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<document id="20B161FBBF6E8DD5C221330517768D77" ID-DOI="10.11646/zootaxa.4088.4.10" ID-GBIF-Dataset="fe977c75-55ca-4b45-b0ad-bbcee05f221e" ID-ISSN="1175-5326" ID-Zenodo-Dep="257763" ID-ZooBank="BA55503E-3A9A-4DB6-BDCE-5AF4E9238FA9" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457686088491" checkinUser="plazi" docAuthor="Hippa, Heikki &amp; Vilkamaa, Pekka" docDate="2016" docId="8B23AF56FFD6157310D11AEEFEA2801D" docLanguage="en" docName="15610-5129-1-PB.pdf.imf" docOrigin="Zootaxa 4088 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Claustropyga modica Hippa &amp; Vilkamaa, 2016, sp. n." docType="treatment" docVersion="11" lastPageNumber="598" masterDocId="771AD72EFFD5157710461938FFBB8100" masterDocTitle="New species of Claustropyga Hippa, Vilkamaa &amp; Mohrig (Diptera, Sciaridae) from the Holarctic region" masterLastPageNumber="600" masterPageNumber="594" pageNumber="597" updateTime="1698178453981" updateUser="plazi">
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<mods:title id="5A7705F0E012A08B9FA1206B0C3E2931">New species of Claustropyga Hippa, Vilkamaa &amp; Mohrig (Diptera, Sciaridae) from the Holarctic region</mods:title>
</mods:titleInfo>
<mods:name id="4050D4724AE2A5ACD99EA641E213AFEE" type="personal">
<mods:role id="1F4011A84A30AFDDC430C6556E32A0F2">
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<mods:namePart id="D37B01BD09EE86A612D1EADF84B06D0F">Hippa, Heikki</mods:namePart>
</mods:name>
<mods:name id="5A1DF9294D6D279332E4E62D059629F7" type="personal">
<mods:role id="44A6B341FA3C62FD7526D5EB30F58EB2">
<mods:roleTerm id="4916252F9781BF14FB83D246924C4405">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C1F95517C98A0C36E69F70BD63653C98">Vilkamaa, Pekka</mods:namePart>
</mods:name>
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<mods:title id="3161A5E80419BD607815569C42704554">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6118E4CDFCABEF6A72464F7362FAF4F4">
<mods:date id="26533E8927AE54BAE11398834D7FC975">2016</mods:date>
<mods:detail id="7E4086CEA2B8EAE66785A21DD6B2F0A3" type="volume">
<mods:number id="124894FF7C82EE0665723063048C6F79">4088</mods:number>
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<mods:detail id="3572BB51AB7A838E3206C7E4DF9A6727" type="issue">
<mods:number id="0107BA98CBD74285308662AC84D65163">4</mods:number>
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<mods:classification id="07576D9C9540C15008AF50A49EB1D7D7">journal article</mods:classification>
<mods:identifier id="94367655D3FA7D007AB119F5B196B425" type="DOI">10.11646/zootaxa.4088.4.10</mods:identifier>
<mods:identifier id="16CE53A55DF14D8652334BDEB8E377BE" type="GBIF-Dataset">fe977c75-55ca-4b45-b0ad-bbcee05f221e</mods:identifier>
<mods:identifier id="19BFE544615E16D315C7C2BEFCA571CC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DC12F496E7F3FF30C755D3590AB12653" type="Zenodo-Dep">257763</mods:identifier>
<mods:identifier id="CD84D7D896BD2B498C78F24D364A9938" type="ZooBank">BA55503E-3A9A-4DB6-BDCE-5AF4E9238FA9</mods:identifier>
</mods:mods>
<treatment id="8B23AF56FFD6157310D11AEEFEA2801D" ID-DOI="http://doi.org/10.5281/zenodo.6088015" ID-GBIF-Taxon="117222044" ID-Zenodo-Dep="6088015" LSID="urn:lsid:plazi:treatment:8B23AF56FFD6157310D11AEEFEA2801D" httpUri="http://treatment.plazi.org/id/8B23AF56FFD6157310D11AEEFEA2801D" lastPageId="4" lastPageNumber="598" pageId="3" pageNumber="597">
<subSubSection id="4B904DCBFFD6157410D11AEEFE5A82F0" box="[151,481,982,1008]" pageId="3" pageNumber="597" type="nomenclature">
<paragraph id="03351E40FFD6157410D11AEEFE5A82F0" blockId="3.[151,481,982,1042]" box="[151,481,982,1008]" pageId="3" pageNumber="597">
<heading id="587DA92CFFD6157410D11AEEFE5A82F0" bold="true" box="[151,481,982,1008]" fontSize="11" level="1" pageId="3" pageNumber="597" reason="1">
