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<document id="4010A02825AC17A5C2945151E3B29E40" ID-DOI="10.3897/zookeys.1212.126536" ID-ZooBank="4A550F74-C788-4282-A731-B8405CA8B8F5" ID-publisher-id="126536" URI-arpha="FC3213C5-BED0-512A-9C14-5F0A39ADDBC7" URI-zoobank="http://zoobank.org/4A550F74-C788-4282-A731-B8405CA8B8F5" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1726239268422" checkinUser="pensoft" docAuthor="Srisonchai, Ruttapon, Likhitrakarn, Natdanai, Sutcharit, Chirasak &amp; Wesener, Thomas" docDate="2024" docId="8B321F4E2E9F58A99B3D07B9612F036A" docLanguage="en" docName="ZooKeys 1212: 29-64" docOrigin="ZooKeys 1212" docSource="https://zookeys.pensoft.net/article/126536/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Zephronia Gray 1832" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="64" masterDocId="FC3213C5BED0512A9C145F0A39ADDBC7" masterDocTitle="Integrative taxonomy reveals two new giant pill-millipedes of the genus Zephronia Gray, 1832 from eastern Thailand (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="64" masterPageNumber="29" pageNumber="29" updateTime="1726249556959" updateUser="admin">
<mods:mods id="0436D56473A63EC05CD93BAE18F86900" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="61510988DBFAB2760ADFBA120187F427">
<mods:title id="2D682F28653FC35D9EA61807A7502DEC">Integrative taxonomy reveals two new giant pill-millipedes of the genus Zephronia Gray, 1832 from eastern Thailand (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
</mods:titleInfo>
<mods:name id="A568CDA954C8F36795BBE13212BFFDCC" type="personal">
<mods:role id="A7975E1B24B53FC6ADF79C721C96E349">
<mods:roleTerm id="05BAFF960A0D8310DB9177C576701282">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="577D68B5F60ACC73B8B951A41DCD97AE">Srisonchai, Ruttapon</mods:namePart>
<mods:nameIdentifier id="6F0A5BEF790383239446B0C85CA568B2" type="ORCID">0000-0002-7142-0999</mods:nameIdentifier>
<mods:affiliation id="32D20CF890962F5CBA1BA78E11C73261">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand</mods:affiliation>
</mods:name>
<mods:name id="28F3D5603607CCAA6D4F546B3AA11D1B" type="personal">
<mods:role id="71CF4F2474F79B46514CAD139F21BAD2">
<mods:roleTerm id="48AE6BDBEDE280B3AC5886690FD3CEBD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="628367B7EB022A01034F5739DD72A52F">Likhitrakarn, Natdanai</mods:namePart>
<mods:nameIdentifier id="AC35879599934C500B7D085E107E4060" type="ORCID">0000-0002-1306-317X</mods:nameIdentifier>
<mods:affiliation id="FAFC69AE8C97927CD4DC40E36EAAB520">Program of Agriculture, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand</mods:affiliation>
</mods:name>
<mods:name id="3AB650CD6981010B5C2001238B14B0C1" type="personal">
<mods:role id="87443A05827CDBCB69600C13F5AE47AB">
<mods:roleTerm id="6756BD4E2D2A62B5BD45922C1D48DA69">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1031B186D8190E051405CDA69F1B1531">Sutcharit, Chirasak</mods:namePart>
<mods:nameIdentifier id="33514CBE3C3F4B9A7293C3EE8DD0CC1C" type="ORCID">0000-0001-7670-9540</mods:nameIdentifier>
<mods:affiliation id="580801D08B3A220E87D82C9CB14DBC9D">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Phayathai Road, Patumwan, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name id="2C63ACA6263A881CA8D69244DFE10AF8" type="personal">
<mods:role id="89FBB6AB3CB6B14013FD1DE45888A8AE">
<mods:roleTerm id="248FBD4DAD640AACB52721DD7559AF34">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F972EC9AA71225FA98F865A524662065">Wesener, Thomas</mods:namePart>
<mods:nameIdentifier id="A036B046D25159329B9A67AA549E6C7D" type="ORCID">0000-0002-2028-3541</mods:nameIdentifier>
<mods:affiliation id="8E2B95B640186E17A40E4712EB6E9399">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Study of Biodiversity Change (LIB), Adenauerallee 160, D- 53113 Bonn, Germany</mods:affiliation>
