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<document id="2B320CD7CB90C70BC977FB499B7B3ED0" ID-CLB-Dataset="29143" ID-DOI="10.11646/zootaxa.4432.1.1" ID-GBIF-Dataset="9f63aafe-4b6d-402b-aae1-602c2c83c1f4" ID-ISSN="1175-5326" ID-Zenodo-Dep="1455946" ID-ZooBank="D44C8383-4070-44B6-91CE-4A0940A0F094" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539204217470" checkinUser="plazi" docAuthor="Ünal, Mustafa" docDate="2018" docId="8B5C87A2FFAFFFF3FF6CFF6108B2FEB8" docLanguage="en" docName="zootaxa.4432.1.1.pdf" docOrigin="Zootaxa 4432 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Mixodusa retusa Ünal 2018, sp. nov." docType="treatment" docVersion="9" lastPageNumber="17" masterDocId="7765FFDAFFA0FFE3FFFBFFF90868FFA5" masterDocTitle="Tettigoniinae (Orthoptera: Tettigoniidae) from Turkey with key to genera and descriptions of six new species" masterLastPageNumber="66" masterPageNumber="1" pageNumber="16" updateTime="1698574244067" updateUser="ExternalLinkService">
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<mods:title id="28A76EA478B2F329EEF1F243046F97BA">Tettigoniinae (Orthoptera: Tettigoniidae) from Turkey with key to genera and descriptions of six new species</mods:title>
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<mods:namePart id="3E936B0458894890EC6D95ACA872D63D">Ünal, Mustafa</mods:namePart>
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<mods:date id="6AE78738194E2675BE8BC3615C323AC7">2018</mods:date>
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<mods:classification id="9993EB6E6DFE50C2B436C6F75A585436">journal article</mods:classification>
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<treatment id="8B5C87A2FFAFFFF3FF6CFF6108B2FEB8" ID-DOI="http://doi.org/10.5281/zenodo.5989319" ID-GBIF-Taxon="148684401" ID-Zenodo-Dep="5989319" LSID="urn:lsid:plazi:treatment:8B5C87A2FFAFFFF3FF6CFF6108B2FEB8" httpUri="http://treatment.plazi.org/id/8B5C87A2FFAFFFF3FF6CFF6108B2FEB8" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection id="4BEF653FFFAFFFECFF6CFF610A88FF76" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="034A36B4FFAFFFECFF6CFF610A79FF17" blockId="15.[151,736,152,211]" box="[151,529,152,178]" pageId="15" pageNumber="16">
<heading id="580281D8FFAFFFECFF6CFF610A79FF17" bold="true" box="[151,529,152,178]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="3181EAA6FFAFFFECFF6CFF610A79FF17" bold="true" box="[151,529,152,178]" pageId="15" pageNumber="16">
<taxonomicName id="C4F54D37FFAFFFECFF6CFF6109CEFF17" ID-CoL="8Q27J" authority="Unal" authorityName="Ünal" authorityYear="2018" box="[151,422,152,178]" class="Insecta" family="Tettigoniidae" genus="Mixodusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="retusa" status="sp. nov.">
<emphasis id="3181EAA6FFAFFFECFF6CFF61090AFF17" bold="true" box="[151,354,152,178]" italics="true" pageId="15" pageNumber="16">Mixodusa retusa</emphasis>
Ünal
</taxonomicName>
,
<taxonomicNameLabel id="2AB257DDFFAFFFECFE4FFF610A79FF17" box="[436,529,152,178]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="034A36B4FFAFFFECFF6CFF430A88FF76" blockId="15.[151,736,152,211]" box="[151,736,186,211]" pageId="15" pageNumber="16">
(
<figureCitation id="9BCE2A31FFAFFFECFF64FF43094EFF76" box="[159,294,186,211]" captionStart="FIGURES 1937" captionStartId="45.[151,264,1845,1867]" captionTargetBox="[166,1425,194,1826]" captionTargetId="figure@45.[163,1427,193,1832]" captionTargetPageId="45" captionText="FIGURES 1937. Habitus lateral view: 19 Pezodrymadusa striolata, male; 20 Pezodrymadusa lata, female; 21 Anadrymadusa (s.str.) adzharica, male; 22 ditto female; 2324. Novadrymadusa karabagi: 23 male end of abdomen; 24 female 7th sternite and subgenital plate; 25 Anadolua burri, male end of abdomen; 26 Mixoduda retusa sp. nov. 7th sternite and subgenital plate; 2729. Frons in male: 27 Paradrymadusa aksirayi; 28 Scotodrymadusa turcica; 29 Mixodusa retusa sp. nov.; 3031. Male end of abdomen: 30 Paradrymadusa aksirayi; 31 Anadolua schwarzi; 3237. Ovipositor: 32, 33 Anadolua schwarzi; 34, 35, 36 Anadolua burri; 37 Scotodrymadusa turcica." httpUri="https://zenodo.org/record/1455950/files/figure.png" pageId="15" pageNumber="16">Figs. 26, 29</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFEC9FF420991FF76" box="[306,505,186,211]" captionStart="FIGURES 3858" captionStartId="46.[151,264,1785,1807]" captionTargetBox="[182,1403,195,1768]" captionTargetId="figure@46.[180,1404,193,1773]" captionTargetPageId="46" captionText="FIGURES 3858. Ovipositor: 38 Scotodrymadusa maculata; 39 Mixodusa retusa sp. nov.; 4041. Female habitus lateral view: 40 Anadolua schwarzi; 41 Phytodrymadusa miramae; 4245. Titillators: 42 Scotodrymadusa turcica; 43 Mixodusa retusa sp. nov.; 44 Phytodrymadusa miramae; 45 Leptodusa expugnata; 4647. Male habitus lateral view: 46 Lithodusa helverseni (from Heller &amp; Korsunovskaya 2009); 47 Leptodusa expugnata; 48 Phytodrymadusa miramae, male end of abdomen; 49 Phytodrymadusa miramae, female pronotum and tegmina, dorso-lateral view; 5051. Leptodusa expugnata: 50 male last tergite; 51 male cercus; 5253. Mixodusa retusa sp. nov.: 52 male last tergite, 53 male cercus; 5455. Female pronotum and tegmina: 54 Leptodusa harzi; 55 Mixodusa retusa sp. nov.; 5657. End of hind tibia, ventral view: 56 Decticus albifrons female; 57 Rhacocleis turcica male; 58 Psorodonotus caucasicus, female fore tibia." httpUri="https://zenodo.org/record/1455952/files/figure.png" pageId="15" pageNumber="16">39, 43, 52, 53, 55</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFDFFFF430A01FF76" box="[516,617,186,211]" captionStart-0="FIGURES 186203" captionStart-1="FIGURES 204221" captionStartId-0="53.[151,264,1835,1857]" captionStartId-1="54.[151,264,1816,1838]" captionTargetBox-0="[191,1407,197,1819]" captionTargetBox-1="[154,1433,196,1789]" captionTargetId-0="figure@53.[186,1410,193,1825]" captionTargetId-1="figure@54.[151,1436,193,1794]" captionTargetPageId-0="53" captionTargetPageId-1="54" captionText-0="FIGURES 186203. Habitus lateral view: 186 Sureyaella bella, male; 187 ditto female; 188 Anterastes burri, male; 189 ditto female; 190 Koroglus disparalatus, male; 191 ditto female; 192193. Male tegmina and pronotum, dorsal view: 192 Anterastes serbicus; 193 Koroglus disparalatus; 194195. Anterastes babadaghi: 194 male left tegmen, inner surface; 195 ditto stridulatory file area; 196197. Koroglus disparalatus: 196 male left tegmen, inner surface; 197 ditto stridulatory file area; 198203. Mixodusa retusa sp. nov.: 198 male habitus, lateral view; 199 male head, pronotum and tegmina, lateral view; 200 ditto dorsal view; 201 male last tergite and cerci; 202 male subgenital plate; 203 female habitus, lateral view." captionText-1="FIGURES 204221. Mixodusa retusa sp. nov.: 204 female head, pronotum and tegmina, lateral view; 205 ditto dorsal; 206 female subgenital plate; 207212. Mixodusa bocquilloni: 207 male head, pronotum and tegmina, lateral view; 208 ditto dorsal view; 209 male last tergite and cerci; 210 female head, pronotum and tegmina, lateral view; 211 ditto dorsal view; 212 female subgenital plate; 213218. Anadolua schwarzi: 213 male habitus, lateral view; 214 male head, pronotum and tegmina, dorsal view; 215 male end of abdomen; 216 female habitus, lateral view; 217 female head and pronotum, dorsal view; 218 female subgenital plate and 7th sternite; 219221. Anadolua burri: 219 male habitus, lateral view; 220 male head, pronotum and tegmina, dorsal view; 221 male end of abdomen." httpUri-0="https://zenodo.org/record/1455966/files/figure.png" httpUri-1="https://zenodo.org/record/1455968/files/figure.png" pageId="15" pageNumber="16">198206</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFD8FFF420AB0FF76" box="[628,728,187,211]" captionStart="FIGURES 312317" captionStartId="60.