1B18E589E1DD6088AC6C194735" colspan="1" rowspan="1">
[The genera included here have a large basal denticle on the tarsungulum (Fig.
<figureCitation id="96CC08076984B9ED0AFA4AC40D888456" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 F</figureCitation>
) or a deep median diastema on the forcipular coxosternite (Fig.
<figureCitation id="3A84061C5462DD688307A99FE27711DA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 G, H</figureCitation>
).]
</td>
<td id="B70BBE0D35CE90F7406E6509090930CE" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="9FB07806C76724FC8B41BA884AD7365C">
<td id="C43DE74B1B7302AF099F62F2D63CC2A0" colspan="1" rowspan="1"></td>
<td id="327EEB641E6C5CC772CF04323099FCDB" colspan="1" rowspan="1">
Forcipular tergite trapezoid, posteriorly wider than anteriorly (Fig.
<figureCitation id="CC8020D885FC5DF75BBD851F770714DB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 A C</figureCitation>
: ft)
</td>
<td id="5631E0A6589A35C2129FB9C659EF77AF" colspan="1" rowspan="1">
<emphasis id="F3DF0A62C7603DFA960E6E8822D0D144" bold="true">5</emphasis>
</td>
</tr>
<tr id="DDFDD76E68D49C9EA5ACEDBB49D8E6F9">
<td id="F775C0DCBBB2F4275C62C483C517A128" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="B821CD33DA270D4C2C136C61765A2EA7" colspan="1" rowspan="1">
[With the exception of
<taxonomicName id="0042178E4D289F9C5EF1EEE5DC4FD69E" authorityName="Brolemann" authorityYear="1909" class="Chilopoda" family="Geophilidae" genus="Gnathoribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="1CBCDF53E06E4E712847250404BB2F7E" italics="true">Gnathoribautia</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="1EF79C22922D4E28DA999BEBEAAF6E6B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 I</figureCitation>
), the basal denticle of the tarsungulum, when existing, is smaller (Fig.
<figureCitation id="5F82B44940DEAF9F5171925D4244B72E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 C, G</figureCitation>
). No deep median diastema on the forcipular coxosternite (Figs
<figureCitation id="2BA96FD7B6253BF2ECC325AF2B7A5CC1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 I</figureCitation>
,
<figureCitation id="253854D3147F5A2D8A8BAF753030136B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 C, G</figureCitation>
).]
</td>
<td id="BB999457619F4D74469799B33BC0118D" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="0C25CE4D2DD9E8FB6F8EA75E9CBC67D4">
<td id="8498505B85FB12C15B987BB9B47D3F8D" colspan="1" rowspan="1">3</td>
<td id="F5067113A9012B9E045E027FB64649F9" colspan="1" rowspan="1">
A large basal denticle on the tarsungulum (Fig.
<figureCitation id="A1C5D2FD8D2E3CA93AC4EE5BAA37609A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 F</figureCitation>
: d). No deep median diastema and no chitin-lines on the forcipular coxosternite (Fig.
<figureCitation id="AE5B335FCDF1C3D6B77E2AEC6500D692" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 F</figureCitation>
)
</td>
<td id="4798EA4EB9F18356BB4F635A36067B61" colspan="1" rowspan="1">
<taxonomicName id="B76B4C2B1BA48490D3D5ADA9D6FFB6F5" authority="Gray, 1843" authorityName="Gray" authorityYear="1843" class="Chilopoda" family="Linotaeniidae" genus="Strigamia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="A5387810C09CA9B78712F2162EF74FB8" bold="true">
<emphasis id="CB9C6A8B763C08A5F2D46A3E4EC5FE2B" bold="true" italics="true">Strigamia</emphasis>
Gray, 1843
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="7FD77EA952C7B3354BC97A8C351E7CB0">
<td id="5FC0B0F194620553B8424F10A260DEFD" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="4D65C41CC061BFEA41F6BE9E2DDA54F5" colspan="1" rowspan="1">[In Europe: many species, across most continental lands.]</td>
<td id="3542C8EFBD6F91B0D369AEF137432254" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="D24574ED3590CF67C5470BD2F0BE6B97">
<td id="423AB4D6EBEBBF0DCB30F9CEAD04D23F" colspan="1" rowspan="1"></td>
<td id="92A1C85180A76B584B8DE282E004B12F" colspan="1" rowspan="1">
No large basal denticle on the tarsungulum (Fig.
<figureCitation id="EA9EDE00A5F8F2702D9A0DC057A2F5CB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 G, H</figureCitation>
). A deep median diastema and distinct chitin-lines on the forcipular coxosternite (Fig.
<figureCitation id="ECA430AEC94A583E0A34B5F8BFD0D2C4" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 G, H</figureCitation>
: cl)
</td>
<td id="E6F2DE83EDFD98E26D9EE82586A47760" colspan="1" rowspan="1">
<emphasis id="F8E319FAC9E0F8F1C122777024DBD53D" bold="true">4</emphasis>
</td>
</tr>
<tr id="367CE6B98DB9160183EDF8189F1B207D">
<td id="76B522BF63F9194B37847842FAE44A96" colspan="1" rowspan="1">4</td>
<td id="088FBBA9B2286F61B3E667140B074954" colspan="1" rowspan="1">
2 denticles on the mesial side of the tarsungulum (Fig.
<figureCitation id="5AD00393ECAEF0A2A33C9C884072EFA2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 G</figureCitation>
: dc). Pore-fields absent
</td>
<td id="328354FDFA36E0ECE206F329361A7513" colspan="1" rowspan="1">
<taxonomicName id="DCF99A7280DC502AA16B943646EA8C10" authority="Meinert, 1870" authorityName="Meinert" authorityYear="1870" class="Chilopoda" family="Dignathodontidae" genus="Dignathodon" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="B5F90120B1118379A794CD793A0B205F" bold="true">
<emphasis id="4AD8D51680E200E418434D0397BB90F6" bold="true" italics="true">Dignathodon</emphasis>
Meinert, 1870
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="7C5A984E80D91A08F2F4844824EFFEDB">
<td id="FC357E30FE3FBC4D3E78A6BE88A2D512" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="59E0BC397891C0937F3D3ECB9B639654" colspan="1" rowspan="1">
[In Europe: only two species,
<taxonomicName id="4FA56A24A8674A6FD3716FF27C0E1A15" authority="(Lucas, 1846)" baseAuthorityName="Lucas" baseAuthorityYear="1846" class="Chilopoda" family="Dignathodontidae" genus="Dignathodon" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="microcephalus">
<emphasis id="056DAAD5F7AA23B8D615078611323D99" italics="true">D. microcephalus</emphasis>
(Lucas, 1846)
</taxonomicName>
(southern Europe; 6589
<abbrev id="ABBRID0ECEBG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
) and
<taxonomicName id="F92CF239D9D805AB896D3EC25D1E3130" authority="(Attems, 1952)" baseAuthorityName="Attems" baseAuthorityYear="1952" class="Chilopoda" family="Dignathodontidae" genus="Dignathodon" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="89DC6251C8B048BC84A63116824260DE" italics="true">D. gracilis</emphasis>
(Attems, 1952)
</taxonomicName>
(only recorded in Andalusia; 5971
<abbrev id="ABBRID0EREBG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
.]
</td>
<td id="940B263005D143AFFA994E993AA5ADD1" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="8769F43D84DBEDCB930BD88D9AA93F03">
<td id="01C713D1F6C33F2616453BB20E272BA0" colspan="1" rowspan="1"></td>
<td id="13B1505ED8D701EDD8D852E58E72B717" colspan="1" rowspan="1">
No denticles on the mesial side of the tarsungulum (Fig.
<figureCitation id="E87CDC9BD49EAAA6F2EAEB6AD35406A0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 H</figureCitation>
). Pore-fields present, sub-circular to longitudinally much elongate
</td>
<td id="BEAD36B6F449E52C1E32A6C0B71890CD" colspan="1" rowspan="1">
<taxonomicName id="70BDF92AA88CA1B5A130EEAC1F363080" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" class="Chilopoda" family="Dignathodontidae" genus="Henia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="286A1130401AA2AFAEF9478CC5E5EFBE" bold="true">
<emphasis id="6CA5896DD2D0F40AF590DF63C2BAD869" bold="true" italics="true">Henia</emphasis>
C. L. Koch, 1847
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="879948D8719BECA7C363BFC4BE872A91">
<td id="1F0BDBDFCEC8DFF63506E71C67F6A2A2" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="AE01E88C6891650B5AE5FEAC74D23603" colspan="1" rowspan="1">[In Europe: many species, across most continental lands and major islands.]</td>
<td id="6163451F017733312BB263BBA19039DA" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="10A4EBF1407F5B9B6EFA6EE0DD592CC7">
<td id="CF3225C0F481375D6DBB0F241C40DB03" colspan="1" rowspan="1">5</td>
<td id="49BBDE528D1D8B6EFE40CF637CE01538" colspan="1" rowspan="1">
Coxal pores grouped in 1, 2, 3, or 4 ventral pits, fossae, or distinct clusters on each coxopleuron, all close to the metasternite or covered by it (Fig.
<figureCitation id="0AD9DBED6D67265F92D8AF65A7C343DD" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Ultimate LBS without distal part of legs. Species A Arctogeophilus inopinatus female B Galliophilus beatensis male C Geophilus flavus female D Pachymerium ferrugineum male E Pleurogeophilus mediterraneus female F Pleurogeophilus mediterraneus male G, H Stenotaenia linearis female I Tuoba poseidonis female. Views: ventral (A G), lateral (H, I). Abbreviations: p pit or fossa (H wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate LBS (medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm." figureDoi="10.3897/zookeys.1213.133171.figure9" httpUri="https://binary.pensoft.net/fig/1142660">9 G I</figureCitation>
), but sometimes accompanied by 1 or 2 lateral or posterior separate pores
</td>
<td id="19263BCE62865FAD929D23D0BA47EC5C" colspan="1" rowspan="1">
<emphasis id="93020A7317F5A475C7EA553317E4908F" bold="true">6</emphasis>
</td>
</tr>
<tr id="7E5CB5211E57B99D0D3B3B33C0324233">
<td id="4C76313213630D7633C433C24CD05F03" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="DD5A140BD7C0C64ADC319E3E4E583CAF" colspan="1" rowspan="1">
[In France and in north-western Europe, with the exception of
<taxonomicName id="7D38FC49A16BBA1B13766F01CB2026B3" authorityName="Lewis, Jones &amp; Keay" authorityYear="1988" class="Chilopoda" family="Geophilidae" genus="Nothogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="A212DE2EC9D6C4894753F25B00BBC872" italics="true">Nothogeophilus</emphasis>
</taxonomicName>
only recorded in England, the labrum of the species of the two other genera present in this area (
<taxonomicName id="5B78C24E49476773EDFCB3D8E78DBED9" authorityName="C. L. Koch" authorityYear="1847" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="763B37770239478C2A3F2DF8472727AD" italics="true">Clinopodes</emphasis>
</taxonomicName>
and
<taxonomicName id="8734358A40FDFEBD81996A47ABF9F11B" authorityName="C. L. Koch" authorityYear="1847" class="Chilopoda" family="Geophilidae" genus="Stenotaenia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="AF0FDAB99F41E1FE1821427FDD9D8029" italics="true">Stenotaenia</emphasis>
</taxonomicName>
) generally has numerous bristles and no tubercles (Fig.
<figureCitation id="E6DC46E6D304BD4AF8C0CEB6A2A2FC9E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 A</figureCitation>
).]