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<mods:title id="16A63EB1DBC7C52838104B7C30BF3088">ZooKeys</mods:title>
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<mods:part id="94452274519B0F25CA66FC2A6870F584">
<mods:date id="045B36001DEE75B08AA31DB139FB1555">2024</mods:date>
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<mods:number id="ED458CE84D92BEE287E39D3FB6E939EA">2024-09-13</mods:number>
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<mods:detail id="3F382032BDD228E85208AAFC61D38157" type="volume">
<mods:number id="03C23367FDEA1CE03A0A67E82DD7B5EE">1212</mods:number>
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<mods:classification id="8156830435C2BCCDE2CA48E9100568C1">journal article</mods:classification>
<mods:identifier id="3CA06A8E51A8909E893027DB91FC9EC6" type="DOI">10.3897/zookeys.1212.126536</mods:identifier>
<mods:identifier id="D6306463CD9048F9AE7D58EBF43B5193" type="ZooBank">4A550F74-C788-4282-A731-B8405CA8B8F5</mods:identifier>
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<treatment id="8B321F4E2E9F58A99B3D07B9612F036A" ID-arpha="8B321F4E-2E9F-58A9-9B3D-07B9612F036A" LSID="urn:lsid:plazi:treatment:8B321F4E2E9F58A99B3D07B9612F036A" httpUri="http://treatment.plazi.org/id/8B321F4E2E9F58A99B3D07B9612F036A">
<subSubSection id="F924A0EAB9AF02023D55BFA12550825C" type="nomenclature">
<paragraph id="C37930575F893A45B703E4B0D13482D3">
<heading id="3C363E2190A07D2CAFC1F358F185F494" reason="label">Genus</heading>
<taxonomicName id="FBE8B3ACD585E06F2C0E9AE5151FEE3D" ID-CoL="7Q6XV" authority="Gray, 1832" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="6A45EFDC6C74290760F9374B4E55D7A7" italics="true">Zephronia</emphasis>
Gray, 1832
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJRBI" type="Differential diagnosis">
<paragraph id="79E671FE77F4DFC4FCF7A3C501B75CAF">
<heading id="23D2A3789986B270996A7F81B4C80523" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="6D1019501767D919862550319CB859FC">
Differs from all other genera of
<taxonomicName id="9EA96F42BFA9889D4908F707FB8B7406" authorityName="Gray" authorityYear="1843" class="Diplopoda" family="Zephroniidae" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="subFamily" subFamily="Zephroniinae">Zephroniinae</taxonomicName>
by the combination of the following characters: 1) Body length
<quantity id="88101FF3D4FB5DE9166C45B07E41A645" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="5.0" metricValueMin="1.8" unit="mm" value="34.0" valueMax="50.0" valueMin="18.0">18–50 mm</quantity>
. 2) Antennae flattened laterally, usually axe-shaped, with numerous (&gt; 4) apical cones. 3) Endotergum (tergite underside) weakly modified: posterior margin (
<abbrev id="ABBRID0E4RBI" xlink_title="posterior margin of endotergum">pm</abbrev>
) usually flat; outer area (os) without setae; marginal bristles arranged in 1–5 rows; middle area (
<abbrev id="ABBRID0EBSBI" xlink_title="middle area of endotergum">ma</abbrev>
) often with a single row of circular cuticular impressions. 4) Tarsi of legs 5–21 often with more than one apical spine (with very few species with only 1) and several ventral spines. 5) Anterior telopods with four podomeres distal to syncoxite; telopoditomere 2 with a large, curved process forming a clamp-like; telopoditomeres 3 and 4 simple. 6) Posterior telopods with four telopoditomeres; immovable finger (process of telopoditomere 2 slender, apically curved; telopoditomeres 3 and 4 with two membranous lobes; telopoditomere 3 also with a row of conspicuous, crenulated teeth, larger than elopoditomere 4; telopoditomere 4 with few (one or two) sclerotized spines. 7) Female vulvae conspicuous, slender; operculum usually round.