[151,264,1934,1956]" captionTargetBox="[193,1417,201,1900]" captionTargetId="figure@60.[177,1421,193,1918]" captionTargetPageId="60" captionText="FIGURES 312317. Mixodusa retusa sp. nov.: 312 male habitus, lateral view; 313 female habitus, lateral view (312 and 313 illustrated by Şeyma Hatip); 314 male last tergite; 315 male cercus; 316 titillators; 317 female 7th sternite and subgenital plate. Scales 1 mm." httpUri="https://zenodo.org/record/1455980/files/figure.png" pageId="15" pageNumber="16">312317</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4BEF653FFFAFFFECFF6CFEFB0946FAD2" pageId="15" pageNumber="16" type="description">
<paragraph id="034A36B4FFAFFFECFF6CFEFB09B5FBF2" blockId="15.[151,1437,258,2011]" pageId="15" pageNumber="16">
<emphasis id="3181EAA6FFAFFFECFF6CFEFB0941FEBE" bold="true" box="[151,297,258,283]" pageId="15" pageNumber="16">Description.</emphasis>
Male (
<typeStatus id="DC4E8816FFAFFFECFE8CFEFB0988FEBE" box="[375,480,258,283]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
): Fastigium of vertex 1.4 times wider than scapus (
<figureCitation id="9BCE2A31FFAFFFECFBECFEFB0C01FEBE" box="[1047,1129,258,283]" captionStart="FIGURES 1937" captionStartId="45.[151,264,1845,1867]" captionTargetBox="[166,1425,194,1826]" captionTargetId="figure@45.[163,1427,193,1832]" captionTargetPageId="45" captionText="FIGURES 1937. Habitus lateral view: 19 Pezodrymadusa striolata, male; 20 Pezodrymadusa lata, female; 21 Anadrymadusa (s.str.) adzharica, male; 22 ditto female; 2324. Novadrymadusa karabagi: 23 male end of abdomen; 24 female 7th sternite and subgenital plate; 25 Anadolua burri, male end of abdomen; 26 Mixoduda retusa sp. nov. 7th sternite and subgenital plate; 2729. Frons in male: 27 Paradrymadusa aksirayi; 28 Scotodrymadusa turcica; 29 Mixodusa retusa sp. nov.; 3031. Male end of abdomen: 30 Paradrymadusa aksirayi; 31 Anadolua schwarzi; 3237. Ovipositor: 32, 33 Anadolua schwarzi; 34, 35, 36 Anadolua burri; 37 Scotodrymadusa turcica." httpUri="https://zenodo.org/record/1455950/files/figure.png" pageId="15" pageNumber="16">Fig. 29</figureCitation>
), frontal part vertical, basal part sloping with smooth surface. Pronotum (
<figureCitation id="9BCE2A31FFAFFFECFD57FEDF0B3CFE9A" box="[684,852,294,319]" captionStart="FIGURES 186203" captionStartId="53.[151,264,1835,1857]" captionTargetBox="[191,1407,197,1819]" captionTargetId="figure@53.[186,1410,193,1825]" captionTargetPageId="53" captionText="FIGURES 186203. Habitus lateral view: 186 Sureyaella bella, male; 187 ditto female; 188 Anterastes burri, male; 189 ditto female; 190 Koroglus disparalatus, male; 191 ditto female; 192193. Male tegmina and pronotum, dorsal view: 192 Anterastes serbicus; 193 Koroglus disparalatus; 194195. Anterastes babadaghi: 194 male left tegmen, inner surface; 195 ditto stridulatory file area; 196197. Koroglus disparalatus: 196 male left tegmen, inner surface; 197 ditto stridulatory file area; 198203. Mixodusa retusa sp. nov.: 198 male habitus, lateral view; 199 male head, pronotum and tegmina, lateral view; 200 ditto dorsal view; 201 male last tergite and cerci; 202 male subgenital plate; 203 female habitus, lateral view." httpUri="https://zenodo.org/record/1455966/files/figure.png" pageId="15" pageNumber="16">Figs. 198200</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFC9FFEDE0BE7FE9A" box="[868,911,295,319]" captionStart="FIGURES 312317" captionStartId="60.[151,264,1934,1956]" captionTargetBox="[193,1417,201,1900]" captionTargetId="figure@60.[177,1421,193,1918]" captionTargetPageId="60" captionText="FIGURES 312317. Mixodusa retusa sp. nov.: 312 male habitus, lateral view; 313 female habitus, lateral view (312 and 313 illustrated by Şeyma Hatip); 314 male last tergite; 315 male cercus; 316 titillators; 317 female 7th sternite and subgenital plate. Scales 1 mm." httpUri="https://zenodo.org/record/1455980/files/figure.png" pageId="15" pageNumber="16">312</figureCitation>
) almost cylindrical along its length dorsally, without median and lateral carinae; 1.8 times longer than its height and 1.1 times longer than fore femur; anterior margin almost straight, posterior margin very broadly rounded; shoulder incision very weak in lateral view. Tegmina (
<figureCitation id="9BCE2A31FFAFFFECFEF1FE6B09D8FE0E" box="[266,432,402,427]" captionStart="FIGURES 186203" captionStartId="53.[151,264,1835,1857]" captionTargetBox="[191,1407,197,1819]" captionTargetId="figure@53.[186,1410,193,1825]" captionTargetPageId="53" captionText="FIGURES 186203. Habitus lateral view: 186 Sureyaella bella, male; 187 ditto female; 188 Anterastes burri, male; 189 ditto female; 190 Koroglus disparalatus, male; 191 ditto female; 192193. Male tegmina and pronotum, dorsal view: 192 Anterastes serbicus; 193 Koroglus disparalatus; 194195. Anterastes babadaghi: 194 male left tegmen, inner surface; 195 ditto stridulatory file area; 196197. Koroglus disparalatus: 196 male left tegmen, inner surface; 197 ditto stridulatory file area; 198203. Mixodusa retusa sp. nov.: 198 male habitus, lateral view; 199 male head, pronotum and tegmina, lateral view; 200 ditto dorsal view; 201 male last tergite and cerci; 202 male subgenital plate; 203 female habitus, lateral view." httpUri="https://zenodo.org/record/1455966/files/figure.png" pageId="15" pageNumber="16">Figs. 198200</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFE45FE6A0982FE0E" box="[446,490,403,427]" captionStart="FIGURES 312317" captionStartId="60.[151,264,1934,1956]" captionTargetBox="[193,1417,201,1900]" captionTargetId="figure@60.[177,1421,193,1918]" captionTargetPageId="60" captionText="FIGURES 312317. Mixodusa retusa sp. nov.: 312 male habitus, lateral view; 313 female habitus, lateral view (312 and 313 illustrated by Şeyma Hatip); 314 male last tergite; 315 male cercus; 316 titillators; 317 female 7th sternite and subgenital plate. Scales 1 mm." httpUri="https://zenodo.org/record/1455980/files/figure.png" pageId="15" pageNumber="16">312</figureCitation>
) brachypterous, reaching slightly beyond end of first abdominal tergite. Fore tibia with 3 dorsal spines on outer side. Hind tibia with 4 apical spurs ventrally. Plantula scale like, 2.6 times shorter than metatarsus. Femora without dorsal spines; outer lobe of fore femur without, but all the other lobes of femora with an indistinct small spine. Hind femur (