<emphasis id="31FEC252FFD6157410D11AEEFE5A82F0" bold="true" box="[151,481,982,1008]" pageId="3" pageNumber="597">
<taxonomicName id="C48A65C3FFD6157410D11AEEFE2C82F0" ID-CoL="8BJ4S" box="[151,407,982,1008]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="modica" status="sp. nov.">
<emphasis id="31FEC252FFD6157410D11AEEFE2C82F0" bold="true" box="[151,407,982,1008]" italics="true" pageId="3" pageNumber="597">Claustropyga modica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2ACD7F29FFD6157411D81AEEFE5A82F0" box="[414,481,982,1008]" pageId="3" pageNumber="597" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4B904DCBFFD6157410D11AC1FEA38511" box="[151,280,1017,1042]" pageId="3" pageNumber="597" type="description">
<paragraph id="03351E40FFD6157410D11AC1FEA38511" blockId="3.[151,481,982,1042]" box="[151,280,1017,1042]" pageId="3" pageNumber="597">
<figureCitation id="9BB102C5FFD6157410D11AC1FF678512" box="[151,220,1017,1042]" captionStart="FIGURE 3" captionStartId="4.[151,250,1073,1094]" captionTargetBox="[194,1390,347,1045]" captionTargetPageId="4" captionText="FIGURE 3. Claustropyga modica sp. n. (holotype). A. Part of hypopygium, ventral. B. Gonostylus, ventral. Scale 0.1 mm." httpUri="https://zenodo.org/record/257766/files/figure.png" pageId="3" pageNumber="597">Figs 3</figureCitation>
A, B
</paragraph>
</subSubSection>
<subSubSection id="4B904DCBFFD6157410D11D79FE2587E5" pageId="3" pageNumber="597" type="materials_examined">
<paragraph id="03351E40FFD6157410D11D79FCA885EA" blockId="3.[151,1437,1089,2017]" pageId="3" pageNumber="597">
<emphasis id="31FEC252FFD6157410D11D79FEDE855A" bold="true" box="[151,357,1089,1114]" pageId="3" pageNumber="597">Material studied</emphasis>
.
<emphasis id="31FEC252FFD61574113A1D79FD938559" box="[380,552,1089,1113]" italics="true" pageId="3" pageNumber="597">
<typeStatus id="DC31A0E2FFD61574113A1D79FE588559" box="[380,483,1089,1113]" pageId="3" pageNumber="597" type="holotype">Holotype</typeStatus>
male
</emphasis>
.
<collectingCountry id="7B9D5ED0FFD6157412791D7AFD06855A" box="[575,701,1090,1114]" name="Finland" pageId="3" pageNumber="597">FINLAND</collectingCountry>
, Regio kuusamoensis, Taivalkoski, Kylmäoja (grid 7275293: 3554865), brook, Malaise trap,
<date id="77343880FFD6157412741D5DFD50857E" box="[562,747,1125,1150]" pageId="3" pageNumber="597" value="2006-06-11" valueMax="2006-07-03" valueMin="2006-06-11">11.vi–3.vii.2006</date>
, J. Salmela (in MZH).
<emphasis id="31FEC252FFD6157414721D5EFB1E857D" box="[1076,1189,1126,1149]" italics="true" pageId="3" pageNumber="597">
<typeStatus id="DC31A0E2FFD6157414721D5EFB1E857D" box="[1076,1189,1126,1149]" pageId="3" pageNumber="597" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectingCountry id="7B9D5ED0FFD6157414841D5EFAFB857E" box="[1218,1344,1126,1150]" name="Finland" pageId="3" pageNumber="597">FINLAND</collectingCountry>
, Regio kuusamoensis, Kuusamo, Uopajanpuro (grid 7362617:3612763), brook, Malaise trap,
<date id="77343880FFD6157414201DB1FB4085A2" box="[1126,1275,1161,1186]" pageId="3" pageNumber="597" value="2005-06-01" valueMax="2005-06-20" valueMin="2005-06-01">1–20.vi.2005</date>
, J. Salmela (
<specimenCount id="158CD5C9FFD6157415D71DB1FF7685C6" pageId="3" pageNumber="597" type="male">1 male</specimenCount>
in MZH); Lapponia kemensis orientalis, Salla, Värriö, Kuntasjoki (grid 7520406:3610772), over brook, Malaise trap,
<date id="77343880FFD6157411761DE9FE7D85EA" box="[304,454,1233,1258]" pageId="3" pageNumber="597" value="2013-06-04" valueMax="2013-06-29" valueMin="2013-06-04">4–29.vi.2013</date>
, J. Salmela (
<specimenCount id="158CD5C9FFD61574121E1DE9FD1B85EA" box="[600,672,1233,1258]" pageId="3" pageNumber="597" type="male">1 male</specimenCount>
in MZH).