</paragraph>
<paragraph id="9289B60C69B76AB7465A5B452B3DA378">
<taxonomicName id="2349CD9E16DD6BF76C760A5A7D7BB742" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="F8ADE0A5CBE7B0C03CEFAFDF10EBB873" italics="true">Zephronia</emphasis>
</taxonomicName>
is one of the most species-rich genera of the family with more than 50 species currently placed in the genus. Numerous species currently placed in the genus are in need of a revision, some only known from the female, and might be placed in separate genera in the future. The posterior telopod in
<taxonomicName id="876256237D44D219647E41DD1AD6B169" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="8CE606F3D427991B60C596099E79A376" italics="true">Zephronia</emphasis>
</taxonomicName>
consists of four podomeres, unlike
<taxonomicName id="FBBB92D85ED334C11D7E9186254F3BB0" authority="Pocock, 1895" authorityName="Pocock" authorityYear="1895" class="Diplopoda" family="Zephroniidae" genus="Castanotherium" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="0ED9918229EBC411B33925BF80563FEB" italics="true">Castanotherium</emphasis>
Pocock, 1895
</taxonomicName>
whose species have three podomeres only. Podomere 4 of the posterior telopod in
<taxonomicName id="6CB3D356551FA8E9A46ADA046ABFA6B1" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="14BA2DF645E0DDD2992F5BE9B78EB935" italics="true">Zephronia</emphasis>
</taxonomicName>
is not strongly curved or overlapping the process of podomere 2 as in
<taxonomicName id="F0238BD06727E9F87BE305AF6B7EF247" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" class="Diplopoda" family="Zephroniidae" genus="Cryxus" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="E648018FACA447D18C6B95FB893C54DD" italics="true">Cryxus</emphasis>
Leach, 1814
</taxonomicName>
. The process of podomere
<quantity id="C78A7C0CC4571835607AA10D1A975CCE" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="EEAA5A78FB5D281C75DF465D5217B85F" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="DB3A2F799BC5A318A87197F73C47ADF9" italics="true">Zephronia</emphasis>
</taxonomicName>
is never apically enlarged or swollen like in
<taxonomicName id="8A29D0DEA3342B4C4D110612117AA38F" authority="Verhoeff, 1924" authorityName="Verhoeff" authorityYear="1924" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="AE37D14ABDF0A6C3263EB2E76E61F205" italics="true">Sphaerobelum</emphasis>
Verhoeff, 1924
</taxonomicName>
. The osterior telopod on podomere
<quantity id="3223302B7FDBD1FDAAFADFD2A8170BB6" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="1D0318FF2E371722D2990EF3EA1C4617" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="C0474FDC5C82892B2A4E9C8C3DBC03FF" italics="true">Zephronia</emphasis>
</taxonomicName>
has crenulated teeth, unlike
<taxonomicName id="655B2FCC12364E5D32DCC49FFE42D740" authority="Attems, 1935" authorityName="Attems" authorityYear="1935" class="Diplopoda" family="Zephroniidae" genus="Kophosphaera" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="7903863BA09BAC1C008043473E16B2FA" italics="true">Kophosphaera</emphasis>
Attems, 1935
</taxonomicName>
which lacks sclerotized teeth or spines entirely. The posterior telopod of
<taxonomicName id="A83E97EA1322343E8102A03C30A365AF">
<emphasis id="0843A56FC91D50583CE9E5DD437E1D0F" italics="true">Zephronia</emphasis>
species
</taxonomicName>
is identical to those of species of
<taxonomicName id="43D53B519F608130EA77D87D47538507" authority="Brandt, 1833" authorityName="Brandt" authorityYear="1833" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="B5EFA84D3ACFD54088FBB2EB7753DC27" italics="true">Sphaeropoeus</emphasis>
Brandt, 1833
</taxonomicName>
,
<taxonomicName id="626775A8C5A2A0D66D82E5528AF7506D" authority="Verhoeff, 1924" authorityName="Verhoeff" authorityYear="1924" class="Diplopoda" family="Zephroniidae" genus="Prionobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="E504A95BBBC5791344B44860F0469824" italics="true">Prionobelum</emphasis>
Verhoeff, 1924
</taxonomicName>
,
<taxonomicName id="B6047D96967AAC58A622016CD7A0943F" authority="Jeekel, 2000" authorityName="Jeekel" authorityYear="2000" class="Diplopoda" family="Zephroniidae" genus="Tigridosphaera" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="50B4DED798DBA083D6D358C68FB26693" italics="true">Tigridosphaera</emphasis>
Jeekel, 2000
</taxonomicName>
, and
<taxonomicName id="29D1F4B6F26B6693545275578E133577" authority="Attems, 1935" authorityName="Attems" authorityYear="1935" class="Diplopoda" family="Zephroniidae" genus="Indosphaera" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="C5230361BC628C4E1A4F627F63306CEC" italics="true">Indosphaera</emphasis>
Attems, 1935
</taxonomicName>
.