<figureCitation id="9BCE2A31FFAFFFECFDB9FE070AC4FDB2" box="[578,684,510,535]" captionStart="FIGURES 186203" captionStartId="53.[151,264,1835,1857]" captionTargetBox="[191,1407,197,1819]" captionTargetId="figure@53.[186,1410,193,1825]" captionTargetPageId="53" captionText="FIGURES 186203. Habitus lateral view: 186 Sureyaella bella, male; 187 ditto female; 188 Anterastes burri, male; 189 ditto female; 190 Koroglus disparalatus, male; 191 ditto female; 192193. Male tegmina and pronotum, dorsal view: 192 Anterastes serbicus; 193 Koroglus disparalatus; 194195. Anterastes babadaghi: 194 male left tegmen, inner surface; 195 ditto stridulatory file area; 196197. Koroglus disparalatus: 196 male left tegmen, inner surface; 197 ditto stridulatory file area; 198203. Mixodusa retusa sp. nov.: 198 male habitus, lateral view; 199 male head, pronotum and tegmina, lateral view; 200 ditto dorsal view; 201 male last tergite and cerci; 202 male subgenital plate; 203 female habitus, lateral view." httpUri="https://zenodo.org/record/1455966/files/figure.png" pageId="15" pageNumber="16">Figs. 198</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFD43FE060A8BFDB2" box="[696,739,511,535]" captionStart="FIGURES 312317" captionStartId="60.[151,264,1934,1956]" captionTargetBox="[193,1417,201,1900]" captionTargetId="figure@60.[177,1421,193,1918]" captionTargetPageId="60" captionText="FIGURES 312317. Mixodusa retusa sp. nov.: 312 male habitus, lateral view; 313 female habitus, lateral view (312 and 313 illustrated by Şeyma Hatip); 314 male last tergite; 315 male cercus; 316 titillators; 317 female 7th sternite and subgenital plate. Scales 1 mm." httpUri="https://zenodo.org/record/1455980/files/figure.png" pageId="15" pageNumber="16">312</figureCitation>
) very long, almost as long as body length (see measurements). Prosternum with 2 very small, narrow triangular projections. Last abdominal tergite (
<figureCitation id="9BCE2A31FFAFFFECFB82FDDB0CB2FD9E" box="[1145,1242,546,571]" captionStart="FIGURES 3858" captionStartId="46.[151,264,1785,1807]" captionTargetBox="[182,1403,195,1768]" captionTargetId="figure@46.[180,1404,193,1773]" captionTargetPageId="46" captionText="FIGURES 3858. Ovipositor: 38 Scotodrymadusa maculata; 39 Mixodusa retusa sp. nov.; 4041. Female habitus lateral view: 40 Anadolua schwarzi; 41 Phytodrymadusa miramae; 4245. Titillators: 42 Scotodrymadusa turcica; 43 Mixodusa retusa sp. nov.; 44 Phytodrymadusa miramae; 45 Leptodusa expugnata; 4647. Male habitus lateral view: 46 Lithodusa helverseni (from Heller &amp; Korsunovskaya 2009); 47 Leptodusa expugnata; 48 Phytodrymadusa miramae, male end of abdomen; 49 Phytodrymadusa miramae, female pronotum and tegmina, dorso-lateral view; 5051. Leptodusa expugnata: 50 male last tergite; 51 male cercus; 5253. Mixodusa retusa sp. nov.: 52 male last tergite, 53 male cercus; 5455. Female pronotum and tegmina: 54 Leptodusa harzi; 55 Mixodusa retusa sp. nov.; 5657. End of hind tibia, ventral view: 56 Decticus albifrons female; 57 Rhacocleis turcica male; 58 Psorodonotus caucasicus, female fore tibia." httpUri="https://zenodo.org/record/1455952/files/figure.png" pageId="15" pageNumber="16">Figs. 52</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFB11FDDA0D7EFD9E" box="[1258,1302,547,571]" captionStart="FIGURES 186203" captionStartId="53.[151,264,1835,1857]" captionTargetBox="[191,1407,197,1819]" captionTargetId="figure@53.[186,1410,193,1825]" captionTargetPageId="53" captionText="FIGURES 186203. Habitus lateral view: 186 Sureyaella bella, male; 187 ditto female; 188 Anterastes burri, male; 189 ditto female; 190 Koroglus disparalatus, male; 191 ditto female; 192193. Male tegmina and pronotum, dorsal view: 192 Anterastes serbicus; 193 Koroglus disparalatus; 194195. Anterastes babadaghi: 194 male left tegmen, inner surface; 195 ditto stridulatory file area; 196197. Koroglus disparalatus: 196 male left tegmen, inner surface; 197 ditto stridulatory file area; 198203. Mixodusa retusa sp. nov.: 198 male habitus, lateral view; 199 male head, pronotum and tegmina, lateral view; 200 ditto dorsal view; 201 male last tergite and cerci; 202 male subgenital plate; 203 female habitus, lateral view." httpUri="https://zenodo.org/record/1455966/files/figure.png" pageId="15" pageNumber="16">201</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFADDFDDA0D39FD9E" box="[1318,1361,547,571]" captionStart="FIGURES 312317" captionStartId="60.[151,264,1934,1956]" captionTargetBox="[193,1417,201,1900]" captionTargetId="figure@60.[177,1421,193,1918]" captionTargetPageId="60" captionText="FIGURES 312317. Mixodusa retusa sp. nov.: 312 male habitus, lateral view; 313 female habitus, lateral view (312 and 313 illustrated by Şeyma Hatip); 314 male last tergite; 315 male cercus; 316 titillators; 317 female 7th sternite and subgenital plate. Scales 1 mm." httpUri="https://zenodo.org/record/1455980/files/figure.png" pageId="15" pageNumber="16">314</figureCitation>
) quite short, with a narrowly rounded posterior incision; posterior lobes very short and rounded at apex. Cerci (
<figureCitation id="9BCE2A31FFAFFFECFAC2FDBF0DFFFDFA" box="[1337,1431,582,607]" captionStart="FIGURES 3858" captionStartId="46.[151,264,1785,1807]" captionTargetBox="[182,1403,195,1768]" captionTargetId="figure@46.[180,1404,193,1773]" captionTargetPageId="46" captionText="FIGURES 3858. Ovipositor: 38 Scotodrymadusa maculata; 39 Mixodusa retusa sp. nov.; 4041. Female habitus lateral view: 40 Anadolua schwarzi; 41 Phytodrymadusa miramae; 4245. Titillators: 42 Scotodrymadusa turcica; 43 Mixodusa retusa sp. nov.; 44 Phytodrymadusa miramae; 45 Leptodusa expugnata; 4647. Male habitus lateral view: 46 Lithodusa helverseni (from Heller &amp; Korsunovskaya 2009); 47 Leptodusa expugnata; 48 Phytodrymadusa miramae, male end of abdomen; 49 Phytodrymadusa miramae, female pronotum and tegmina, dorso-lateral view; 5051. Leptodusa expugnata: 50 male last tergite; 51 male cercus; 5253. Mixodusa retusa sp. nov.: 52 male last tergite, 53 male cercus; 5455. Female pronotum and tegmina: 54 Leptodusa harzi; 55 Mixodusa retusa sp. nov.; 5657. End of hind tibia, ventral view: 56 Decticus albifrons female; 57 Rhacocleis turcica male; 58 Psorodonotus caucasicus, female fore tibia." httpUri="https://zenodo.org/record/1455952/files/figure.png" pageId="15" pageNumber="16">Figs. 53</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFF6CFD9208ABFD26" box="[151,195,619,643]" captionStart="FIGURES 186203" captionStartId="53.[151,264,1835,1857]" captionTargetBox="[191,1407,197,1819]" captionTargetId="figure@53.[186,1410,193,1825]" captionTargetPageId="53" captionText="FIGURES 186203. Habitus lateral view: 186 Sureyaella bella, male; 187 ditto female; 188 Anterastes burri, male; 189 ditto female; 190 Koroglus disparalatus, male; 191 ditto female; 192193. Male tegmina and pronotum, dorsal view: 192 Anterastes serbicus; 193 Koroglus disparalatus; 194195. Anterastes babadaghi: 194 male left tegmen, inner surface; 195 ditto stridulatory file area; 196197. Koroglus disparalatus: 196 male left tegmen, inner surface; 197 ditto stridulatory file area; 198203. Mixodusa retusa sp. nov.: 198 male habitus, lateral view; 199 male head, pronotum and tegmina, lateral view; 200 ditto dorsal view; 201 male last tergite and cerci; 202 male subgenital plate; 203 female habitus, lateral view." httpUri="https://zenodo.org/record/1455966/files/figure.png" pageId="15" pageNumber="16">201</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFF2EFD930968FD26" box="[213,256,618,643]" captionStart="FIGURES 312317" captionStartId="60.[151,264,1934,1956]" captionTargetBox="[193,1417,201,1900]" captionTargetId="figure@60.[177,1421,193,1918]" captionTargetPageId="60" captionText="FIGURES 312317. Mixodusa retusa sp. nov.: 312 male habitus, lateral view; 313 female habitus, lateral view (312 and 313 illustrated by Şeyma Hatip); 314 male last tergite; 315 male cercus; 316 titillators; 317 female 7th sternite and subgenital plate. Scales 1 mm." httpUri="https://zenodo.org/record/1455980/files/figure.png" pageId="15" pageNumber="16">315</figureCitation>