</paragraph>
<paragraph id="03351E40FFD6157410811DCDFE2D87C0" blockId="3.[151,1437,1089,2017]" pageId="3" pageNumber="597">
Male.
<emphasis id="31FEC252FFD6157411481DCEFEF6840D" bold="true" box="[270,333,1270,1293]" pageId="3" pageNumber="597">Head</emphasis>
. Brown, antenna paler, maxillary palpus very pale brown. Eye bridge 2 facets wide. Face with 2–8 scattered longer and shorter setae. Clypeus with 1 setae or non-setose. Maxillary palpus with 2 or 3 segments; segment 1 longer than 2 and 3, these subequal in length; segment 1 with 2 long sharp setae, with a dorsal group of sensilla; segment 2 with 1 long sharp seta and 3–4 shorter truncate setae, segment 3 with 4–5 short truncate setae. Body of antennal flagellomere
<geoCoordinate id="66BE7887FFD6157411B71CBEFD92849E" box="[497,553,1414,1438]" pageId="3" pageNumber="597">4 2.7</geoCoordinate>
x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere.
<emphasis id="31FEC252FFD6157411D11C92FE4B84C1" bold="true" box="[407,496,1450,1473]" pageId="3" pageNumber="597">Thorax</emphasis>
. Anterior pronotum with 1–4 setae. Episternum 1 with 4–5 setae.
<emphasis id="31FEC252FFD6157414921C91FAA784C2" bold="true" box="[1236,1308,1449,1474]" pageId="3" pageNumber="597">Wing.</emphasis>
Length
<quantity id="C472B3A5FFD61574153F1C92FF7D84E5" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="597" unit="mm" value="1.9">1.9 mm</quantity>
. Width/length 0.45. R1/R 0.60–0.95. c/
<geoCoordinate id="66BE7887FFD6157412391CF5FD6984E6" box="[639,722,1485,1510]" direction="west" orientation="longitude" pageId="3" pageNumber="597" precision="555" value="-0.8">w 0.80</geoCoordinate>
. r-m and bM non-setose. Halter pale brown.
<emphasis id="31FEC252FFD6157414931CF6FAB684E5" bold="true" box="[1237,1293,1486,1509]" pageId="3" pageNumber="597">Legs</emphasis>
. Front tibial organ with vestiture in indistinct patch. Front tibial spur slightly longer than the tibial width.
<emphasis id="31FEC252FFD6157414D81CCCFAA9870D" bold="true" box="[1182,1298,1524,1549]" pageId="3" pageNumber="597">Abdomen</emphasis>
. Setae dark, rather short.
<emphasis id="31FEC252FFD6157411611F20FE798731" bold="true" box="[295,450,1560,1585]" pageId="3" pageNumber="597">Hypopygium</emphasis>
(
<figureCitation id="9BB102C5FFD6157411921F20FDAC8730" box="[468,535,1560,1585]" captionStart="FIGURE 3" captionStartId="4.[151,250,1073,1094]" captionTargetBox="[194,1390,347,1045]" captionTargetPageId="4" captionText="FIGURE 3. Claustropyga modica sp. n. (holotype). A. Part of hypopygium, ventral. B. Gonostylus, ventral. Scale 0.1 mm." httpUri="https://zenodo.org/record/257766/files/figure.png" pageId="3" pageNumber="597">Fig. 3</figureCitation>
A). Intercoxal area not lobe-like, medially much shorter than gonocoxite at the base of gonostylus, membraneous area richly setose; gonocoxa slightly longer than gonostylus, with normal setosity; gonostylus (
<figureCitation id="9BB102C5FFD6157411CE1F58FE708778" box="[392,459,1632,1657]" captionStart="FIGURE 3" captionStartId="4.[151,250,1073,1094]" captionTargetBox="[194,1390,347,1045]" captionTargetPageId="4" captionText="FIGURE 3. Claustropyga modica sp. n. (holotype). A. Part of hypopygium, ventral. B. Gonostylus, ventral. Scale 0.1 mm." httpUri="https://zenodo.org/record/257766/files/figure.png" pageId="3" pageNumber="597">Fig. 3</figureCitation>
B) swollen, medially concave, with normal setosity, bearing an apical tooth, and 4–5 apical-subapical megasetae, megasetae slender, straight, subequal in length. Tegmen longer than broad, with a distinct apical process.