</td>
<td id="C358F793A9C8DA1D097E95B7C9C1404E" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="CBD910118F291151D1F87425A90FE25C">
<td id="022F02B2C0A8FBF5B90CD83B5D173CF5" colspan="1" rowspan="1"></td>
<td id="66C12EDAC651114D98DBE2D38850FECB" colspan="1" rowspan="1">
All coxal pores separate, not in pits, fossae, or distinct clusters (Figs
<figureCitation id="6037E45CB482534754287C545293E79E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 E, F</figureCitation>
,
<figureCitation id="5B34B49BC3983C7F57FD430EFF435159" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endogeophilus alberti sp. nov., holotype A head and forcipular segment B antennae C distal articles of left antenna D right telopodite of second maxillae E head and forcipular segment F right forcipule G LBS 15 and anterior part of 16 H ultimate LBS without right leg. Views: ventral (D H), dorsal (A C). Abbreviations: cls club-like sensilla, cmx claw of the second maxillae, cp coxopleural pores, cs carpophagus-structure. Scale bars: 200 µm (A B, E, G H); 50 µm (C D, F)." figureDoi="10.3897/zookeys.1213.133171.figure3" httpUri="https://binary.pensoft.net/fig/1142654">3 H</figureCitation>
,
<figureCitation id="3ECC3FEA14AA881DD698985C5C758E2D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Ultimate LBS without distal part of legs. Species A Arctogeophilus inopinatus female B Galliophilus beatensis male C Geophilus flavus female D Pachymerium ferrugineum male E Pleurogeophilus mediterraneus female F Pleurogeophilus mediterraneus male G, H Stenotaenia linearis female I Tuoba poseidonis female. Views: ventral (A G), lateral (H, I). Abbreviations: p pit or fossa (H wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate LBS (medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm." figureDoi="10.3897/zookeys.1213.133171.figure9" httpUri="https://binary.pensoft.net/fig/1142660">9 A F</figureCitation>
,
<figureCitation id="432D48612429C13FA8C5278DC7CEEAC2" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Legs of the ultimate pair A, C right leg B, D distal articles of the right leg E G both legs H left leg. Species A Arctogeophilus inopinatus male B Arenophilus peregrinus female C Eurygeophilus pinguis female D Geophilus fucorum female E Geophilus richardi female F Pachymerium ferrugineum female G Pleurogeophilus mediterraneus male H Pleurogeophilus mediterraneus female I Acanthogeophilus spiniger female. Views: ventral (A-E, I), lateral (F) and dorsal (G-H). Abbreviations: c claw, t tubercle with spines. Scale bars: 300 µm (A, C, F); 50 µm (B); 100 µm (D, E, I); 1 mm (G H)." figureDoi="10.3897/zookeys.1213.133171.figure10" httpUri="https://binary.pensoft.net/fig/1142661">10 E</figureCitation>
)
</td>
<td id="132F527A237C1496CD393A32095F73F7" colspan="1" rowspan="1">
<emphasis id="8F200A191D074B778F23D628CBAC9DA2" bold="true">12</emphasis>
</td>
</tr>
<tr id="D602CC312CD85E72251FE60429D7CE52">
<td id="8A0A6335090D2043831A6D0FA1E67DD4" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="F9E4A75A4FFD70F33FAF5AF9FCD15C2A" colspan="1" rowspan="1">
[In France and in north-western Europe, the genera included here have one to several tubercles on the intermediate part and bristles on the side pieces of the labrum (Figs
<figureCitation id="D0A8F34FD3B2C7F83451F0206027A5BB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 A</figureCitation>
,
<figureCitation id="E8656F08DAFF251D87DD925961FAF52F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 B</figureCitation>
).]
</td>
<td id="31487EE599FD9D45B22547B3C947E7EA" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="ED0B0AE6CD03D7E984BB7DEE5C24B07F">
<td id="B2715D5C37AD14DD4CD3FD02ECB5F73E" colspan="1" rowspan="1">6</td>
<td id="0A0994DCBD8B45DEAAA90365BC6E4691" colspan="1" rowspan="1">
Forcipular coxosternite with a pair of distinctly sclerotised denticles (Fig.
<figureCitation id="13ED7028B5648BD34C2C09967F45EA10" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 E</figureCitation>
)
</td>
<td id="C2F5AF8FBA22502CAB546A51F7E03ED7" colspan="1" rowspan="1">
<emphasis id="087BC1A9DC89E800CD1443D4AEF1E74B" bold="true">7</emphasis>
</td>
</tr>
<tr id="8F259C67CE8CBBE0A0BF00617F6D1F12">
<td id="5C92AEB39C6661C731B889CF563CC5FC" colspan="1" rowspan="1"></td>
<td id="8B0AA6B5B89FF4F3EA8A231B155C68CF" colspan="1" rowspan="1">
Forcipular coxosternite without distinctly sclerotised denticles (e. g., Fig.
<figureCitation id="781A9426F064F44B3FF6686D176BE8FC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 G</figureCitation>
)
</td>
<td id="2F57C131A1DAE6470DE3F16F70E2F6AD" colspan="1" rowspan="1">
<emphasis id="B9B5E797FDC55C270637D8085F5225F7" bold="true">8</emphasis>
</td>
</tr>
<tr id="3D62E775DE334788F4BFDA338C4ECCB6">
<td id="AD9F08BC4E07816CB8485DC74F852556" colspan="1" rowspan="1">7</td>
<td id="14EE9B90357EDF8FA102EE627206BDF6" colspan="1" rowspan="1">
Carpophagus structures absent (e. g., Fig.
<figureCitation id="8AD69C07058900A83D8E6FAAF9E78FAB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 G, H</figureCitation>
). Coxal pores mostly or entirely in 1 ventral pit on each coxopleuron (Fig.
<figureCitation id="C599DC98A05C49A46DE95D6CAED18C42" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Ultimate LBS without distal part of legs. Species A Arctogeophilus inopinatus female B Galliophilus beatensis male C Geophilus flavus female D Pachymerium ferrugineum male E Pleurogeophilus mediterraneus female F Pleurogeophilus mediterraneus male G, H Stenotaenia linearis female I Tuoba poseidonis female. Views: ventral (A G), lateral (H, I). Abbreviations: p pit or fossa (H wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate LBS (medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm." figureDoi="10.3897/zookeys.1213.133171.figure9" httpUri="https://binary.pensoft.net/fig/1142660">9 I</figureCitation>
)
</td>
<td id="F511C34115CC29AB5C20F5F840345AE0" colspan="1" rowspan="1">
<taxonomicName id="0B6DBF183C763E44E3AD00D7A509A870" authority="Bonato, Zapparoli &amp; Minelli, 2008" authorityName="Bonato, Zapparoli &amp; Minelli" authorityYear="2008" class="Chilopoda" family="Geophilidae" genus="Diphyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="787CF9A9E15BDFA37E184D81EE188002" bold="true">
<emphasis id="33DE7B7A8294ECF98C745B6B9518551F" bold="true" italics="true">Diphyonyx</emphasis>
Bonato, Zapparoli &amp; Minelli, 2008
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="A5E4E3438268AAA95001D9E636BF0173">
<td id="22B3B986913BD6F8429FBF72C1AB37AC" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="BBE5A927A97BF27A03BCC8571C2C8BA1" colspan="1" rowspan="1">[In Europe: a few species, from the Balkan peninsula to Caucasus. Claws of many anterior legs bearing an enlarged accessory spine.]</td>
<td id="6E0E190AB2960895DFFF2468998EBE19" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="F891F8988E667A8FCBA2385EC6A71196">
<td id="20BDE7BA573968167C85C527FE11ADB8" colspan="1" rowspan="1"></td>
<td id="AB7E9124688358239CFB739B12EE6F1C" colspan="1" rowspan="1">
Carpophagus structures present (Fig.
<figureCitation id="D10051355103CAD1314E7B8DD4D5A641" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 A, B</figureCitation>
: cs). Coxal pores mostly or entirely in 24 ventral pits or clusters (e. g., Fig.
<figureCitation id="94F598D1F8D5C395FE744BC623345877" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Ultimate LBS without distal part of legs. Species A Arctogeophilus inopinatus female B Galliophilus beatensis male C Geophilus flavus female D Pachymerium ferrugineum male E Pleurogeophilus mediterraneus female F Pleurogeophilus mediterraneus male G, H Stenotaenia linearis female I Tuoba poseidonis female. Views: ventral (A G), lateral (H, I). Abbreviations: p pit or fossa (H wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate LBS (medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm." figureDoi="10.3897/zookeys.1213.133171.figure9" httpUri="https://binary.pensoft.net/fig/1142660">9 G</figureCitation>
)
</td>
<td id="340DCA6D2CBEB2D9CE2C0AB67C95624D" colspan="1" rowspan="1">
<taxonomicName id="8077AAC096328DBEE06D34D7B5380854" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="DE8ABCAB19F360E5E45E26003078B5CC" bold="true">
<emphasis id="84797C95D1FB228B724AEA44A084EEFE" bold="true" italics="true">Clinopodes</emphasis>
C. L. Koch, 1847
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="F6E1C3390F32BE6BA29F29993E792918">
<td id="FEA855153620ADA7EE955CCA0BE49AF9" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="A33F1037831D88EF2A592D10179BAF87" colspan="1" rowspan="1">[In Europe: many species, across most of central and eastern Europe.]</td>
<td id="3D7A948B56FE119BF922F6C53CF6CE34" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="2C2882BD0632BCB9FF11CED4A1B58558">
<td id="2F97B2C6DFA8287BE38AC5B61BE0F29D" colspan="1" rowspan="1">8</td>
<td id="EC248A638DE66AA5FC7432E745C11753" colspan="1" rowspan="1">No pore-fields. Labrum without bristles and without tubercles</td>
<td id="B75435998E7C6CA47FB3240522C92116" colspan="1" rowspan="1">
<taxonomicName id="0EA2BCCAEB7248B9AD4E624280F00F4C" authority="Verhoeff, 1941" authorityName="Verhoeff" authorityYear="1941" class="Chilopoda" family="Geophilidae" genus="Bebekium" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="F76E5C134BC2F4C58B70A4A4E2C45D4B" bold="true">
<emphasis id="81098C930B97072205BC1A12E5118FE9" bold="true" italics="true">Bebekium</emphasis>
<bibRefCitation id="73D3F3C44EBBC1ECC65DB09170450AE2" author="Verhoeff KW" journalOrPublisher="Zoologischer Anzeiger" pagination="35-41" refId="B45" refString="Verhoeff KW (1941) Untersuchungen uber die Fauna des Hologaeum: Chilopoden, Diplopoden und Land-Isopoden. Zoologischer Anzeiger 135: 35 - 41." title="Untersuchungen über die Fauna des Hologaeum: Chilopoden, Diplopoden und Land-Isopoden." volume="135" year="1941">Verhoeff, 1941</bibRefCitation>
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="68008177C51B082DF19D017CD6AE0463">
<td id="D68A6F37D195916DF530849E484E8479" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="59773981CFA71C37699FE7A7D17D8AEA" colspan="1" rowspan="1">
[In Europe: a single species,
<taxonomicName id="7C9CF902BD746FDF5A0C05E2FA903E29" authority="Verhoeff, 1941" authorityName="Verhoeff" authorityYear="1941" class="Chilopoda" family="Geophilidae" genus="Bebekium" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="mirabile">
<emphasis id="70060881B76CEABA852BACB775603186" italics="true">B. mirabile</emphasis>
<bibRefCitation id="213662B15025A8E811DE61CE1F8D6066" author="Verhoeff KW" journalOrPublisher="Zoologischer Anzeiger" pagination="35-41" refId="B45" refString="Verhoeff KW (1941) Untersuchungen uber die Fauna des Hologaeum: Chilopoden, Diplopoden und Land-Isopoden. Zoologischer Anzeiger 135: 35 - 41." title="Untersuchungen über die Fauna des Hologaeum: Chilopoden, Diplopoden und Land-Isopoden." volume="135" year="1941">Verhoeff, 1941</bibRefCitation>
</taxonomicName>
; recorded only in the eastern part of the Balkan peninsula; 3941
<abbrev id="ABBRID0E2MBG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
.]
</td>
<td id="4971BCF43A81E8348F0E40CA412D0ED3" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="40EC061EF4663069263A6F459628050A">
<td id="8A73D1621ABE15E9BA237EC7DDC4AFD2" colspan="1" rowspan="1"></td>
<td id="CDF9CF13FE3185BD1438DEA77DC88CD8" colspan="1" rowspan="1">
Pore-fields present (Fig.
<figureCitation id="8C3BE4C02A459FD23C447D639E3118F5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 D, E, G, H</figureCitation>
: pf). Labrum with bristles, sometimes also with tubercles
</td>
<td id="B272574BE97FAA7D3EDF488AEF757AA3" colspan="1" rowspan="1">
<emphasis id="C681E5748BFC0A6639BF150123070851" bold="true">9</emphasis>
</td>
</tr>
<tr id="C96EE2BABFA61418BFE25CE486360D0D">
<td id="D7F63950909C0250FA028676B6173FB5" colspan="1" rowspan="1">9</td>
<td id="00FDFE3A9FD4614ECDDED2E3C5B533FE" colspan="1" rowspan="1">
All coxal pores in 2 or 3 ventral pits on each coxopleuron (Fig.
<figureCitation id="409282CAC4547FD62207FE993EB616C6" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Ultimate LBS without distal part of legs. Species A Arctogeophilus inopinatus female B Galliophilus beatensis male C Geophilus flavus female D Pachymerium ferrugineum male E Pleurogeophilus mediterraneus female F Pleurogeophilus mediterraneus male G, H Stenotaenia linearis female I Tuoba poseidonis female. Views: ventral (A G), lateral (H, I). Abbreviations: p pit or fossa (H wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate LBS (medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm." figureDoi="10.3897/zookeys.1213.133171.figure9" httpUri="https://binary.pensoft.net/fig/1142660">9 G, H</figureCitation>
). Pore-fields oval or sub-trapezoid (Fig.