) short and broad; proximal part before inner tooth wide and almost cylindrical, distal part narrow triangular and slightly recurved, with a distinct, sharp inner tooth, not longer than distal part of cercus. Subgenital plate (
<figureCitation id="9BCE2A31FFAFFFECFF19FD4B092EFD6E" box="[226,326,690,715]" captionStart="FIGURES 186203" captionStartId="53.[151,264,1835,1857]" captionTargetBox="[191,1407,197,1819]" captionTargetId="figure@53.[186,1410,193,1825]" captionTargetPageId="53" captionText="FIGURES 186203. Habitus lateral view: 186 Sureyaella bella, male; 187 ditto female; 188 Anterastes burri, male; 189 ditto female; 190 Koroglus disparalatus, male; 191 ditto female; 192193. Male tegmina and pronotum, dorsal view: 192 Anterastes serbicus; 193 Koroglus disparalatus; 194195. Anterastes babadaghi: 194 male left tegmen, inner surface; 195 ditto stridulatory file area; 196197. Koroglus disparalatus: 196 male left tegmen, inner surface; 197 ditto stridulatory file area; 198203. Mixodusa retusa sp. nov.: 198 male habitus, lateral view; 199 male head, pronotum and tegmina, lateral view; 200 ditto dorsal view; 201 male last tergite and cerci; 202 male subgenital plate; 203 female habitus, lateral view." httpUri="https://zenodo.org/record/1455966/files/figure.png" pageId="15" pageNumber="16">Fig. 202</figureCitation>
) with a triangular posterior median incision, styli longer then the depth of posterior incision. Titillators (
<figureCitation id="9BCE2A31FFAFFFECFEEFFD2F091BFD4A" box="[276,371,726,751]" captionStart="FIGURES 3858" captionStartId="46.[151,264,1785,1807]" captionTargetBox="[182,1403,195,1768]" captionTargetId="figure@46.[180,1404,193,1773]" captionTargetPageId="46" captionText="FIGURES 3858. Ovipositor: 38 Scotodrymadusa maculata; 39 Mixodusa retusa sp. nov.; 4041. Female habitus lateral view: 40 Anadolua schwarzi; 41 Phytodrymadusa miramae; 4245. Titillators: 42 Scotodrymadusa turcica; 43 Mixodusa retusa sp. nov.; 44 Phytodrymadusa miramae; 45 Leptodusa expugnata; 4647. Male habitus lateral view: 46 Lithodusa helverseni (from Heller &amp; Korsunovskaya 2009); 47 Leptodusa expugnata; 48 Phytodrymadusa miramae, male end of abdomen; 49 Phytodrymadusa miramae, female pronotum and tegmina, dorso-lateral view; 5051. Leptodusa expugnata: 50 male last tergite; 51 male cercus; 5253. Mixodusa retusa sp. nov.: 52 male last tergite, 53 male cercus; 5455. Female pronotum and tegmina: 54 Leptodusa harzi; 55 Mixodusa retusa sp. nov.; 5657. End of hind tibia, ventral view: 56 Decticus albifrons female; 57 Rhacocleis turcica male; 58 Psorodonotus caucasicus, female fore tibia." httpUri="https://zenodo.org/record/1455952/files/figure.png" pageId="15" pageNumber="16">Figs. 43</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFE7BFD2F09C4FD4A" box="[384,428,726,751]" captionStart="FIGURES 312317" captionStartId="60.[151,264,1934,1956]" captionTargetBox="[193,1417,201,1900]" captionTargetId="figure@60.[177,1421,193,1918]" captionTargetPageId="60" captionText="FIGURES 312317. Mixodusa retusa sp. nov.: 312 male habitus, lateral view; 313 female habitus, lateral view (312 and 313 illustrated by Şeyma Hatip); 314 male last tergite; 315 male cercus; 316 titillators; 317 female 7th sternite and subgenital plate. Scales 1 mm." httpUri="https://zenodo.org/record/1455980/files/figure.png" pageId="15" pageNumber="16">316</figureCitation>
) with long apical arms with a row of sharp teeth along its outer side; inner side of apical arms with a distinct projection at the base; basal arms short and strongly upcurved, reaching to half of apical arms. Female: Fastigium of vertex 1.75 times wider than scapus. Pronotum (
<figureCitation id="9BCE2A31FFAFFFECFC46FCE70C75FC92" box="[957,1053,798,823]" captionStart="FIGURES 3858" captionStartId="46.[151,264,1785,1807]" captionTargetBox="[182,1403,195,1768]" captionTargetId="figure@46.[180,1404,193,1773]" captionTargetPageId="46" captionText="FIGURES 3858. Ovipositor: 38 Scotodrymadusa maculata; 39 Mixodusa retusa sp. nov.; 4041. Female habitus lateral view: 40 Anadolua schwarzi; 41 Phytodrymadusa miramae; 4245. Titillators: 42 Scotodrymadusa turcica; 43 Mixodusa retusa sp. nov.; 44 Phytodrymadusa miramae; 45 Leptodusa expugnata; 4647. Male habitus lateral view: 46 Lithodusa helverseni (from Heller &amp; Korsunovskaya 2009); 47 Leptodusa expugnata; 48 Phytodrymadusa miramae, male end of abdomen; 49 Phytodrymadusa miramae, female pronotum and tegmina, dorso-lateral view; 5051. Leptodusa expugnata: 50 male last tergite; 51 male cercus; 5253. Mixodusa retusa sp. nov.: 52 male last tergite, 53 male cercus; 5455. Female pronotum and tegmina: 54 Leptodusa harzi; 55 Mixodusa retusa sp. nov.; 5657. End of hind tibia, ventral view: 56 Decticus albifrons female; 57 Rhacocleis turcica male; 58 Psorodonotus caucasicus, female fore tibia." httpUri="https://zenodo.org/record/1455952/files/figure.png" pageId="15" pageNumber="16">Figs. 55</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFBD1FCE70CE5FC92" box="[1066,1165,798,823]" captionStart-0="FIGURES 186203" captionStart-1="FIGURES 204221" captionStartId-0="53.[151,264,1835,1857]" captionStartId-1="54.[151,264,1816,1838]" captionTargetBox-0="[191,1407,197,1819]" captionTargetBox-1="[154,1433,196,1789]" captionTargetId-0="figure@53.[186,1410,193,1825]" captionTargetId-1="figure@54.[151,1436,193,1794]" captionTargetPageId-0="53" captionTargetPageId-1="54" captionText-0="FIGURES 186203. Habitus lateral view: 186 Sureyaella bella, male; 187 ditto female; 188 Anterastes burri, male; 189 ditto female; 190 Koroglus disparalatus, male; 191 ditto female; 192193. Male tegmina and pronotum, dorsal view: 192 Anterastes serbicus; 193 Koroglus disparalatus; 194195. Anterastes babadaghi: 194 male left tegmen, inner surface; 195 ditto stridulatory file area; 196197. Koroglus disparalatus: 196 male left tegmen, inner surface; 197 ditto stridulatory file area; 198203. Mixodusa retusa sp. nov.: 198 male habitus, lateral view; 199 male head, pronotum and tegmina, lateral view; 200 ditto dorsal view; 201 male last tergite and cerci; 202 male subgenital plate; 203 female habitus, lateral view." captionText-1="FIGURES 204221. Mixodusa retusa sp. nov.: 204 female head, pronotum and tegmina, lateral view; 205 ditto dorsal; 206 female subgenital plate; 207212. Mixodusa bocquilloni: 207 male head, pronotum and tegmina, lateral view; 208 ditto dorsal view; 209 male last tergite and cerci; 210 female head, pronotum and tegmina, lateral view; 211 ditto dorsal view; 212 female subgenital plate; 213218. Anadolua schwarzi: 213 male habitus, lateral view; 214 male head, pronotum and tegmina, dorsal view; 215 male end of abdomen; 216 female habitus, lateral view; 217 female head and pronotum, dorsal view; 218 female subgenital plate and 7th sternite; 219221. Anadolua burri: 219 male habitus, lateral view; 220 male head, pronotum and tegmina, dorsal view; 221 male end of abdomen." httpUri-0="https://zenodo.org/record/1455966/files/figure.png" httpUri-1="https://zenodo.org/record/1455968/files/figure.png" pageId="15" pageNumber="16">203205</figureCitation>