<taxonomicName id="06CB4C76EE1F3EEC6B2F03AE9E660A5E">
<emphasis id="C3C70E87AFEA1549B864767DF2E6376B" italics="true">Zephronia</emphasis>
species
</taxonomicName>
differ from species of
<taxonomicName id="A285F8FF78063BE4CCA7B6899F8C9A89" authorityName="Brandt" authorityYear="1833" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="638AFF49BBA22EB0FA2641EA077FE295" italics="true">Sphaeropoeus</emphasis>
</taxonomicName>
and
<taxonomicName id="9AD5C985EFAEC648478C16B029E44CEB" authorityName="Verhoeff" authorityYear="1924" class="Diplopoda" family="Zephroniidae" genus="Prionobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="E092027B23024E34CCC92C881112F1F9" italics="true">Prionobelum</emphasis>
</taxonomicName>
in the anterior telopods, lacking the characteristic processes on joints three (
<taxonomicName id="64C58EC80C6078270161E1822E9682CB" authorityName="Brandt" authorityYear="1833" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="4B7A5F54B5221E199BA3291E8E2A2DBF" italics="true">Sphaeropoeus</emphasis>
</taxonomicName>
) and four (
<taxonomicName id="FEB9E5ED31C648E522C446F6E45DA6E4" authorityName="Verhoeff" authorityYear="1924" class="Diplopoda" family="Zephroniidae" genus="Prionobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="7BEFAC9EB8573AA9FF07EBA3416F6CCC" italics="true">Prionobelum</emphasis>
</taxonomicName>
).
<taxonomicName id="628218F3DFF8F44A435DBC1B5D62740E">
<emphasis id="F8960272684F8DABD4773AE9BE99790B" italics="true">Zephronia</emphasis>
species
</taxonomicName>
differ from
<taxonomicName id="AD8AF31E0819FC787614AF0243C1794D" authorityName="Attems" authorityYear="1935" class="Diplopoda" family="Zephroniidae" genus="Indosphaera" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="042E876BB966DD6E389901B7D5661A3C" italics="true">Indosphaera</emphasis>
</taxonomicName>
in the coxae of leg
<quantity id="99F29DDD3120E2EF5A5AFC774CFF6229" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
females being separate, not fused.
<taxonomicName id="323D18BBAA2D7233F3370AD2AF747194">
<emphasis id="A7F27FB161D4AA939B85A641411D268B" italics="true">Zephronia</emphasis>
species
</taxonomicName>
are currently impossible to distinguish from species of
<taxonomicName id="F118926A2DC6D1F67EB644996C6EDE7C" authorityName="Jeekel" authorityYear="2000" class="Diplopoda" family="Zephroniidae" genus="Tigridosphaera" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="C2F3474706F902F31C645470E2ABEDD2" italics="true">Tigridosphaera</emphasis>
</taxonomicName>
as that species-poor genus is in need of revision. Females of
<taxonomicName id="327354854999B55D22CBF89685872CC5" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="4D79240BBC18B6E6B0170CD834C44404" italics="true">Zephronia</emphasis>
</taxonomicName>
differ in their shape of the operculum of the vulva from species of
<taxonomicName id="1F3854FD81252DCD8C0EDCA12F6ECA71" authorityName="Attems" authorityYear="1935" class="Diplopoda" family="Zephroniidae" genus="Indosphaera" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="B70005B96935ABE0CB9DFDAA11B0A093" italics="true">Indosphaera</emphasis>
</taxonomicName>
, where it is much lower than in
<taxonomicName id="DBD902F2CDAC1D0FF8AAA4B8F41C249E">
<emphasis id="71C90806402678331205A173DAA14F8B" italics="true">Zephronia</emphasis>
species
</taxonomicName>
, as well as from species of
<taxonomicName id="87FD5406AD01128AAC1C85E05BD41BF7" authorityName="Brandt" authorityYear="1833" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="EEB7C47AC396FBF45C7E7B34EF5EF49C" italics="true">Sphaeropoeus</emphasis>
</taxonomicName>
and
<taxonomicName id="88CED9EA47FBC33AF9C1FC8AE408C875" authorityName="Verhoeff" authorityYear="1924" class="Diplopoda" family="Zephroniidae" genus="Prionobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="853A0FBB928988D816A1B00898BFE8CB" italics="true">Prionobelum</emphasis>
</taxonomicName>
where the operculum is greatly enlarged and almost rectangular in shape.