<figureCitation id="9D188D7C443254A26ED9B060DE28FC42" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 G</figureCitation>
: pf)
</td>
<td id="F23E29B8033AE6748C160C748E899E69" colspan="1" rowspan="1">
<taxonomicName id="0E86614E61681EC1EE1FC841DB94C378" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" class="Chilopoda" family="Geophilidae" genus="Stenotaenia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="AC07752BCEA41DD66115CBDDADC4A2B1" bold="true">
<emphasis id="83E2E3DFD9EE0A7B835563C543847995" bold="true" italics="true">Stenotaenia</emphasis>
C. L. Koch, 1847
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="60C0E75649BA3B31ED6A2428624065E5">
<td id="18471F5DDD9E82044099B81A9739D97A" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="EFE1F52B38947DDD363AE6D7379FBF6A" colspan="1" rowspan="1">[In Europe: many species, across most of central and eastern Europe.]</td>
<td id="4F85D67CA625C6560C684552B3F43A00" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="E8B24314F42AA7A4E6D0C251E8F60571">
<td id="13F2C6E62663D7599BB1DA540D039AB5" colspan="1" rowspan="1"></td>
<td id="DBEAA0F50D75CE6632C97544A70FE4F7" colspan="1" rowspan="1">
All coxal pores in 1 ventral pit on each coxopleuron (Fig.
<figureCitation id="1486E5230BCBBCD7D90310F26FAE1FB6" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Ultimate LBS without distal part of legs. Species A Arctogeophilus inopinatus female B Galliophilus beatensis male C Geophilus flavus female D Pachymerium ferrugineum male E Pleurogeophilus mediterraneus female F Pleurogeophilus mediterraneus male G, H Stenotaenia linearis female I Tuoba poseidonis female. Views: ventral (A G), lateral (H, I). Abbreviations: p pit or fossa (H wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate LBS (medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm." figureDoi="10.3897/zookeys.1213.133171.figure9" httpUri="https://binary.pensoft.net/fig/1142660">9 I</figureCitation>
). Pore-fields sub-triangular (e. g., Fig.
<figureCitation id="8894E6C1CAC517F52FF1C463CE98DFD1" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 D, E</figureCitation>
: pf) or reniform or in a transverse band (Fig.
<figureCitation id="2DF9C62303AB4F7E75A42D3A94A0774B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 H</figureCitation>
: pf)
</td>
<td id="44665E24CBF73A6DB8FDD7FF4C4C7B2F" colspan="1" rowspan="1">
<emphasis id="AA3E8CACBC17131764AF540C6E92CD9B" bold="true">10</emphasis>
</td>
</tr>
<tr id="E6FB054C943BB090A9D60B2D986B0411">
<td id="983422A5F4DECAD14FC3630137DDDE5D" colspan="1" rowspan="1">10</td>
<td id="0004086B0AB3DAF1BD48144FDC89DCBC" colspan="1" rowspan="1">
Carpophagus structures present (Fig.
<figureCitation id="F1848C40C0030CA388BBA2C7AFB165BB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 E</figureCitation>
: cs). Pore-fields sub-triangular (e. g., Fig.
<figureCitation id="BE7C49445FCDCD50B4393D70E94AE21D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 D, E</figureCitation>
: pf). 5771
<abbrev id="ABBRID0E4PBG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
</td>
<td id="B70CB8A03A272C419CAD0554CB5510CC" colspan="1" rowspan="1">
<taxonomicName id="7DCE757C93BFCB917FA5297D4B1774CD" authority="Brolemann, 1925" authorityName="Brolemann" authorityYear="1925" class="Chilopoda" family="Geophilidae" genus="Algerophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="5078E0CB016A2DD71666A353773C6877" bold="true">
<emphasis id="A219B8C533B55DA8298AEF4F9652D156" bold="true" italics="true">Algerophilus</emphasis>
Brolemann, 1925
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="7F43AB65AFE3111A4087B12AD5832630">
<td id="E5FA589A0699A478A01236BA1DE18C64" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="6F9A2A7BCD1A5ABCDEE9CA1026F088C8" colspan="1" rowspan="1">
[In Europe: a single species,
<taxonomicName id="8EDE3F31159158C376B3FC8AFF12284D" authority="(Meinert, 1870)" baseAuthorityName="Meinert" baseAuthorityYear="1870" class="Chilopoda" family="Geophilidae" genus="Algerophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="BD80B7AFA7E7EF8EA61BA96C225DBB6C" italics="true">Algerophilus hispanicus</emphasis>
(Meinert, 1870)
</taxonomicName>
; only from southern Iberian peninsula.]
</td>
<td id="318A07A7FA9B21079359E2E282591A83" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="24FB38CA23BB62C750193624F39D48BF">
<td id="E10516EF3C26D03D2A4BC6F6943FE510" colspan="1" rowspan="1"></td>
<td id="E05E2F55618B0CD990A2102D7A530333" colspan="1" rowspan="1">
Carpophagus structures absent (Fig.
<figureCitation id="1CB5B15F4D16427AD590CF1704DC0B54" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 H</figureCitation>
). Pore-fields reniform or in a transverse band (Fig.
<figureCitation id="520C64E830BFE300AEB7786D3D939CDA" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 H</figureCitation>
: pf); 3755
<abbrev id="ABBRID0EORBG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
</td>
<td id="193FB708ADA5B2EB07FC1F45BD5A6011" colspan="1" rowspan="1">
<emphasis id="F0D06DD4678B821710559AEED4E38B5D" bold="true">11</emphasis>
</td>
</tr>
<tr id="63E83E084D744ED9F99C3675EA23888B">
<td id="5C6B78E41EEA91CDC592A38913C96728" colspan="1" rowspan="1">11</td>
<td id="3C3F3FC637AA0565A104D0D2C97FB491" colspan="1" rowspan="1">
Pore-fields more or less reniform, ≤ 1 / 2 of the width of the metasternite; 3739
<abbrev id="ABBRID0E6RBG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
</td>
<td id="0390D9B1847400552371C16707DE22CC" colspan="1" rowspan="1">
<taxonomicName id="368638A25BB3E8F928A4482A7356AEF1" authority="Lewis, Jones &amp; Keay, 1988" authorityName="Lewis, Jones &amp; Keay" authorityYear="1988" class="Chilopoda" family="Geophilidae" genus="Nothogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="2C6552765B4FD6B23C9EACBA76D0804D" bold="true">
<emphasis id="C93FF1125B2D0CD7C5D284515C6EEDD5" bold="true" italics="true">Nothogeophilus</emphasis>
<bibRefCitation id="9476C1EB90AA2E3ADB1850585AD9075B" DOI="10.1080/00222938800771031" author="Lewis JGE &amp; Jones RE &amp; Keay AN" journalOrPublisher="Journal of Natural History" pagination="1657-1663" refId="B36" refString="Lewis JGE, Jones RE, Keay AN (1988) On a new genus and species of centipede (Chilopoda Geophilomorpha Chilenophilidae) from the British Isles. Journal of Natural History 22: 1657 - 1663. https: // doi. org / 10.1080 / 00222938800771031" title="On a new genus and species of centipede (Chilopoda Geophilomorpha Chilenophilidae) from the British Isles." volume="22" year="1988">Lewis, Jones &amp; Keay, 1988</bibRefCitation>
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="430102DA0DC44B1EBC62A5DFFDD13E1A">
<td id="C8387F7AC5FE09A9B64FEBC51FC1ADC0" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="4AED76CA4A72A912E595D04162B14E7E" colspan="1" rowspan="1">
[In Europe: a single species,
<taxonomicName id="09E59B40E03039C3624486D4E0781A1B" authority="Lewis, Jones &amp; Keay, 1988" authorityName="Lewis, Jones &amp; Keay" authorityYear="1988" class="Chilopoda" family="Geophilidae" genus="Nothogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="turki">
<emphasis id="59611F1FF82252D8A36E043DDA649CB3" italics="true">N. turki</emphasis>
<bibRefCitation id="DABFFD6CEE010D20B6268B253347BC88" DOI="10.1080/00222938800771031" author="Lewis JGE &amp; Jones RE &amp; Keay AN" journalOrPublisher="Journal of Natural History" pagination="1657-1663" refId="B36" refString="Lewis JGE, Jones RE, Keay AN (1988) On a new genus and species of centipede (Chilopoda Geophilomorpha Chilenophilidae) from the British Isles. Journal of Natural History 22: 1657 - 1663. https: // doi. org / 10.1080 / 00222938800771031" title="On a new genus and species of centipede (Chilopoda Geophilomorpha Chilenophilidae) from the British Isles." volume="22" year="1988">Lewis, Jones &amp; Keay, 1988</bibRefCitation>
</taxonomicName>
; only recorded from the isles of Scilly and Wight (southern England). Body length &lt;15 mm.]
</td>
<td id="876BF9FCDE5FDCE81B3EEC490674738C" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="1F9BD8FE4D40C11AB290D14E084BE865">
<td id="5EC6DA1AE06C93DAD1A390A03BC86130" colspan="1" rowspan="1"></td>
<td id="264504F027FE2E7D53FA9C21C5947A60" colspan="1" rowspan="1">
Pore-fields not reniform,&gt; 1 / 2 of the width of the metasternite (Fig.
<figureCitation id="D96918BF954554030A35A2531FC0AFC0" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 H</figureCitation>
); 4755
<abbrev id="ABBRID0EMTBG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
</td>
<td id="034ACF9EA842BD24A4463EA318BDB075" colspan="1" rowspan="1">
<taxonomicName id="38F40F9BEC8A126B4325ECF031D156AF" authority="Chamberlin, 1920" authorityName="Chamberlin" authorityYear="1920" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="21F1FA007FBCC7720E249698739DAF8C" bold="true">
<emphasis id="504DF7D83E876C2994D1F40BA007A7C4" bold="true" italics="true">Tuoba</emphasis>
Chamberlin, 1920
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="4D831EC53ACDFB5730380E626D5ED9C0">
<td id="C5ABA8AD66EFCD7912EE5E43CDD38A00" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="4E9EF341B8AA364CF2D900A4809640D8" colspan="1" rowspan="1">
[In Europe: only two species,
<taxonomicName id="44FF521644C8F31940FFB3C41461DD81" authority="(Verhoeff, 1901)" baseAuthorityName="Verhoeff" baseAuthorityYear="1901" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="poseidonis">
<emphasis id="CB3B6148C7FAC8C38BA3011DBAF38913" italics="true">T. poseidonis</emphasis>
(Verhoeff, 1901)
</taxonomicName>
(with 4955
<abbrev id="ABBRID0ENUBG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
; strictly halophilic, only present on the Mediterranean seashores) and
<taxonomicName id="6FBFB582C832D8ED71DA8454ADF60F1E" authority="(Brolemann, 1900)" baseAuthorityName="Brolemann" baseAuthorityYear="1900" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="zograffi">
<emphasis id="A7E86CFE2B10B925BABB1BBE41E4BB86" italics="true">T. zograffi</emphasis>
(Brolemann, 1900)
</taxonomicName>
(47
<abbrev id="ABBRID0E3UBG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
; not halophilic; only recorded in the Canary islands).]
</td>
<td id="7F8520E38A73F000834CDF7394191A81" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="EA19B1DD0E12334F083FF73A5062F32D">
<td id="96E1D2F10FD9474CA5975394B14264BD" colspan="1" rowspan="1">12</td>
<td id="71EE39172613018EFDC60BC17F5D65DB" colspan="1" rowspan="1">
More than 90
<abbrev id="ABBRID0EIVBG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
. Pore-fields sub-ovoid / sub-triangular, approx. as long as wide or slightly longer than wide (Fig.