) almost cylindrical, 1.8 times longer than its height, and 1.1 times longer than fore femur; anterior margin slightly concave, posterior margin very broadly rounded; shoulder incision more distict than that of male. Tegmina (
<figureCitation id="9BCE2A31FFAFFFECFB46FC9F0D48FCDA" box="[1213,1312,870,895]" captionStart="FIGURES 3858" captionStartId="46.[151,264,1785,1807]" captionTargetBox="[182,1403,195,1768]" captionTargetId="figure@46.[180,1404,193,1773]" captionTargetPageId="46" captionText="FIGURES 3858. Ovipositor: 38 Scotodrymadusa maculata; 39 Mixodusa retusa sp. nov.; 4041. Female habitus lateral view: 40 Anadolua schwarzi; 41 Phytodrymadusa miramae; 4245. Titillators: 42 Scotodrymadusa turcica; 43 Mixodusa retusa sp. nov.; 44 Phytodrymadusa miramae; 45 Leptodusa expugnata; 4647. Male habitus lateral view: 46 Lithodusa helverseni (from Heller &amp; Korsunovskaya 2009); 47 Leptodusa expugnata; 48 Phytodrymadusa miramae, male end of abdomen; 49 Phytodrymadusa miramae, female pronotum and tegmina, dorso-lateral view; 5051. Leptodusa expugnata: 50 male last tergite; 51 male cercus; 5253. Mixodusa retusa sp. nov.: 52 male last tergite, 53 male cercus; 5455. Female pronotum and tegmina: 54 Leptodusa harzi; 55 Mixodusa retusa sp. nov.; 5657. End of hind tibia, ventral view: 56 Decticus albifrons female; 57 Rhacocleis turcica male; 58 Psorodonotus caucasicus, female fore tibia." httpUri="https://zenodo.org/record/1455952/files/figure.png" pageId="15" pageNumber="16">Figs. 55</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFACAFC9F0DFCFCDA" box="[1329,1428,870,895]" captionStart-0="FIGURES 186203" captionStart-1="FIGURES 204221" captionStartId-0="53.[151,264,1835,1857]" captionStartId-1="54.[151,264,1816,1838]" captionTargetBox-0="[191,1407,197,1819]" captionTargetBox-1="[154,1433,196,1789]" captionTargetId-0="figure@53.[186,1410,193,1825]" captionTargetId-1="figure@54.[151,1436,193,1794]" captionTargetPageId-0="53" captionTargetPageId-1="54" captionText-0="FIGURES 186203. Habitus lateral view: 186 Sureyaella bella, male; 187 ditto female; 188 Anterastes burri, male; 189 ditto female; 190 Koroglus disparalatus, male; 191 ditto female; 192193. Male tegmina and pronotum, dorsal view: 192 Anterastes serbicus; 193 Koroglus disparalatus; 194195. Anterastes babadaghi: 194 male left tegmen, inner surface; 195 ditto stridulatory file area; 196197. Koroglus disparalatus: 196 male left tegmen, inner surface; 197 ditto stridulatory file area; 198203. Mixodusa retusa sp. nov.: 198 male habitus, lateral view; 199 male head, pronotum and tegmina, lateral view; 200 ditto dorsal view; 201 male last tergite and cerci; 202 male subgenital plate; 203 female habitus, lateral view." captionText-1="FIGURES 204221. Mixodusa retusa sp. nov.: 204 female head, pronotum and tegmina, lateral view; 205 ditto dorsal; 206 female subgenital plate; 207212. Mixodusa bocquilloni: 207 male head, pronotum and tegmina, lateral view; 208 ditto dorsal view; 209 male last tergite and cerci; 210 female head, pronotum and tegmina, lateral view; 211 ditto dorsal view; 212 female subgenital plate; 213218. Anadolua schwarzi: 213 male habitus, lateral view; 214 male head, pronotum and tegmina, dorsal view; 215 male end of abdomen; 216 female habitus, lateral view; 217 female head and pronotum, dorsal view; 218 female subgenital plate and 7th sternite; 219221. Anadolua burri: 219 male habitus, lateral view; 220 male head, pronotum and tegmina, dorsal view; 221 male end of abdomen." httpUri-0="https://zenodo.org/record/1455966/files/figure.png" httpUri-1="https://zenodo.org/record/1455968/files/figure.png" pageId="15" pageNumber="16">203205</figureCitation>
) reduced, contiguous at dorsum, reaching to 1/3 of first abdominal tergite, slightly protruded under pronotum. Legs (
<figureCitation id="9BCE2A31FFAFFFECFF64FC570929FC62" box="[159,321,942,967]" captionStart-0="FIGURES 186203" captionStart-1="FIGURES 204221" captionStartId-0="53.[151,264,1835,1857]" captionStartId-1="54.[151,264,1816,1838]" captionTargetBox-0="[191,1407,197,1819]" captionTargetBox-1="[154,1433,196,1789]" captionTargetId-0="figure@53.[186,1410,193,1825]" captionTargetId-1="figure@54.[151,1436,193,1794]" captionTargetPageId-0="53" captionTargetPageId-1="54" captionText-0="FIGURES 186203. Habitus lateral view: 186 Sureyaella bella, male; 187 ditto female; 188 Anterastes burri, male; 189 ditto female; 190 Koroglus disparalatus, male; 191 ditto female; 192193. Male tegmina and pronotum, dorsal view: 192 Anterastes serbicus; 193 Koroglus disparalatus; 194195. Anterastes babadaghi: 194 male left tegmen, inner surface; 195 ditto stridulatory file area; 196197. Koroglus disparalatus: 196 male left tegmen, inner surface; 197 ditto stridulatory file area; 198203. Mixodusa retusa sp. nov.: 198 male habitus, lateral view; 199 male head, pronotum and tegmina, lateral view; 200 ditto dorsal view; 201 male last tergite and cerci; 202 male subgenital plate; 203 female habitus, lateral view." captionText-1="FIGURES 204221. Mixodusa retusa sp. nov.: 204 female head, pronotum and tegmina, lateral view; 205 ditto dorsal; 206 female subgenital plate; 207212. Mixodusa bocquilloni: 207 male head, pronotum and tegmina, lateral view; 208 ditto dorsal view; 209 male last tergite and cerci; 210 female head, pronotum and tegmina, lateral view; 211 ditto dorsal view; 212 female subgenital plate; 213218. Anadolua schwarzi: 213 male habitus, lateral view; 214 male head, pronotum and tegmina, dorsal view; 215 male end of abdomen; 216 female habitus, lateral view; 217 female head and pronotum, dorsal view; 218 female subgenital plate and 7th sternite; 219221. Anadolua burri: 219 male habitus, lateral view; 220 male head, pronotum and tegmina, dorsal view; 221 male end of abdomen." httpUri-0="https://zenodo.org/record/1455966/files/figure.png" httpUri-1="https://zenodo.org/record/1455968/files/figure.png" pageId="15" pageNumber="16">Figs. 203213</figureCitation>
) as in male. Prosternum with 2 small, triangular projections. Last abdominal tergite very short, with a deep triangular incision. Subgenital plate fused with 7th sternite (