</paragraph>
<paragraph id="F067F8384EEF66F8EF8E71719CEE1259">
<title id="59D90B2341F342C2808860D05F7DA009">
<taxonomicName id="D088367589F1064E6FB55FA6C61A4665" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="4DD03A68C84F2317B7F70252B6F6A91B" italics="true">Zephronia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A06D8ABE1CA2918446BE6912AA6E07D7" sensu="stricto">s. s.</taxonomicNameLabel>
</title>
</paragraph>
<paragraph id="CB6AF2673F8ED476A96DAED173ADA833">
Differing from all
<taxonomicName id="4E67F812574FB07B60450E864B387E21" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="6AC29C1D3CE01069676DEE52845AF871" italics="true">Zephronia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D73BAA5774D1281CBA1B31AC831ED01E" sensu="lato">s. l.</taxonomicNameLabel>
by the position of the organ of Tömösváry, which is located at the brim and not inside the antennal groove as in all other known
<taxonomicName id="D0C634B0CFFCDAC2B301A48A4C2E0666" authorityName="Brandt" authorityYear="1833" class="Diplopoda" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="order">Sphaerotheriida</taxonomicName>
.
<taxonomicName id="FFC4FA206ED96C7E17A4427F7212A249" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="EE0B84B41C6D05DDE559B5D567778AE6" italics="true">Zephronia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D82484D4A97EBD1E918DACCF94A7356F" sensu="stricto">s. s.</taxonomicNameLabel>
includes the
<typeStatus id="751C945C3019C5AC958136305D95B90B">type</typeStatus>
species of the genus,
<taxonomicName id="389F99D51649A51F240BDF5FA5033B76" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Diplopoda" family="Zephroniidae" genus="Cryxus" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="3097C9A4C6E2A6BB1116066772F79164" italics="true">Z. ovalis</emphasis>
</taxonomicName>
, as well as
<taxonomicName id="E542CB68C62E483112A3736BADE68EA8" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="chantaburiensis" status="sp. nov.">
<emphasis id="27D611C3599D19D394E6AF41D017AB0D" italics="true">Z. chantaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33D2C518DF86B648A7EE1AAD280834A0" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="EF43C0122B3DE2AE171A55FB456BF01A" authorityName="Bhansali &amp; Wesener" authorityYear="2022" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="chrysomallos">
<emphasis id="E24C944D87CD31704C0D7945C60C5890" italics="true">Z. chrysomallos</emphasis>
</taxonomicName>
,
<taxonomicName id="7A2AD52872F6BD803DBB7B551553CED3" authorityName="Attems" authorityYear="1953" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="dawydoffi">
<emphasis id="9158E86FD308AAFE73C757AF79F1BFA6" italics="true">Z. dawydoffi</emphasis>
</taxonomicName>
,
<taxonomicName id="FD93FEF3B822EC0B0B72DEAE36C6EF16" authorityName="Bhansali &amp; Wesener" authorityYear="2022" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="erawani">
<emphasis id="3CB5AFE1055EC58AF70EB24829C5340D" italics="true">Z. erawani</emphasis>
</taxonomicName>
,
<taxonomicName id="65CC5B8562D453498E1D888D323BF657" authorityName="Srisonchai &amp; Likhitrakarn" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="enghoffi">
<emphasis id="2ED80681E7AA829099F48B12BAA9D65E" italics="true">Z. enghoffi</emphasis>
</taxonomicName>
,
<taxonomicName id="8004D908752431EA0951F78ABC80A45A" authorityName="Srisonchai, Sutcharit &amp; Likhitrakarn" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="4A4D137D6458D1F4B167B84163227A12" italics="true">Z. golovatchi</emphasis>
</taxonomicName>
,
<taxonomicName id="123932FB00502FE3D43824B98426E1F4" authority="Liu &amp; Wesener, 2022" authorityName="Liu &amp; Wesener" authorityYear="2022" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="hui">