<figureCitation id="68844635C103691F30910EA4D6AA04CF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Endogeophilus alberti sp. nov., holotype A labrum B first maxillae C right part of the forcipular segment D LBS 18 E ultimate LBS without distal part of legs F left part of the ultimate LBS without distal part of leg. Views: ventral (A E), dorsal (F). Scale bars: 20 µm (A); 40 µm (B); 100 µm (C F)." figureDoi="10.3897/zookeys.1213.133171.figure2" httpUri="https://binary.pensoft.net/fig/1142653">2 D</figureCitation>
)
</td>
<td id="07B459B27BA4B24DD277BE030F09C2A4" colspan="1" rowspan="1">
<taxonomicName id="7110278D3287C5CC322DBA888B793581" authority="Bonato, Zapparoli, Drago &amp; Minelli, 2016" authorityName="Bonato, Zapparoli, Drago &amp; Minelli" authorityYear="2016" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="CD509CC30DA7F80DEC0D8E6F4546D528" bold="true">
<emphasis id="7426E2FA1E1C9046FA639FF32E54837A" bold="true" italics="true">Endogeophilus</emphasis>
<bibRefCitation id="DEE0404E7D6C4A11AF1A2CF04D1BD993" DOI="10.5852/ejt.2016.231" author="Bonato L &amp; Zapparoli M &amp; Drago L &amp; Minelli A" journalOrPublisher="European Journal of Taxonomy" pagination="1-19" refId="B14" refString="Bonato L, Zapparoli M, Drago L, Minelli A (2016) An unusually elongate endogeic centipede from Sardinia (Chilopoda: Geophilidae). European Journal of Taxonomy 231: 1 - 19. https: // doi. org / 10.5852 / ejt. 2016.231" title="An unusually elongate endogeic centipede from Sardinia (Chilopoda: Geophilidae)." volume="231" year="2016">Bonato, Zapparoli, Drago &amp; Minelli, 2016</bibRefCitation>
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="B1350F04ED40CD528ED81A2E2755C319">
<td id="8B75BFCF2D3964C2B1781373A2AB5AE8" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="BC2FC323993D4CCC2E84CBFF972CC660" colspan="1" rowspan="1">
[In Europe: only 2 species,
<taxonomicName id="1F21A6336BFBAA50F048B522A637D6BA" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="ichnusae">
<emphasis id="470E712B5C68DE9E1B483D3DAA1DAE96" italics="true">E. ichnusae</emphasis>
</taxonomicName>
from Sardinia and
<taxonomicName id="A1FAFA09647494F2656DBD8828FA359E" authorityName="Iorio &amp; Bonato" authorityYear="2024" class="Chilopoda" family="Geophilidae" genus="Endogeophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
<emphasis id="E74DC1A4BD491D16ECE183266FA53A08" italics="true">E. alberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9757956E06254957444D19C4DFDC16B0" rank="species">sp. nov.</taxonomicNameLabel>
from southern Provence.]
</td>
<td id="10966F990DB9F2A13EA05816573041C2" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="CEA8A4E06D7A97BC2C619A7D38EE304B">
<td id="DEC12CC14D62DBF5D2CF1A34C1DFDAF0" colspan="1" rowspan="1"></td>
<td id="DBA16ADB78D57B8482F14A9FF7999387" colspan="1" rowspan="1">
Usually &lt;90
<abbrev id="ABBRID0EBXBG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
<superScript id="1CEAECE32C37EB24D726DADA6719DB28">
<xref id="634EBDD31D288934D111BD5A740C897E" ref-type="fn" rid="en1">1</xref>
</superScript>
. Pore-fields not sub-ovoid / sub-triangular (Fig.
<figureCitation id="47887CA158A55B38B608C8690D945A77" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 C F</figureCitation>
: pf) or absent (Fig.
<figureCitation id="E075E614391C71180861FB55E9EAEAB1" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 B</figureCitation>
)
</td>
<td id="0EAC37A8E957FA4072EE1304CE737B14" colspan="1" rowspan="1">
<emphasis id="CA478AC7FCCA4E961F6DE26F27FC726C" bold="true">13</emphasis>
</td>
</tr>
<tr id="EF5930724249F4FC1B1A275B10B37447">
<td id="AF425BE6FC87A5ED236BE754A3440DAD" colspan="1" rowspan="1">13</td>
<td id="8AD5AA46EEC1AB079A0C5B729CBF2D3C" colspan="1" rowspan="1">
Legs of the ultimate pair without an apical claw (Fig.
<figureCitation id="1CCDC48C4F521122C3CC4A7DFFFDEF22" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Legs of the ultimate pair A, C right leg B, D distal articles of the right leg E G both legs H left leg. Species A Arctogeophilus inopinatus male B Arenophilus peregrinus female C Eurygeophilus pinguis female D Geophilus fucorum female E Geophilus richardi female F Pachymerium ferrugineum female G Pleurogeophilus mediterraneus male H Pleurogeophilus mediterraneus female I Acanthogeophilus spiniger female. Views: ventral (A-E, I), lateral (F) and dorsal (G-H). Abbreviations: c claw, t tubercle with spines. Scale bars: 300 µm (A, C, F); 50 µm (B); 100 µm (D, E, I); 1 mm (G H)." figureDoi="10.3897/zookeys.1213.133171.figure10" httpUri="https://binary.pensoft.net/fig/1142661">10 A, G, H</figureCitation>
). Pore-fields either absent (Fig.
<figureCitation id="EE4B273CE0F64BB8F1402A4B82D26CC9" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 B</figureCitation>
) or small, sub-circular to slightly elliptical (Fig.
<figureCitation id="502AF0254350551B0423FADCB6D2737D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 F</figureCitation>
: pf)
</td>
<td id="FAAE556FACCD7B5623E8C8B3A1B6501E" colspan="1" rowspan="1">
<emphasis id="E4F5519D19D059F348864B29586EB2DB" bold="true">14</emphasis>
</td>
</tr>
<tr id="AD06C398C40F6AFD71476568D5E366DA">
<td id="8893577D759C83A8EF1F6AAD06CA5272" colspan="1" rowspan="1"></td>
<td id="FA3D574AA8B0EDA523D40A12A3706069" colspan="1" rowspan="1">
Legs of the ultimate pair with an apical claw, usually well-developed
<superScript id="F19946449202EEB6E73C44DD26DC7C41">
<xref id="D07D7CBF847969A18010844601783B09" ref-type="fn" rid="en2">2</xref>
</superScript>
(Fig.
<figureCitation id="5AE80D5142D211C7A11945F9A1EBCD0C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Legs of the ultimate pair A, C right leg B, D distal articles of the right leg E G both legs H left leg. Species A Arctogeophilus inopinatus male B Arenophilus peregrinus female C Eurygeophilus pinguis female D Geophilus fucorum female E Geophilus richardi female F Pachymerium ferrugineum female G Pleurogeophilus mediterraneus male H Pleurogeophilus mediterraneus female I Acanthogeophilus spiniger female. Views: ventral (A-E, I), lateral (F) and dorsal (G-H). Abbreviations: c claw, t tubercle with spines. Scale bars: 300 µm (A, C, F); 50 µm (B); 100 µm (D, E, I); 1 mm (G H)." figureDoi="10.3897/zookeys.1213.133171.figure10" httpUri="https://binary.pensoft.net/fig/1142661">10 C, D</figureCitation>
: c, E, F: c, I). Pore-fields usually present
<superScript id="155BF98F38E126882B02F86C1BBF6795">
<xref id="2477AE2E8C72BD942E2F5FF984CBC3E1" ref-type="fn" rid="en3">3</xref>
</superScript>
, sub-diamond or in a tranverse band (Fig.
<figureCitation id="4D5CB6A5F2CD10A1A47FF7ABEC00772A" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 D, E</figureCitation>
: pf)
</td>
<td id="FEF66C51DBEFC00B9268C41A517B84B3" colspan="1" rowspan="1">
<emphasis id="68CA8EAADE25DAEE3E9FDC1CE54D721F" bold="true">17</emphasis>
</td>
</tr>
<tr id="E639CE41481E4114D8008B19CB3271E8">
<td id="472D5CE0EC7BD6DC312E5A17261D3A2C" colspan="1" rowspan="1">14</td>
<td id="7B220BA1F7B829C0D22E88516C3459D7" colspan="1" rowspan="1">
Pore-fields present, sub-circular to slightly elliptical (Fig.
<figureCitation id="48A68EF03AC201EF4FB6390CD543B1DB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 F</figureCitation>
: pf). Legs of the ultimate pair very elongated,&gt; 2 × as long as the penultimate legs (Fig.
<figureCitation id="E8FDEA4864F49A2A686ADA6C85983A66" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Legs of the ultimate pair A, C right leg B, D distal articles of the right leg E G both legs H left leg. Species A Arctogeophilus inopinatus male B Arenophilus peregrinus female C Eurygeophilus pinguis female D Geophilus fucorum female E Geophilus richardi female F Pachymerium ferrugineum female G Pleurogeophilus mediterraneus male H Pleurogeophilus mediterraneus female I Acanthogeophilus spiniger female. Views: ventral (A-E, I), lateral (F) and dorsal (G-H). Abbreviations: c claw, t tubercle with spines. Scale bars: 300 µm (A, C, F); 50 µm (B); 100 µm (D, E, I); 1 mm (G H)." figureDoi="10.3897/zookeys.1213.133171.figure10" httpUri="https://binary.pensoft.net/fig/1142661">10 G, H</figureCitation>
)
</td>
<td id="7F0C52C01D92200F8DF26EE15668F8DE" colspan="1" rowspan="1">
<taxonomicName id="0DB8E94825A608DA69A8C8BFF390F78A" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" class="Chilopoda" family="Geophilidae" genus="Pleurogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="A7E9101BB08B65808948992FA4DDE26D" bold="true">
<emphasis id="2FCED16D0ED1426D8F3E11ACDA40B175" bold="true" italics="true">Pleurogeophilus</emphasis>
Verhoeff, 1901
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="5446133CCA782AC0B3792374428CEBC9">
<td id="D753FBAF98474A3014381DCE380313AE" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="3F9C0052B7A944228C6FDAE0455896B6" colspan="1" rowspan="1">
[In Europe: at least one species,
<taxonomicName id="6F844C6BB2F3F2BAE7699C145C905A03" authority="(Meinert, 1870)" baseAuthorityName="Meinert" baseAuthorityYear="1870" class="Chilopoda" family="Geophilidae" genus="Pleurogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="mediterraneus">
<emphasis id="2E9F9A2616A2FF743F89300458782351" italics="true">P. mediterraneus</emphasis>
(Meinert, 1870)
</taxonomicName>
; in south-eastern Europe; 6185
<abbrev id="ABBRID0E61BG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
; numerous coxal pores (frequently&gt; 25 on each coxopleuron in adults) and the ultimate metasternite longer than wide (Fig.
<figureCitation id="33D0B057512D9F7918990A7012CCE20A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Ultimate LBS without distal part of legs. Species A Arctogeophilus inopinatus female B Galliophilus beatensis male C Geophilus flavus female D Pachymerium ferrugineum male E Pleurogeophilus mediterraneus female F Pleurogeophilus mediterraneus male G, H Stenotaenia linearis female I Tuoba poseidonis female. Views: ventral (A G), lateral (H, I). Abbreviations: p pit or fossa (H wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate LBS (medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm." figureDoi="10.3897/zookeys.1213.133171.figure9" httpUri="https://binary.pensoft.net/fig/1142660">9 E, F</figureCitation>
).]
</td>
<td id="405EEC37517CBF097517010A8F4B4920" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="246ED0BF6D3AFE1C64FF43524AD92DC2">
<td id="77322A8566C0810C5E91DB8D265AD371" colspan="1" rowspan="1"></td>
<td id="2B0588D3911F6B754F80E147FC5C2EE4" colspan="1" rowspan="1">
No pore-fields (Fig.
<figureCitation id="834BF2532A358932524CBF7E176FE8FE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 B</figureCitation>
). Legs of the ultimate pair usually &lt;2 × as long as the penultimate legs
<superScript id="7504CEE0816670B9966B59D72F29B169">
<xref id="059A78B3C531865E461214CB7CE1446F" ref-type="fn" rid="en4">4</xref>
</superScript>
</td>
<td id="463AEEBDBCA6A103A7EFC17CCE11ABBC" colspan="1" rowspan="1">
<emphasis id="9146CCEB2FA76C9D1FC626CB69FE3C69" bold="true">15</emphasis>
</td>
</tr>
<tr id="BB8955427DA65D7CC693DFB90DA87830">
<td id="79CA3890FA68CB890F8992C345863577" colspan="1" rowspan="1">15</td>
<td id="97BDA08859D819AA5CD90A38E2702B73" colspan="1" rowspan="1">
Forcipular trochanteroprefemur ~ as long as wide and without denticle (Fig.
<figureCitation id="5AC916C583A94A58AC9CED919C3E3A77" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 F</figureCitation>
: tp). Metasternite of the ultimate leg-bearing segment wider than long (Fig.