<figureCitation id="9BCE2A31FFAFFFECFC6AFC2B0B87FC4E" box="[913,1007,978,1003]" captionStart="FIGURES 1937" captionStartId="45.[151,264,1845,1867]" captionTargetBox="[166,1425,194,1826]" captionTargetId="figure@45.[163,1427,193,1832]" captionTargetPageId="45" captionText="FIGURES 1937. Habitus lateral view: 19 Pezodrymadusa striolata, male; 20 Pezodrymadusa lata, female; 21 Anadrymadusa (s.str.) adzharica, male; 22 ditto female; 2324. Novadrymadusa karabagi: 23 male end of abdomen; 24 female 7th sternite and subgenital plate; 25 Anadolua burri, male end of abdomen; 26 Mixoduda retusa sp. nov. 7th sternite and subgenital plate; 2729. Frons in male: 27 Paradrymadusa aksirayi; 28 Scotodrymadusa turcica; 29 Mixodusa retusa sp. nov.; 3031. Male end of abdomen: 30 Paradrymadusa aksirayi; 31 Anadolua schwarzi; 3237. Ovipositor: 32, 33 Anadolua schwarzi; 34, 35, 36 Anadolua burri; 37 Scotodrymadusa turcica." httpUri="https://zenodo.org/record/1455950/files/figure.png" pageId="15" pageNumber="16">Figs. 26</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFC07FC2B0C41FC4E" box="[1020,1065,978,1003]" captionStart="FIGURES 204221" captionStartId="54.[151,264,1816,1838]" captionTargetBox="[154,1433,196,1789]" captionTargetId="figure@54.[151,1436,193,1794]" captionTargetPageId="54" captionText="FIGURES 204221. Mixodusa retusa sp. nov.: 204 female head, pronotum and tegmina, lateral view; 205 ditto dorsal; 206 female subgenital plate; 207212. Mixodusa bocquilloni: 207 male head, pronotum and tegmina, lateral view; 208 ditto dorsal view; 209 male last tergite and cerci; 210 female head, pronotum and tegmina, lateral view; 211 ditto dorsal view; 212 female subgenital plate; 213218. Anadolua schwarzi: 213 male habitus, lateral view; 214 male head, pronotum and tegmina, dorsal view; 215 male end of abdomen; 216 female habitus, lateral view; 217 female head and pronotum, dorsal view; 218 female subgenital plate and 7th sternite; 219221. Anadolua burri: 219 male habitus, lateral view; 220 male head, pronotum and tegmina, dorsal view; 221 male end of abdomen." httpUri="https://zenodo.org/record/1455968/files/figure.png" pageId="15" pageNumber="16">206</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFBCEFC2A0C0BFC4E" box="[1077,1123,979,1003]" captionStart="FIGURES 312317" captionStartId="60.[151,264,1934,1956]" captionTargetBox="[193,1417,201,1900]" captionTargetId="figure@60.[177,1421,193,1918]" captionTargetPageId="60" captionText="FIGURES 312317. Mixodusa retusa sp. nov.: 312 male habitus, lateral view; 313 female habitus, lateral view (312 and 313 illustrated by Şeyma Hatip); 314 male last tergite; 315 male cercus; 316 titillators; 317 female 7th sternite and subgenital plate. Scales 1 mm." httpUri="https://zenodo.org/record/1455980/files/figure.png" pageId="15" pageNumber="16">317</figureCitation>
); longer than wide with 7th sternite, lateral margins strongly bent upwards, with a deep and wide triangular posterior incision; 7th sternite triangular with smooth surface. Ovipositor (
<figureCitation id="9BCE2A31FFAFFFECFD63FBE30A91FB96" box="[664,761,1050,1075]" captionStart="FIGURES 3858" captionStartId="46.[151,264,1785,1807]" captionTargetBox="[182,1403,195,1768]" captionTargetId="figure@46.[180,1404,193,1773]" captionTargetPageId="46" captionText="FIGURES 3858. Ovipositor: 38 Scotodrymadusa maculata; 39 Mixodusa retusa sp. nov.; 4041. Female habitus lateral view: 40 Anadolua schwarzi; 41 Phytodrymadusa miramae; 4245. Titillators: 42 Scotodrymadusa turcica; 43 Mixodusa retusa sp. nov.; 44 Phytodrymadusa miramae; 45 Leptodusa expugnata; 4647. Male habitus lateral view: 46 Lithodusa helverseni (from Heller &amp; Korsunovskaya 2009); 47 Leptodusa expugnata; 48 Phytodrymadusa miramae, male end of abdomen; 49 Phytodrymadusa miramae, female pronotum and tegmina, dorso-lateral view; 5051. Leptodusa expugnata: 50 male last tergite; 51 male cercus; 5253. Mixodusa retusa sp. nov.: 52 male last tergite, 53 male cercus; 5455. Female pronotum and tegmina: 54 Leptodusa harzi; 55 Mixodusa retusa sp. nov.; 5657. End of hind tibia, ventral view: 56 Decticus albifrons female; 57 Rhacocleis turcica male; 58 Psorodonotus caucasicus, female fore tibia." httpUri="https://zenodo.org/record/1455952/files/figure.png" pageId="15" pageNumber="16">Figs. 39</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFCF2FBE20B5EFB96" box="[777,822,1051,1075]" captionStart="FIGURES 186203" captionStartId="53.[151,264,1835,1857]" captionTargetBox="[191,1407,197,1819]" captionTargetId="figure@53.[186,1410,193,1825]" captionTargetPageId="53" captionText="FIGURES 186203. Habitus lateral view: 186 Sureyaella bella, male; 187 ditto female; 188 Anterastes burri, male; 189 ditto female; 190 Koroglus disparalatus, male; 191 ditto female; 192193. Male tegmina and pronotum, dorsal view: 192 Anterastes serbicus; 193 Koroglus disparalatus; 194195. Anterastes babadaghi: 194 male left tegmen, inner surface; 195 ditto stridulatory file area; 196197. Koroglus disparalatus: 196 male left tegmen, inner surface; 197 ditto stridulatory file area; 198203. Mixodusa retusa sp. nov.: 198 male habitus, lateral view; 199 male head, pronotum and tegmina, lateral view; 200 ditto dorsal view; 201 male last tergite and cerci; 202 male subgenital plate; 203 female habitus, lateral view." httpUri="https://zenodo.org/record/1455966/files/figure.png" pageId="15" pageNumber="16">203</figureCitation>
,
<figureCitation id="9BCE2A31FFAFFFECFCBDFBE20B19FB96" box="[838,881,1051,1075]" captionStart="FIGURES 312317" captionStartId="60.[151,264,1934,1956]" captionTargetBox="[193,1417,201,1900]" captionTargetId="figure@60.[177,1421,193,1918]" captionTargetPageId="60" captionText="FIGURES 312317. Mixodusa retusa sp. nov.: 312 male habitus, lateral view; 313 female habitus, lateral view (312 and 313 illustrated by Şeyma Hatip); 314 male last tergite; 315 male cercus; 316 titillators; 317 female 7th sternite and subgenital plate. Scales 1 mm." httpUri="https://zenodo.org/record/1455980/files/figure.png" pageId="15" pageNumber="16">313</figureCitation>
) slender, thin and long, as long as hind femur, downcurved along its length.
</paragraph>
<paragraph id="034A36B4FFAFFFECFF3CFB9B0946FAD2" blockId="15.[151,1437,258,2011]" pageId="15" pageNumber="16">
<emphasis id="3181EAA6FFAFFFECFF3CFB9B0978FBDE" bold="true" box="[199,272,1122,1147]" pageId="15" pageNumber="16">Color.</emphasis>
Body greyish-brown with black, brown and milky brown markings. Ventral 3/4 of head unicolor milky brown; frons with a weak black band between eyes; dorsal surface of head greyish brown. Antenna light brown. Pronotum bicolored, ventral half of paranota milky brown, above it bordered by a thin blackish band, remaining part and dorsal surface of pronotum greyish brown. Tegmina brown, with darkened main veins and some creamish veinlets. Fore and mid legs with black, greyish brown and creamish irregular spots in 2 rows dorso-anteriorly; outer surface with a black stripe in male. Abdomen greish brown with small irregular black spots. In one female body uniformly light greyish brown. Ovipositor with homogeneously dense blackish spots. Ventral surface of body milky brown.