<emphasis id="0EBA3A1E5F71C8CCA837543F4EE19067" italics="true">Z. hui</emphasis>
Liu &amp; Wesener, 2022
</taxonomicName>
,
<taxonomicName id="81A3F2EF3DE2DDBAC9D961332B0C095A" authority="Semenyuk, Golovatch &amp; Wesener, 2018" authorityName="Semenyuk, Golovatch &amp; Wesener" authorityYear="2018" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="konkakinhensis">
<emphasis id="14A281D60D1EC789F09BCDD42F60703E" italics="true">Z. konkakinhensis</emphasis>
<bibRefCitation id="5C47AA94265F5AC3FEEE56FC3545DC40" DOI="10.11646/zootaxa.4459.3.7" author="Semenyuk I &amp; Golovatch SI &amp; Wesener T" issue="3" journalOrPublisher="Zootaxa" pagination="535-550" refId="B27" refString="Semenyuk I, Golovatch SI, Wesener T (2018) Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae). Zootaxa 4459 (3): 535 - 550. https: // doi. org / 10.11646 / zootaxa. 4459.3. 7" title="Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae)." volume="4459" year="2018">Semenyuk, Golovatch &amp; Wesener, 2018</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="CD6ABCCE6786EBBFC5834E1C781C3DDF" authorityName="Likhitrakarn &amp; Golovatch" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="lannaensis">
<emphasis id="20241FB282ABFD1B11495E8CB3532BD8" italics="true">Z. lannaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="9F21DBFFC7855A8F089B695DF02FEAE8" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="laotica">
<emphasis id="6E4C0AF10272247FECE1E780AA577478" italics="true">Z. laotica</emphasis>
</taxonomicName>
,
<taxonomicName id="0258EA2D3757D46B602DECED629CA5D4" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="B722D6627C86D258B8123E81DDEC656F" italics="true">Z. macula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F36A09629B2D1FA3BABBDE9E1C34D668" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="850125977C289E470B5D9811002FACA0" authority="Zhao &amp; Liu, 2022" authorityName="Zhao &amp; Liu" authorityYear="2022" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="medongensis">
<emphasis id="25D055000899702A707797B04E4BD352" italics="true">Z. medongensis</emphasis>
<bibRefCitation id="7A3242C4A0D78A1299E1CC8F2C40EB88" DOI="10.1080/00379271.2022.2137059" author="Zhao Y &amp; Hu L &amp; Wesener T &amp; Liu W" journalOrPublisher="Annales de la Société entomologique de France" pagination="499-524" refId="B39" refString="Zhao Y, Hu L, Wesener T, Liu W (2022) Five new species of the giant pill-millipedes of the genera Zephronia and Sphaerobelum, from China (Diplopoda: Sphaerotheriida: Zephroniidae). Annales de la Societe entomologique de France 58: 499 - 524. https: // doi. org / 10.1080 / 00379271.2022. 2137059" title="Five new species of the giant pill-millipedes of the genera Zephronia and Sphaerobelum, from China (Diplopoda: Sphaerotheriida: Zephroniidae)." volume="58" year="2022">Zhao &amp; Liu, 2022</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="0023DBD5FCD8525A296F65EFBB2850FF" authority="Semenyuk, Golovatch &amp; Wesener, 2018" authorityName="Semenyuk, Golovatch &amp; Wesener" authorityYear="2018" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="montis">
<emphasis id="DB8B17511ABB5BF670A97EE6593EFABC" italics="true">Z. montis</emphasis>
<bibRefCitation id="F5C396FD91A24F09ACAAB2AB6D058D23" DOI="10.11646/zootaxa.4459.3.7" author="Semenyuk I &amp; Golovatch SI &amp; Wesener T" issue="3" journalOrPublisher="Zootaxa" pagination="535-550" refId="B27" refString="Semenyuk I, Golovatch SI, Wesener T (2018) Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae). Zootaxa 4459 (3): 535 - 550. https: // doi. org / 10.11646 / zootaxa. 4459.3. 7" title="Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae)." volume="4459" year="2018">Semenyuk, Golovatch &amp; Wesener, 2018</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="381FED0BA4AE0FF18D70160CAE9A95CE" authorityName="Srisonchai, Sutcharit &amp; Likhitrakarn" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="3FAF96E983F231EDB1ABCC2DFF105DF9" italics="true">Z. panhai</emphasis>