<figureCitation id="42D474F17E8312B24F062992346EDB16" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Ultimate LBS without distal part of legs. Species A Arctogeophilus inopinatus female B Galliophilus beatensis male C Geophilus flavus female D Pachymerium ferrugineum male E Pleurogeophilus mediterraneus female F Pleurogeophilus mediterraneus male G, H Stenotaenia linearis female I Tuoba poseidonis female. Views: ventral (A G), lateral (H, I). Abbreviations: p pit or fossa (H wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate LBS (medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm." figureDoi="10.3897/zookeys.1213.133171.figure9" httpUri="https://binary.pensoft.net/fig/1142660">9 B</figureCitation>
)
</td>
<td id="48447B0C4F13D683996906B6DD00389E" colspan="1" rowspan="1">
<taxonomicName id="EBB29F6797B3415A3A3566F73B904708" authority="Ribaut &amp; Brolemann, 1927" authorityName="Ribaut &amp; Brolemann" authorityYear="1927" class="Chilopoda" family="Geophilidae" genus="Galliophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="030BFAA6CD1D9305B4766D9F0196BD01" bold="true">
<emphasis id="21B2F7BB7B10F29EEC60B367FD166CEF" bold="true" italics="true">Galliophilus</emphasis>
Ribaut &amp; Brolemann, 1927
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="D15FFA81719B9F3281BBB1606A69755B">
<td id="AE9D468EA8848D4C87460676ECAFFA8E" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="8EE421024A675E02D792F5C41B9FDA4D" colspan="1" rowspan="1">
[In Europe: only one species,
<taxonomicName id="7130A7C25CC76F4ACA53CF0D1BB0E533" authorityName="Ribaut &amp; Brolemann" authorityYear="1927" class="Chilopoda" family="Geophilidae" genus="Galliophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="beatensis">
<emphasis id="69319CE025DCB418116D88C9D8F51110" italics="true">G. beatensis</emphasis>
</taxonomicName>
, recorded confidently only in the eastern Pyrenees; ~ 8185
<abbrev id="ABBRID0EL4BG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
; tarsungulum sharply compressed in the internal side, making almost a right angle in horizontal view (Fig.
<figureCitation id="9B1F3C86393CC8D8D6D91D5D6FD296B0" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 F</figureCitation>
).]
</td>
<td id="DC4B73D03E9866220B8ABDABA50AECE6" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="E30837D4962137617F8D47418EAC14D1">
<td id="2ADCF9397802B6D0A9C97957276C63D5" colspan="1" rowspan="1"></td>
<td id="71023017AA34B4B5D600B380CFFD8ABD" colspan="1" rowspan="1">
Forcipular trochanteroprefemur much longer than wide and with a denticle (Figs
<figureCitation id="6FC93B995F06A26203DAAD6F578C82F4" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 I</figureCitation>
,
<figureCitation id="3519C051036478DE6EE7C07BEE25676D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 C</figureCitation>
: tp). Metasternite of the ultimate leg-bearing segment longer than wide (Fig.
<figureCitation id="C5DE9A1F846102487F956DAA46C3D96A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Ultimate LBS without distal part of legs. Species A Arctogeophilus inopinatus female B Galliophilus beatensis male C Geophilus flavus female D Pachymerium ferrugineum male E Pleurogeophilus mediterraneus female F Pleurogeophilus mediterraneus male G, H Stenotaenia linearis female I Tuoba poseidonis female. Views: ventral (A G), lateral (H, I). Abbreviations: p pit or fossa (H wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate LBS (medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm." figureDoi="10.3897/zookeys.1213.133171.figure9" httpUri="https://binary.pensoft.net/fig/1142660">9 A</figureCitation>
)
</td>
<td id="9622EE48F72E6B55EC619813AF0322CC" colspan="1" rowspan="1">
<emphasis id="A7BA853B2F8B822E126CF88986363F6F" bold="true">16</emphasis>
</td>
</tr>
<tr id="14875F7AF6A5C758B3335F23E97BC60A">
<td id="D8EA9ACE17EA1E23146C60CDE26094CC" colspan="1" rowspan="1">16</td>
<td id="E915E5C725EA08514F3961DF36327ADC" colspan="1" rowspan="1">
3941
<abbrev id="ABBRID0EV5BG" xlink_title="leg-bearing segment(s)">LBS</abbrev>
. Forcipular coxosternite slightly wider than long. All forcipular articles with a denticle (Fig.
<figureCitation id="74E6C1FBBF88A6A8EF1295B8058D2E45" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 C</figureCitation>
)
</td>
<td id="ABD51B706D4EB63D1C633C33C3B008C9" colspan="1" rowspan="1">
<taxonomicName id="4DC7BF0EC4297B35D4F47F0C8F329E05" authority="Attems, 1909" authorityName="Attems" authorityYear="1909" class="Chilopoda" family="Geophilidae" genus="Arctogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="C0C866B98A62D437391C033E9DB1763D" bold="true">
<emphasis id="B6677BCC208D9348D88AF75C4E055415" bold="true" italics="true">Arctogeophilus</emphasis>
Attems, 1909
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="D1B23F32BED55BA9C4BF25E9240A538A">
<td id="707050DA966EBA1A9F597B070588BA9D" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="E53092B53B803AABEB2C3EACA397C82C" colspan="1" rowspan="1">
[In Europe: three species,
<taxonomicName id="507817FE1E12C2FA6F14809E7AAA88AE" authority="(Ribaut, 1911)" baseAuthorityName="Ribaut" baseAuthorityYear="1911" class="Chilopoda" family="Geophilidae" genus="Arctogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="inopinatus">
<emphasis id="E04379ABD90F2EDEAC0C33038344629B" italics="true">A. inopinatus</emphasis>
(Ribaut, 1911)
</taxonomicName>
(western and central France),
<taxonomicName id="3B32C80FCF23C3FC378BCB5D8CAD94C2" authority="Folkmanova, 1956" authorityName="Folkmanova" authorityYear="1956" class="Chilopoda" family="Geophilidae" genus="Arctogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="attemsi">
<emphasis id="7A2F0F1D1636ACBA037EFFCBEB964768" italics="true">A. attemsi</emphasis>
Folkmanová, 1956
</taxonomicName>
, and
<taxonomicName id="7F995E238205A60C64EF1642C53F696F" authority="Folkmanova &amp; Dobroruka, 1960" authorityName="Folkmanova &amp; Dobroruka" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Arctogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="macrocephalus">
<emphasis id="20B82065518179A49A0A45D65795FE1E" italics="true">A. macrocephalus</emphasis>
Folkmanova &amp; Dobroruka, 1960
</taxonomicName>
(both only recorded in Ukraine).]
</td>
<td id="5F0D6FF36C6567795CA15B872EF0EFD5" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="C354CDAD3EEC01B66099EC6FF626B654">
<td id="1187CB3BCC7F6DD75783DEA7357304A0" colspan="1" rowspan="1"></td>
<td id="1D418DED5008DB7E943D73882A21C4BD" colspan="1" rowspan="1">
6787
<abbrev id="ABBRID0E1AAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
. Forcipular coxosternite slightly longer than wide. Forcipular trochanteroprefemur and tarsungulum with a denticle, intermediate forcipular articles without denticle (Fig.
<figureCitation id="8C00C10506AF73CEB3CAEBCB93546EBD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and forcipular segment. Species A Pachymerium ferrugineum B Gnathoribautia bonensis C Pleurogeophilus mediterraneus D Strigamia carniolensis E Henia vesuviana F Strigamia carniolensis G Dignathodon microcephalus H Henia bicarinata I Gnathoribautia bonensis. Views: dorsal (A E), ventral (F I). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (A F, H-I); 300 µm (G)." figureDoi="10.3897/zookeys.1213.133171.figure6" httpUri="https://binary.pensoft.net/fig/1142657">6 I</figureCitation>
)
</td>
<td id="A6B26D8C6E4345320EBE4C38473E68FC" colspan="1" rowspan="1">
<taxonomicName id="F6AB93B6B98EABDB84F89EC41F4FD319" authority="Brolemann, 1909" authorityName="Brolemann" authorityYear="1909" class="Chilopoda" family="Geophilidae" genus="Gnathoribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="C562335084F75D0C36AFED968EEDF30A" bold="true">
<emphasis id="381082F9D41EA49A73E77C35773D7FFB" bold="true" italics="true">Gnathoribautia</emphasis>
Brolemann, 1909
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="F565A54473C100E1D311CD5C3C3384DD">
<td id="9386E74094D8AAB767AA762C0915BA6E" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="F2A91452A0D877DF4241AB6BCDCFEE91" colspan="1" rowspan="1">
[In Europe: only 2 species,
<taxonomicName id="9A49DF88B2A8CFE6DCF61F70F330AB43" baseAuthorityName="Meinert" baseAuthorityYear="1870" class="Chilopoda" family="Geophilidae" genus="Gnathoribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="bonensis">
<emphasis id="585898D9139250723E9243425E465285" italics="true">G. bonensis</emphasis>
</taxonomicName>
(6783
<abbrev id="ABBRID0EACAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
; body length ≤ 7 cm; Macaronesia, Iberian peninsula, and Sicily) and
<taxonomicName id="0D3ED4054AB69FF18E706EDC84C3CF74" authority="(Attems, 1903)" baseAuthorityName="Attems" baseAuthorityYear="1903" class="Chilopoda" family="Geophilidae" genus="Gnathoribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="syriaca">
<emphasis id="1E9CADD5B620298D2B1ECC7398C22F04" italics="true">G. syriaca</emphasis>
(Attems, 1903)
</taxonomicName>
(~ 87
<abbrev id="ABBRID0EPCAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
; body length ≤ 11 cm; Aegean islands]
</td>
<td id="CCBCAF9B7236B1DA0620B952B144C47B" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="EDA4627782805CE561D2B8D5ACBDE189">
<td id="871D98B444D86AD965E4E9CFA486B554" colspan="1" rowspan="1">17</td>
<td id="4025C7A7853930EC0C32C2B004EB2373" colspan="1" rowspan="1">
Metasternite of the ultimate leg-bearing segment longer than wide (Figs
<figureCitation id="87F45A3A087FA1DACC9BB5EEF7572D51" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Ultimate LBS without distal part of legs. Species A Arctogeophilus inopinatus female B Galliophilus beatensis male C Geophilus flavus female D Pachymerium ferrugineum male E Pleurogeophilus mediterraneus female F Pleurogeophilus mediterraneus male G, H Stenotaenia linearis female I Tuoba poseidonis female. Views: ventral (A G), lateral (H, I). Abbreviations: p pit or fossa (H wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate LBS (medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm." figureDoi="10.3897/zookeys.1213.133171.figure9" httpUri="https://binary.pensoft.net/fig/1142660">9 D</figureCitation>
,
<figureCitation id="690DE36CF377924F5049B26EFD294E44" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Legs of the ultimate pair A, C right leg B, D distal articles of the right leg E G both legs H left leg. Species A Arctogeophilus inopinatus male B Arenophilus peregrinus female C Eurygeophilus pinguis female D Geophilus fucorum female E Geophilus richardi female F Pachymerium ferrugineum female G Pleurogeophilus mediterraneus male H Pleurogeophilus mediterraneus female I Acanthogeophilus spiniger female. Views: ventral (A-E, I), lateral (F) and dorsal (G-H). Abbreviations: c claw, t tubercle with spines. Scale bars: 300 µm (A, C, F); 50 µm (B); 100 µm (D, E, I); 1 mm (G H)." figureDoi="10.3897/zookeys.1213.133171.figure10" httpUri="https://binary.pensoft.net/fig/1142661">10 I</figureCitation>
)
</td>
<td id="B470E462620DE05755021970142F816C" colspan="1" rowspan="1">
<emphasis id="2D37D449ACA51B6689EEC7C49FFEA90C" bold="true">18</emphasis>
</td>
</tr>
<tr id="181B354E16EA310416EB9E1C9DE4A5CA">
<td id="2518D19102AB601D7DAD13BA15A6DC74" colspan="1" rowspan="1"></td>
<td id="F15C5B3DA26E3A41621C8EF49C5D02F4" colspan="1" rowspan="1">
Metasternite of the ultimate leg-bearing segment wider than long (Fig.
<figureCitation id="D4E2ED4F5B3674D14417C0A8953C8E53" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Ultimate LBS without distal part of legs. Species A Arctogeophilus inopinatus female B Galliophilus beatensis male C Geophilus flavus female D Pachymerium ferrugineum male E Pleurogeophilus mediterraneus female F Pleurogeophilus mediterraneus male G, H Stenotaenia linearis female I Tuoba poseidonis female. Views: ventral (A G), lateral (H, I). Abbreviations: p pit or fossa (H wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate LBS (medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm." figureDoi="10.3897/zookeys.1213.133171.figure9" httpUri="https://binary.pensoft.net/fig/1142660">9 C</figureCitation>
)
</td>
<td id="47D59026FE24829CC1030D956A072788" colspan="1" rowspan="1">
<emphasis id="C67D29CBCB2D41D58EFC3B00E53A6D9B" bold="true">19</emphasis>
</td>
</tr>
<tr id="FCCAA72CC77A076E9BFD50ACA7A86603">
<td id="3C2359B1D0EB95909157829C7D640D31" colspan="1" rowspan="1">18</td>
<td id="21CAF2497F853A212731322A88BE05E7" colspan="1" rowspan="1">
Forcipular trochanteroprefemur and tarsungulum each with a denticle. Legs of the ultimate leg-bearing segment without elongate projections (Fig.