</paragraph>
<paragraph id="03351E40FFD6157410811FF4FE2587E5" blockId="3.[151,1437,1089,2017]" box="[199,414,1740,1765]" pageId="3" pageNumber="597">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="4B904DCBFFD6157310811FC8FB4181D5" lastPageId="4" lastPageNumber="598" pageId="3" pageNumber="597" type="discussion">
<paragraph id="03351E40FFD6157310811FC8FB4181D5" blockId="3.[151,1437,1089,2017]" lastBlockId="4.[151,1436,151,285]" lastPageId="4" lastPageNumber="598" pageId="3" pageNumber="597">
<emphasis id="31FEC252FFD6157410811FC8FEFE8609" bold="true" box="[199,325,1776,1801]" pageId="3" pageNumber="597">Discussion</emphasis>
.
<taxonomicName id="C48A65C3FFD6157411121FC8FDFC8608" box="[340,583,1776,1800]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="modica">
<emphasis id="31FEC252FFD6157411121FC8FDFC8608" box="[340,583,1776,1800]" italics="true" pageId="3" pageNumber="597">Claustropyga modica</emphasis>
</taxonomicName>
is one of the few species in the genus lacking any indication of the ventral intercoxal lobe of the hypopygium. The other similar species are
<taxonomicName id="C48A65C3FFD6157413D41E2CFB50862D" authority="Hippa et al. 2003" authorityName="Hippa et al." authorityYear="2003" box="[914,1259,1812,1837]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="aperta">
<emphasis id="31FEC252FFD6157413D41E2CFC44862C" box="[914,1023,1812,1836]" italics="true" pageId="3" pageNumber="597">C. aperta</emphasis>
(Hippa
<emphasis id="31FEC252FFD6157414241E2DFB25862C" box="[1122,1182,1812,1836]" italics="true" pageId="3" pageNumber="597">et al.</emphasis>
2003)
</taxonomicName>
,
<taxonomicName id="C48A65C3FFD6157414BC1E2CFE4F8651" authority="Mohrig &amp; Roschmann, 1993" authorityName="Mohrig &amp; Roschmann" authorityYear="1993" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="janetscheki">
<emphasis id="31FEC252FFD6157414BC1E2CFA27862C" box="[1274,1436,1812,1836]" italics="true" pageId="3" pageNumber="597">C. janetscheki</emphasis>
(Mohrig &amp; Röschmann, 1993)
</taxonomicName>
,
<taxonomicName id="C48A65C3FFD6157412441E00FC318651" authority="Lengersdorf, 1930" authorityName="Lengersdorf" authorityYear="1930" box="[514,906,1848,1873]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="refrigerata">
<emphasis id="31FEC252FFD6157412441E00FD258650" box="[514,670,1848,1872]" italics="true" pageId="3" pageNumber="597">C. refrigerata</emphasis>
(Lengersdorf, 1930)
</taxonomicName>
and
<taxonomicName id="C48A65C3FFD6157413831E00FA238651" authority="Mohrig &amp; Antonova, 1978" authorityName="Mohrig &amp; Antonova" authorityYear="1978" box="[965,1432,1848,1873]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="sajanica">
<emphasis id="31FEC252FFD6157413831E00FBFC8650" box="[965,1095,1848,1872]" italics="true" pageId="3" pageNumber="597">C. sajanica</emphasis>
(Mohrig &amp; Antonova, 1978)
</taxonomicName>
.