</taxonomicName>
,
<taxonomicName id="ED5C9AF68EFEF8CE88A41CDB3724C03E" authorityName="Hirst" authorityYear="1907" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="DE640DD50CEC0E9A25CE148E2EC8C404" italics="true">Z. siamensis</emphasis>
</taxonomicName>
and
<taxonomicName id="58897AC6D6ECDA083B1204D5A60BF5DD" authority="Zhao &amp; Liu, 2022" authorityName="Zhao &amp; Liu" authorityYear="2022" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="zhouae">
<emphasis id="C4CB7FEE74CD70D03050B88FDFD3E06F" italics="true">Z. zhouae</emphasis>
<bibRefCitation id="586498828473E2FE034DA8B17951D436" DOI="10.1080/00379271.2022.2137059" author="Zhao Y &amp; Hu L &amp; Wesener T &amp; Liu W" journalOrPublisher="Annales de la Société entomologique de France" pagination="499-524" refId="B39" refString="Zhao Y, Hu L, Wesener T, Liu W (2022) Five new species of the giant pill-millipedes of the genera Zephronia and Sphaerobelum, from China (Diplopoda: Sphaerotheriida: Zephroniidae). Annales de la Societe entomologique de France 58: 499 - 524. https: // doi. org / 10.1080 / 00379271.2022. 2137059" title="Five new species of the giant pill-millipedes of the genera Zephronia and Sphaerobelum, from China (Diplopoda: Sphaerotheriida: Zephroniidae)." volume="58" year="2022">Zhao &amp; Liu, 2022</bibRefCitation>
</taxonomicName>
. See more details for the genus in
<bibRefCitation id="0789B60CF75DBEB69917324A47E49EC6" author="Wesener T" firstAuthor="Wesener" journalOrPublisher="Bonn Zoological Bulletin Supplementum," pagination="1-104" refId="B32" refString="Wesener T (2016) The giant pill-millipedes, order Sphaerotheriida – An annotated species catalogue with morphological atlas and list of apomorphies (Arthropoda: Diplopoda). Bonn Zoological Bulletin Supplementum, 63: 1–104." title="The giant pill-millipedes, order Sphaerotheriida - An annotated species catalogue with morphological atlas and list of apomorphies (Arthropoda: Diplopoda)." volume="63" year="2016">Wesener (2016)</bibRefCitation>
and
<bibRefCitation id="21C89C3C1163370DB2000CE580A82274" DOI="10.11646/zootaxa.4459.3.7" author="Semenyuk I &amp; Golovatch SI &amp; Wesener T" etAl="et al." firstAuthor="Semenyuk" issue="3" journalOrPublisher="Zootaxa" pagination="535-550" refId="B27" refString="Semenyuk I, Golovatch SI, Wesener T (2018) Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae). Zootaxa 4459 (3): 535–550. https://doi.org/10.11646/zootaxa.4459.3.7" title="Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae)." volume="4459" year="2018">Semenyuk et al. (2018)</bibRefCitation>
, and a recent update of Thai species in
<bibRefCitation id="5ABEF6A82028CE66385316EC1C07BBA8" DOI="10.11646/zootaxa.5105.3.2" author="Bhansali S &amp; Wesener T" firstAuthor="Bhansali" issue="3" journalOrPublisher="Zootaxa" pagination="357-380" refId="B2" refString="Bhansali S, Wesener T (2022) New Thai Giant Pill-Millipede species, with genetic barcoding data for Thai species (Diplopoda, Sphaerotheriida, Zephroniidae). Zootaxa 5105 (3): 357–380. https://doi.org/10.11646/zootaxa.5105.3.2" title="New Thai Giant Pill-Millipede species, with genetic barcoding data for Thai species (Diplopoda, Sphaerotheriida, Zephroniidae)." volume="5105" year="2022">Bhansali and Wesener (2022)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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