<figureCitation id="178733EBC625BFC8C072AA80DB5E84AA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Legs of the ultimate pair A, C right leg B, D distal articles of the right leg E G both legs H left leg. Species A Arctogeophilus inopinatus male B Arenophilus peregrinus female C Eurygeophilus pinguis female D Geophilus fucorum female E Geophilus richardi female F Pachymerium ferrugineum female G Pleurogeophilus mediterraneus male H Pleurogeophilus mediterraneus female I Acanthogeophilus spiniger female. Views: ventral (A-E, I), lateral (F) and dorsal (G-H). Abbreviations: c claw, t tubercle with spines. Scale bars: 300 µm (A, C, F); 50 µm (B); 100 µm (D, E, I); 1 mm (G H)." figureDoi="10.3897/zookeys.1213.133171.figure10" httpUri="https://binary.pensoft.net/fig/1142661">10 F</figureCitation>
)
</td>
<td id="B31CE22FA487256B83A89F86368345DE" colspan="1" rowspan="1">
<taxonomicName id="B1B76E9811D063043895AE9B8E3B0DB5" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="C5CAE35364CF42474175689388CA0C23" bold="true">
<emphasis id="650FBF37D2A8B23AF49ECFC555761496" bold="true" italics="true">Pachymerium</emphasis>
C. L. Koch, 1847
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3FF13A3109FC871CF4FEA8184135F47F">
<td id="D4050FF3B4961F7E880A1CE463248EA4" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="2ECF28E2BF2DF482EBF929D41E3153DD" colspan="1" rowspan="1">
[In Europe: few species, across most of Europe. Numerous coxal pores,&gt; 20 widely distributed on each coxopleuron in adults (Fig.
<figureCitation id="A2F431F4D0C667BCC320737BB6E30046" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Ultimate LBS without distal part of legs. Species A Arctogeophilus inopinatus female B Galliophilus beatensis male C Geophilus flavus female D Pachymerium ferrugineum male E Pleurogeophilus mediterraneus female F Pleurogeophilus mediterraneus male G, H Stenotaenia linearis female I Tuoba poseidonis female. Views: ventral (A G), lateral (H, I). Abbreviations: p pit or fossa (H wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate LBS (medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm." figureDoi="10.3897/zookeys.1213.133171.figure9" httpUri="https://binary.pensoft.net/fig/1142660">9 D</figureCitation>
).]
</td>
<td id="E10400881E07AAF787BDB237D6AACEEE" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="C4E9ACF75589FF39AB39FE6CE04DA126">
<td id="BE3621485D6A68E35EDCB69EF4D130EE" colspan="1" rowspan="1"></td>
<td id="7251BF531D3F5D7F9FE6008F738CC899" colspan="1" rowspan="1">
Forcipules without denticles. Legs of the ultimate leg-bearing segment with elongate projections (Fig.
<figureCitation id="9225974B6141FBEAF47602E5C2E0AB0A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Legs of the ultimate pair A, C right leg B, D distal articles of the right leg E G both legs H left leg. Species A Arctogeophilus inopinatus male B Arenophilus peregrinus female C Eurygeophilus pinguis female D Geophilus fucorum female E Geophilus richardi female F Pachymerium ferrugineum female G Pleurogeophilus mediterraneus male H Pleurogeophilus mediterraneus female I Acanthogeophilus spiniger female. Views: ventral (A-E, I), lateral (F) and dorsal (G-H). Abbreviations: c claw, t tubercle with spines. Scale bars: 300 µm (A, C, F); 50 µm (B); 100 µm (D, E, I); 1 mm (G H)." figureDoi="10.3897/zookeys.1213.133171.figure10" httpUri="https://binary.pensoft.net/fig/1142661">10 I</figureCitation>
)
</td>
<td id="580CF6F31729428C8652EAF1651DA541" colspan="1" rowspan="1">
<taxonomicName id="893927D548D81430754EA0E70B326C6D" authority="Minelli, 1982" authorityName="Minelli" authorityYear="1982" class="Chilopoda" family="Geophilidae" genus="Acanthogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="198DDE7D7C41B053EBBD622440E76524" bold="true">
<emphasis id="68175FAD6936CF6C535EE470DC52F638" bold="true" italics="true">Acanthogeophilus</emphasis>
<bibRefCitation id="CAB783DB1D5EE0614C97A9B06468E726" author="Minelli A" journalOrPublisher="Bollettino del Museo Civico di Storia Naturale di Verona" pagination="235-240" refId="B40" refString="Minelli A (1982) Un nuovo chilopodo italiano Acanthogeophilus dentifer, n. g. n. sp. (Geophilomorpha Geophilidae). Bollettino del Museo Civico di Storia Naturale di Verona 8: 235 - 240." title="Un nuovo chilopodo italiano Acanthogeophilus dentifer, n. g. n. sp. (Geophilomorpha Geophilidae)." volume="8" year="1982">Minelli, 1982</bibRefCitation>
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="A7ADBA50365ABBABB58448EB2DF12B65">
<td id="A7B1E800837CDE0990A35E766496441A" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="CB374F2D155E85C454B108E9EAD4D0EB" colspan="1" rowspan="1">
[In Europe: only 1 species,
<taxonomicName id="08BDD60DDBBFE54A9C7DD9F73A9D8866" authority="Minelli, 1982" authorityName="Minelli" authorityYear="1982" class="Chilopoda" family="Geophilidae" genus="Acanthogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dentifer">
<emphasis id="E318479A9D3BD9042DF5A47105EC8031" italics="true">A. dentifer</emphasis>
<bibRefCitation id="F3582F3F12C61C8D1C9A2E3F1D8EF29B" author="Minelli A" journalOrPublisher="Bollettino del Museo Civico di Storia Naturale di Verona" pagination="235-240" refId="B40" refString="Minelli A (1982) Un nuovo chilopodo italiano Acanthogeophilus dentifer, n. g. n. sp. (Geophilomorpha Geophilidae). Bollettino del Museo Civico di Storia Naturale di Verona 8: 235 - 240." title="Un nuovo chilopodo italiano Acanthogeophilus dentifer, n. g. n. sp. (Geophilomorpha Geophilidae)." volume="8" year="1982">Minelli, 1982</bibRefCitation>
</taxonomicName>
, only recorded in the Italian peninsula; ~ 67
<abbrev id="ABBRID0EIGAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
.]
</td>
<td id="FB4AB555F72A8E4068C1E44E55FAFE90" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="F3384742E45C6B42D41A4E83DE003268">
<td id="F56B00DB6F1D5506EE140EA572086197" colspan="1" rowspan="1">19</td>
<td id="2D1D5C2E36BA27C12E3FD2D6701CEAEA" colspan="1" rowspan="1">
Forcipular coxosternite&gt; 2 × as wide as long (Fig.
<figureCitation id="426B8275380DCBDDABD7F8560E7A5C2C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 D</figureCitation>
). Tarsungulum strongly narrowing near the base, very narrow for most of its length and distinctly compressed also along the external side (Fig.
<figureCitation id="2484CEEF9F747D946BF654B4F871DCDB" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 D</figureCitation>
)
</td>
<td id="44E89BFEFBEE63A8703993A84645C926" colspan="1" rowspan="1">
<taxonomicName id="81B19CFA7DB0FF315C2D56A8B951805F" authority="Verhoeff, 1899" authorityName="Verhoeff" authorityYear="1899" class="Chilopoda" family="Geophilidae" genus="Eurygeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="639AEB6E5446427DB2D09D806B89E3FB" bold="true">
<emphasis id="50FA152D2059749D5EF8B197D1027F64" bold="true" italics="true">Eurygeophilus</emphasis>
Verhoeff, 1899
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="5A2C318C5F7960557C9315BC32DE4CA9">
<td id="DC82ED366E7BD7B032BFCAC03DBD539C" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="822D58A126B28489BDC0F45FC312099C" colspan="1" rowspan="1">
[In Europe: only 2 species,
<taxonomicName id="7917B073F40A2660C2CB32207116FDB6" authority="(Brolemann, 1898)" baseAuthorityName="Brolemann" baseAuthorityYear="1898" class="Chilopoda" family="Geophilidae" genus="Eurygeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pinguis">
<emphasis id="F7E96E6947A02506183A9A843F733C68" italics="true">E. pinguis</emphasis>
(Brolemann, 1898)
</taxonomicName>
(&lt;50
<abbrev id="ABBRID0E1HAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
) and
<taxonomicName id="F70B07A4AD1171E18C4BA55A741B86D1" authority="(Verhoeff, 1899)" baseAuthorityName="Verhoeff" baseAuthorityYear="1899" class="Chilopoda" family="Geophilidae" genus="Eurygeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="multistiliger">
<emphasis id="3C469D62628583D8EA21A184FD69058A" italics="true">E. multistiliger</emphasis>
(Verhoeff, 1899)
</taxonomicName>
(&gt; 50
<abbrev id="ABBRID0EJIAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
). Pore-fields in a very stretched transverse band in both species (Fig.
<figureCitation id="9661DA08A02F7D53862BB5F9725AB938" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 C</figureCitation>
: pf).]
</td>
<td id="9A44E9958F8422C4EF012FE84A26DFF4" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
<tr id="8BBAAB6DCEE740C7905D3E771BFA8753">
<td id="C5DCA23F3D81DDB3D07437EB94EF7EAB" colspan="1" rowspan="1"></td>
<td id="5BCA4A0A8704C56EDC04B0C0419722B4" colspan="1" rowspan="1">
Forcipular coxosternite &lt;2 × as wide as long (Fig.
<figureCitation id="5C945A981FD7CF5D8A1E97FD6F6091F1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 G</figureCitation>
; also F). Tarsungulum gradually narrowing and compressed only along the internal side (Fig.
<figureCitation id="0AFB5B6E69F3B3D29BDCE2131FB649F3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B labrum C, D, F, G forcipular segment E distal part of the forcipular coxosternite. Species A Clinopodes vesubiensis B Geophilus studeri C Arctogeophilus inopinatus D Eurygeophilus pinguis E Clinopodes vesubiensis F Galliophilus beatensis G Geophilus gavoyi. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (A, B); 300 µm (C G)." figureDoi="10.3897/zookeys.1213.133171.figure7" httpUri="https://binary.pensoft.net/fig/1142658">7 G</figureCitation>
)
</td>
<td id="B8F1386CC99EE0CE54101FFA018B1C87" colspan="1" rowspan="1">
<taxonomicName id="2CBF64CC57AF4791582308A900050737" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="84693F5E0C9B8B0E6831F4BE36E564F5" bold="true">
<emphasis id="0B72295D5463138CFE843D4928DBE0D7" bold="true" italics="true">Geophilus</emphasis>
Leach, 1814
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="D729F31CE00ED21B0E7048C71D427987">
<td id="EADD322DB7F37CAF0D486B6DFD1C6379" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="5AF55A3D796657531BC53E4D29775DA9" colspan="1" rowspan="1">
[In Europe: many species. With the exception of
<taxonomicName id="9FF3696297A69402780F6A384CD9FA63" authority="Leach, 1815" authorityName="Leach" authorityYear="1815" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="carpophagus">
<emphasis id="D382F50F0DBBE4AABF3195A8CD8D27AE" italics="true">G. carpophagus</emphasis>
Leach, 1815
</taxonomicName>
, which has pore-fields in a very stretched transverse band, the other species have either stretched sub-triangular / sub-diamond pore-fields (Fig.
<figureCitation id="FAD9B1CD47A5B760D29CAF561B32F5BA" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Metasternite of an anterior LBS. Species A Clinopodes vesubiensis LBS 16 B Galliophilus beatensis LBS 19 C Eurygeophilus pinguis LBS 10 D Geophilus osquidatum LBS 11 E Geophilus electricus LBS 11 F Pleurogeophilus mediterraneus LBS 15 G Stenotaenia linearis LBS 20 H Tuoba poseidonis LBS 12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm." figureDoi="10.3897/zookeys.1213.133171.figure8" httpUri="https://binary.pensoft.net/fig/1142659">8 D, E</figureCitation>
: pf) or no pore-fields
<superScript id="941A287DC389C0B5E7A1646DA0788F7D">3</superScript>
.]