</paragraph>
</subSubSection>
<subSubSection id="4BEF653FFFAFFFECFF3CFA7B0902F8EE" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="034A36B4FFAFFFECFF3CFA7B0902F8EE" blockId="15.[151,1437,258,2011]" pageId="15" pageNumber="16">
<emphasis id="3181EAA6FFAFFFECFF3CFA7B0929FA3E" bold="true" box="[199,321,1410,1435]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
This new species is most closely related to
<taxonomicName id="C4F54D37FFAFFFECFCD1FA7D0CE9FA3E" authority="(Uvarov, 1917)" baseAuthorityName="Uvarov" baseAuthorityYear="1917" box="[810,1153,1410,1435]" class="Insecta" family="Tettigoniidae" genus="Mixodusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bocquilloni">
<emphasis id="3181EAA6FFAFFFECFCD1FA7D0BA6FA3E" box="[810,974,1410,1435]" italics="true" pageId="15" pageNumber="16">M. bocquilloni</emphasis>
(
<bibRefCitation id="67644B45FFAFFFECFC25FA7A0C10FA3E" box="[990,1144,1411,1435]" pageId="15" pageNumber="16" refId="ref48957" refString="Uvarov, B. P. (1917) Materialia ad conitionem Orthopterorum Caucasi et confinium. II. Diagnoses specierum et subspecierum novarum ex collectione Musei Caucasici. Izvestiya Kavkaszskago Muzeya (Bulletin du Musee du Caucause), 11, 281 - 298." type="journal article">Uvarov, 1917</bibRefCitation>
)
</taxonomicName>
by the similar male cerci and titillators. But it is separated in the male by the last tergite with very short and rounded lobes, with shallow and rounded posterior incision (lobes long and pointed with deep and triangular posterior incision in
<taxonomicName id="C4F54D37FFAFFFECFB15FA350DFCFA46" baseAuthorityName="Uvarov" baseAuthorityYear="1917" box="[1262,1428,1482,1507]" class="Insecta" family="Tettigoniidae" genus="Mixodusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bocquilloni">
<emphasis id="3181EAA6FFAFFFECFB15FA350DFCFA46" box="[1262,1428,1482,1507]" italics="true" pageId="15" pageNumber="16">M. bocquilloni</emphasis>
</taxonomicName>
,
<figureCitation id="9BCE2A31FFAFFFECFF6CFA170892F9A2" box="[151,250,1518,1543]" captionStart="FIGURES 204221" captionStartId="54.[151,264,1816,1838]" captionTargetBox="[154,1433,196,1789]" captionTargetId="figure@54.[151,1436,193,1794]" captionTargetPageId="54" captionText="FIGURES 204221. Mixodusa retusa sp. nov.: 204 female head, pronotum and tegmina, lateral view; 205 ditto dorsal; 206 female subgenital plate; 207212. Mixodusa bocquilloni: 207 male head, pronotum and tegmina, lateral view; 208 ditto dorsal view; 209 male last tergite and cerci; 210 female head, pronotum and tegmina, lateral view; 211 ditto dorsal view; 212 female subgenital plate; 213218. Anadolua schwarzi: 213 male habitus, lateral view; 214 male head, pronotum and tegmina, dorsal view; 215 male end of abdomen; 216 female habitus, lateral view; 217 female head and pronotum, dorsal view; 218 female subgenital plate and 7th sternite; 219221. Anadolua burri: 219 male habitus, lateral view; 220 male head, pronotum and tegmina, dorsal view; 221 male end of abdomen." httpUri="https://zenodo.org/record/1455968/files/figure.png" pageId="15" pageNumber="16">Fig. 209</figureCitation>
), the female subgenital plate with almost smooth surface (with a distinct U-shaped carica in the end of posterior incision in
<taxonomicName id="C4F54D37FFAFFFECFE7EF9ED0A45F98E" baseAuthorityName="Uvarov" baseAuthorityYear="1917" box="[389,557,1554,1579]" class="Insecta" family="Tettigoniidae" genus="Mixodusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bocquilloni">
<emphasis id="3181EAA6FFAFFFECFE7EF9ED0A45F98E" box="[389,557,1554,1579]" italics="true" pageId="15" pageNumber="16">M. bocquilloni</emphasis>
</taxonomicName>
,
<figureCitation id="9BCE2A31FFAFFFECFDC6F9EB0AC9F98E" box="[573,673,1554,1579]" captionStart="FIGURES 204221" captionStartId="54.[151,264,1816,1838]" captionTargetBox="[154,1433,196,1789]" captionTargetId="figure@54.[151,1436,193,1794]" captionTargetPageId="54" captionText="FIGURES 204221. Mixodusa retusa sp. nov.: 204 female head, pronotum and tegmina, lateral view; 205 ditto dorsal; 206 female subgenital plate; 207212. Mixodusa bocquilloni: 207 male head, pronotum and tegmina, lateral view; 208 ditto dorsal view; 209 male last tergite and cerci; 210 female head, pronotum and tegmina, lateral view; 211 ditto dorsal view; 212 female subgenital plate; 213218. Anadolua schwarzi: 213 male habitus, lateral view; 214 male head, pronotum and tegmina, dorsal view; 215 male end of abdomen; 216 female habitus, lateral view; 217 female head and pronotum, dorsal view; 218 female subgenital plate and 7th sternite; 219221. Anadolua burri: 219 male habitus, lateral view; 220 male head, pronotum and tegmina, dorsal view; 221 male end of abdomen." httpUri="https://zenodo.org/record/1455968/files/figure.png" pageId="15" pageNumber="16">Fig. 212</figureCitation>
), female 7th sternite with smooth surface (with a longitudinal low carina in
<taxonomicName id="C4F54D37FFAFFFECFEFAF9CE09CEF9EA" baseAuthorityName="Uvarov" baseAuthorityYear="1917" box="[257,422,1590,1615]" class="Insecta" family="Tettigoniidae" genus="Mixodusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bocquilloni">
<emphasis id="3181EAA6FFAFFFECFEFAF9CE09CEF9EA" box="[257,422,1590,1615]" italics="true" pageId="15" pageNumber="16">M. bocquilloni</emphasis>
</taxonomicName>
,
<figureCitation id="9BCE2A31FFAFFFECFE4EF9CF0A7FF9EA" box="[437,535,1590,1615]" captionStart="FIGURES 204221" captionStartId="54.[151,264,1816,1838]" captionTargetBox="[154,1433,196,1789]" captionTargetId="figure@54.[151,1436,193,1794]" captionTargetPageId="54" captionText="FIGURES 204221. Mixodusa retusa sp. nov.: 204 female head, pronotum and tegmina, lateral view; 205 ditto dorsal; 206 female subgenital plate; 207212. Mixodusa bocquilloni: 207 male head, pronotum and tegmina, lateral view; 208 ditto dorsal view; 209 male last tergite and cerci; 210 female head, pronotum and tegmina, lateral view; 211 ditto dorsal view; 212 female subgenital plate; 213218. Anadolua schwarzi: 213 male habitus, lateral view; 214 male head, pronotum and tegmina, dorsal view; 215 male end of abdomen; 216 female habitus, lateral view; 217 female head and pronotum, dorsal view; 218 female subgenital plate and 7th sternite; 219221. Anadolua burri: 219 male habitus, lateral view; 220 male head, pronotum and tegmina, dorsal view; 221 male end of abdomen." httpUri="https://zenodo.org/record/1455968/files/figure.png" pageId="15" pageNumber="16">Fig. 212</figureCitation>
), pronotum with more truncate, broadly rounded posterior margin (strongly and narrowly rounded in
<taxonomicName id="C4F54D37FFAFFFECFE7EF9A50A42F9D6" baseAuthorityName="Uvarov" baseAuthorityYear="1917" box="[389,554,1626,1651]" class="Insecta" family="Tettigoniidae" genus="Mixodusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bocquilloni">
<emphasis id="3181EAA6FFAFFFECFE7EF9A50A42F9D6" box="[389,554,1626,1651]" italics="true" pageId="15" pageNumber="16">M. bocquilloni</emphasis>
</taxonomicName>
,
<figureCitation id="9BCE2A31FFAFFFECFDC2F9A30B10F9D6" box="[569,888,1626,1651]" captionStart="FIGURES 204221" captionStartId="54.[151,264,1816,1838]" captionTargetBox="[154,1433,196,1789]" captionTargetId="figure@54.[151,1436,193,1794]" captionTargetPageId="54" captionText="FIGURES 204221. Mixodusa retusa sp. nov.: 204 female head, pronotum and tegmina, lateral view; 205 ditto dorsal; 206 female subgenital plate; 207212. Mixodusa bocquilloni: 207 male head, pronotum and tegmina, lateral view; 208 ditto dorsal view; 209 male last tergite and cerci; 210 female head, pronotum and tegmina, lateral view; 211 ditto dorsal view; 212 female subgenital plate; 213218. Anadolua schwarzi: 213 male habitus, lateral view; 214 male head, pronotum and tegmina, dorsal view; 215 male end of abdomen; 216 female habitus, lateral view; 217 female head and pronotum, dorsal view; 218 female subgenital plate and 7th sternite; 219221. Anadolua burri: 219 male habitus, lateral view; 220 male head, pronotum and tegmina, dorsal view; 221 male end of abdomen." httpUri="https://zenodo.org/record/1455968/files/figure.png" pageId="15" pageNumber="16">Figs. 207208 and 210211</figureCitation>
), shorter tegmina (in
<taxonomicName id="C4F54D37FFAFFFECFB91F9A50D67F9D6" baseAuthorityName="Uvarov" baseAuthorityYear="1917" box="[1130,1295,1626,1651]" class="Insecta" family="Tettigoniidae" genus="Mixodusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bocquilloni">