<taxonomicName id="C48A65C3FFD6157410D11E64FE318674" box="[151,394,1884,1908]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="modica">
<emphasis id="31FEC252FFD6157410D11E64FE318674" box="[151,394,1884,1908]" italics="true" pageId="3" pageNumber="597">Claustropyga modica</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="C48A65C3FFD6157412C81E64FC948674" box="[654,815,1884,1908]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="janetscheki">
<emphasis id="31FEC252FFD6157412C81E64FC948674" box="[654,815,1884,1908]" italics="true" pageId="3" pageNumber="597">C. janetscheki</emphasis>
</taxonomicName>
and
<taxonomicName id="C48A65C3FFD61574132C1E64FC6D8674" box="[874,982,1884,1908]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="aperta">
<emphasis id="31FEC252FFD61574132C1E64FC6D8674" box="[874,982,1884,1908]" italics="true" pageId="3" pageNumber="597">C. aperta</emphasis>
</taxonomicName>
in having only 4 or 5 megasetae on its gonostylus, instead of nine or more, from
<taxonomicName id="C48A65C3FFD61574123F1EB8FCAD8698" box="[633,790,1920,1944]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="refrigerata">
<emphasis id="31FEC252FFD61574123F1EB8FCAD8698" box="[633,790,1920,1944]" italics="true" pageId="3" pageNumber="597">C. refrigerata</emphasis>
</taxonomicName>
in lacking a group of lateral subapical megasetae on the gonostylus and from
<taxonomicName id="C48A65C3FFD6157411C31E9CFDBE86BC" box="[389,517,1956,1980]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="sajanica">
<emphasis id="31FEC252FFD6157411C31E9CFDBE86BC" box="[389,517,1956,1980]" italics="true" pageId="3" pageNumber="597">C. sajanica</emphasis>
</taxonomicName>
in the rounded, not angular, posterolateral shoulders of the tegmen and by having all the ventral gonostylar setae long and gently curved (in
<taxonomicName id="C48A65C3FFD6157413671EF0FC1A86E0" box="[801,929,1992,2016]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="sajanica">
<emphasis id="31FEC252FFD6157413671EF0FC1A86E0" box="[801,929,1992,2016]" italics="true" pageId="3" pageNumber="597">C. sajanica</emphasis>
</taxonomicName>
there is a posterior area of short and strongly curved setae). By its flattened apicoventral portion of the gonostylus,
<taxonomicName id="C48A65C3FFD1157313DB19A0FBAB81AF" box="[925,1040,151,176]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="598" phylum="Arthropoda" rank="species" species="modica">
<emphasis id="31FEC252FFD1157313DB19A0FBAB81AF" box="[925,1040,151,176]" italics="true" pageId="4" pageNumber="598">C. modica</emphasis>
</taxonomicName>
differs from all
<taxonomicName id="C48A65C3FFD11573148219AFFAE781AF" box="[1220,1372,151,175]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="598" phylum="Arthropoda" rank="genus">
<emphasis id="31FEC252FFD11573148219AFFAE781AF" box="[1220,1372,151,175]" italics="true" pageId="4" pageNumber="598">Claustropyga</emphasis>
</taxonomicName>
other than
<taxonomicName id="C48A65C3FFD1157310881984FD7681D5" authority="Mohrig &amp; Krivosheina, 1985" authorityName="Mohrig &amp; Krivosheina" authorityYear="1985" box="[206,717,188,213]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="598" phylum="Arthropoda" rank="species" species="subcorticis">
<emphasis id="31FEC252FFD1157310881984FED181D4" box="[206,362,188,212]" italics="true" pageId="4" pageNumber="598">C. subcorticis</emphasis>
(Mohrig &amp; Krivosheina, 1985)
</taxonomicName>
. The two species are otherwise greatly dissimilar.
</paragraph>
</subSubSection>
<subSubSection id="4B904DCBFFD11573108119D8FEA2801D" pageId="4" pageNumber="598" type="etymology">
<paragraph id="03351E40FFD11573108119D8FEA2801D" blockId="4.[151,1436,151,285]" pageId="4" pageNumber="598">
<emphasis id="31FEC252FFD11573108119D8FEFC81F7" bold="true" box="[199,327,224,247]" pageId="4" pageNumber="598">Etymology</emphasis>
. The name is Latin,
<taxonomicName id="C48A65C3FFD11573127519E7FD3D81F7" box="[563,646,223,247]" class="Insecta" family="Sciaridae" genus="Claustropyga" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="598" phylum="Arthropoda" rank="species" species="modica">
<emphasis id="31FEC252FFD11573127519E7FD3D81F7" box="[563,646,223,247]" italics="true" pageId="4" pageNumber="598">modica</emphasis>
</taxonomicName>
(modest), referring to the unmodified structure of the hypopygium of the species.
</paragraph>
</subSubSection>
</treatment>
</document>
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