</td>
<td id="F340BECB75CD50259D6AD0C35504904B" colspan="1" isEmpty="true" rowspan="1">-</td>
</tr>
</table-wrap>
</tbody>
</table>
</paragraph>
<caption id="F73254F598E5ACE5E6CD99101CD43EDF" ID-DOI="10.3897/zookeys.1213.133171.figure6" ID-arpha="65394539-469C-5487-8B1A-4BF7A2353820" httpUri="https://binary.pensoft.net/fig/1142657" startId="F6">
<paragraph id="FC945AF60F272B205ABC1D1D8D401C58">
<label id="CCD999C11769122646D2667B5A6B693E">Figure 6.</label>
</paragraph>
<paragraph id="9F4A7CFC2DEECAE55F1A7BDD99C76C38">
Head and forcipular segment. Species
<emphasis id="7F9A1009E3EB8A42CB342353EC3767E7" bold="true">A</emphasis>
<taxonomicName id="E973845360C204D86768B8F43E58445F" baseAuthorityName="C. L. Koch" baseAuthorityYear="1835" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="ferrugineum">
<emphasis id="4B9C06A77D9BBB2024A26351D193CD40" italics="true">Pachymerium ferrugineum</emphasis>
</taxonomicName>
<emphasis id="2F8AA111FDF7938A957B4DAE5E1648CC" bold="true">B</emphasis>
<taxonomicName id="9B87416692EFA5E99C9E70BF1E4B7968" baseAuthorityName="Meinert" baseAuthorityYear="1870" class="Chilopoda" family="Geophilidae" genus="Gnathoribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="bonensis">
<emphasis id="6F0A21D87F00781A80578F9AE932153C" italics="true">Gnathoribautia bonensis</emphasis>
</taxonomicName>
<emphasis id="F8F9999DEEF3ECD2EF23CC612073CD39" bold="true">C</emphasis>
<taxonomicName id="8102A362188067DFC345342789922811" baseAuthorityName="Meinert" baseAuthorityYear="1870" class="Chilopoda" family="Geophilidae" genus="Pleurogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="mediterraneus">
<emphasis id="4AA55A66399331F7E1AED73CA661641D" italics="true">Pleurogeophilus mediterraneus</emphasis>
</taxonomicName>
<emphasis id="3435E00727BA8E06920B802B9DBD6F90" bold="true">D</emphasis>
<taxonomicName id="E54EB3A619DEDB2CB311FB786B9512FE" baseAuthorityName="Verhoeff" baseAuthorityYear="1895" class="Chilopoda" family="Linotaeniidae" genus="Strigamia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="carniolensis">
<emphasis id="4C4F97568B8E5A5F47BA9712D95E49A9" italics="true">Strigamia carniolensis</emphasis>
</taxonomicName>
<emphasis id="65140DB2332A5E3AED8812462A5B9B24" bold="true">E</emphasis>
<taxonomicName id="E5A6118417E7EB05E6FAB28B59088B4E" authorityName="Newport" authorityYear="1845" class="Chilopoda" family="Dignathodontidae" genus="Henia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="vesuviana">
<emphasis id="C83A96D5AE27BDDEDF863D53BA3A95D2" italics="true">Henia vesuviana</emphasis>
</taxonomicName>
<emphasis id="E7F87D2D4076005E820B7377BB4C60AF" bold="true">F</emphasis>
<taxonomicName id="0C93687ABF1F441E7CF902218D2A5B5C" baseAuthorityName="Verhoeff" baseAuthorityYear="1895" class="Chilopoda" family="Linotaeniidae" genus="Strigamia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="carniolensis">
<emphasis id="F2C53DD4797DFC102991702088FE0CD8" italics="true">Strigamia carniolensis</emphasis>
</taxonomicName>
<emphasis id="5DB1FA18EA404047DC0A25A613DD60E5" bold="true">G</emphasis>
<taxonomicName id="9847B1D441BD6891BF91AE8286ED40C2" baseAuthorityName="Lucas" baseAuthorityYear="1846" class="Chilopoda" family="Dignathodontidae" genus="Dignathodon" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="microcephalus">
<emphasis id="B6B72DF87435ECB9DE822ADD4561129C" italics="true">Dignathodon microcephalus</emphasis>
</taxonomicName>
<emphasis id="52A424B964DFEF7F1577DD8FB2FD768F" bold="true">H</emphasis>
<taxonomicName id="34BCE867E3F5A060B1863DCE1F5AA8A5" baseAuthorityName="Meinert" baseAuthorityYear="1870" class="Chilopoda" family="Dignathodontidae" genus="Henia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="bicarinata">
<emphasis id="B4AE360A6B0EE86CDAB3D92E3272BDB7" italics="true">Henia bicarinata</emphasis>
</taxonomicName>
<emphasis id="4AD2FEB98E28556CA6D46919383BDF8E" bold="true">I</emphasis>
<taxonomicName id="3E10E64C336197D1A2023B627EDDCF7F" baseAuthorityName="Meinert" baseAuthorityYear="1870" class="Chilopoda" family="Geophilidae" genus="Gnathoribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="bonensis">
<emphasis id="228780B2F7D46E42E4F4714515992AEB" italics="true">Gnathoribautia bonensis</emphasis>
</taxonomicName>
. Views: dorsal (
<emphasis id="8AE11B48E8D7C3422DF6AF94396A6092" bold="true">A E</emphasis>
), ventral (
<emphasis id="10AFD42D9B0E4C7D59F0BA266499AFEE" bold="true">F I</emphasis>
). Abbreviations: cl chitin-lines, d forcipular denticle, dc denticles of the mesal side of the forcipular tarsungulum, ft forcipular tergite. Arrow: median diastema of the forcipular coxosternite. Scale bars: 400 µm (
<emphasis id="F41657E0336EAA38EDB6E0F29780A567" bold="true">A F, H-I</emphasis>
); 300 µm (
<emphasis id="D00FD807E110947F4014CB55F583D450" bold="true">G</emphasis>
).
</paragraph>
</caption>
<caption id="A982F8BE5594FC6D659BF1E4DF1AD3D0" ID-DOI="10.3897/zookeys.1213.133171.figure7" ID-arpha="3499865B-AA43-51B3-B0BB-7778CE923FF9" httpUri="https://binary.pensoft.net/fig/1142658" startId="F7">
<paragraph id="68E9A886DC58556EF27F159B5C5B9A12">
<label id="58A6B4F42C7BE394C42407308F7B3FFA">Figure 7.</label>
</paragraph>
<paragraph id="F9EF5150728562505782F6B07ED9B9AF">
<emphasis id="33ACE8440C2B3D0B2750ED3FA5EC1BC5" bold="true">A, B</emphasis>
labrum
<emphasis id="7630736BF446C012D1ACF404B40B9538" bold="true">C, D, F, G</emphasis>
forcipular segment
<emphasis id="E7C6A0F807FEDDEEEDE4457DBFE8D3BF" bold="true">E</emphasis>
distal part of the forcipular coxosternite. Species
<emphasis id="B023FA1CB04364C3BEFDCEE06EEEF883" bold="true">A</emphasis>
<taxonomicName id="3F227236EF32D3B65EE6A755ACE0BADA" authorityName="Bonato, Iorio &amp; Minelli" authorityYear="2011" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="vesubiensis">
<emphasis id="836B261132D2200F39B2FD1BC2BAD7A2" italics="true">Clinopodes vesubiensis</emphasis>
</taxonomicName>
<emphasis id="FBF61AE69E3CF37222E94360B3DE4AF5" bold="true">B</emphasis>
<taxonomicName id="6D2CBB188C903BC59E5D46CD0F6F593D" authorityName="Rothenbuhler" authorityYear="1899" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="studeri">
<emphasis id="E6F64221A25862DB8411C5ACBECA277F" italics="true">Geophilus studeri</emphasis>
</taxonomicName>
<emphasis id="19B86E067769FB80E1DC7E94C29E6BFE" bold="true">C</emphasis>
<taxonomicName id="26F9548B6A716A1932D83DDFAAD4ECE6" baseAuthorityName="Ribaut" baseAuthorityYear="1911" class="Chilopoda" family="Geophilidae" genus="Arctogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="inopinatus">
<emphasis id="C4FD272F541B11AC049509D8AA890377" italics="true">Arctogeophilus inopinatus</emphasis>
</taxonomicName>
<emphasis id="31E1AFB8F44F1511F38EE22DF0C1ECD7" bold="true">D</emphasis>
<taxonomicName id="88664EDFA83A3CF04A2CF121D6AA4840" baseAuthorityName="Brolemann" baseAuthorityYear="1898" class="Chilopoda" family="Geophilidae" genus="Eurygeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pinguis">
<emphasis id="5D8CB9B1967CDF29C51625200EB9364A" italics="true">Eurygeophilus pinguis</emphasis>
</taxonomicName>
<emphasis id="B0FBB3F48792257819509A2E0B6479EB" bold="true">E</emphasis>
<taxonomicName id="45230AA7CC23208053765E626E45CC3A" authorityName="Bonato, Iorio &amp; Minelli" authorityYear="2011" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="vesubiensis">
<emphasis id="91F6705E6B8D7D0D80069ECE77DB5967" italics="true">Clinopodes vesubiensis</emphasis>
</taxonomicName>
<emphasis id="5B7861C40887A5AA2CB1916C36EE9548" bold="true">F</emphasis>
<taxonomicName id="80DD11241337D60726132B84655FA29C" authorityName="Ribaut &amp; Brolemann" authorityYear="1927" class="Chilopoda" family="Geophilidae" genus="Galliophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="beatensis">
<emphasis id="DADE634F824E8C693B88B6F68291F688" italics="true">Galliophilus beatensis</emphasis>
</taxonomicName>
<emphasis id="9315B790EBE4A22A7BC61FAD9FC4C48D" bold="true">G</emphasis>
<taxonomicName id="C0A8B763D11D0F2DD9DE14936F5A2155" authorityName="Chalande" authorityYear="1910" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="gavoyi">
<emphasis id="A01AF7D5DCE9963D42319CA8E5635566" italics="true">Geophilus gavoyi</emphasis>
</taxonomicName>
. Views: all ventral. Abbreviations: cl chitin-lines, d forcipular denticle, tp trochanteroprefemur. Arrow: denticles on the forcipular coxosternite. Scale bars: 50 µm (
<emphasis id="C38336F6D7F12BE75F24452C43B4858A" bold="true">A, B</emphasis>
); 300 µm (
<emphasis id="542308CEAF7223DE038D15F4D9ECDE20" bold="true">C G</emphasis>
).
</paragraph>
</caption>
<caption id="E4343C49A4DFFFE6C7BFAEAE8B827FE6" ID-DOI="10.3897/zookeys.1213.133171.figure8" ID-arpha="362EA273-21A6-532C-A9E5-BEE0CD3C45D4" httpUri="https://binary.pensoft.net/fig/1142659" startId="F8">
<paragraph id="2C52FE5193C24FF1528D25F80E533C3F">
<label id="F9F90D483C478776C0187744C1915ED4">Figure 8.</label>
</paragraph>
<paragraph id="6A76BEA55D2C5701920343D7307BDAC5">
Metasternite of an anterior
<abbrev id="ABBRID0EJSAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
. Species
<emphasis id="E043141ED0381EAD07861E961B965416" bold="true">A</emphasis>
<taxonomicName id="486E9861C57FC179BC8381DD59D544C4" authority="LBS" authorityName="LBS" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="vesubiensis">
<emphasis id="E214DBCA6A3A0C370842195698070A16" italics="true">Clinopodes vesubiensis</emphasis>
<abbrev id="ABBRID0E1SAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
</taxonomicName>
16
<emphasis id="53A04782AF236A840AC7F23490589B6A" bold="true">B</emphasis>
<taxonomicName id="9B85A9A05C1624031DB983F5075CC6EA" authority="LBS" authorityName="LBS" class="Chilopoda" family="Geophilidae" genus="Galliophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="beatensis">
<emphasis id="0F367388F8BEA6632620B4DA6D7318C7" italics="true">Galliophilus beatensis</emphasis>
<abbrev id="ABBRID0ELTAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
</taxonomicName>
19
<emphasis id="8150784720FB7BA4B08052195EFFC047" bold="true">C</emphasis>
<taxonomicName id="BE9667CB9ACE28C997A9178231B78EA8" authority="LBS" authorityName="LBS" class="Chilopoda" family="Geophilidae" genus="Eurygeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pinguis">
<emphasis id="2F9617DF063B98C25A8B9DF509962E50" italics="true">Eurygeophilus pinguis</emphasis>
<abbrev id="ABBRID0E3TAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
</taxonomicName>
10
<emphasis id="B230A8A79FDFE1DF3B3EDC42599A8B55" bold="true">D</emphasis>
<taxonomicName id="6B3EE01F3C7DE29B073FBFB841D5F992" authority="LBS" authorityName="LBS" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="osquidatum">
<emphasis id="97D5CF295D212A483D9B2D2C17D0481E" italics="true">Geophilus osquidatum</emphasis>
<abbrev id="ABBRID0ENUAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
</taxonomicName>
11
<emphasis id="AD6061F292B14635B0B63517026CEFB6" bold="true">E</emphasis>
<taxonomicName id="3EE0B606EF9BCF838CF39385A3E3C7F1" authority="LBS" authorityName="LBS" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="electricus">
<emphasis id="61A266BF233A6CD3EEAADD39935BDC8B" italics="true">Geophilus electricus</emphasis>
<abbrev id="ABBRID0E5UAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
</taxonomicName>
11
<emphasis id="65EF2BCC9F4B894D2C5AEC49DE1AC8D0" bold="true">F</emphasis>
<taxonomicName id="00C37994F21D20A3DB56911A3715EBEA" authority="LBS" authorityName="LBS" class="Chilopoda" family="Geophilidae" genus="Pleurogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="mediterraneus">
<emphasis id="BF5332B3E713867B9C073162C914CF10" italics="true">Pleurogeophilus mediterraneus</emphasis>
<abbrev id="ABBRID0EPVAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
</taxonomicName>
15
<emphasis id="514FC134AC93AB85574EEBFAB4136321" bold="true">G</emphasis>
<taxonomicName id="6207E0010245D1863CE5A583DA84400C" authority="LBS" authorityName="LBS" class="Chilopoda" family="Geophilidae" genus="Stenotaenia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="linearis">
<emphasis id="633472D211EF4D1665E7EBEFC7120899" italics="true">Stenotaenia linearis</emphasis>
<abbrev id="ABBRID0EAWAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
</taxonomicName>
20
<emphasis id="881A476C1FE5A5E5904FE8AD9E8E5EA2" bold="true">H</emphasis>
<taxonomicName id="A432542117933149208BA75C9F3F6595" authority="LBS" authorityName="LBS" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="poseidonis">
<emphasis id="FFBDDF68A13E344D75225335721B1517" italics="true">Tuoba poseidonis</emphasis>