<emphasis id="3181EAA6FFAFFFECFB91F9A50D67F9D6" box="[1130,1295,1626,1651]" italics="true" pageId="15" pageNumber="16">M. bocquilloni</emphasis>
</taxonomicName>
, reaching to half-end of 2nd tergite in male,
<figureCitation id="9BCE2A31FFAFFFECFDF9F9870ACEF932" box="[514,678,1662,1687]" captionStart="FIGURES 204221" captionStartId="54.[151,264,1816,1838]" captionTargetBox="[154,1433,196,1789]" captionTargetId="figure@54.[151,1436,193,1794]" captionTargetPageId="54" captionText="FIGURES 204221. Mixodusa retusa sp. nov.: 204 female head, pronotum and tegmina, lateral view; 205 ditto dorsal; 206 female subgenital plate; 207212. Mixodusa bocquilloni: 207 male head, pronotum and tegmina, lateral view; 208 ditto dorsal view; 209 male last tergite and cerci; 210 female head, pronotum and tegmina, lateral view; 211 ditto dorsal view; 212 female subgenital plate; 213218. Anadolua schwarzi: 213 male habitus, lateral view; 214 male head, pronotum and tegmina, dorsal view; 215 male end of abdomen; 216 female habitus, lateral view; 217 female head and pronotum, dorsal view; 218 female subgenital plate and 7th sternite; 219221. Anadolua burri: 219 male habitus, lateral view; 220 male head, pronotum and tegmina, dorsal view; 221 male end of abdomen." httpUri="https://zenodo.org/record/1455968/files/figure.png" pageId="15" pageNumber="16">Figs. 207208</figureCitation>
, and to half of first tergite in female,
<figureCitation id="9BCE2A31FFAFFFECFBA8F9870C9FF932" box="[1107,1271,1662,1687]" captionStart="FIGURES 204221" captionStartId="54.[151,264,1816,1838]" captionTargetBox="[154,1433,196,1789]" captionTargetId="figure@54.[151,1436,193,1794]" captionTargetPageId="54" captionText="FIGURES 204221. Mixodusa retusa sp. nov.: 204 female head, pronotum and tegmina, lateral view; 205 ditto dorsal; 206 female subgenital plate; 207212. Mixodusa bocquilloni: 207 male head, pronotum and tegmina, lateral view; 208 ditto dorsal view; 209 male last tergite and cerci; 210 female head, pronotum and tegmina, lateral view; 211 ditto dorsal view; 212 female subgenital plate; 213218. Anadolua schwarzi: 213 male habitus, lateral view; 214 male head, pronotum and tegmina, dorsal view; 215 male end of abdomen; 216 female habitus, lateral view; 217 female head and pronotum, dorsal view; 218 female subgenital plate and 7th sternite; 219221. Anadolua burri: 219 male habitus, lateral view; 220 male head, pronotum and tegmina, dorsal view; 221 male end of abdomen." httpUri="https://zenodo.org/record/1455968/files/figure.png" pageId="15" pageNumber="16">Figs. 210211</figureCitation>
; see also
<bibRefCitation id="67644B45FFAFFFECFA9DF98708BCF91E" pageId="15" pageNumber="16" refId="ref49479" refString="Unal, M. (2012) Tettigoniidae (Orthoptera) from Turkey and the Middle East II. Transactions of the American Entomological Society, 1382 (1 - 4), 21 - 54." type="journal article">Ünal 2012</bibRefCitation>
: Figs. 912 and 8891). It is different from
<taxonomicName id="C4F54D37FFAFFFECFD36F95D0B2AF91E" baseAuthorityName="Uvarov" baseAuthorityYear="1929" box="[717,834,1700,1723]" class="Insecta" family="Tettigoniidae" genus="Mixodusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="siazovi">
<emphasis id="3181EAA6FFAFFFECFD36F95D0B2AF91E" box="[717,834,1700,1723]" italics="true" pageId="15" pageNumber="16">M. siazovi</emphasis>
</taxonomicName>
by the shape of cerci with much shorter inner tooth, male last tergite, shorter tegmina in both sexes (in
<taxonomicName id="C4F54D37FFAFFFECFD1FF93E0B31F97B" baseAuthorityName="Uvarov" baseAuthorityYear="1929" box="[740,857,1735,1758]" class="Insecta" family="Tettigoniidae" genus="Mixodusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="siazovi">
<emphasis id="3181EAA6FFAFFFECFD1FF93E0B31F97B" box="[740,857,1735,1758]" italics="true" pageId="15" pageNumber="16">M. siazovi</emphasis>
</taxonomicName>
, male tegmina reaching to half-end of 3rd tergite, female tegmina reaching to almost half of 2nd tergite), the fused female 7th sternite with smooth surface (7th sternite separated from subgenital plate in
<taxonomicName id="C4F54D37FFAFFFECFD79F8F60A90F883" baseAuthorityName="Uvarov" baseAuthorityYear="1929" box="[642,760,1807,1830]" class="Insecta" family="Tettigoniidae" genus="Mixodusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="siazovi">
<emphasis id="3181EAA6FFAFFFECFD79F8F60A90F883" box="[642,760,1807,1830]" italics="true" pageId="15" pageNumber="16">M. siazovi</emphasis>
</taxonomicName>
with a high conical acute projection, see
<bibRefCitation id="67644B45FFAFFFECFB20F8F70D3FF882" box="[1243,1367,1806,1831]" pageId="15" pageNumber="16" refId="ref49479" refString="Unal, M. (2012) Tettigoniidae (Orthoptera) from Turkey and the Middle East II. Transactions of the American Entomological Society, 1382 (1 - 4), 21 - 54." type="journal article">Ünal 2012</bibRefCitation>
: Figs. 1316 and 9195).
</paragraph>
</subSubSection>
<subSubSection id="4BEF653FFFAFFFECFF3CF8AF0C62F812" pageId="15" pageNumber="16" type="description">
<paragraph id="034A36B4FFAFFFECFF3CF8AF0C62F812" blockId="15.[151,1437,258,2011]" pageId="15" pageNumber="16">
<emphasis id="3181EAA6FFAFFFECFF3CF8AF09A0F8CA" bold="true" box="[199,456,1878,1903]" pageId="15" pageNumber="16">Measurements (mm).</emphasis>
<typeStatus id="DC4E8816FFAFFFECFE29F8AF0A57F8CA" box="[466,575,1878,1903]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
: body 28.1; pronotum 9.2; tegmina 5.3; hind femur 28.8.
<typeStatus id="DC4E8816FFAFFFECFB25F8AE0D27F8CA" box="[1246,1359,1879,1903]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
: body: male 27.631.4, female 24.830.6; pronotum: male 8.79.8, female 8.49.5; tegmina: male 5.25.6, female 2.83.7; hind femur: male 26.830.4, female 27.130.6; ovipositor: 25.328.2.
</paragraph>
</subSubSection>
<subSubSection id="4BEF653FFFAFFFF3FF3CF83B0948FF70" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="034A36B4FFAFFFF3FF3CF83B0948FF70" blockId="15.[151,1437,258,2011]" lastBlockId="16.[151,1436,151,285]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="3181EAA6FFAFFFECFF3CF83B09DCF87E" bold="true" box="[199,436,1986,2011]" pageId="15" pageNumber="16">Material examined.</emphasis>
<collectingCountry id="7BE27624FFAFFFECFE44F83A0A5BF87F" box="[447,563,1987,2010]" name="Turkey" pageId="15" pageNumber="16">TURKEY</collectingCountry>
:
<collectingRegion id="C131F856FFAFFFECFDBBF83B0AE4F87E" box="[576,652,1986,2011]" country="Turkey" name="Sirnak" pageId="15" pageNumber="16">Şırnak</collectingRegion>
, Beytüşşebab, Günyüzü Köyü,
<quantity id="C40D9B51FFAFFFECFC06F83A0C30F87F" box="[1021,1112,1987,2011]" metricMagnitude="3" metricUnit="m" metricValue="1.374" pageId="15" pageNumber="16" unit="m" value="1374.0">1374 m</quantity>
, 37.27.382 N, 43.12.123 E,
<date id="774B1074FFB0FFF3FF6CFF6E0893FF15" box="[151,251,151,176]" pageId="16" pageNumber="17" value="2013-09-06">6.9.2013</date>
,
<specimenCount id="15F3FD3DFFB0FFF3FEF2FF610909FF15" box="[265,353,151,176]" pageId="16" pageNumber="17" type="male">2 males</specimenCount>
(including
<typeStatus id="DC4E8816FFB0FFF3FE1CFF6E0A25FF15" box="[487,589,151,176]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
),
<specimenCount id="15F3FD3DFFB0FFF3FD99FF610ABBFF15" box="[610,723,151,176]" pageId="16" pageNumber="17" type="female">2 females</specimenCount>
, plus
<specimenCount id="15F3FD3DFFB0FFF3FCE3FF610B01FF15" box="[792,873,151,176]" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
,
<specimenCount id="15F3FD3DFFB0FFF3FC8DFF610B8DFF15" box="[886,997,151,176]" pageId="16" pageNumber="17" type="female">3 females</specimenCount>
in alcohol (leg. M. Ünal &amp; A. Erden) (AİBÜEM).
</paragraph>
</subSubSection>
<subSubSection id="4BEF653FFFB0FFF3FF3CFF2608B2FEB8" pageId="16" pageNumber="17" type="etymology">
<paragraph id="034A36B4FFB0FFF3FF3CFF2608B2FEB8" blockId="16.[151,1436,151,285]" pageId="16" pageNumber="17">
<emphasis id="3181EAA6FFB0FFF3FF3CFF260926FF5D" bold="true" box="[199,334,223,248]" pageId="16" pageNumber="17">Etymology.</emphasis>
“retusus” means blunt in Latin, relating to the males last tergite with blunt, rounded posterior lobes.
</paragraph>
</subSubSection>
</treatment>
</document>