<abbrev id="ABBRID0ERWAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
</taxonomicName>
12. All ventral views. Abbreviations: cs carpophagus-structure, pf pore-field. Scale bars: 200 µm.
</paragraph>
</caption>
<caption id="C7BCD5F056383D4F5C4B8148B0FCA4E4" ID-DOI="10.3897/zookeys.1213.133171.figure9" ID-arpha="8CFA0167-F7D6-587B-9335-9D357CA187B0" httpUri="https://binary.pensoft.net/fig/1142660" startId="F9">
<paragraph id="FB1CC2E6A414550D456DCDF23E523787">
<label id="42B91040C2A375D692F7467148AAC316">Figure 9.</label>
</paragraph>
<paragraph id="C888C972CB0624972B487CF01B25A7DE">
Ultimate
<abbrev id="ABBRID0ELXAI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
without distal part of legs. Species
<emphasis id="4E6D47791E9DC566220D5F15E37C36D0" bold="true">A</emphasis>
<taxonomicName id="5C8E225E6CD120BD16F24AE78EF4E49D" baseAuthorityName="Ribaut" baseAuthorityYear="1911" class="Chilopoda" family="Geophilidae" genus="Arctogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="inopinatus">
<emphasis id="4CA47C0B50D26342773F5F978082485F" italics="true">Arctogeophilus inopinatus</emphasis>
</taxonomicName>
female
<emphasis id="449DE321363D32F5470AEF69AE1BF55F" bold="true">B</emphasis>
<taxonomicName id="8E2F346B50C5327791BEB0929900A35E" class="Chilopoda" family="Geophilidae" genus="Galliophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="beatensis">
<emphasis id="DDB0574946316911B8DE244BEFD2ECB0" italics="true">Galliophilus beatensis</emphasis>
</taxonomicName>
male
<emphasis id="DBA826C1A6EE91C996ACEFA155A80222" bold="true">C</emphasis>
<taxonomicName id="2664B9DECB9D7029DA050404AAD546F7" baseAuthorityName="De Geer" baseAuthorityYear="1778" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="flavus">
<emphasis id="57D6D1D850684210A7DBEFF775AF7F88" italics="true">Geophilus flavus</emphasis>
</taxonomicName>
female
<emphasis id="F2A860440A5681885A11543645F30DAE" bold="true">D</emphasis>
<taxonomicName id="437CDB474FF3103F928F8DEC4678D574" baseAuthorityName="C. L. Koch" baseAuthorityYear="1835" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="ferrugineum">
<emphasis id="F3344607E0E16711B9B5C01872874FD6" italics="true">Pachymerium ferrugineum</emphasis>
</taxonomicName>
male
<emphasis id="A7A61E76DA4CEC84DCD34687C3B66C96" bold="true">E</emphasis>
<taxonomicName id="E58F88DF5817C8DF807799E3324B6E98" class="Chilopoda" family="Geophilidae" genus="Pleurogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="mediterraneus">
<emphasis id="D4277BF160A380A29C2C72E6A4E54D88" italics="true">Pleurogeophilus mediterraneus</emphasis>
</taxonomicName>
female
<emphasis id="C227B33F3ED94CB0EE2FC4C91C62BB7B" bold="true">F</emphasis>
<taxonomicName id="EBFA03C3DB1B881AB0AE1E46F4401D4F" class="Chilopoda" family="Geophilidae" genus="Pleurogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="mediterraneus">
<emphasis id="6EADF3347F5452E5ACE39EB85E2E7B95" italics="true">Pleurogeophilus mediterraneus</emphasis>
</taxonomicName>
male
<emphasis id="C74CD7938B289F158E250F1FCF18903B" bold="true">G</emphasis>
,
<emphasis id="5DEB9C2E0EF0AC492AEF07AD029F9B22" bold="true">H</emphasis>
<taxonomicName id="49FADBD737A7C73E38FDECFB08DD53DE" class="Chilopoda" family="Geophilidae" genus="Stenotaenia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="linearis">
<emphasis id="3D9B3F1CE13EBFADC491BA824F085C23" italics="true">Stenotaenia linearis</emphasis>
</taxonomicName>
female
<emphasis id="1E939D39D8E1133FABCD7A498F2D7C37" bold="true">I</emphasis>
<taxonomicName id="0DA366F8DE7C4E98B245C849F9D6A3E0" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="poseidonis">
<emphasis id="79C224C4E2669334592D9AA48BF1485C" italics="true">Tuoba poseidonis</emphasis>
</taxonomicName>
female. Views: ventral (
<emphasis id="B4A051A779C47CE8B61DC78ECEE75EAB" bold="true">A G</emphasis>
), lateral (
<emphasis id="C242C216263268400A728016B17F2746" bold="true">H, I</emphasis>
). Abbreviations: p pit or fossa (
<emphasis id="957D543DDDB2136D7FF7C3EA5CF275B3" bold="true">H</emphasis>
wide fossa from dorsal to ventral side, indicated by dotted lines), um metasternite of the ultimate
<abbrev id="ABBRID0E61AI" xlink_title="leg-bearing segment(s)">LBS</abbrev>
(medial length and maximum width are indicated by dotted lines). Scale bars: 300 µm.
</paragraph>
</caption>
<caption id="16D42C563A2A1DA0C20852DB8318BBDD" ID-DOI="10.3897/zookeys.1213.133171.figure10" ID-arpha="C89112AD-E76F-5EF5-84EB-7B08C57C5F13" httpUri="https://binary.pensoft.net/fig/1142661" startId="F10">
<paragraph id="8AFAB739B2E6D7A06955395A63BD9138">
<label id="3A3A5BC06DC3110A25054285B8527DA5">Figure 10.</label>
</paragraph>
<paragraph id="F97293FC468D52DA9FCC83318C0312CA">
Legs of the ultimate pair
<emphasis id="CC1DC33AB203499549CFAB2E46CF20C1" bold="true">A, C</emphasis>
right leg
<emphasis id="8E1553EEB8D5C908B1E79ADCEE2475FC" bold="true">B, D</emphasis>
distal articles of the right leg
<emphasis id="3DEF4BE7A2129AA5696D3F23D0D4493A" bold="true">E G</emphasis>
both legs
<emphasis id="400AF1117982B931EDA641CBE107BA4C" bold="true">H</emphasis>
left leg. Species
<emphasis id="FFC785150F304D5BB72C41F3C33612C6" bold="true">A</emphasis>
<taxonomicName id="43140F6B518F0A82EC89AD53C5B3BC44" baseAuthorityName="Ribaut" baseAuthorityYear="1911" class="Chilopoda" family="Geophilidae" genus="Arctogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="inopinatus">
<emphasis id="BA6AFB637FE39B4798A0133CEFAB3276" italics="true">Arctogeophilus inopinatus</emphasis>
</taxonomicName>
male
<emphasis id="96CA68DD969F90CFA5F8C70C75FA4EEE" bold="true">B</emphasis>
<taxonomicName id="8002189DBF2CC1105B576339FCD30CDF" authorityName="Jones" authorityYear="1989" class="Chilopoda" family="Geophilidae" genus="Arenophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="363DBE7C3030E43D321075F1BA49605B" italics="true">Arenophilus peregrinus</emphasis>
</taxonomicName>
female
<emphasis id="A39690EA8ED8B260A0281A010B29BD98" bold="true">C</emphasis>
<taxonomicName id="CBD8BC08B8D27667A3925384AD780ADC" class="Chilopoda" family="Geophilidae" genus="Eurygeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="pinguis">
<emphasis id="B58586C61EB341AE01C1C58920A915A2" italics="true">Eurygeophilus pinguis</emphasis>
</taxonomicName>
female
<emphasis id="5C1D79DD30574CFB6273C1B2E266F014" bold="true">D</emphasis>
<taxonomicName id="A6A481AD5C4DFC44E69ABEB02D412259" authorityName="Brolemann" authorityYear="1909" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="fucorum">
<emphasis id="95CF9FC5D6666DEBF42CDB009CC3B752" italics="true">Geophilus fucorum</emphasis>
</taxonomicName>
female
<emphasis id="1151A7CDD3E53591D8BA50A973FFB24E" bold="true">E</emphasis>
<taxonomicName id="A850B88322129EE25F2A458189C9EB26" authorityName="Brolemann" authorityYear="1904" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="richardi">
<emphasis id="319E748CCE1B3A4AD30AFE5577DE304D" italics="true">Geophilus richardi</emphasis>
</taxonomicName>
female
<emphasis id="BB6A835BB40E8E26AC04731A2A9CFBCB" bold="true">F</emphasis>
<taxonomicName id="CF4B1889CC089C8868F973855AE7C7E8" baseAuthorityName="C. L. Koch" baseAuthorityYear="1835" class="Chilopoda" family="Geophilidae" genus="Pachymerium" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="ferrugineum">
<emphasis id="DB5695630E724C913C8A4B34F7AA6739" italics="true">Pachymerium ferrugineum</emphasis>
</taxonomicName>
female
<emphasis id="CE8544C1D68CD149C95E8F3C89F16F1E" bold="true">G</emphasis>
<taxonomicName id="173195FB7DCD4F0994FB4EA393515482" class="Chilopoda" family="Geophilidae" genus="Pleurogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="mediterraneus">
<emphasis id="22025A5806DC4187CEEC3A466529A30F" italics="true">Pleurogeophilus mediterraneus</emphasis>
</taxonomicName>
male
<emphasis id="D672CB7FFEEF356D866DE7B54EE57605" bold="true">H</emphasis>
<taxonomicName id="E4A47AB1E91BBDAF8F4B218A0DDA342C" class="Chilopoda" family="Geophilidae" genus="Pleurogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="mediterraneus">
<emphasis id="F0F6AAA4660C9D100EE567D6FC85A440" italics="true">Pleurogeophilus mediterraneus</emphasis>
</taxonomicName>
female
<emphasis id="C2AA7D2555109D4B453546812289408C" bold="true">I</emphasis>
<taxonomicName id="AAF0BDED9B11D0DBA33E4CDFBDF0197D" baseAuthorityName="Meinert" baseAuthorityYear="1870" class="Chilopoda" family="Geophilidae" genus="Acanthogeophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="4619898B1133A2789AE1956DC406FE02" italics="true">Acanthogeophilus spiniger</emphasis>
</taxonomicName>
female. Views: ventral (
<emphasis id="C9817AE3E5B1E65D32EACF5116420037" bold="true">A-E, I</emphasis>
), lateral (
<emphasis id="E661A963CF36E61691EDDA849FCC5420" bold="true">F</emphasis>
) and dorsal (
<emphasis id="FF36832F3B6E369362D9C219DA3663C4" bold="true">G-H</emphasis>
). Abbreviations: c claw, t tubercle with spines. Scale bars: 300 µm (
<emphasis id="C17AC3BEF1EFFF41ECEE45F3479923F9" bold="true">A, C, F</emphasis>
); 50 µm (
<emphasis id="39F1CBA8A73C5C016A4E28041DCF7140" bold="true">B</emphasis>
); 100 µm (
<emphasis id="63EC645C475358331D718C3E9E70F8D4" bold="true">D, E, I</emphasis>
); 1 mm (
<emphasis id="DA66E147F95A0C28E31CCF36E9F67AA3" bold="true">G H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>